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<document id="0EBBB6EFA44C1C788B49CFB8E2390AE5" ID-CLB-Dataset="21333" ID-DOI="10.5852/ejt.2020.700" ID-GBIF-Dataset="5fcc010d-8df1-418f-87b1-b29e649ed604" ID-Zenodo-Dep="3981069" ID-ZooBank="17D38562-BF4F-448F-9969-8524CEACC4B2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597240089656" checkinUser="valdenar" docAuthor="Maetens, Heleen, Steenberge, Maarten Van, Snoeks, Jos &amp; Decru, Eva" docDate="2020" docId="03B287EEF85FF30EFD90D6845856C948" docLanguage="en" docName="ejt-700_maetens_van-steenberge_snoeks.pdf" docOrigin="European Journal of Taxonomy 700" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Enteromius alberti" docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FF8BFF96F851F31FFFCED72E5D09CC1A" masterDocTitle="Revalidation of Enteromius alberti and presence of Enteromius cf. mimus (Cypriniformes: Cyprinidae) in the Lake Edward system, East Africa" masterLastPageNumber="28" masterPageNumber="1" pageNumber="15" updateTime="1698844288485" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4E58DD0B9CC8796C66E3DB116F891AC3">Revalidation of Enteromius alberti and presence of Enteromius cf. mimus (Cypriniformes: Cyprinidae) in the Lake Edward system, East Africa</mods:title>
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<mods:namePart id="02AF87875B34A98390FA6C5A104555C7">Maetens, Heleen</mods:namePart>
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<mods:affiliation id="9919A454FC0F21BEA2212036C1377CE8">Section Vertebrates, Biology Department, Royal Museum for Central Africa, Leuvensesteenweg 13, 3080 Tervuren, Belgium. Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, University of Leuven, Ch. Deberiotstraat 32, 3000 Leuven, Belgium. Operational Directorate Taxonomy and Phylogeny, Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000 Brussels, Belgium. Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:namePart id="DF8BD1E7B1A27070C5A26BB9F23F73DA">Steenberge, Maarten Van</mods:namePart>
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<mods:affiliation id="6A719594633591DDB573789A31BF5C69">Section Vertebrates, Biology Department, Royal Museum for Central Africa, Leuvensesteenweg 13, 3080 Tervuren, Belgium. Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, University of Leuven, Ch. Deberiotstraat 32, 3000 Leuven, Belgium. Operational Directorate Taxonomy and Phylogeny, Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000 Brussels, Belgium. Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:namePart id="09E35C745ADA3C5D065BFB95D51595B8">Snoeks, Jos</mods:namePart>
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<mods:affiliation id="8FE33918B91BA88E607CFD9F0F436B08">Section Vertebrates, Biology Department, Royal Museum for Central Africa, Leuvensesteenweg 13, 3080 Tervuren, Belgium. Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, University of Leuven, Ch. Deberiotstraat 32, 3000 Leuven, Belgium. Operational Directorate Taxonomy and Phylogeny, Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000 Brussels, Belgium. Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:namePart id="FE4281B1FFB54E4F668A0635C964DAB1">Decru, Eva</mods:namePart>
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<mods:affiliation id="E95BDD12E78EFA08E9377E34E01676D2">Section Vertebrates, Biology Department, Royal Museum for Central Africa, Leuvensesteenweg 13, 3080 Tervuren, Belgium. Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, University of Leuven, Ch. Deberiotstraat 32, 3000 Leuven, Belgium. Operational Directorate Taxonomy and Phylogeny, Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000 Brussels, Belgium. Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<treatment id="03B287EEF85FF30EFD90D6845856C948" ID-DOI="http://doi.org/10.5281/zenodo.4327983" ID-GBIF-Taxon="166239951" ID-Zenodo-Dep="4327983" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948" httpUri="http://treatment.plazi.org/id/03B287EEF85FF30EFD90D6845856C948" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="C3016573F85FF311FD90D6845EDCCDDE" box="[606,981,425,452]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BA436F8F85FF311FD90D6845EDCCDDE" blockId="14.[606,981,425,487]" box="[606,981,425,452]" pageId="14" pageNumber="15">
<heading id="D0EC8194F85FF311FD90D6845EDCCDDE" box="[606,981,425,452]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C1B4D7BF85FF311FD90D6845EDCCDDE" ID-CoL="KMWJ" authority="(Poll, 1939)" baseAuthorityName="Poll" baseAuthorityYear="1939" box="[606,981,425,452]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85FF311FD90D6845E48CDDE" bold="true" box="[606,833,425,452]" italics="true" pageId="14" pageNumber="15">Enteromius alberti</emphasis>
(Poll, 1939)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016573F85FF311FD7ED6E35E8DCDFD" box="[688,900,461,487]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BA436F8F85FF311FD7ED6E35E8DCDFD" blockId="14.[606,981,425,487]" box="[688,900,461,487]" pageId="14" pageNumber="15">
<figureCitation id="13202A7DF85FF311FD7ED6E35FFDCDFD" box="[688,756,461,487]" captionStart="Fig" captionStartId="14.[189,232,1956,1982]" captionTargetBox="[189,1398,1090,1915]" captionTargetId="figure@14.[189,1398,1090,1918]" captionTargetPageId="14" captionText="Fig. 7. (a) Lectotype of Enteromius alberti (Poll, 1939) (MRAC 64723) with 74.3 mm SL. (b) Fresh specimen of E. alberti (RMCA 2018.008.P.0248 HP 3246) with 62.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3981083" httpUri="https://zenodo.org/record/3981083/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
ab,
<tableCitation id="C6990343F85FF311FCE3D6E35E8DCDFD" box="[813,900,461,487]" captionStart="Table 2" captionStartId="12.[179,248,531,559]" captionText="Table 2 (continued on the next page). Measurements and meristics of Enteromius alberti (Poll, 1939) (n = 47, *= 46, ** = 45), the types of Enteromius mimus (Boulenger, 1912) (n = 9), and Enteromius cf. mimus (n = 76, * = 75, ** = 74, *** = 73)." pageId="14" pageNumber="15">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3016573F85FF311FF73D5215EE6CE33" box="[189,1007,526,553]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BA436F8F85FF311FF73D5215EE6CE33" blockId="14.[189,1007,526,553]" box="[189,1007,526,553]" pageId="14" pageNumber="15">
<taxonomicName id="4C1B4D7BF85FF311FF73D5215F81CE33" ID-CoL="6FDCM" authority="(Whitehead, 1960)" baseAuthorityName="Whitehead" baseAuthorityYear="1960" box="[189,648,526,553]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="cercops">
<emphasis id="B96FEAEAF85FF311FF73D5215CADCE32" box="[189,420,527,552]" italics="true" pageId="14" pageNumber="15">Enteromius cercops</emphasis>
(
<bibRefCitation id="EF8A4B09F85FF311FE7AD5205F76CE33" author="Whitehead P. J. P." box="[436,639,526,553]" firstAuthor="Whitehead" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="14" pageNumber="15" pagination="106 - 119" part="62" refId="ref17994" refString="Whitehead P. J. P. 1960. Three new cyprinid fishes of the genus Barbus from the Lake Victoria basin. Revue de Zoologie et de Botanique africaines 62 (1 - 2): 106 - 119." title="Three new cyprinid fishes of the genus Barbus from the Lake Victoria basin" type="journal article" year="1960">Whitehead, 1960</bibRefCitation>
)
</taxonomicName>
(here synonymised). syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="C3016573F85FF310FF73D57E5C8CC9EF" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BA436F8F85FF311FF73D57E5C3FCE70" blockId="14.[189,310,592,618]" box="[189,310,592,618]" pageId="14" pageNumber="15">
<heading id="D0EC8194F85FF311FF73D57E5C3FCE70" bold="true" box="[189,310,592,618]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B96FEAEAF85FF311FF73D57E5C3FCE70" bold="true" box="[189,310,592,618]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436F8F85FF310FF73D5515C8CC9EF" blockId="14.[189,1399,638,1017]" lastBlockId="15.[190,1400,267,1525]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<taxonomicName id="4C1B4D7BF85FF311FF73D5515CA9CE82" box="[189,416,638,664]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85FF311FF73D5515CA9CE82" box="[189,416,638,664]" italics="true" pageId="14" pageNumber="15">Enteromius alberti</emphasis>
</taxonomicName>
(
<figureCitation id="13202A7DF85FF311FE7BD5505CF6CE83" box="[437,511,638,665]" captionStart="Fig" captionStartId="14.[189,232,1956,1982]" captionTargetBox="[189,1398,1090,1915]" captionTargetId="figure@14.[189,1398,1090,1918]" captionTargetPageId="14" captionText="Fig. 7. (a) Lectotype of Enteromius alberti (Poll, 1939) (MRAC 64723) with 74.3 mm SL. (b) Fresh specimen of E. alberti (RMCA 2018.008.P.0248 HP 3246) with 62.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3981083" httpUri="https://zenodo.org/record/3981083/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
ab) belongs to the group of species of
<taxonomicName id="4C1B4D7BF85FF311FC34D551598BCE82" authorityName="Cope" authorityYear="1867" box="[1018,1154,639,664]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEAF85FF311FC34D551598BCE82" box="[1018,1154,639,664]" italics="true" pageId="14" pageNumber="15">Enteromius</emphasis>
</taxonomicName>
with a flexible last unbranched dorsal fin ray that lacks serrations along its posterior edge. A comparison was made with other species of
<taxonomicName id="4C1B4D7BF85FF311FEB8D5E85CF7CEC5" authorityName="Cope" authorityYear="1867" box="[374,510,710,735]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEAF85FF311FEB8D5E85CF7CEC5" box="[374,510,710,735]" italics="true" pageId="14" pageNumber="15">Enteromius</emphasis>
</taxonomicName>
from this group from the East Coast and Nilo-Sudan ichthyofaunal provinces (list made based on information available in, e.g.,
<bibRefCitation id="EF8A4B09F85FF311FCEAD5C65EFECF18" accessDate="2018-11-21" author="Greenwood P. H." box="[804,1015,744,770]" firstAuthor="Greenwood" journalOrPublisher="Bulletin of the Natural History Museum, Zoology" pageId="14" pageNumber="15" pagination="151 - 208" part="4" publicationUrl="https://www.biodiversitylibrary.org/part/118628#/summary" refId="ref15784" refString="Greenwood P. H. 1962. A revision of certain Barbus (Pisces, Cyprinidae) from East, Central and South Africa. Bulletin of the Natural History Museum, Zoology series 8 (4): 151 - 208. Available from https: // www. biodiversitylibrary. org / part / 118628 # / summary [accessed 21 Nov. 2018]." title="A revision of certain Barbus (Pisces, Cyprinidae) from East, Central and South Africa" type="journal article" year="1962">Greenwood 1962</bibRefCitation>
;
<bibRefCitation id="EF8A4B09F85FF311FBCBD5C65805CF18" accessDate="2018-11-21" author="Okaronon J. O. &amp; Katunzi E. F. B. &amp; Asila A. A." box="[1029,1292,744,770]" etAl="et al." firstAuthor="Okaronon" journalOrPublisher="Lake Victoria Fisheries Research Project Phase II, Technical Document 1, Jinja, Uganda" pageId="14" pageNumber="15" publicationUrl="http://aquaticcommons.org/4150/" refId="ref16698" refString="Okaronon J. O., Katunzi E. F. B. &amp; Asila A. A. 1997. Fish Species Identification Guide for Lake Victoria, Enumerator's Field Guide. Lake Victoria Fisheries Research Project Phase II, Technical Document 1, Jinja, Uganda. Available from http: // aquaticcommons. org / 4150 / [accessed 21 Nov. 2018]." title="Fish Species Identification Guide for Lake Victoria, Enumerator's Field Guide" type="url" year="1997">
Okaronon
<emphasis id="B96FEAEAF85FF311FB4BD5C759CACF18" box="[1157,1219,744,770]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="EF8A4B09F85FF311FAD4D5C65C37CF3F" author="Seegers L. &amp; De Vos L. &amp; Okeyo D. O." etAl="et al." firstAuthor="Seegers" journalOrPublisher="Journal of East African Natural History" pageId="14" pageNumber="15" pagination="11 - 47" part="92" refId="ref17474" refString="Seegers L., De Vos L. &amp; Okeyo D. O. 2003. Annotated checklist of the freshwater fishes of Kenya (excluding the lacustrine haplochromines from Lake Victoria). Journal of East African Natural History 92: 11 - 47. https: // doi. org / 10.2982 / 0012 - 8317 (2003) 92 % 5 B 11: ACOTFF % 5 D 2.0. CO; 2" title="Annotated checklist of the freshwater fishes of Kenya (excluding the lacustrine haplochromines from Lake Victoria)" type="journal article" year="2003">
Seegers
<emphasis id="B96FEAEAF85FF311FF73D4225DF1CF3F" box="[189,248,779,805]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2003
</bibRefCitation>
;
<bibRefCitation id="EF8A4B09F85FF311FE84D4255F3CCF3F" author="Schmidt R. C. &amp; Bart Jr. H. L. &amp; Nyingi W. D." box="[330,565,779,805]" etAl="et al." firstAuthor="Schmidt" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="14" pageNumber="15" pagination="35 - 43" part="111" refId="ref17181" refString="Schmidt R. C., Bart Jr. H. L. &amp; Nyingi W. D. 2017. Multi-locus phylogeny reveals instances of mitochondrial introgression and unrecognized diversity in Kenyan barbs (Cyprininae: Smiliogastrini). Molecular Phylogenetics and Evolution 111: 35 - 43. https: // doi. org / 10.1016 / j. ympev. 2017.03.015" title="Multi-locus phylogeny reveals instances of mitochondrial introgression and unrecognized diversity in Kenyan barbs (Cyprininae: Smiliogastrini)" type="journal article" year="2017">
Schmidt
<emphasis id="B96FEAEAF85FF311FE7AD4225CE6CF3F" box="[436,495,779,805]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2017
</bibRefCitation>
;
<bibRefCitation id="EF8A4B09F85FF311FD8FD4225E49CF3F" author="Froese R. &amp; Pauly D." box="[577,832,779,806]" firstAuthor="Froese" journalOrPublisher="Available from www. fishbase. org" pageId="14" pageNumber="15" refId="ref15726" refString="Froese R. &amp; Pauly D. (eds) 2018. FishBase. Available from www. fishbase. org [accessed 21 Nov. 2018]." title="FishBase" type="book" year="2018">Froese &amp; Pauly 2018</bibRefCitation>
).
<taxonomicName id="4C1B4D7BF85FF311FC9DD4225938CF3F" box="[851,1073,779,805]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85FF311FC9DD4225938CF3F" box="[851,1073,779,805]" italics="true" pageId="14" pageNumber="15">Enteromius alberti</emphasis>
</taxonomicName>
can easily be distinguished from the other species of this group from the East Coast and Nilo-Sudan ichthyofaunal regions by the following combination of characteristics (data from original description, unless otherwise stated): a complete lateral line vs an incomplete lateral line in
<taxonomicName id="4C1B4D7BF85FF311FCD1D4585954CF8A" authority="(Bailey, 1969)" baseAuthorityName="Bailey" baseAuthorityYear="1969" box="[799,1117,885,912]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="atkinsoni">
<emphasis id="B96FEAEAF85FF311FCD1D4585EA5CF95" box="[799,940,885,911]" italics="true" pageId="14" pageNumber="15">E. atkinsoni</emphasis>
(Bailey, 1969)
</taxonomicName>
,
<taxonomicName id="4C1B4D7BF85FF311FBA9D4585C0CCFA8" authority="(Boulenger, 1901)" baseAuthorityName="Boulenger" baseAuthorityYear="1901" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B96FEAEAF85FF311FBA9D45859ECCF95" box="[1127,1253,885,911]" italics="true" pageId="14" pageNumber="15">E. pumilus</emphasis>
(Boulenger, 1901)
</taxonomicName>
,
<taxonomicName id="4C1B4D7BF85FF311FEDED4B75F70CFA8" authority="(Farm, 2000)" baseAuthorityName="Farm" baseAuthorityYear="2000" box="[272,633,920,946]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="serengetiensis">
<emphasis id="B96FEAEAF85FF311FEDED4B75CDFCFA8" box="[272,470,921,946]" italics="true" pageId="14" pageNumber="15">E. serengetiensis</emphasis>
(Farm, 2000)
</taxonomicName>
,
<taxonomicName id="4C1B4D7BF85FF311FD4AD4B75EE1CFA9" authority="(Rendahl, 1935)" baseAuthorityName="Rendahl" baseAuthorityYear="1935" box="[644,1000,920,947]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="tongaensis">
<emphasis id="B96FEAEAF85FF311FD4AD4B75E2ACFA8" box="[644,803,921,946]" italics="true" pageId="14" pageNumber="15">E. tongaensis</emphasis>
(Rendahl, 1935)
</taxonomicName>
and
<taxonomicName id="4C1B4D7BF85FF311FBD0D4B7587BCFA9" authority="(Boulenger, 1916)" baseAuthorityName="Boulenger" baseAuthorityYear="1916" box="[1054,1394,920,947]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="toppini">
<emphasis id="B96FEAEAF85FF311FBD0D4B7599BCFA8" box="[1054,1170,921,946]" italics="true" pageId="14" pageNumber="15">E. toppini</emphasis>
(
<bibRefCitation id="EF8A4B09F85FF311FB6ED4B65860CFA9" author="Boulenger G. A." box="[1184,1385,920,947]" firstAuthor="Boulenger" journalOrPublisher="British Museum (Natural History), London, UK" pageId="14" pageNumber="15" refId="ref15374" refString="Boulenger G. A. 1916. Catalogue of the Fresh-Water Fishes of Africa in the British museum (Natural History), Vol. IV. British Museum (Natural History), London, UK. https: // doi. org / 10.5962 / bhl. title. 8869" title="Catalogue of the Fresh-Water Fishes of Africa in the British museum (Natural History), Vol. IV" type="book" year="1916">Boulenger, 1916</bibRefCitation>
)
</taxonomicName>
; two pairs of barbels vs one pair in
<taxonomicName id="4C1B4D7BF85FF311FDB9D4925919CFCC" authority="(Seegers, 1996)" baseAuthorityName="Seegers" baseAuthorityYear="1996" box="[631,1040,955,982]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pseudotoppini">
<emphasis id="B96FEAEAF85FF311FDB9D4925E4DCFCF" box="[631,836,955,981]" italics="true" pageId="14" pageNumber="15">E. pseudotoppini</emphasis>
(Seegers, 1996)
</taxonomicName>
, and no barbels in
<taxonomicName id="4C1B4D7BF85FF311FACDD4925C9ECFE3" authority="(Boulenger, 1903)" baseAuthorityName="Boulenger" baseAuthorityYear="1903" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="anema">
<emphasis id="B96FEAEAF85FF311FACDD492587ECFCF" box="[1283,1399,956,981]" italics="true" pageId="14" pageNumber="15">E. anema</emphasis>
(
<bibRefCitation id="EF8A4B09F85FF311FF0BD4F05C87CFE3" author="Boulenger G. A." box="[197,398,990,1017]" firstAuthor="Boulenger" journalOrPublisher="Annals and Magazine of Natural History" pageId="14" pageNumber="15" pagination="532 - 533" part="12" refId="ref15223" refString="Boulenger G. A. 1903. Diagnosis of four new species of Barbus from the Nile. Annals and Magazine of Natural History, Series 7 12: 532 - 533. https: // doi. org / 10.1080 / 00222930308678890" title="Diagnosis of four new species of Barbus from the Nile" type="journal article" year="1903">Boulenger, 1903</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1B4D7BF85FF311FE1CD4F15E69CFE3" authority="(Greenwood, 1970)" baseAuthorityName="Greenwood" baseAuthorityYear="1970" box="[466,864,990,1017]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="profundus">
<emphasis id="B96FEAEAF85FF311FE1CD4F15F63CFE2" box="[466,618,990,1016]" italics="true" pageId="14" pageNumber="15">E. profundus</emphasis>
(
<bibRefCitation id="EF8A4B09F85FF311FDB5D4F05E5ECFE3" author="Greenwood P. H." box="[635,855,990,1017]" firstAuthor="Greenwood" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="14" pageNumber="15" pagination="1 - 13" part="82" refId="ref15870" refString="Greenwood P. H. 1970. A revision of the cyprinid species Barbus (Enteromius) radiatus Peters, 1853, with a note on the synonymy of the subgenera Beirabarbus and Enteromius. Revue de Zoologie et de Botanique africaines 82: 1 - 13." title="A revision of the cyprinid species Barbus (Enteromius) radiatus Peters, 1853, with a note on the synonymy of the subgenera Beirabarbus and Enteromius" type="journal article" year="1970">Greenwood, 1970</bibRefCitation>
)
</taxonomicName>
; one to three dark spots on the flanks, which sometimes fuse into a mid-lateral line in preserved specimens, starting posterior to the operculum vs a dark line running from the tip of the snout to the caudal fin base in
<taxonomicName id="4C1B4D7BF85EF310FC3DD601584CCD53" authority="(Fowler, 1935)" baseAuthorityName="Fowler" baseAuthorityYear="1935" box="[1011,1349,302,329]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="bifrenatus">
<emphasis id="B96FEAEAF85EF310FC3DD6015985CD53" box="[1011,1164,303,329]" italics="true" pageId="15" pageNumber="16">E. bifrenatus</emphasis>
(Fowler, 1935)
</taxonomicName>
and
<taxonomicName id="4C1B4D7BF85EF310FF70D67D5F1DCD77" authority="(Whitehead, 1960)" baseAuthorityName="Whitehead" baseAuthorityYear="1960" box="[190,532,338,365]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="yongei">
<emphasis id="B96FEAEAF85EF310FF70D67D5C24CD76" box="[190,301,339,364]" italics="true" pageId="15" pageNumber="16">E. yongei</emphasis>
(
<bibRefCitation id="EF8A4B09F85EF310FEF5D67C5F03CD77" author="Whitehead P. J. P." box="[315,522,338,365]" firstAuthor="Whitehead" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="15" pageNumber="16" pagination="106 - 119" part="62" refId="ref17994" refString="Whitehead P. J. P. 1960. Three new cyprinid fishes of the genus Barbus from the Lake Victoria basin. Revue de Zoologie et de Botanique africaines 62 (1 - 2): 106 - 119." title="Three new cyprinid fishes of the genus Barbus from the Lake Victoria basin" type="journal article" year="1960">Whitehead, 1960</bibRefCitation>
)
</taxonomicName>
, and a thin dark line from the beginning of the operculum to the caudal fin base in
<taxonomicName id="4C1B4D7BF85EF310FED6D6585F55CD95" authority="(Weber, 1897)" baseAuthorityName="Weber" baseAuthorityYear="1897" box="[280,604,373,399]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="viviparus">
<emphasis id="B96FEAEAF85EF310FED6D6585CAECD95" box="[280,423,374,399]" italics="true" pageId="15" pageNumber="16">E. viviparus</emphasis>
(Weber, 1897)
</taxonomicName>
; 12 scales around the caudal peduncle, with one aberrant specimen in the
<typeStatus id="54A0885AF85EF310FEC8D6B75C71CDA9" box="[262,376,409,435]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C1B4D7BF85EF310FE50D6B75F10CDA8" baseAuthorityName="Whitehead" baseAuthorityYear="1960" box="[414,537,409,434]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="cercops">
<emphasis id="B96FEAEAF85EF310FE50D6B75F10CDA8" box="[414,537,409,434]" italics="true" pageId="15" pageNumber="16">E. cercops</emphasis>
</taxonomicName>
(16), vs
<quantity id="4CE39B1DF85EF310FD4ED6B65FA4CDA8" box="[640,685,408,434]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="15" pageNumber="16" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="4C1B4D7BF85EF310FD7DD6B7592ACDA9" authority="(Boulenger, 1915)" baseAuthorityName="Boulenger" baseAuthorityYear="1915" box="[691,1059,408,435]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leonensis">
<emphasis id="B96FEAEAF85EF310FD7DD6B75E48CDA8" box="[691,833,408,434]" italics="true" pageId="15" pageNumber="16">E. leonensis</emphasis>
(Boulenger, 1915)
</taxonomicName>
,
<quantity id="4CE39B1DF85EF310FBE0D6B6598ECDA8" box="[1070,1159,408,434]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="15" pageNumber="16" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">910 in</quantity>
<taxonomicName id="4C1B4D7BF85EF310FB43D6B75C0FCDCF" authority="(Bailey, 1980)" baseAuthorityName="Bailey" baseAuthorityYear="1980" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="venustus">
<emphasis id="B96FEAEAF85EF310FB43D6B7581DCDA8" box="[1165,1300,409,434]" italics="true" pageId="15" pageNumber="16">E. venustus</emphasis>
(Bailey, 1980)
</taxonomicName>
, and
<quantity id="4CE39B1DF85EF310FE8CD6955C77CDCF" box="[322,382,443,469]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="16" unit="in" value="10.0">10 in</quantity>
<taxonomicName id="4C1B4D7BF85EF310FE4AD6925E1ACDCC" authority="(Boulenger, 1906)" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[388,787,443,470]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="magdalenae">
<emphasis id="B96FEAEAF85EF310FE4AD6925F3CCDCF" box="[388,565,443,469]" italics="true" pageId="15" pageNumber="16">E. magdalenae</emphasis>
(Boulenger, 1906)
</taxonomicName>
and
<taxonomicName id="4C1B4D7BF85EF310FC85D6925999CDCC" authority="(Johnsen, 1926)" baseAuthorityName="Johnsen" baseAuthorityYear="1926" box="[843,1168,443,470]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="yeiensis">
<emphasis id="B96FEAEAF85EF310FC85D6925EC0CDCF" box="[843,969,444,469]" italics="true" pageId="15" pageNumber="16">E. yeiensis</emphasis>
(Johnsen, 1926)
</taxonomicName>
; 4.5 scales between the dorsal fin base and the lateral line vs
<quantity id="4CE39B1DF85EF310FD7FD6F05FFFCDE2" box="[689,758,478,505]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" pageId="15" pageNumber="16" unit="in" value="3.5">3.5 in</quantity>
<taxonomicName id="4C1B4D7BF85EF310FD31D6F1582CCDE3" authority="(Peters, 1853) (Greenwood 1970)" authorityName="Greenwood" authorityYear="1970" baseAuthorityName="Peters" baseAuthorityYear="1853" box="[767,1317,478,505]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="radiatus">
<emphasis id="B96FEAEAF85EF310FD31D6F15E8CCDE3" box="[767,901,479,505]" italics="true" pageId="15" pageNumber="16">E. radiatus</emphasis>
(Peters, 1853) (
<bibRefCitation id="EF8A4B09F85EF310FB89D6F05812CDE3" author="Greenwood P. H." box="[1095,1307,478,505]" firstAuthor="Greenwood" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="15" pageNumber="16" pagination="1 - 13" part="82" refId="ref15870" refString="Greenwood P. H. 1970. A revision of the cyprinid species Barbus (Enteromius) radiatus Peters, 1853, with a note on the synonymy of the subgenera Beirabarbus and Enteromius. Revue de Zoologie et de Botanique africaines 82: 1 - 13." title="A revision of the cyprinid species Barbus (Enteromius) radiatus Peters, 1853, with a note on the synonymy of the subgenera Beirabarbus and Enteromius" type="journal article" year="1970">Greenwood 1970</bibRefCitation>
)
</taxonomicName>
,
<quantity id="4CE39B1DF85EF310FAFCD6F0587ECDE2" box="[1330,1399,478,505]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" pageId="15" pageNumber="16" unit="in" value="5.5">5.5 in</quantity>
<taxonomicName id="4C1B4D7BF85EF310FF70D52D5C7ACE06" baseAuthorityName="Günther" baseAuthorityYear="1867" box="[190,371,515,540]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="unitaeniatus">
<emphasis id="B96FEAEAF85EF310FF70D52D5C7ACE06" box="[190,371,515,540]" italics="true" pageId="15" pageNumber="16">E. unitaeniatus</emphasis>
</taxonomicName>
(G̹nther, 1867), and
<quantity id="4CE39B1DF85EF310FD4ED52C5FA6CE06" box="[640,687,514,540]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="15" pageNumber="16" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="4C1B4D7BF85EF310FD76D52D5937CE06" authority="(Lonnberg, 1907)" baseAuthorityName="Lonnberg" baseAuthorityYear="1907" box="[696,1086,514,540]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="usambarae">
<emphasis id="B96FEAEAF85EF310FD76D52D5E56CE06" box="[696,863,514,540]" italics="true" pageId="15" pageNumber="16">E. usambarae</emphasis>
(Lönnberg, 1907)
</taxonomicName>
; a dorsal fin length which is larger than the head length vs a dorsal fin length equal to the head length in
<taxonomicName id="4C1B4D7BF85EF310FBB1D5085C0ECE79" authority="(Pfeffer, 1896)" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="innocens">
<emphasis id="B96FEAEAF85EF310FBB1D5085802CE25" box="[1151,1291,550,575]" italics="true" pageId="15" pageNumber="16">E. innocens</emphasis>
(Pfeffer, 1896)
</taxonomicName>
; a body depth which is larger than the head length vs a body depth which is equal to the head length in
<taxonomicName id="4C1B4D7BF85EF310FEE5D5425FA3CE9F" authority="(Boulenger, 1903)" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[299,682,619,645]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nigeriensis">
<emphasis id="B96FEAEAF85EF310FEE5D5425CC4CE9F" box="[299,461,620,645]" italics="true" pageId="15" pageNumber="16">E. nigeriensis</emphasis>
(
<bibRefCitation id="EF8A4B09F85EF310FE15D5455FABCE9F" author="Boulenger G. A." box="[475,674,619,645]" firstAuthor="Boulenger" journalOrPublisher="Annals and Magazine of Natural History" pageId="15" pageNumber="16" pagination="532 - 533" part="12" refId="ref15223" refString="Boulenger G. A. 1903. Diagnosis of four new species of Barbus from the Nile. Annals and Magazine of Natural History, Series 7 12: 532 - 533. https: // doi. org / 10.1080 / 00222930308678890" title="Diagnosis of four new species of Barbus from the Nile" type="journal article" year="1903">Boulenger, 1903</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1B4D7BF85EF310FD2FD542598DCE9F" authority="(Boulenger, 1902)" baseAuthorityName="Boulenger" baseAuthorityYear="1902" box="[737,1156,619,645]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="trispilopleura">
<emphasis id="B96FEAEAF85EF310FD2FD5425EAACE9F" box="[737,931,619,645]" italics="true" pageId="15" pageNumber="16">E. trispilopleura</emphasis>
(Boulenger, 1902)
</taxonomicName>
; a pectoral fin length which is ⁵⁄₆ of the head length vs ¾ in
<taxonomicName id="4C1B4D7BF85EF310FDB3D5A1593DCEB3" authority="(Boulenger, 1903)" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[637,1076,654,681]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B96FEAEAF85EF310FDB3D5A15E5ACEB3" box="[637,851,655,681]" italics="true" pageId="15" pageNumber="16">E. lineomaculatus</emphasis>
(
<bibRefCitation id="EF8A4B09F85EF310FCACD5A05922CEB3" author="Boulenger G. A." box="[866,1067,654,681]" firstAuthor="Boulenger" journalOrPublisher="Annals and Magazine of Natural History" pageId="15" pageNumber="16" pagination="532 - 533" part="12" refId="ref15223" refString="Boulenger G. A. 1903. Diagnosis of four new species of Barbus from the Nile. Annals and Magazine of Natural History, Series 7 12: 532 - 533. https: // doi. org / 10.1080 / 00222930308678890" title="Diagnosis of four new species of Barbus from the Nile" type="journal article" year="1903">Boulenger, 1903</bibRefCitation>
)
</taxonomicName>
, and ¾ to ¼ in
<taxonomicName id="4C1B4D7BF85EF310FB29D5A15F78CED6" authority="(Boulenger, 1903) (Boulenger 1907)" authorityName="Boulenger" authorityYear="1907" baseAuthorityName="Boulenger" baseAuthorityYear="1903" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="neglectus">
<emphasis id="B96FEAEAF85EF310FB29D5A1587ECEB3" box="[1255,1399,655,681]" italics="true" pageId="15" pageNumber="16">E. neglectus</emphasis>
(
<bibRefCitation id="EF8A4B09F85EF310FF08D59C5C84CED6" author="Boulenger G. A." box="[198,397,690,716]" firstAuthor="Boulenger" journalOrPublisher="Annals and Magazine of Natural History" pageId="15" pageNumber="16" pagination="532 - 533" part="12" refId="ref15223" refString="Boulenger G. A. 1903. Diagnosis of four new species of Barbus from the Nile. Annals and Magazine of Natural History, Series 7 12: 532 - 533. https: // doi. org / 10.1080 / 00222930308678890" title="Diagnosis of four new species of Barbus from the Nile" type="journal article" year="1903">Boulenger, 1903</bibRefCitation>
) (
<bibRefCitation id="EF8A4B09F85EF310FE6AD59C5F61CED6" author="Boulenger G. A." box="[420,616,690,716]" firstAuthor="Boulenger" journalOrPublisher="Published for the Egyptian Government by Hugh Rees, London" pageId="15" pageNumber="16" refId="ref15266" refString="Boulenger G. A. 1907. Zoology of Egypt: the Fishes of the Nile. Published for the Egyptian Government by Hugh Rees, London. https: // doi. org / 10.5962 / bhl. title. 51710" title="Zoology of Egypt: the Fishes of the Nile" type="book" year="1907">Boulenger 1907</bibRefCitation>
)
</taxonomicName>
; a maximum caudal peduncle depth which is ³⁄₅ of the head length vs ²⁄
<quantity id="4CE39B1DF85EF310FF3FD5FB5C1ACEF5" box="[241,275,725,751]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="15" pageNumber="16" unit="in" value="5.0">₅ in</quantity>
<taxonomicName id="4C1B4D7BF85EF310FED7D5F85FB8CEEA" authority="(Pfeffer, 1896)" baseAuthorityName="Pfeffer" baseAuthorityYear="1896" box="[281,689,725,752]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="quadripunctatus">
<emphasis id="B96FEAEAF85EF310FED7D5F85CF2CEF5" box="[281,507,725,751]" italics="true" pageId="15" pageNumber="16">E. quadripunctatus</emphasis>
(Pfeffer, 1896)
</taxonomicName>
.
<taxonomicName id="4C1B4D7BF85EF310FD75D5F85E9ECEF5" box="[699,919,725,751]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85EF310FD75D5F85E9ECEF5" box="[699,919,725,751]" italics="true" pageId="15" pageNumber="16">Enteromius alberti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1B4D7BF85EF310FBE1D5F859AECEF5" baseAuthorityName="Rüppell" baseAuthorityYear="1835" box="[1071,1191,726,751]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="perince">
<emphasis id="B96FEAEAF85EF310FBE1D5F859AECEF5" box="[1071,1191,726,751]" italics="true" pageId="15" pageNumber="16">E. perince</emphasis>
</taxonomicName>
by a combination of a smaller body depth [21.731.2% SL (mean 26.9) vs 33.837.5 (35.0)], a smaller minimum caudal peduncle depth [10.013.6% SL (mean 12.0) vs 15.016.4 (15.5)], and a smaller maximum caudal peduncle depth [11.815.5% SL (mean 13.6) vs 17.719.4 (18.8)].
<taxonomicName id="4C1B4D7BF85EF310FC37D41159D2CF43" box="[1017,1243,831,857]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85EF310FC37D41159D2CF43" box="[1017,1243,831,857]" italics="true" pageId="15" pageNumber="16">Enteromius alberti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1B4D7BF85EF310FF70D44D5C85CF66" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[190,396,867,892]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="stigmatopygus">
<emphasis id="B96FEAEAF85EF310FF70D44D5C85CF66" box="[190,396,867,892]" italics="true" pageId="15" pageNumber="16">E. stigmatopygus</emphasis>
</taxonomicName>
by a combination of a higher number of lateral line scales [2734 (median 31) vs 2025 (21)], a smaller pre-dorsal distance [45.751.3% SL (mean 49.0) vs 51.955.8 (53.8)] and a smaller pre-occipital distance [16.6-21.8% SL (mean 18.6) vs 22.127.3 (24.0)].
<taxonomicName id="4C1B4D7BF85EF310FB59D487587ECFD8" box="[1175,1399,936,962]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85EF310FB59D487587ECFD8" box="[1175,1399,936,962]" italics="true" pageId="15" pageNumber="16">Enteromius alberti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1B4D7BF85EF310FE99D4E25CCECFFF" baseAuthorityName="Boulenger" baseAuthorityYear="1912" box="[343,455,972,997]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="mimus">
<emphasis id="B96FEAEAF85EF310FE99D4E25CCECFFF" box="[343,455,972,997]" italics="true" pageId="15" pageNumber="16">E. mimus</emphasis>
</taxonomicName>
by a higher number of lateral line scales [2734 (median 31) vs 2427 (25)], a smaller pre-anal distance [65.173.0% SL (average 69.3) vs 68.474.8 (72.4)], a larger post-anal distance [16.324.2% SL (average 21.1) vs 16.319.3 (17.8)], a smaller body depth [21.731.2% SL (average 26.9) vs 26.534.3 (30.1)], a smaller head depth [15.119.0% SL (average 17.3) vs 17.821.8 (20.2)] and a smaller pelvic fin length [15.020.9% SL (average 18.0) vs 18.222.4 (20.4)] (
<tableCitation id="C6990343F85EF310FAC2D3765860C868" box="[1292,1385,1112,1138]" captionStart="Table 2" captionStartId="12.[179,248,531,559]" captionText="Table 2 (continued on the next page). Measurements and meristics of Enteromius alberti (Poll, 1939) (n = 47, *= 46, ** = 45), the types of Enteromius mimus (Boulenger, 1912) (n = 9), and Enteromius cf. mimus (n = 76, * = 75, ** = 74, *** = 73)." pageId="15" pageNumber="16">Table 2</tableCitation>
). Specimens of
<taxonomicName id="4C1B4D7BF85EF310FEA8D3525CDFC88F" box="[358,470,1147,1173]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85EF310FEA8D3525CDFC88F" box="[358,470,1147,1173]" italics="true" pageId="15" pageNumber="16">E. alberti</emphasis>
</taxonomicName>
differ from the population of
<taxonomicName id="4C1B4D7BF85EF310FCF5D3525EC6C88F" baseAuthorityName="Boulenger" baseAuthorityYear="1912" box="[827,975,1147,1173]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="mimus">
<emphasis id="B96FEAEAF85EF310FCF5D3525E5CC88F" box="[827,853,1148,1173]" italics="true" pageId="15" pageNumber="16">E.</emphasis>
cf.
<emphasis id="B96FEAEAF85EF310FC4CD3525EC6C88F" box="[898,975,1148,1173]" italics="true" pageId="15" pageNumber="16">mimus</emphasis>
</taxonomicName>
from the Lake Edward system by a smaller head depth. In general, a specimen with a head depth smaller than 19% of the standard length can be assigned to
<taxonomicName id="4C1B4D7BF85EF310FE69D3ED5F10C8C6" box="[423,537,1218,1244]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85EF310FE69D3ED5F10C8C6" box="[423,537,1218,1244]" italics="true" pageId="15" pageNumber="16">E. alberti</emphasis>
</taxonomicName>
(only one specimen had a higher value). Specimens with a head depth larger than 19% of the standard length can be assigned to
<taxonomicName id="4C1B4D7BF85EF310FCA8D3C85EF0C8E5" baseAuthorityName="Boulenger" baseAuthorityYear="1912" box="[870,1017,1253,1279]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="mimus">
<emphasis id="B96FEAEAF85EF310FCA8D3C85E89C8E5" box="[870,896,1254,1279]" italics="true" pageId="15" pageNumber="16">E.</emphasis>
cf.
<emphasis id="B96FEAEAF85EF310FC62D3C85EF0C8E5" box="[940,1017,1254,1279]" italics="true" pageId="15" pageNumber="16">mimus</emphasis>
</taxonomicName>
(only one specimen had a lower value). Other characters to distinguish
<taxonomicName id="4C1B4D7BF85EF310FD4BD2275FFDC938" box="[645,756,1288,1314]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85EF310FD4BD2275FFDC938" box="[645,756,1288,1314]" italics="true" pageId="15" pageNumber="16">E. alberti</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1B4D7BF85EF310FCF4D2275EC5C938" baseAuthorityName="Boulenger" baseAuthorityYear="1912" box="[826,972,1288,1314]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="mimus">
<emphasis id="B96FEAEAF85EF310FCF4D2275E5DC938" box="[826,852,1289,1314]" italics="true" pageId="15" pageNumber="16">E.</emphasis>
cf.
<emphasis id="B96FEAEAF85EF310FCB1D2275EC5C938" box="[895,972,1289,1314]" italics="true" pageId="15" pageNumber="16">mimus</emphasis>
</taxonomicName>
are the higher number of lateral line scales [2734 (median 31) vs 2131 (26)], the smaller interorbital width [5.88.3% SL (mean 7.3) vs 6.99.9 (8.7)], the smaller pre-pelvic distance [44.853.3% SL (mean 49,2) vs 48.455.4 (52.1)], the smaller body depth [21.731.2% SL (mean 26.9) vs 23.535.8 (31.0)], the smaller maximum [11.8 15.5% SL (mean 13.6) vs 8.719.0 (16)] and minimum [10.013.6% SL (mean 12) vs 7.115.9 (13.7)] caudal peduncle depth, and the smaller head width [10.414.2% SL (mean 12) vs 11.614.5 (13.2)] (
<tableCitation id="C6990343F85EF310FF08D2F55C2BC9EF" box="[198,290,1499,1525]" captionStart="Table 2" captionStartId="12.[179,248,531,559]" captionText="Table 2 (continued on the next page). Measurements and meristics of Enteromius alberti (Poll, 1939) (n = 47, *= 46, ** = 45), the types of Enteromius mimus (Boulenger, 1912) (n = 9), and Enteromius cf. mimus (n = 76, * = 75, ** = 74, *** = 73)." pageId="15" pageNumber="16">Table 2</tableCitation>
,
<figureCitation id="13202A7DF85EF310FEE0D2F55C7FC9EF" box="[302,374,1499,1525]" captionStart="Fig" captionStartId="8.[189,232,1849,1875]" captionTargetBox="[189,1398,274,1802]" captionTargetId="figure@8.[189,1398,274,1802]" captionTargetPageId="8" captionText="Fig. 3. Scatterplots of (a) interorbital width (IOW), (b) pre-pelvic distance (PrPelD), (c) body depth (BD), (d) maximum caudal peduncle depth (MxCPD), (e) minimum caudal peduncle depth (MnCPD), (f) head width (HW), and (g) head depth (HD) in % SL against SL (in mm) on 72 specimens of Enteromius Cope, 1867. The genetic groups A (●) (n = 22) and B (■) (n = 50) are indicated separately. The dashed lines indicate the size class used for the MWU tests." figureDoi="http://doi.org/10.5281/zenodo.3981075" httpUri="https://zenodo.org/record/3981075/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF646670F85FF311FF73D08A594ECBFB" ID-DOI="http://doi.org/10.5281/zenodo.3981083" ID-Zenodo-Dep="3981083" httpUri="https://zenodo.org/record/3981083/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1956,1982]" targetBox="[189,1398,1090,1915]" targetPageId="14">
<paragraph id="8BA436F8F85FF311FF73D08A594ECBFB" blockId="14.[189,1398,1956,2018]" pageId="14" pageNumber="15">
<emphasis id="B96FEAEAF85FF311FF73D08A5C31CBA4" bold="true" box="[189,312,1956,1982]" pageId="14" pageNumber="15">Fig. 7. (a)</emphasis>
Lectotype of
<taxonomicName id="4C1B4D7BF85FF311FE2FD08B5E5CCBA4" authority="(Poll, 1939)" baseAuthorityName="Poll" baseAuthorityYear="1939" box="[481,853,1956,1982]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85FF311FE2FD08B5FB7CBA4" box="[481,702,1956,1982]" italics="true" pageId="14" pageNumber="15">Enteromius alberti</emphasis>
(Poll, 1939)
</taxonomicName>
(MRAC 64723) with 74.3 mm SL.
<emphasis id="B96FEAEAF85FF311FAC6D08A5825CBA4" bold="true" box="[1288,1324,1956,1982]" pageId="14" pageNumber="15">(b)</emphasis>
Fresh specimen of
<taxonomicName id="4C1B4D7BF85FF311FE9DD0E65CC8CBFB" box="[339,449,1991,2017]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85FF311FE9DD0E65CC8CBFB" box="[339,449,1991,2017]" italics="true" pageId="14" pageNumber="15">E. alberti</emphasis>
</taxonomicName>
(RMCA 2018.008.P.0248 HP 3246) with 62.4 mm SL.
</paragraph>
</caption>
<subSubSection id="C3016573F85EF310FF70D1335C3CCAB6" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BA436F8F85EF310FF70D1335C4CCA2D" blockId="15.[190,325,1565,1591]" box="[190,325,1565,1591]" pageId="15" pageNumber="16">
<heading id="D0EC8194F85EF310FF70D1335C4CCA2D" bold="true" box="[190,325,1565,1591]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96FEAEAF85EF310FF70D1335C4CCA2D" bold="true" box="[190,325,1565,1591]" pageId="15" pageNumber="16">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436F8F85EF310FF70D1655C3CCAB6" blockId="15.[190,1399,1611,1708]" pageId="15" pageNumber="16">
Etymology not explained in the original description. Probably, the species epithet
<taxonomicName id="4C1B4D7BF85EF310FBBCD16559C9CA7F" box="[1138,1216,1611,1637]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF85EF310FBBCD16559C9CA7F" box="[1138,1216,1611,1637]" italics="true" pageId="15" pageNumber="16">alberti</emphasis>
</taxonomicName>
is derived from the name of the former Parc National Albert (now Virunga National Park) in which the
<typeStatus id="54A0885AF85EF310FB2AD141581ECA93" box="[1252,1303,1647,1673]" pageId="15" pageNumber="16">type</typeStatus>
locality is located.
</paragraph>
</subSubSection>
<subSubSection id="C3016573F85EF30FFF70D1FD593FC9EB" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BA436F8F85EF310FF70D1FD5CA2CAF7" blockId="15.[190,427,1747,1773]" box="[190,427,1747,1773]" pageId="15" pageNumber="16">
<heading id="D0EC8194F85EF310FF70D1FD5CA2CAF7" bold="true" box="[190,427,1747,1773]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96FEAEAF85EF310FF70D1FD5CA2CAF7" bold="true" box="[190,427,1747,1773]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436F8F85EF310FF25D02C5F30CB07" blockId="15.[190,1399,1794,1891]" box="[235,569,1794,1821]" pageId="15" pageNumber="16">
<emphasis id="B96FEAEAF85EF310FF25D02C5C6ECB06" bold="true" box="[235,359,1794,1820]" pageId="15" pageNumber="16">
<typeStatus id="54A0885AF85EF310FF25D02C5C6ECB06" box="[235,359,1794,1820]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8BA436F8F85EF310FF70D00B5ECCCB79" blockId="15.[190,1399,1794,1891]" pageId="15" pageNumber="16">
<materialsCitation id="3B733CA5F85EF310FF70D00B5ECCCB79" ID-GBIF-Occurrence="2830976304" collectingDate="1934-11-05" collectingDateMax="1934-11-11" collectingDateMin="1934-11-05" collectionCode="SM" collectorName="de Witte" country="Democratic Republic of the Congo" location="Rutshuru River" pageId="15" pageNumber="16" specimenCode="MRAC 64723" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="F30C7668F85EF310FF70D00B5FFACB25" box="[190,755,1829,1855]" name="Democratic Republic of the Congo" pageId="15" pageNumber="16">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
<location id="8EC46023F85EF310FCC1D00B5ECECB25" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F85EF310FCC1D00B5ECECB25" box="[783,967,1829,1855]" country="Democratic Republic of the Congo" name="Rutshuru River" pageId="15" pageNumber="16">Rutshuru River</location>
,
<location id="8EC46023F85EF310FC1AD00B5989CB25" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F85EF310FC1AD00B5989CB25" box="[980,1152,1829,1855]" country="Democratic Republic of the Congo" name="May-Ya-Moto" pageId="15" pageNumber="16">May-Ya-Moto</location>
;
<date id="FFA51038F85EF310FB43D00B5847CB25" box="[1165,1358,1829,1856]" pageId="15" pageNumber="16" value="1934-11-05" valueMax="1934-11-11" valueMin="1934-11-05">
<collectingDate id="EFE1E9D0F85EF310FB43D00B5847CB25" box="[1165,1358,1829,1856]" pageId="15" pageNumber="16" value="1934-11-05" valueMax="1934-11-11" valueMin="1934-11-05">511 Nov. 1934</collectingDate>
</date>
;
<collectorName id="26EE532EF85EF310FA95D00B5DF7CB78" pageId="15" pageNumber="16">de Witte</collectorName>
leg.;
<specimenCode id="DBBD9E83F85EF310FE8ED0665CEFCB78" box="[320,486,1864,1890]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="15" pageNumber="16" type="Museum">MRAC 64723</specimenCode>
(Supplementary file SM.01: Fiche-209).
</materialsCitation>
</paragraph>
<paragraph id="8BA436F8F85EF310FF25D0A45C91CBBE" blockId="15.[190,1399,1930,2027]" box="[235,408,1930,1956]" pageId="15" pageNumber="16">
<emphasis id="B96FEAEAF85EF310FF25D0A45C91CBBE" bold="true" box="[235,408,1930,1956]" pageId="15" pageNumber="16">
<typeStatus id="54A0885AF85EF310FF25D0A45C91CBBE" box="[235,408,1930,1956]" pageId="15" pageNumber="16" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA436F8F85EF310FF70D0835FECCBF1" blockId="15.[190,1399,1930,2027]" pageId="15" pageNumber="16">
<materialsCitation id="3B733CA5F85EF310FF70D0835FE8CBF1" ID-GBIF-Occurrence="2830976307" collectingDate="1934-11-05" collectingDateMax="1934-11-11" collectingDateMin="1934-11-05" collectionCode="SM" collectorName="de Witte" country="Democratic Republic of the Congo" location="DEMOCRATIC REPUBLIC OF THE CONGO" pageId="15" pageNumber="16" specimenCode="MRAC 64722" specimenCount="1" typeStatus="paralectotype">
<collectingCountry id="F30C7668F85EF310FF70D0835FE4CBDD" box="[190,749,1965,1991]" name="Democratic Republic of the Congo" pageId="15" pageNumber="16">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
<specimenCount id="9D1DFD71F85EF310FCC8D0835E52CBDD" box="[774,859,1965,1991]" pageId="15" pageNumber="16" type="generic">1 spec.</specimenCount>
; same collection data as for lectotype;
<specimenCode id="DBBD9E83F85EF310FAEFD0835C0ECBF0" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="15" pageNumber="