589 lines
85 KiB
XML
589 lines
85 KiB
XML
<document id="B2DC020DFBBA33EC6A67C593A8403360" ID-CLB-Dataset="3777" ID-DOI="10.11646/zootaxa.5060.2.1" ID-GBIF-Dataset="ed98a754-c3ac-49ff-9973-b8d9839993ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="5627213" ID-ZooBank="F6BFFABC-7F88-473C-9517-D031485140DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635492351830" checkinUser="plazi" docAuthor="Moravec, Jiří & Trýzna, Miloš" docDate="2021" docId="03B477221135FFE7DCF8D83C2BF6FB0B" docLanguage="en" docName="zootaxa.5060.2.1.pdf" docOrigin="Zootaxa 5060 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Physodeutera (Toxoma) dubia" docType="treatment" docVersion="9" lastPageNumber="164" masterDocId="FF8D0F5A113CFFEADC6FD91A2A22FFAA" masterDocTitle="New or rare Madagascar tiger beetles- 23. Physodeutera (Toxoma) lokobensis sp nov., a new species close to Ph. (T.) conturbata Moravec, Ph. (T.) sulcoprothoracica (W. Horn) and Ph. (T.) dubia (Mařan), with revised type designation of the latter (Coleoptera: Cicindelidae)" masterLastPageNumber="182" masterPageNumber="151" pageNumber="160" updateTime="1699467157693" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3CDCFBA642A2578DE3966AD793818A8E">New or rare Madagascar tiger beetles- 23. Physodeutera (Toxoma) lokobensis sp nov., a new species close to Ph. (T.) conturbata Moravec, Ph. (T.) sulcoprothoracica (W. Horn) and Ph. (T.) dubia (Mařan), with revised type designation of the latter (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="C516C70C250D4E5DBA6D15D478B72329">Moravec, Jiří</mods:namePart>
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<mods:namePart id="8F7A5B17245717628B4CCBBAB3463345">Trýzna, Miloš</mods:namePart>
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<mods:title id="AA2C2D533A4D3775526DB57AE59C2DF4">Zootaxa</mods:title>
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<taxonomicName id="4C1DBDB71135FFE3DCF8D83C289CFEEA" ID-CoL="8MB3N" authority="(Maran, 1942)" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[151,702,294,321]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
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<emphasis id="B9691A261135FFE3DCF8D83C289CFEEA" bold="true" box="[151,702,294,321]" pageId="9" pageNumber="160">
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<emphasis id="B9691A261135FFE3DCF8D83C2B1EFEEB" bold="true" box="[151,316,294,321]" italics="true" pageId="9" pageNumber="160">Physodeutera</emphasis>
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(
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<emphasis id="B9691A261135FFE3DD22D83D2B8CFEEB" bold="true" box="[333,430,295,321]" italics="true" pageId="9" pageNumber="160">Toxoma</emphasis>
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)
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<emphasis id="B9691A261135FFE3DDD0D83C2821FEEB" bold="true" box="[447,515,294,321]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
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(
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<bibRefCitation id="EF8CBBC51135FFE3DE7CD83C2897FEEA" author="Maran, J." box="[531,693,294,320]" pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan, 1942</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30795BF1135FFE3DCF8D8512B7DFECC" box="[151,351,330,358]" pageId="9" pageNumber="160" type="description">
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<paragraph id="8BA2C6341135FFE3DCF8D8512B7DFECC" blockId="9.[151,702,294,358]" box="[151,351,330,358]" pageId="9" pageNumber="160">
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(
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<figureCitation id="1326DAB11135FFE3DCF0D8512B02FECF" box="[159,288,330,357]" captionStart-0="FIGURES 24–32" captionStart-1="FIGURES 33–43" captionStart-2="FIGURES 44–45" captionStartId-0="10.[151,264,1867,1891]" captionStartId-1="11.[151,264,1867,1891]" captionStartId-2="12.[151,264,1018,1042]" captionTargetBox-0="[157,1431,195,1835]" captionTargetBox-1="[158,1429,188,1839]" captionTargetBox-2="[164,1425,191,978]" captionTargetId-0="figure-31@10.[151,1435,181,1844]" captionTargetId-1="figure-17@11.[151,1435,181,1844]" captionTargetId-2="figure-31@12.[151,1435,181,993]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." captionText-1="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." captionText-2="FIGURES 44–45. Physodeutera (Toxoma) dubia Mařan, all in SDEI, labels. 44—HT; 45—PT." figureDoi-0="http://doi.org/10.5281/zenodo.5627221" figureDoi-1="http://doi.org/10.5281/zenodo.5627223" figureDoi-2="http://doi.org/10.5281/zenodo.5627225" httpUri-0="https://zenodo.org/record/5627221/files/figure.png" httpUri-1="https://zenodo.org/record/5627223/files/figure.png" httpUri-2="https://zenodo.org/record/5627225/files/figure.png" pageId="9" pageNumber="160">Figs 24–45</figureCitation>
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,
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<figureCitation id="1326DAB11135FFE3DD43D8512B75FECC" box="[300,343,331,358]" captionStart="FIGURE 106" captionStartId="27.[151,250,1561,1585]" captionTargetBox="[271,1319,572,1531]" captionTargetId="figure-167@27.[262,1326,561,1538]" captionTargetPageId="27" captionText="FIGURE 106. Distribution of four Physodeutera (Toxoma) species in northern part of Madagascar. Adopted from the map of preserved areas in Madagascar according to Moat & Smith (2007), adapted and published previously (Moravec 2010) under permission of the trustee of the Royal Botanic Gardens, Kew, GB." figureDoi="http://doi.org/10.5281/zenodo.5627247" httpUri="https://zenodo.org/record/5627247/files/figure.png" pageId="9" pageNumber="160">106</figureCitation>
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)
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</paragraph>
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<subSubSection id="C30795BF1135FFE3DCF8D8952951FE4F" pageId="9" pageNumber="160" type="reference_group">
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<paragraph id="8BA2C6341135FFE3DCF8D89528A4FE0D" blockId="9.[151,883,399,485]" box="[151,646,399,423]" pageId="9" pageNumber="160">
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<treatmentCitationGroup id="AB0DE11A1135FFE3DCF8D89528A4FE0D" box="[151,646,399,423]" pageId="9" pageNumber="160">
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<treatmentCitation id="0ABCE0251135FFE3DCF8D89528A3FE0D" author="Maran, J." box="[151,641,399,423]" page="70" pageId="9" pageNumber="160" year="1942">
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<taxonomicName id="4C1DBDB71135FFE3DCF8D89528A3FE0D" ID-CoL="8LMH3" authority="Maran, 1942: 70" authorityName="Maran" authorityPageNumber="70" authorityYear="1942" box="[151,641,399,423]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Megalomma">
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<emphasis id="B9691A261135FFE3DCF8D8952AD8FE0D" box="[151,250,399,423]" italics="true" pageId="9" pageNumber="160">Prothyma</emphasis>
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(
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<emphasis id="B9691A261135FFE3DD66D8952BAAFE0D" box="[265,392,399,423]" italics="true" pageId="9" pageNumber="160">Megalomma</emphasis>
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)
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<emphasis id="B9691A261135FFE3DDF9D8952BF2FE0D" box="[406,464,399,423]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
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<bibRefCitation id="EF8CBBC51135FFE3DDB9D89528A3FE0D" author="Maran, J." box="[470,641,399,423]" pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan, 1942: 70</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BA2C6341135FFE3DCF8D8B428D2FE6C" blockId="9.[151,883,399,485]" box="[151,752,430,454]" pageId="9" pageNumber="160">
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<treatmentCitation id="0ABCE0251135FFE3DCF8D8B428CFFE6C" author="Moravec, J." box="[151,749,430,454]" page="169" pageId="9" pageNumber="160" year="1998">
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<taxonomicName id="4C1DBDB71135FFE3DCF8D8B428CFFE6C" ID-CoL="8MB3N" authority=": Moravec 1998: 169" authorityName="Moravec" authorityPageNumber="169" authorityYear="1998" box="[151,749,430,454]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="subSpecies" species="sikorai" subGenus="Toxoma" subSpecies="dubia">
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<emphasis id="B9691A261135FFE3DCF8D8B42B00FE6C" box="[151,290,430,454]" italics="true" pageId="9" pageNumber="160">Physodeutera</emphasis>
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(
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<emphasis id="B9691A261135FFE3DD5ED8B52BA2FE6C" box="[305,384,431,454]" italics="true" pageId="9" pageNumber="160">Toxoma</emphasis>
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)
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<emphasis id="B9691A261135FFE3DDE0D8B42834FE6C" box="[399,534,430,454]" italics="true" pageId="9" pageNumber="160">sikorai dubia</emphasis>
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:
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<bibRefCitation id="EF8CBBC51135FFE3DE4CD8B428CFFE6C" author="Moravec, J." box="[547,749,430,454]" pageId="9" pageNumber="160" pagination="167 - 186" refId="ref15377" refString="Moravec, J. (1998) Revision within the genus Physodeutera - neglected species described by Maran and related taxa (Coleop- tera: Cicindelidae). Folia Heyrovskyana, 6, 167 - 186." type="journal article" year="1998">Moravec 1998: 169</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BA2C6341135FFE3DCF8D8D62951FE4F" blockId="9.[151,883,399,485]" box="[151,883,460,485]" pageId="9" pageNumber="160">
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<treatmentCitationGroup id="AB0DE11A1135FFE3DCF8D8D62951FE4F" box="[151,883,460,485]" pageId="9" pageNumber="160">
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<treatmentCitation id="0ABCE0251135FFE3DCF8D8D6288BFE4E" author="Moravec, J." box="[151,681,460,485]" page="115" pageId="9" pageNumber="160" year="2002">
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<taxonomicName id="4C1DBDB71135FFE3DCF8D8D6288BFE4E" ID-CoL="8MB3N" authority=": Moravec 2002 a: 115" authorityName="Moravec" authorityPageNumber="115" authorityYear="2002" box="[151,681,460,485]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
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<emphasis id="B9691A261135FFE3DCF8D8D62B00FE4E" box="[151,290,460,484]" italics="true" pageId="9" pageNumber="160">Physodeutera</emphasis>
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(
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<emphasis id="B9691A261135FFE3DD5ED8D72BA2FE4E" box="[305,384,461,484]" italics="true" pageId="9" pageNumber="160">Toxoma</emphasis>
|
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)
|
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<emphasis id="B9691A261135FFE3DDE0D8D62BEBFE4E" box="[399,457,460,484]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
|
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:
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<bibRefCitation id="EF8CBBC51135FFE3DDB9D8D7288BFE4E" author="Moravec, J." box="[470,681,460,485]" pageId="9" pageNumber="160" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a: 115</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 303–308, 723.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30795BF1135FFE3DCF8DB09292FFB3B" pageId="9" pageNumber="160" type="materials_examined">
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<paragraph id="8BA2C6341135FFE3DCF8DB092EAEFD84" blockId="9.[151,1437,530,1997]" box="[151,1164,530,558]" pageId="9" pageNumber="160">
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<materialsCitation id="3B75CC691135FFE3DCF8DB092EAAFD84" ID-GBIF-Occurrence="3395513311" box="[151,1160,530,558]" country="Madagascar" location="Vohemar" municipality="Northeastern" pageId="9" pageNumber="160" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9691A261135FFE3DCF8DB092B12FD86" bold="true" box="[151,304,530,557]" pageId="9" pageNumber="160">
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<typeStatus id="54A678961135FFE3DCF8DB092AF0FD87" box="[151,210,531,557]" pageId="9" pageNumber="160">Type</typeStatus>
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locality
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</emphasis>
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||
. “Ampanefena” (about
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south of
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<location id="8EC290EF1135FFE3DE84DB092970FD87" LSID="urn:lsid:plazi:treatment:03B477221135FFE7DCF8D83C2BF6FB0B:8EC290EF1135FFE3DE84DB092970FD87" box="[747,850,531,557]" country="Madagascar" municipality="Northeastern" name="Vohemar" pageId="9" pageNumber="160">Vohémar</location>
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,
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||
<collectingMunicipality id="6BC65C4E1135FFE3DF33DB0929CFFD87" box="[860,1005,531,557]" pageId="9" pageNumber="160">Northeastern</collectingMunicipality>
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<collectingCountry id="F30A86A41135FFE3DF9BDB092EA3FD84" box="[1012,1153,531,558]" name="Madagascar" pageId="9" pageNumber="160">Madagascar</collectingCountry>
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)
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||
</materialsCitation>
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.
|
||
</paragraph>
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<paragraph id="8BA2C6341135FFE3DCA8DB2D2FB0FCAF" blockId="9.[151,1437,530,1997]" pageId="9" pageNumber="160">
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<emphasis id="B9691A261135FFE3DCA8DB2D2B52FDFA" bold="true" box="[199,368,566,593]" pageId="9" pageNumber="160">Type material</emphasis>
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.
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<typeStatus id="54A678961135FFE3DDEFDB2D2BCAFDFB" box="[384,488,567,593]" pageId="9" pageNumber="160" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D1B0DBD1135FFE3DD9EDB2D2824FDFB" box="[497,518,567,593]" pageId="9" pageNumber="160" type="male">♂</specimenCount>
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in
|
||
<collectionCode id="ED0C5EF11135FFE3DE42DB2D285DFDFB" box="[557,639,567,593]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="160">NMPC</collectionCode>
|
||
, labelled: “
|
||
<collectingCountry id="F30A86A41135FFE3DE90DB2D29A9FDFB" box="[767,907,567,593]" name="Madagascar" pageId="9" pageNumber="160">Madagascar</collectingCountry>
|
||
, Ampanefena, Mus. Praha” [printed]; “Typus” [red, printed]; “
|
||
<taxonomicName id="4C1DBDB71135FFE3DD50DB412BD1FDDF" authorityName="Maran" authorityYear="1942" box="[319,499,603,629]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia">Prothyma dubia</taxonomicName>
|
||
m. Dr. Mařan det.” [handwritten]; “
|
||
<typeStatus id="54A678961135FFE3DF15DB4129CAFDDF" box="[890,1000,603,629]" pageId="9" pageNumber="160" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<taxonomicName id="4C1DBDB71135FFE3DF9EDB412AF0FD33" authority="Maran, 1942" authorityName="Maran" authorityYear="1942" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Megalomma">
|
||
Prothyma (Megalomma) dubia
|
||
<bibRefCitation id="EF8CBBC51135FFE3D923DB412AF0FD33" author="Maran, J." pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan, 1942</bibRefCitation>
|
||
</taxonomicName>
|
||
, J.
|
||
<bibRefCitation id="EF8CBBC51135FFE3DC98DB652BB8FD33" author="Moravec, J." box="[247,410,639,665]" pageId="9" pageNumber="160" pagination="167 - 186" refId="ref15377" refString="Moravec, J. (1998) Revision within the genus Physodeutera - neglected species described by Maran and related taxa (Coleop- tera: Cicindelidae). Folia Heyrovskyana, 6, 167 - 186." type="journal article" year="1998">Moravec 1998</bibRefCitation>
|
||
design.” [red, printed].
|
||
<typeStatus id="54A678961135FFE3DEC6DB652929FD33" box="[681,779,639,665]" pageId="9" pageNumber="160" type="paratype">Paratype</typeStatus>
|
||
<specimenCount id="9D1B0DBD1135FFE3DF7DDB65291FFD33" box="[786,829,639,665]" pageId="9" pageNumber="160" type="male">1 ♂</specimenCount>
|
||
in
|
||
<collectionCode id="ED0C5EF11135FFE3DF0EDB652992FD33" box="[865,944,639,665]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="160">NMPC</collectionCode>
|
||
“Vohemar, III, 1938” [printed]; “CoTypus” [red, printed, the “Co” handwritten]; “
|
||
<typeStatus id="54A678961135FFE3DE2CDBB92889FD17" box="[579,683,675,701]" pageId="9" pageNumber="160" type="paratype">Paratype</typeStatus>
|
||
,
|
||
<taxonomicName id="4C1DBDB71135FFE3DED9DBB92E88FD17" authority="Maran, 1942" authorityName="Maran" authorityYear="1942" box="[694,1194,675,702]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Megalomma">
|
||
Prothyma (Megalomma) dubia
|
||
<bibRefCitation id="EF8CBBC51135FFE3D878DBB92E88FD17" author="Maran, J." box="[1047,1194,675,701]" pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan, 1942</bibRefCitation>
|
||
</taxonomicName>
|
||
, J.
|
||
<bibRefCitation id="EF8CBBC51135FFE3D8A1DBB92F53FD17" author="Moravec, J." box="[1230,1393,675,701]" pageId="9" pageNumber="160" pagination="167 - 186" refId="ref15377" refString="Moravec, J. (1998) Revision within the genus Physodeutera - neglected species described by Maran and related taxa (Coleop- tera: Cicindelidae). Folia Heyrovskyana, 6, 167 - 186." type="journal article" year="1998">Moravec 1998</bibRefCitation>
|
||
design.” [red, printed]. The
|
||
<specimenCount id="9D1B0DBD1135FFE3DDD9DBDD283AFD4B" box="[438,536,711,737]" pageId="9" pageNumber="160" type="generic">two type</specimenCount>
|
||
specimens provided with labels “
|
||
<taxonomicName id="4C1DBDB71135FFE3DFFDDBDD2ED7FD4B" baseAuthorityName="W. Horn" baseAuthorityYear="1896" box="[914,1269,711,737]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="sikorai" subGenus="Toxoma">Physodeutera (Toxoma) sikorai</taxonomicName>
|
||
<taxonomicName id="4C1DBDB71135FFE3D892DBDD2AF9FCAF" authority="(Maran, 1942)" baseAuthorityName="Maran" baseAuthorityYear="1942" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia">
|
||
dubia (
|
||
<bibRefCitation id="EF8CBBC51135FFE3D925DBDD2AF1FCAF" author="Maran, J." pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan, 1942</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, J.
|
||
<bibRefCitation id="EF8CBBC51135FFE3DC90DBF12B80FCAF" author="Moravec, J." box="[255,418,747,773]" pageId="9" pageNumber="160" pagination="167 - 186" refId="ref15377" refString="Moravec, J. (1998) Revision within the genus Physodeutera - neglected species described by Maran and related taxa (Coleop- tera: Cicindelidae). Folia Heyrovskyana, 6, 167 - 186." type="journal article" year="1998">Moravec 1998</bibRefCitation>
|
||
det.” and “
|
||
<taxonomicName id="4C1DBDB71135FFE3DE71DBF12E39FCAF" authority="(Maran, 1942)" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[542,1051,747,773]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
Physodeutera (Toxoma) dubia (
|
||
<bibRefCitation id="EF8CBBC51135FFE3DFEEDBF12E31FCAF" author="Maran, J." box="[897,1043,747,773]" pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan, 1942</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
det. J. Moravec, 2001” [printed].
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341135FFE3DCA8DA15292FFB3B" blockId="9.[151,1437,530,1997]" pageId="9" pageNumber="160">
|
||
<emphasis id="B9691A261135FFE3DCA8DA152BFAFC82" bold="true" box="[199,472,782,809]" pageId="9" pageNumber="160">Note to the typification</emphasis>
|
||
.
|
||
<bibRefCitation id="EF8CBBC51135FFE3DD8ADA15285DFC83" author="Maran, J." box="[485,639,783,809]" pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan (1942)</bibRefCitation>
|
||
based the original description of this species on
|
||
<specimenCount id="9D1B0DBD1135FFE3D8F5DA152F2DFC83" box="[1178,1295,783,809]" pageId="9" pageNumber="160" type="male">two males</specimenCount>
|
||
, each from a different locality, both deposited in
|
||
<collectionCode id="ED0C5EF11135FFE3DE49DA29285BFCE7" box="[550,633,819,845]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="160">NMPC</collectionCode>
|
||
. He stated it as: “
|
||
<emphasis id="B9691A261135FFE3DF52DA292873FCDA" italics="true" pageId="9" pageNumber="160">
|
||
Habitat:
|
||
<collectingCountry id="F30A86A41135FFE3DFCDDA292E0DFCE7" box="[930,1071,819,845]" name="Madagascar" pageId="9" pageNumber="160">Madagascar</collectingCountry>
|
||
, Ampanefena
|
||
<specimenCount id="9D1B0DBD1135FFE3D8A2DA2E2EC0FCE5" box="[1229,1250,820,847]" pageId="9" pageNumber="160" type="male">♂</specimenCount>
|
||
typus; Vohémar,
|
||
<date id="FFA3E0F41135FFE3DCF8DA4D2AD6FCDB" box="[151,244,855,881]" pageId="9" pageNumber="160" value="1934-03">III 1934</date>
|
||
, Lamberton leg.
|
||
<specimenCount id="9D1B0DBD1135FFE3DDC2DA4D2BF1FCD9" box="[429,467,855,883]" pageId="9" pageNumber="160" type="male">1 ♂</specimenCount>
|
||
—
|
||
<specimenCount id="9D1B0DBD1135FFE3DD9FDA422821FCD9" box="[496,515,856,883]" pageId="9" pageNumber="160" type="female">♀</specimenCount>
|
||
ignota
|
||
</emphasis>
|
||
”. It means that the male from Ampanefena was designated by Mařan as type and in accordance labelled by him with the red label “Typus” (see
|
||
<figureCitation id="1326DAB11135FFE3DF19DA6129E6FC3F" box="[886,964,891,917]" captionStart="FIGURES 44–45" captionStartId="12.[151,264,1018,1042]" captionTargetBox="[164,1425,191,978]" captionTargetId="figure-31@12.[151,1435,181,993]" captionTargetPageId="12" captionText="FIGURES 44–45. Physodeutera (Toxoma) dubia Mařan, all in SDEI, labels. 44—HT; 45—PT." figureDoi="http://doi.org/10.5281/zenodo.5627225" httpUri="https://zenodo.org/record/5627225/files/figure.png" pageId="9" pageNumber="160">Fig. 44</figureCitation>
|
||
here), while the male from Vohémar bears the same red printed label but rewritten by hand as “CoType” (see
|
||
<figureCitation id="1326DAB11135FFE3DF17DA8529E4FC13" box="[888,966,926,953]" captionStart="FIGURES 44–45" captionStartId="12.[151,264,1018,1042]" captionTargetBox="[164,1425,191,978]" captionTargetId="figure-31@12.[151,1435,181,993]" captionTargetPageId="12" captionText="FIGURES 44–45. Physodeutera (Toxoma) dubia Mařan, all in SDEI, labels. 44—HT; 45—PT." figureDoi="http://doi.org/10.5281/zenodo.5627225" httpUri="https://zenodo.org/record/5627225/files/figure.png" pageId="9" pageNumber="160">Fig. 45</figureCitation>
|
||
here). This statement was previously misinterpreted by the first author (
|
||
<bibRefCitation id="EF8CBBC51135FFE3DD9DDAD928BAFC77" author="Moravec, J." box="[498,664,963,989]" pageId="9" pageNumber="160" pagination="167 - 186" refId="ref15377" refString="Moravec, J. (1998) Revision within the genus Physodeutera - neglected species described by Maran and related taxa (Coleop- tera: Cicindelidae). Folia Heyrovskyana, 6, 167 - 186." type="journal article" year="1998">Moravec 1998</bibRefCitation>
|
||
) who examined the male from Vohémar and erroneously designated and illustrated it as
|
||
<typeStatus id="54A678961135FFE3DD16DAFD2BFDFBAB" box="[377,479,999,1025]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
. This error was later rectified in the monograph of the genus (
|
||
<bibRefCitation id="EF8CBBC51135FFE3D8C8DAFD2FB6FBAB" author="Moravec, J." box="[1191,1428,998,1025]" pageId="9" pageNumber="160" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a: 115</bibRefCitation>
|
||
) where the male from Ampanefena was correctly designated as the
|
||
<typeStatus id="54A678961135FFE3DFEDDD1129C6FB8F" box="[898,996,1035,1061]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
according to the original type designation by
|
||
<bibRefCitation id="EF8CBBC51135FFE3DC85DD352BA4FBE3" author="Maran, J." box="[234,390,1071,1097]" pageId="9" pageNumber="160" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan (1942)</bibRefCitation>
|
||
. Nonetheless, the illustrations remained in error the same, thus the picture of the
|
||
<typeStatus id="54A678961135FFE3D96FDD352F42FBE3" box="[1280,1376,1071,1097]" pageId="9" pageNumber="160" type="paratype">paratype</typeStatus>
|
||
from Vohémar was presented as
|
||
<typeStatus id="54A678961135FFE3DDA4DD49280FFBC7" box="[459,557,1107,1133]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="EF8CBBC51135FFE3DE52DD4928D3FBC7" author="Moravec, J." box="[573,753,1107,1133]" pageId="9" pageNumber="160" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
|
||
, figs 304–308, 723). The true
|
||
<typeStatus id="54A678961135FFE3D824DD492E8FFBC7" box="[1099,1197,1107,1133]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
from Ampanefena is illustrated here in
|
||
<figureCitation id="1326DAB11135FFE3DD0DDD6D2817FB3B" box="[354,565,1143,1169]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="9" pageNumber="160">Figs 24, 26, 28, 31</figureCitation>
|
||
,
|
||
<figureCitation id="1326DAB11135FFE3DE2EDD6D292BFB3B" box="[577,777,1142,1169]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">33, 35, 37, 40–41</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30795BF1135FFE3DCA8DD802ED0F8B3" pageId="9" pageNumber="160" type="diagnosis">
|
||
<paragraph id="8BA2C6341135FFE3DCA8DD802F64FA27" blockId="9.[151,1437,530,1997]" pageId="9" pageNumber="160">
|
||
<emphasis id="B9691A261135FFE3DCA8DD802BE4FB1E" bold="true" box="[199,454,1178,1204]" pageId="9" pageNumber="160">Differential diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C1DBDB71135FFE3DDBBDD812905FB1F" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[468,807,1179,1205]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3DDBBDD81284CFB1F" box="[468,622,1179,1205]" italics="true" pageId="9" pageNumber="160">Physodeutera</emphasis>
|
||
(
|
||
<emphasis id="B9691A261135FFE3DE10DD8128F5FB1E" box="[639,727,1179,1204]" italics="true" pageId="9" pageNumber="160">Toxoma</emphasis>
|
||
)
|
||
<emphasis id="B9691A261135FFE3DE88DD812905FB1F" box="[743,807,1179,1205]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
|
||
</taxonomicName>
|
||
shares similar, rather uniformly bright green-blue elytra with only lateral areas violaceous (
|
||
<figureCitation id="1326DAB11135FFE3DE4BDDA52885FB73" box="[548,679,1214,1241]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="9" pageNumber="160">Figs 24–25</figureCitation>
|
||
,
|
||
<figureCitation id="1326DAB11135FFE3DEDBDDA528DEFB73" box="[692,764,1215,1241]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">40–43</figureCitation>
|
||
) with
|
||
<taxonomicName id="4C1DBDB71135FFE3DF29DDA52E3EFB73" authority="Moravec, 2021" authorityName="Moravec" authorityYear="2021" box="[838,1052,1215,1241]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="lokobensis" status="sp. nov." subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3DF29DDA52947FB73" box="[838,869,1215,1241]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3DF11DDA529ACFB72" box="[894,910,1215,1240]" italics="true" pageId="9" pageNumber="160">T</emphasis>
|
||
.)
|
||
<emphasis id="B9691A261135FFE3DFC9DDA52E3EFB73" box="[934,1052,1215,1241]" italics="true" pageId="9" pageNumber="160">lokobensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9691A261135FFE3D84ADDA52EA2FB73" bold="true" box="[1061,1152,1215,1241]" pageId="9" pageNumber="160">
|
||
<taxonomicNameLabel id="A25AA75D1135FFE3D84ADDA52EA2FB73" box="[1061,1152,1215,1241]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Diagnostic for both species are also the ivory to ochre-testaceous two basal thirds of the antennomere
|
||
<quantity id="4CE56BD11135FFE3D862DDF92E1BFB57" box="[1037,1081,1251,1277]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="160" unit="in" value="4.0">4 in</quantity>
|
||
male (
|
||
<figureCitation id="1326DAB11135FFE3D8EADDF92F2BFB54" box="[1157,1289,1251,1278]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="9" pageNumber="160">Figs 26–27</figureCitation>
|
||
), but
|
||
<taxonomicName id="4C1DBDB71135FFE3D927DDF92AF5FA8B" baseAuthorityName="Maran" baseAuthorityYear="1942" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3D927DDF92F45FB57" box="[1352,1383,1251,1277]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3D911DDF92FB6FB56" box="[1406,1428,1251,1276]" italics="true" pageId="9" pageNumber="160">T.</emphasis>
|
||
)
|
||
<emphasis id="B9691A261135FFE3DCF8DC1D2AF5FA8B" box="[151,215,1287,1313]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
|
||
</taxonomicName>
|
||
differs from the new species in having shorter antennae, more arcuate-curved labial palpus in male (
|
||
<figureCitation id="1326DAB11135FFE3D953DC1D2FADFA8B" box="[1340,1423,1287,1313]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="9" pageNumber="160">Fig. 30</figureCitation>
|
||
), coarser, sharpened rugae on whole surface of the pronotal disc (
|
||
<figureCitation id="1326DAB11135FFE3DF0EDC3129C1FAEF" box="[865,995,1323,1349]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="9" pageNumber="160">Figs 31–32</figureCitation>
|
||
), larger elytral punctures also on posterior elytral half (
|
||
<figureCitation id="1326DAB11135FFE3DD3EDC552BF6FAC3" box="[337,468,1359,1385]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">Figs 40–43</figureCitation>
|
||
), shorter antennae, slightly longer aedeagus apex, which is more rounded, particularly ventrally and on its top (
|
||
<figureCitation id="1326DAB11135FFE3DDC7DC692809FA27" box="[424,555,1395,1421]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">Figs 37–39</figureCitation>
|
||
), and somewhat different sclerites within the internal sac (
|
||
<figureCitation id="1326DAB11135FFE3D8D8DC692F18FA27" box="[1207,1338,1395,1421]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">Figs 38–39</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341135FFE3DCA8DC8D2B1CFA7F" blockId="9.[151,1437,530,1997]" pageId="9" pageNumber="160">
|
||
Moreover, the elytral apex in male is rounded in
|
||
<taxonomicName id="4C1DBDB71135FFE3DE9DDC8D29ADFA1B" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[754,911,1431,1457]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3DE9DDC8D2933FA1B" box="[754,785,1431,1457]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3DF46DC8D291CFA1A" box="[809,830,1431,1456]" italics="true" pageId="9" pageNumber="160">T.</emphasis>
|
||
)
|
||
<emphasis id="B9691A261135FFE3DF20DC8D29ADFA1B" box="[847,911,1431,1457]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1326DAB11135FFE3DFF1DC8D2E03FA1B" box="[926,1057,1431,1457]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">Figs 40–42</figureCitation>
|
||
), while subacute in
|
||
<taxonomicName id="4C1DBDB71135FFE3D96BDC8D2AFFFA7F" authority="Moravec, 2021" authorityName="Moravec" authorityYear="2021" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="lokobensis" status="sp. nov." subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3D96BDC8D2F01FA1B" box="[1284,1315,1431,1457]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3D954DC8D2F69FA1A" box="[1339,1355,1431,1456]" italics="true" pageId="9" pageNumber="160">T</emphasis>
|
||
.)
|
||
<emphasis id="B9691A261135FFE3D90CDC8D2AFFFA7F" italics="true" pageId="9" pageNumber="160">lokobensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9691A261135FFE3DC8BDCA12B1CFA7F" bold="true" box="[228,318,1467,1493]" pageId="9" pageNumber="160">
|
||
<taxonomicNameLabel id="A25AA75D1135FFE3DC8BDCA12B1CFA7F" box="[228,318,1467,1493]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341135FFE3DCA8DCC52ED0F8B3" blockId="9.[151,1437,530,1997]" pageId="9" pageNumber="160">
|
||
Nevertheless, one of the characters considered diagnostic for
|
||
<taxonomicName id="4C1DBDB71135FFE3DFECDCC52E03FA53" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[899,1057,1503,1529]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3DFECDCC52980FA53" box="[899,930,1503,1529]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3DFD5DCC529F2FA52" box="[954,976,1503,1528]" italics="true" pageId="9" pageNumber="160">T.</emphasis>
|
||
)
|
||
<emphasis id="B9691A261135FFE3DF8EDCC52E03FA53" box="[993,1057,1503,1529]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
|
||
</taxonomicName>
|
||
previously (
|
||
<bibRefCitation id="EF8CBBC51135FFE3D8C0DCC52F44FA53" author="Moravec, J." box="[1199,1382,1503,1529]" pageId="9" pageNumber="160" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
|
||
), the anteriad-prolonged, triangular median tooth in the anteromedian lobe of the labrum (
|
||
<figureCitation id="1326DAB11135FFE3D820DF192E81F9B4" box="[1103,1187,1539,1566]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">Fig. 36</figureCitation>
|
||
), proved to be present only in the
|
||
<typeStatus id="54A678961135FFE3DD73DF3D2B5EF9EB" box="[284,380,1575,1601]" pageId="9" pageNumber="160" type="paratype">paratype</typeStatus>
|
||
(which was misinterpreted previously as the
|
||
<typeStatus id="54A678961135FFE3DFE8DF3D29CCF9E8" box="[903,1006,1575,1602]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
). In contrast, the true
|
||
<typeStatus id="54A678961135FFE3D884DF3D2F6FF9EB" box="[1259,1357,1575,1601]" pageId="9" pageNumber="160" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1DBDB71135FFE3D919DF3D2B24F9CF" baseAuthorityName="Maran" baseAuthorityYear="1942" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3D919DF3D2FB7F9EB" box="[1398,1429,1575,1601]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3DCCEDF512A94F9CE" box="[161,182,1611,1636]" italics="true" pageId="9" pageNumber="160">T.</emphasis>
|
||
)
|
||
<emphasis id="B9691A261135FFE3DCA9DF512B24F9CF" box="[198,262,1611,1637]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
|
||
</taxonomicName>
|
||
has the anterior margin of its labrum almost subtruncate (
|
||
<figureCitation id="1326DAB11135FFE3DFF6DF5129CFF9CF" box="[921,1005,1610,1637]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">Fig. 35</figureCitation>
|
||
), thus somewhat similar to that in
|
||
<taxonomicName id="4C1DBDB71135FFE3D919DF512B1DF923" authority="Moravec, 2021" authorityName="Moravec" authorityYear="2021" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="lokobensis" status="sp. nov." subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3D919DF512FB7F9CF" box="[1398,1429,1611,1637]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3DCCEDF752A93F922" box="[161,177,1647,1672]" italics="true" pageId="9" pageNumber="160">T</emphasis>
|
||
.)
|
||
<emphasis id="B9691A261135FFE3DCA6DF752B1DF923" box="[201,319,1647,1673]" italics="true" pageId="9" pageNumber="160">lokobensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9691A261135FFE3DD27DF752B81F923" bold="true" box="[328,419,1647,1673]" pageId="9" pageNumber="160">
|
||
<taxonomicNameLabel id="A25AA75D1135FFE3DD27DF752B81F923" box="[328,419,1647,1673]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Despite the significant biodiversity in
|
||
<collectingCountry id="F30A86A41135FFE3DF0FDF7529CBF923" box="[864,1001,1647,1673]" name="Madagascar" pageId="9" pageNumber="160">Madagascar</collectingCountry>
|
||
, we consider here these differences in the shape of the male labrum to be exceptional variability, also due to the fact that the two only known males come very probably from the same locality (see “Distribution” below). Notwithstanding, as also the body of the
|
||
<typeStatus id="54A678961135FFE3D953DFAD2FBEF97B" box="[1340,1436,1719,1745]" pageId="9" pageNumber="160" type="paratype">paratype</typeStatus>
|
||
is notably wider, particularly across the elytra (
|
||
<figureCitation id="1326DAB11135FFE3DECDDFC128D7F95F" box="[674,757,1754,1781]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="9" pageNumber="160">Fig. 25</figureCitation>
|
||
), a possibility that the
|
||
<typeStatus id="54A678961135FFE3DF9EDFC12E73F95F" box="[1009,1105,1755,1781]" pageId="9" pageNumber="160" type="paratype">paratype</typeStatus>
|
||
may in fact represent another undescribed species cannot be excluded. Unfortunately, female of
|
||
<taxonomicName id="4C1DBDB71135FFE3DF11DFE52E3EF8B3" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[894,1052,1791,1817]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261135FFE3DF11DFE529BFF8B3" box="[894,925,1791,1817]" italics="true" pageId="9" pageNumber="160">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261135FFE3DFDADFE529E7F8B2" box="[949,965,1791,1816]" italics="true" pageId="9" pageNumber="160">T</emphasis>
|
||
.)
|
||
<emphasis id="B9691A261135FFE3DFB3DFE52E3EF8B3" box="[988,1052,1791,1817]" italics="true" pageId="9" pageNumber="160">dubia</emphasis>
|
||
</taxonomicName>
|
||
remains unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30795BF1135FFE7DCA8DE382E8EFCBF" lastPageId="13" lastPageNumber="164" pageId="9" pageNumber="160" type="description">
|
||
<paragraph id="8BA2C6341135FFE3DCA8DE382B63F8CB" blockId="9.[151,1437,530,1997]" pageId="9" pageNumber="160">
|
||
<emphasis id="B9691A261135FFE3DCA8DE382B4FF896" bold="true" box="[199,365,1826,1852]" pageId="9" pageNumber="160">Redescription</emphasis>
|
||
(male). Body (
|
||
<figureCitation id="1326DAB11135FFE3DE70DE392880F897" box="[543,674,1826,1853]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="9" pageNumber="160">Figs 24–25</figureCitation>
|
||
) medium-sized, length 8.80–9.00 (HT 8.80) mm, width 2.70–2.90 (HT 2.70) mm.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341135FFE3DCA8DE712EE1F82F" blockId="9.[151,1437,530,1997]" box="[199,1219,1899,1925]" pageId="9" pageNumber="160">
|
||
Head (
|
||
<figureCitation id="1326DAB11135FFE3DD7EDE712BB0F82F" box="[273,402,1899,1925]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="9" pageNumber="160">Figs 33–34</figureCitation>
|
||
) with large eyes, only slightly narrower than body, width
|
||
<quantity id="4CE56BD11135FFE3D876DE712E9DF82F" box="[1049,1215,1899,1925]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="9" pageNumber="160" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.60–2.70 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341135FFE3DCA8DE952FBFF867" blockId="9.[151,1437,530,1997]" pageId="9" pageNumber="160">Frons deep metallic blue, convex in middle, nearly smooth on anteromedian area, laterally parallel-wrinkled; supraantennal plates uneven, flat; frons-vertex fold rounded merging fluently with vertex, finely vermicular-wrinkled.</paragraph>
|
||
<caption id="DF6296BC1136FFE0DCF8DE512F11F801" ID-DOI="http://doi.org/10.5281/zenodo.5627221" ID-Zenodo-Dep="5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,264,1867,1891]" targetBox="[157,1431,195,1835]" targetPageId="10">
|
||
<paragraph id="8BA2C6341136FFE0DCF8DE512F11F801" blockId="10.[151,1436,1867,1964]" pageId="10" pageNumber="161">
|
||
<emphasis id="B9691A261136FFE0DCF8DE512B77F8CE" bold="true" box="[151,341,1867,1892]" pageId="10" pageNumber="161">FIGURES 24–32.</emphasis>
|
||
<taxonomicName id="4C1DBDB71136FFE0DD33DE5628F9F8CE" authority="Maran" authorityName="Maran" box="[348,731,1868,1892]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261136FFE0DD33DE562BC5F8CE" box="[348,487,1868,1892]" italics="true" pageId="10" pageNumber="161">Physodeutera</emphasis>
|
||
(
|
||
<emphasis id="B9691A261136FFE0DD99DE562867F8C9" box="[502,581,1868,1891]" italics="true" pageId="10" pageNumber="161">Toxoma</emphasis>
|
||
)
|
||
<emphasis id="B9691A261136FFE0DE3ADE5628ADF8CE" box="[597,655,1868,1892]" italics="true" pageId="10" pageNumber="161">dubia</emphasis>
|
||
Mařan
|
||
</taxonomicName>
|
||
, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vohémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from
|
||
<bibRefCitation id="EF8CBBC51136FFE0DE5EDE8E28F1F806" author="Moravec, J." box="[561,723,1940,1964]" pageId="10" pageNumber="161" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
|
||
); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6296BC1137FFE1DCF8DE512BC9F865" ID-DOI="http://doi.org/10.5281/zenodo.5627223" ID-Zenodo-Dep="5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,264,1867,1891]" targetBox="[158,1429,188,1839]" targetPageId="11">
|
||
<paragraph id="8BA2C6341137FFE1DCF8DE512BC9F865" blockId="11.[151,1437,1867,2000]" pageId="11" pageNumber="162">
|
||
<emphasis id="B9691A261137FFE1DCF8DE512B77F8CE" bold="true" box="[151,341,1867,1892]" pageId="11" pageNumber="162">FIGURES 33–43.</emphasis>
|
||
<taxonomicName id="4C1DBDB71137FFE1DD33DE5628FEF8CE" authority="Maran" authorityName="Maran" box="[348,732,1868,1892]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261137FFE1DD33DE562BC5F8CE" box="[348,487,1868,1892]" italics="true" pageId="11" pageNumber="162">Physodeutera</emphasis>
|
||
(
|
||
<emphasis id="B9691A261137FFE1DD98DE562864F8C9" box="[503,582,1868,1891]" italics="true" pageId="11" pageNumber="162">Toxoma</emphasis>
|
||
)
|
||
<emphasis id="B9691A261137FFE1DE39DE5628B2F8CE" box="[598,656,1868,1892]" italics="true" pageId="11" pageNumber="162">dubia</emphasis>
|
||
Mařan
|
||
</taxonomicName>
|
||
, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohémar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6296BC1130FFE6DCF8DAE02E7CFBB8" ID-DOI="http://doi.org/10.5281/zenodo.5627225" ID-Zenodo-Dep="5627225" box="[151,1118,1018,1042]" httpUri="https://zenodo.org/record/5627225/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,264,1018,1042]" targetBox="[164,1425,191,978]" targetPageId="12">
|
||
<paragraph id="8BA2C6341130FFE6DCF8DAE02E7CFBB8" blockId="12.[151,1118,1018,1042]" box="[151,1118,1018,1042]" pageId="12" pageNumber="163">
|
||
<emphasis id="B9691A261130FFE6DCF8DAE02B76FBB8" bold="true" box="[151,340,1018,1042]" pageId="12" pageNumber="163">FIGURES 44–45.</emphasis>
|
||
<taxonomicName id="4C1DBDB71130FFE6DD35DAE028F5FBB8" authority="Maran" authorityName="Maran" box="[346,727,1018,1042]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261130FFE6DD35DAE02BC7FBB8" box="[346,485,1018,1042]" italics="true" pageId="12" pageNumber="163">Physodeutera</emphasis>
|
||
(
|
||
<emphasis id="B9691A261130FFE6DD9BDAE12861FBB8" box="[500,579,1019,1042]" italics="true" pageId="12" pageNumber="163">Toxoma</emphasis>
|
||
)
|
||
<emphasis id="B9691A261130FFE6DE3DDAE028AEFBB8" box="[594,652,1018,1042]" italics="true" pageId="12" pageNumber="163">dubia</emphasis>
|
||
Mařan
|
||
</taxonomicName>
|
||
, all in SDEI, labels. 44—HT; 45—PT.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA2C6341130FFE6DCA8DD5B2FB5FB6F" blockId="12.[151,1437,1088,2039]" pageId="12" pageNumber="163">Vertex bright blue in middle, with bronze and cupreous lustre in lateral and posterior areas, with deep transverse anterior impression crossing juxtaorbital areas, which are striate-rugose, but rugae interrupted by two shallower impressions; median area longitudinally vermicular to wavy-rugose, posterior area vermicular-rugose; divergent sublateral rugae coarser, wavy, passing onto temples; posterior and occipital areas bronze-green, vermicular-rugulose.</paragraph>
|
||
<paragraph id="8BA2C6341130FFE6DCA8DDD528DCFB42" blockId="12.[151,1437,1088,2039]" box="[199,766,1230,1257]" pageId="12" pageNumber="163">Genae metallic green-blue, finely striate-wrinkled.</paragraph>
|
||
<paragraph id="8BA2C6341130FFE6DCA8DDE82BB8FA31" blockId="12.[151,1437,1088,2039]" pageId="12" pageNumber="163">
|
||
Labrum (male)
|
||
<quantity id="4CE56BD11130FFE6DD11DDE82806FAA6" box="[382,548,1266,1293]" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="10.5" metricValueMin="9.0" pageId="12" pageNumber="163" unit="mm" value="0.975" valueMax="1.05" valueMin="0.9">0.90–1.05 mm</quantity>
|
||
long,
|
||
<quantity id="4CE56BD11130FFE6DE1DDDE828FBFAA6" box="[626,729,1266,1292]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="163" unit="mm" value="1.1">1.10 mm</quantity>
|
||
wide, basically shaped and coloured as in preceding species except for the shape of the anteromedian lobe, which differs in the two known males: while the
|
||
<typeStatus id="54A678961130FFE6D88DDC0F2F66FA85" box="[1250,1348,1301,1327]" pageId="12" pageNumber="163" type="holotype">holotype</typeStatus>
|
||
has the anteromedian lobe notably shorter, subtruncate and with rounded lateral margins (
|
||
<figureCitation id="1326DAB11130FFE6D84DDC232E57FAFE" box="[1058,1141,1337,1364]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="12" pageNumber="163">Fig. 35</figureCitation>
|
||
), the anteromedian lobe of the
|
||
<typeStatus id="54A678961130FFE6DCAFDC472B02FADD" box="[192,288,1373,1399]" pageId="12" pageNumber="163" type="paratype">paratype</typeStatus>
|
||
possesses two right-angled, indistinctly pointed teeth in either side of notably protruding, right-angled median tooth (
|
||
<figureCitation id="1326DAB11130FFE6DD55DC9A2BACFA31" box="[314,398,1408,1435]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="12" pageNumber="163">Fig. 36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341130FFE6DCA8DCB92928FA17" blockId="12.[151,1437,1088,2039]" box="[199,778,1443,1469]" pageId="12" pageNumber="163">Clypeus metallic-blue with greenish lustre, smooth.</paragraph>
|
||
<paragraph id="8BA2C6341130FFE6DCA8DCDD2B70F982" blockId="12.[151,1437,1088,2039]" pageId="12" pageNumber="163">
|
||
Mandibles (
|
||
<figureCitation id="1326DAB11130FFE6DD23DCDD2BEEFA4B" box="[332,460,1479,1505]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="12" pageNumber="163">Figs 33–34</figureCitation>
|
||
) subsymmetrical, normally shaped with four teeth and basal molar, brown with mahogany tinge and blackened margins, inner teeth gradually smaller towards basal molar, fourth tooth markedly distant from third tooth.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341130FFE6DCA8DF2B2ECBF938" blockId="12.[151,1437,1088,2039]" pageId="12" pageNumber="163">
|
||
Palpi (male) ivory testaceous, maxillary palpi with terminal palpomeres brownish-testaceous with darkened subapical area (
|
||
<figureCitation id="1326DAB11130FFE6DD27DF4E2BEEF9C5" box="[328,460,1620,1647]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="12" pageNumber="163">Figs 28–29</figureCitation>
|
||
); labial palpi with dark brown terminal palpomeres, outer margin of penultimate (longest) palpomeres arcuate, inner margin straight or slightly emarginated, ochre-testaceous (
|
||
<figureCitation id="1326DAB11130FFE6D8E7DF622EFEF938" box="[1160,1244,1656,1682]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="12" pageNumber="163">Fig. 30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341130FFE6DCA8DF8629BAF956" blockId="12.[151,1437,1088,2039]" pageId="12" pageNumber="163">
|
||
Antennae (
|
||
<figureCitation id="1326DAB11130FFE6DD2DDF812BE0F91C" box="[322,450,1691,1718]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="12" pageNumber="163">Figs 26–27</figureCitation>
|
||
) rather long, passing elytral half; scape ivory or testaceous with only apical seta; antennomeres 2–3 metallic black, antennomere 4 black-brown with two basal thirds ivory-testaceous to yellowish, antennomeres 5–9 light brownish testaceous, 10–11 moderately darkened.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341130FFE7DCA8DE1C28BDFF7F" blockId="12.[151,1437,1088,2039]" lastBlockId="13.[151,1437,151,1185]" lastPageId="13" lastPageNumber="164" pageId="12" pageNumber="163">
|
||
Thorax. Pronotum (
|
||
<figureCitation id="1326DAB11130FFE6DDCDDE1C2803F88A" box="[418,545,1798,1824]" captionStart="FIGURES 24–32" captionStartId="10.[151,264,1867,1891]" captionTargetBox="[157,1431,195,1835]" captionTargetId="figure-31@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURES 24–32. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 24–25—habitus: 24—♂, Ampanefena, HT; 25—♂, Vo- hémar, PT; 26—Antennomeres 1–5, ♂, HT; 27—Antennomeres 1–6, ♂, PT; 28–29—maxillary palpus: 28—♂, HT; 29—♂, PT; 30—labial palpus, ♂, PT (adopted from Moravec 2002a); 31–32—pronotum: 31—♂, HT; 32—♂, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627221" httpUri="https://zenodo.org/record/5627221/files/figure.png" pageId="12" pageNumber="163">Figs 31–32</figureCitation>
|
||
) elongate, length
|
||
<quantity id="4CE56BD11130FFE6DE88DE1C29AEF88A" box="[743,908,1798,1825]" metricMagnitude="-3" metricUnit="m" metricValue="1.875" metricValueMax="1.9" metricValueMin="1.85" pageId="12" pageNumber="163" unit="mm" value="1.875" valueMax="1.9" valueMin="1.85">1.85–1.90 mm</quantity>
|
||
, width
|
||
<quantity id="4CE56BD11130FFE6DFB3DE1C2EA0F88A" box="[988,1154,1798,1825]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.55" metricValueMin="1.5" pageId="12" pageNumber="163" unit="mm" value="1.525" valueMax="1.55" valueMin="1.5">1.50–1.55 mm</quantity>
|
||
; metallic blue-green with indistinct bronze-cupreous and violet lustre on discal median area and intense violaceous lustre within well-pronounced anterior and posterior sulci; anterior lobe markedly narrower than posterior lobe and disc; lateral margins of disc almost parallel in middle, slightly constricted anteriad; notopleural sutures barely visible from above; discal surface distinctly transversely wavy-rugose, rugae coarser on median area; median line thin but running throughout; posterior lobe with distinct basal rim, dorsolateral bulges pulvinate-elevated, almost smooth, with strongly violaceous lustre, gradually passing to smooth or finely wrinkled median area with faint greenish lustre; prosternum and proepisterna metallic blue-green, finely wrinkled; mesepisterna black-blue with purple lustre; metasternum and metepisterna metallic green with bronze tinge.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341131FFE7DCA8D9C52822FDB3" blockId="13.[151,1437,151,1185]" pageId="13" pageNumber="164">
|
||
Elytra (
|
||
<figureCitation id="1326DAB11131FFE7DD74D9C52BBCFF53" box="[283,414,223,249]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="13" pageNumber="164">Figs 40–43</figureCitation>
|
||
), elongate,
|
||
<quantity id="4CE56BD11131FFE7DE71D9C428E3FF53" box="[542,705,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="5.324999999999999" metricValueMax="5.35" metricValueMin="5.3" pageId="13" pageNumber="164" unit="mm" value="5.324999999999999" valueMax="5.35" valueMin="5.3">5.30–5.35 mm</quantity>
|
||
long; juxtahumeral impressions deep, short and wide, basodiscal convexity and discal impression distinct, apical impressions moderate; lateral margins subparallel, anteapical angles arcuate, apices rounded, more distinctly so in female, with only faint emargination towards indistinct, blunt sutural spine; limited baso-humeral area smooth and shiny, other elytral surface rather coarsely punctate, punctures notably larger on anterior elytral third, largest within humeral impressions, occasionally arranged in chains but not entirely anastomosing, their mostly narrow intervals optically forming irregular reticulum; elytral coloration bright metallic blue-green, more cyaneous green-blue within humeral and discal impression, with faint bronze-greenish lustre on posterior declivity and deep violaceous on outer lateral areas and apices; whitish to ivory maculation consisting of only one, large humeral macula.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341131FFE7DCA8DB3929F8FDC8" blockId="13.[151,1437,151,1185]" pageId="13" pageNumber="164">Abdomen. Abdominal ventrites dark metallic-blue with violaceous reflections, their surface smooth and glabrous (except for sensory seta on either side of posterior ventrite margins).</paragraph>
|
||
<paragraph id="8BA2C6341131FFE7DCA8DB712E69FD2F" blockId="13.[151,1437,151,1185]" box="[199,1099,619,645]" pageId="13" pageNumber="164">
|
||
Legs of HT similarly coloured as in
|
||
<emphasis id="B9691A261131FFE7DE32DB71285EFD2F" box="[605,636,619,645]" italics="true" pageId="13" pageNumber="164">Ph</emphasis>
|
||
.
|
||
<taxonomicName id="4C1DBDB71131FFE7DEE5DB712914FD2F" box="[650,822,619,645]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
|
||
(
|
||
<emphasis id="B9691A261131FFE7DEFBDB712886FD2E" box="[660,676,619,644]" italics="true" pageId="13" pageNumber="164">T</emphasis>
|
||
.)
|
||
<emphasis id="B9691A261131FFE7DED4DB712914FD2F" box="[699,822,619,645]" italics="true" pageId="13" pageNumber="164">conturbata</emphasis>
|
||
</taxonomicName>
|
||
, but notably paler in PT.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341131FFE7DCA8DB952E8EFCBF" blockId="13.[151,1437,151,1185]" pageId="13" pageNumber="164">
|
||
Aedeagus short (
|
||
<figureCitation id="1326DAB11131FFE7DDE7DB95282EFD03" box="[392,524,655,681]" captionStart="FIGURES 33–43" captionStartId="11.[151,264,1867,1891]" captionTargetBox="[158,1429,188,1839]" captionTargetId="figure-17@11.[151,1435,181,1844]" captionTargetPageId="11" captionText="FIGURES 33–43. Physodeutera (Toxoma) dubia Mařan, all in SDEI. 33–34—head: 33—♂, Ampanefena, HT; 34—♂, Vohé- mar, PT; 35–36—labrum: 35—♂, HT; 36—♂, PT; 37–39—male aedeagus: 37—HT; 38–39—showing internal sac in left and right lateral view, PT; 40–43—elytron: 40—♂, HT; 41—ditto, detail of elytral disc; 42—♂, PT; 43— ditto, detail of posterior area of elytral disc. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627223" httpUri="https://zenodo.org/record/5627223/files/figure.png" pageId="13" pageNumber="164">Figs 37–39</figureCitation>
|
||
), length
|
||
<quantity id="4CE56BD11131FFE7DE1EDB94293AFD03" box="[625,792,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.225" metricValueMax="2.25" metricValueMin="2.2" pageId="13" pageNumber="164" unit="mm" value="2.225" valueMax="2.25" valueMin="2.2">2.20–2.25 mm</quantity>
|
||
, width
|
||
<quantity id="4CE56BD11131FFE7DF01DB9429F4FD03" box="[878,982,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="164" unit="mm" value="0.45">0.45 mm</quantity>
|
||
, ventral outline straight or slightly bent ventrad apically, dorsally conically attenuated towards narrow, rounded apex; sclerites within internal sac consisting of distinct arciform piece with dilates apex, thin satellite piece which appears more distinct in right lateral view, membranous plate and large, longitudinal membranous upper-ventral piece with obtuse apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30795BF1131FFE7DCA8DA042E36FC63" pageId="13" pageNumber="164" type="distribution">
|
||
<paragraph id="8BA2C6341131FFE7DCA8DA042E36FC63" blockId="13.[151,1437,151,1185]" pageId="13" pageNumber="164">
|
||
<emphasis id="B9691A261131FFE7DCA8DA042B7BFC92" bold="true" box="[199,345,798,824]" pageId="13" pageNumber="164">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1326DAB11131FFE7DD07DA052BE8FC90" box="[360,458,799,826]" captionStart="FIGURE 106" captionStartId="27.[151,250,1561,1585]" captionTargetBox="[271,1319,572,1531]" captionTargetId="figure-167@27.[262,1326,561,1538]" captionTargetPageId="27" captionText="FIGURE 106. Distribution of four Physodeutera (Toxoma) species in northern part of Madagascar. Adopted from the map of preserved areas in Madagascar according to Moat & Smith (2007), adapted and published previously (Moravec 2010) under permission of the trustee of the Royal Botanic Gardens, Kew, GB." figureDoi="http://doi.org/10.5281/zenodo.5627247" httpUri="https://zenodo.org/record/5627247/files/figure.png" pageId="13" pageNumber="164">Fig. 106</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1DBDB71131FFE7DDB2DA05290DFC93" box="[477,815,799,825]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261131FFE7DDB2DA052855FC93" box="[477,631,799,825]" italics="true" pageId="13" pageNumber="164">Physodeutera</emphasis>
|
||
(
|
||
<emphasis id="B9691A261131FFE7DEE8DA0528FDFC92" box="[647,735,799,824]" italics="true" pageId="13" pageNumber="164">Toxoma</emphasis>
|
||
)
|
||
<emphasis id="B9691A261131FFE7DE80DA05290DFC93" box="[751,815,799,825]" italics="true" pageId="13" pageNumber="164">dubia</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<specimenCount id="9D1B0DBD1131FFE7D82FDA052E93FC93" box="[1088,1201,799,825]" pageId="13" pageNumber="164" type="male">two males</specimenCount>
|
||
caught in Northeastern
|
||
<collectingCountry id="F30A86A41131FFE7DCADDA592B69FCF7" box="[194,331,835,861]" name="Madagascar" pageId="13" pageNumber="164">Madagascar</collectingCountry>
|
||
. Ampanefena lies about
|
||
<quantity id="4CE56BD11131FFE7DE30DA592885FCF7" box="[607,679,835,861]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="164" unit="km" value="70.0">70 km</quantity>
|
||
south of the city of Vohemar (also spelled Vohémar). However, the name on the label probably means the whole district, which is very large, and therefore we may suppose that the
|
||
<specimenCount id="9D1B0DBD1131FFE7DCF8DA912B29FC0F" box="[151,267,907,933]" pageId="13" pageNumber="164" type="male">two males</specimenCount>
|
||
come from the same locality (
|
||
<bibRefCitation id="EF8CBBC51131FFE7DE06DA912903FC0F" author="Moravec, J." box="[617,801,907,933]" pageId="13" pageNumber="164" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
|
||
). Although no collector name is indicated on the labels, according to
|
||
<bibRefCitation id="EF8CBBC51131FFE7DD46DAB52BE4FC63" author="Maran, J." box="[297,454,943,969]" pageId="13" pageNumber="164" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan (1942)</bibRefCitation>
|
||
, the
|
||
<specimenCount id="9D1B0DBD1131FFE7DD94DAB5287EFC63" box="[507,604,943,969]" pageId="13" pageNumber="164" type="generic" typeStatus="type">two type</specimenCount>
|
||
specimens were caught by Lamberton.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30795BF1131FFE7DCA8DAC92BF6FB0B" pageId="13" pageNumber="164" type="discussion">
|
||
<paragraph id="8BA2C6341131FFE7DCA8DAC928EDFBF3" blockId="13.[151,1437,151,1185]" pageId="13" pageNumber="164">
|
||
<emphasis id="B9691A261131FFE7DCA8DAC92B16FC47" bold="true" box="[199,308,979,1005]" pageId="13" pageNumber="164">Remarks</emphasis>
|
||
. This taxon, originally described as
|
||
<taxonomicName id="4C1DBDB71131FFE7DE86DAC92E6FFC47" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[745,1101,979,1005]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="dubia" subGenus="Megalomma">
|
||
<emphasis id="B9691A261131FFE7DE86DAC92974FC47" box="[745,854,979,1005]" italics="true" pageId="13" pageNumber="164">Prothyma</emphasis>
|
||
(
|
||
<emphasis id="B9691A261131FFE7DF04DAC929D5FC47" box="[875,1015,979,1005]" italics="true" pageId="13" pageNumber="164">Megalomma</emphasis>
|
||
)
|
||
<emphasis id="B9691A261131FFE7D862DAC92E6FFC47" box="[1037,1101,979,1005]" italics="true" pageId="13" pageNumber="164">dubia</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8CBBC51131FFE7D8EFDAC92F00FC47" author="Maran, J." box="[1152,1314,979,1005]" pageId="13" pageNumber="164" pagination="63 - 73" refId="ref15317" refString="Maran, J. (1942) De novis generis Prothyma Hope speciebus formisque insullae Madagascariensis. Acta Entomologica Musei Nationalis Pragae, 2, 63 - 73." type="journal article" year="1942">Mařan (1942)</bibRefCitation>
|
||
, was later (
|
||
<bibRefCitation id="EF8CBBC51131FFE7DCF0DAED2B67FBBB" author="Moravec, J." box="[159,325,1015,1041]" pageId="13" pageNumber="164" pagination="167 - 186" refId="ref15377" refString="Moravec, J. (1998) Revision within the genus Physodeutera - neglected species described by Maran and related taxa (Coleop- tera: Cicindelidae). Folia Heyrovskyana, 6, 167 - 186." type="journal article" year="1998">Moravec 1998</bibRefCitation>
|
||
) treated as a subspecies of
|
||
<emphasis id="B9691A261131FFE7DE11DAED28BFFBBB" box="[638,669,1015,1041]" italics="true" pageId="13" pageNumber="164">Ph</emphasis>
|
||
.
|
||
<taxonomicName id="4C1DBDB71131FFE7DEC2DAED290EFBBB" baseAuthorityName="W. Horn" baseAuthorityYear="1896" box="[685,812,1015,1041]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="sikorai" subGenus="Toxoma">
|
||
(
|
||
<emphasis id="B9691A261131FFE7DED9DAED28E4FBBA" box="[694,710,1015,1040]" italics="true" pageId="13" pageNumber="164">T</emphasis>
|
||
.)
|
||
<emphasis id="B9691A261131FFE7DEB1DAED290EFBBB" box="[734,812,1015,1041]" italics="true" pageId="13" pageNumber="164">sikorai</emphasis>
|
||
</taxonomicName>
|
||
. However, as further examinations have confirmed the above-mentioned characters, this taxon was treated (
|
||
<bibRefCitation id="EF8CBBC51131FFE7DE8FDD0129B0FB9F" author="Moravec, J." box="[736,914,1051,1077]" pageId="13" pageNumber="164" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
|
||
) as a natural member of the subgenus
|
||
<taxonomicName id="4C1DBDB71131FFE7D951DD012FB4FB9E" authorityName="Rivalier" authorityYear="1967" box="[1342,1430,1051,1076]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="subGenus" subGenus="Toxoma">
|
||
<emphasis id="B9691A261131FFE7D951DD012FB4FB9E" box="[1342,1430,1051,1076]" italics="true" pageId="13" pageNumber="164">Toxoma</emphasis>
|
||
</taxonomicName>
|
||
. Only male specimens of
|
||
<taxonomicName id="4C1DBDB71131FFE7DDC1DD252869FBF3" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[430,587,1087,1113]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
|
||
<emphasis id="B9691A261131FFE7DDC1DD252BEFFBF3" box="[430,461,1087,1113]" italics="true" pageId="13" pageNumber="164">Ph</emphasis>
|
||
. (
|
||
<emphasis id="B9691A261131FFE7DD8BDD252BD6FBF2" box="[484,500,1087,1112]" italics="true" pageId="13" pageNumber="164">T</emphasis>
|
||
.)
|
||
<emphasis id="B9691A261131FFE7DE64DD252869FBF3" box="[523,587,1087,1113]" italics="true" pageId="13" pageNumber="164">dubia</emphasis>
|
||
</taxonomicName>
|
||
are known.
|
||
</paragraph>
|
||
<paragraph id="8BA2C6341131FFE7DCA8DD792BF6FB0B" blockId="13.[151,1437,151,1185]" pageId="13" pageNumber="164">
|
||
For the previous confusion of the
|
||
<typeStatus id="54A678961131FFE7DE24DD792859FBD7" box="[587,635,1123,1149]" pageId="13" pageNumber="164">type</typeStatus>
|
||
designation see “Note to the typification” above and the discussion in “Introduction” of this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |