244 lines
25 KiB
XML
244 lines
25 KiB
XML
<document id="7320BD5076EECEB4A820BC089B2F3388" ID-DOI="10.11646/zootaxa.5284.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7923218" ID-ZooBank="2A865757-B6B4-48CD-A9AD-334F7E7B508B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683785948286" checkinUser="plazi" docAuthor="Anichtchenko, Alexander & Valainis, Uldis" docDate="2023" docId="03B46F27FFD3FF8A6DC3F8C3FB389336" docLanguage="en" docName="zootaxa.5284.2.1.pdf" docOrigin="Zootaxa 5284 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5284.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omophron (Omophron) capense subsp. pumilum Anichtchenko & Valainis 2023, ssp. n." docType="treatment" docVersion="2" lastPageNumber="225" masterDocId="FF8D175FFFC4FF926D54FFFCFF8B900F" masterDocTitle="Revision of the subgenus Omophron (s. str.) Latreille, 1802 (Coleoptera: Carabidae Omophron) of the Afrotropical region" masterLastPageNumber="246" masterPageNumber="201" pageNumber="224" updateTime="1683786577624" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D50A3507356BC8D5F9DC0679DD3F2C2B">Revision of the subgenus Omophron (s. str.) Latreille, 1802 (Coleoptera: Carabidae Omophron) of the Afrotropical region</mods:title>
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<mods:name id="FF1B00F2CEDB99D08030616EA6B9C1AA" type="personal">
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<mods:namePart id="8EB2C6EDA2C001F6EF30BB4242ADC130">Anichtchenko, Alexander</mods:namePart>
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<mods:nameIdentifier id="5432255F64CC14A255CBC033F85CEA4F" type="email">alexander.anishchenko@du.lv</mods:nameIdentifier>
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<mods:namePart id="6C7B40D77CC4463C8D6779589F67EF85">Valainis, Uldis</mods:namePart>
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<mods:title id="FA60065B0F7185829A7566F064C543AD">Zootaxa</mods:title>
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<mods:part id="8BD3C2BD281170502946F26992327EC3">
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<mods:date id="43417BC55762BEF81F095CFC3B5EBECD">2023</mods:date>
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<mods:number id="3655DD919C5D99E4D463634C9F8EED2B">2023-05-11</mods:number>
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<mods:number id="917357F074BF548915DD8F7C6328B886">5284</mods:number>
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<mods:start id="C4DBE7B56460899580AE5E2F31A1DCEA">201</mods:start>
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<mods:url id="B11F8B36343B0ED504219181CB758876">http://dx.doi.org/10.11646/zootaxa.5284.2.1</mods:url>
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<mods:classification id="F890EC439F5F04D971C21327CD4E6528">journal article</mods:classification>
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<mods:identifier id="15808681751F5C6713DC2E548B6D3A46" type="DOI">10.11646/zootaxa.5284.2.1</mods:identifier>
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<mods:identifier id="B97D1B46F1434D5C3AC6F6C682167D97" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B46F27FFD3FF8A6DC3F8C3FB389336" ID-DOI="http://doi.org/10.5281/zenodo.7923270" ID-Zenodo-Dep="7923270" LSID="urn:lsid:plazi:treatment:03B46F27FFD3FF8A6DC3F8C3FB389336" httpUri="http://treatment.plazi.org/id/03B46F27FFD3FF8A6DC3F8C3FB389336" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224">
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<subSubSection id="C3078DBAFFD3FF856DC3F8C3FD609755" box="[151,747,1855,1882]" pageId="23" pageNumber="224" type="nomenclature">
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<paragraph id="8BA2DE31FFD3FF856DC3F8C3FD609755" blockId="23.[151,747,1855,1882]" box="[151,747,1855,1882]" pageId="23" pageNumber="224">
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<heading id="D0EA695DFFD3FF856DC3F8C3FD609755" bold="true" box="[151,747,1855,1882]" fontSize="11" level="1" pageId="23" pageNumber="224" reason="1">
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<emphasis id="B9690223FFD3FF856DC3F8C3FD609755" bold="true" box="[151,747,1855,1882]" pageId="23" pageNumber="224">
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<taxonomicName id="4C1DA5B2FFD3FF856DC3F8C3FD1D9755" authority="Anichtchenko & Valainis, 2023" authorityName="Anichtchenko & Valainis" authorityYear="2023" box="[151,662,1855,1882]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="subSpecies" species="capense" status="ssp. n." subGenus="Omophron" subSpecies="pumilum">
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<emphasis id="B9690223FFD3FF856DC3F8C3FD1D9755" bold="true" box="[151,662,1855,1882]" italics="true" pageId="23" pageNumber="224">Omophron (Omophron) capense pumilum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25ABF58FFD3FF856FC9F8BCFD609755" box="[669,747,1856,1882]" pageId="23" pageNumber="224" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C3078DBAFFD3FF8A6DC3F874FF779112" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224" type="materials_examined">
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<emphasis id="B9690223FFD3FF856DC3F874FECD97AD" bold="true" box="[151,326,1928,1954]" pageId="23" pageNumber="224">Type material.</emphasis>
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<materialsCitation id="3B75D46CFFD3FF8A6C1AF874FF779112" collectingDate="2010-01-13" collectingDateMax="2019-11-21" collectingDateMin="2010-01-13" collectionCode="DUBC, RKC, PSC, BMNH" collectorName="Namibia & Rudolf Kmeco & Zambia & Western & P. Schule & Angola & Kangumbe" country="Namibia" elevation="950" lastPageId="24" lastPageNumber="225" latitude="-12.58449" location="Langue-Bungo River" longLatPrecision="1" longitude="18.66771" municipality="Chissamba Mission" pageId="23" pageNumber="224" specimenCount="956" specimenCount-female="6" stateProvince="Bie" typeStatus="holotype">
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<emphasis id="B9690223FFD3FF856C1AF874FE5C97AD" bold="true" box="[334,471,1928,1954]" pageId="23" pageNumber="224">
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<collectingCountry id="F30A9EA1FFD3FF856C1AF874FE5997AD" box="[334,466,1928,1954]" name="Namibia" pageId="23" pageNumber="224">NAMIBIA</collectingCountry>
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:
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♁
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, Caprivi reg.,
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</quantity>
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.
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</specimenCount>
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,
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<locationDeviation id="9A10C713FFD3FF856E9DF874FB6997AD" box="[969,1250,1928,1955]" location="Katima Mulilo" pageId="23" pageNumber="224">
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<quantity id="4CE573D4FFD3FF856E9DF874FC7197AD" box="[969,1018,1928,1954]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="224" unit="km" value="8.0">8km</quantity>
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E of Katima Mulilo
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</locationDeviation>
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at Zambezi riv.,
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<date id="FFA3F8F1FFD3FF856DC3F850FE9F97C9" box="[151,276,1964,1990]" pageId="23" pageNumber="224" value="2011-12-08">
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<collectingDate id="EFE70119FFD3FF856DC3F850FE9F97C9" box="[151,276,1964,1990]" pageId="23" pageNumber="224" value="2011-12-08">8.XII.2011</collectingDate>
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</date>
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,
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<collectorName id="26E8BBE7FFD3FF856C75F850FE4397C8" box="[289,456,1964,1991]" pageId="23" pageNumber="224">Rudolf Kmeco</collectorName>
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leg. [
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,
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] (8 ♁,
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).
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:
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,
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<collectingDate id="EFE70119FFD3FF856DC3F82CFE8197E5" box="[151,266,2000,2026]" pageId="23" pageNumber="224" value="2010-01-13">13.1.2010</collectingDate>
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</date>
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,
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NW Sesheke
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, S20°17.08, E24°08.49 [erroroneous coordinates, should be S
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24°09´08],
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leg. [
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<emphasis id="B9690223FFDCFF8A6F00FF6BFDD690BE" bold="true" box="[596,605,151,177]" pageId="24" pageNumber="225">]</emphasis>
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(1 ♁,
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).
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<collectingCountry id="F30A9EA1FFDCFF8A6FABFF6AFCF590BF" box="[767,894,150,176]" name="Angola" pageId="24" pageNumber="225">ANGOLA</collectingCountry>
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:
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</emphasis>
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<collectingDate id="EFE70119FFDCFF8A6EDAFF6BFB8990BE" box="[910,1026,151,177]" pageId="24" pageNumber="225" value="2011-11-09">9.XI.2011</collectingDate>
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,
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<collectorName id="26E8BBE7FFDCFF8A695BFF6BFBEF90BE" box="[1039,1124,151,177]" pageId="24" pageNumber="225">Angola</collectorName>
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,
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<collectingRegion id="49D910D3FFDCFF8A6927FF6BFA8090BE" box="[1139,1291,151,177]" country="Angola" name="Bie" pageId="24" pageNumber="225">Bié Province</collectingRegion>
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,
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NEE Kuito
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</locationDeviation>
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,
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<collectingMunicipality id="6BC6444BFFDCFF8A6DB0FF47FE4990DA" box="[228,450,187,213]" pageId="24" pageNumber="225">Chissamba Mission</collectingMunicipality>
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,
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<collectorName id="26E8BBE7FFDCFF8A6C99FF47FDB990DA" box="[461,562,187,213]" pageId="24" pageNumber="225">P. Schüle</collectorName>
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leg. [
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,
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<geoCoordinate id="EE29B8F6FFDCFF8A6FABFF46FCF290DA" box="[767,889,186,213]" degrees="17.335280" direction="east" orientation="longitude" pageId="24" pageNumber="225" precision="1" value="17.33528">17.335280</geoCoordinate>
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] (1 ♁,
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<collectionCode id="ED0C46F4FFDCFF8A6E91FF47FC7090DA" box="[965,1019,187,213]" pageId="24" pageNumber="225">PSC</collectionCode>
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);
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<collectorName id="26E8BBE7FFDCFF8A6959FF47FBE990DA" box="[1037,1122,187,213]" pageId="24" pageNumber="225">Angola</collectorName>
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,
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<collectorName id="26E8BBE7FFDCFF8A6938FF47FB6190DA" box="[1132,1258,187,213]" pageId="24" pageNumber="225">Kangumbe</collectorName>
|
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,
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<location id="8EC288EAFFDCFF8A69A0FF47FF5C90F6" LSID="urn:lsid:plazi:treatment:03B46F27FFD3FF8A6DC3F8C3FB389336:8EC288EAFFDCFF8A69A0FF47FF5C90F6" country="Namibia" latitude="-12.58449" longLatPrecision="1" longitude="18.66771" municipality="Chissamba Mission" name="Langue-Bungo River" pageId="24" pageNumber="225" stateProvince="Bie">Langué-Bungo River</location>
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,
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,
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<geoCoordinate id="EE29B8F6FFDCFF8A6C33FF23FE6F90F6" box="[359,484,223,249]" degrees="18.66771" direction="east" orientation="longitude" pageId="24" pageNumber="225" precision="1" value="18.66771">18.66771E</geoCoordinate>
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,
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<quantity id="4CE573D4FFDCFF8A6CBBFF23FDB490F6" box="[495,575,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.298" pageId="24" pageNumber="225" unit="m" value="1298.0">
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</quantity>
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,
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|
||
<collectingDate id="EFE70119FFDCFF8A6F1EFF23FD6490F6" box="[586,751,223,249]" pageId="24" pageNumber="225" value="2019-11-18" valueMax="2019-11-21" valueMin="2019-11-18">18–21.xi.2019</collectingDate>
|
||
</date>
|
||
”, “NHMUK014373480-85” [
|
||
<geoCoordinate id="EE29B8F6FFDCFF8A696CFF22FB2490F6" box="[1080,1199,222,249]" degrees="12.58449" direction="south" orientation="latitude" pageId="24" pageNumber="225" precision="1" value="-12.58449">-12.58449</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE29B8F6FFDCFF8A69EEFF23FAAD90F6" box="[1210,1318,223,249]" degrees="18.66771" direction="east" orientation="longitude" pageId="24" pageNumber="225" precision="1" value="18.66771">18.66771</geoCoordinate>
|
||
] (3 ♁,
|
||
<specimenCount id="9D1B15B8FFDCFF8A6826FF23FA1D90F6" box="[1394,1430,223,249]" pageId="24" pageNumber="225" type="female">3♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED0C46F4FFDCFF8A6DC3FEFFFF609112" box="[151,235,259,285]" pageId="24" pageNumber="225">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3078DBAFFDCFF8A6D93FEDAFC5A91DE" pageId="24" pageNumber="225" type="diagnosis">
|
||
<paragraph id="8BA2DE31FFDCFF8A6D93FEDAFC5A91DE" blockId="24.[151,1437,150,825]" pageId="24" pageNumber="225">
|
||
<emphasis id="B9690223FFDCFF8A6D93FEDAFE45914F" bold="true" box="[199,462,294,320]" pageId="24" pageNumber="225">Differential diagnosis.</emphasis>
|
||
Members of this subspecies differ from those of the nominative subspecies in having a small size and very round body shape. The yellow pattern on the elytra in most specimens is more developed than in other subspecies. The sides of the pronotum are almost straight, similar to
|
||
<taxonomicName id="4C1DA5B2FFDCFF8A6EA6FE93FB4F9186" authorityName="Anichtchenko & Valainis" authorityYear="2023" box="[1010,1220,367,393]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="capense" subSpecies="kmecoi">
|
||
<emphasis id="B9690223FFDCFF8A6EA6FE93FB4F9186" box="[1010,1220,367,393]" italics="true" pageId="24" pageNumber="225">O. capense kmecoi</emphasis>
|
||
</taxonomicName>
|
||
ssp. n., while in
|
||
<taxonomicName id="4C1DA5B2FFDCFF8A68D5FE93FF7A91A3" authorityName="Gory" authorityYear="1833" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="capense">
|
||
<emphasis id="B9690223FFDCFF8A68D5FE93FF7A91A3" italics="true" pageId="24" pageNumber="225">O. capense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25ABF58FFDCFF8A6DADFE6FFEA691A2" box="[249,301,403,429]" pageId="24" pageNumber="225" sensu="stricto">s.str.</taxonomicNameLabel>
|
||
they are slightly rounded. The apex of the aedeagus is narrower and more strongly curved ventrally. The sides of aedeagus taper sharply, forming the shape of a “bottle neck”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3078DBAFFDCFF8A6D93FE26FEBD92C2" pageId="24" pageNumber="225" type="description">
|
||
<paragraph id="8BA2DE31FFDCFF8A6D93FE26FB6391FA" blockId="24.[151,1437,150,825]" box="[199,1256,474,501]" pageId="24" pageNumber="225">
|
||
<emphasis id="B9690223FFDCFF8A6D93FE26FED291FB" bold="true" box="[199,345,474,500]" pageId="24" pageNumber="225">Description.</emphasis>
|
||
Body length 4.0–
|
||
<quantity id="4CE573D4FFDCFF8A6F76FE26FDF191FA" box="[546,634,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="24" pageNumber="225" unit="mm" value="4.5">4.5 mm</quantity>
|
||
; width
|
||
<quantity id="4CE573D4FFDCFF8A6F98FE27FCDD91FA" box="[716,854,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="24" pageNumber="225" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8–3.2 mm</quantity>
|
||
. Habitus (
|
||
<figureCitation id="1326C2B4FFDCFF8A6E93FE27FB9791FA" box="[967,1052,475,501]" captionStart="FIGURES 27–34" captionStartId="20.[152,270,1909,1933]" captionTargetBox="[223,1371,183,1883]" captionTargetId="figure-36@20.[215,1372,181,1884]" captionTargetPageId="20" captionText="FIGURES 27–34. Habitus of subspecies of O. (O.) capense. dorsal view: 27—O. (O.) capense s.str. (Type); 28—O. (O.) capense s.str. (f. suturale); 29—O. (O.) capense s.str. (inland form, RSA); 30—O. dominicense Chaudoir, 1868 syn. n. (Type); 31—O. (O.) capense congoense stat. n. (Paratype); 32—O. (O.) capense pumilum ssp. n. (Paratype); 33—O. (O.) capense kmecoi ssp. n. (Holotype); 34—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7923252" httpUri="https://zenodo.org/record/7923252/files/figure.png" pageId="24" pageNumber="225">Fig. 32</figureCitation>
|
||
). Venter (
|
||
<figureCitation id="1326C2B4FFDCFF8A69D3FE27FB5091FA" box="[1159,1243,475,501]" captionStart="FIGURES 35–38" captionStartId="22.[152,270,1943,1967]" captionTargetBox="[258,1328,190,1899]" captionTargetId="figure-36@22.[243,1345,181,1919]" captionTargetPageId="22" captionText="FIGURES 35–38. Habitus of subspecies of O. (O.) capense. ventral view: 35—O. (O.) capense s.str.; 36—O. (O.) capense pumilum ssp. n. (Paratype); 37—O. (O.) capense kmecoi ssp. n. (Paratype); 38—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7923256" httpUri="https://zenodo.org/record/7923256/files/figure.png" pageId="24" pageNumber="225">Fig. 36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2DE31FFDCFF8A6D93FE03FEBD92C2" blockId="24.[151,1437,150,825]" pageId="24" pageNumber="225">
|
||
General taxon characteristics correspond to those given in the description of the nominate subspecies. Distinguishing features are indicated in the differential diagnosis and key.Aedeagus (
|
||
<figureCitation id="1326C2B4FFDCFF8A6913FDDFFB1C9232" box="[1095,1175,547,573]" captionStart="FIGURES 39–45" captionStartId="23.[152,270,1246,1270]" captionTargetBox="[156,1422,190,1209]" captionTargetId="figure-26@23.[151,1436,181,1222]" captionTargetPageId="23" captionText="FIGURES 39–45. Aedeagus structure. a—right lateral view; b—apex, dorsal view:: 39—O. (O.) capense s.str. (RSA); 40— O. dominicense Chaudoir, 1868 syn. n. (Type); 41—O. (O.) capense s.str. (f. suturale); 42—O. (O.) capense kmecoi ssp. n. (Holotype); 43—O. (O.) capense congoense stat. n. (Paratype); 44—O. (O.) capense pumilum ssp. n. (Paratype); 45—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7923262" httpUri="https://zenodo.org/record/7923262/files/figure.png" pageId="24" pageNumber="225">Fig. 44</figureCitation>
|
||
) Endophallus is typical for the species: with two groups of medium sized spines. One is large and situated in distal part tubular formation, extended to basal orifice of median lobe, and covered with scales. The second group of spines is much smaller, located more apically and to the left of the first and extended apically beyond the first group by about half its length (
|
||
<figureCitation id="1326C2B4FFDCFF8A6DCBFD4FFEA192C2" box="[159,298,690,717]" captionStart="FIGURES 46–51" captionStartId="26.[152,270,966,990]" captionTargetBox="[155,1422,185,939]" captionTargetId="figure-36@26.[151,1436,181,942]" captionTargetPageId="26" captionText="FIGURES 46–51. Endofallus structure of subspecies of O. (O.) capense. dorsal view: 46—O. (O.) capense s.str. (f. suturale); 47—O. (O.) capense kmecoi ssp. n. (Holotype); 48—O. (O.) capense congoense stat. n. (Paratype); 49—O. (O.) capense pumilum ssp. n. (Paratype; Angola: Kangube); 50—O. (O.) capense pumilum ssp. n. (Paratype; Namibia: Caprivi); 51—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7923266" httpUri="https://zenodo.org/record/7923266/files/figure.png" pageId="24" pageNumber="225">Figs. 49–50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3078DBAFFDCFF8A6D93FD2BFDB3931A" pageId="24" pageNumber="225" type="etymology">
|
||
<paragraph id="8BA2DE31FFDCFF8A6D93FD2BFDB3931A" blockId="24.[151,1437,150,825]" pageId="24" pageNumber="225">
|
||
<emphasis id="B9690223FFDCFF8A6D93FD2BFEC592FE" bold="true" box="[199,334,727,753]" pageId="24" pageNumber="225">Etymology.</emphasis>
|
||
The specific epithet
|
||
<emphasis id="B9690223FFDCFF8A6F11FD2BFD2C92FE" box="[581,679,727,753]" italics="true" pageId="24" pageNumber="225">pumilum</emphasis>
|
||
is derived from the Latin word “pumilus”, meaning ‘dwarf’, and refers to the small size of subspecies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3078DBAFFDCFF8A6D93FCE2FB389336" box="[199,1203,798,825]" pageId="24" pageNumber="225" type="distribution">
|
||
<paragraph id="8BA2DE31FFDCFF8A6D93FCE2FB389336" blockId="24.[151,1437,150,825]" box="[199,1203,798,825]" pageId="24" pageNumber="225">
|
||
<emphasis id="B9690223FFDCFF8A6D93FCE2FEEA9337" bold="true" box="[199,353,798,824]" pageId="24" pageNumber="225">Distribution.</emphasis>
|
||
The new subspecies is known from
|
||
<collectingCountry id="F30A9EA1FFDCFF8A6FA3FCE3FCC79336" box="[759,844,799,825]" name="Angola" pageId="24" pageNumber="225">Angola</collectingCountry>
|
||
,
|
||
<collectingCountry id="F30A9EA1FFDCFF8A6E0CFCE3FC339336" box="[856,952,799,825]" name="Namibia" pageId="24" pageNumber="225">Namibia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F30A9EA1FFDCFF8A6EBAFCE3FBCF9336" box="[1006,1092,799,825]" name="Zambia" pageId="24" pageNumber="225">Zambia</collectingCountry>
|
||
(Map. 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |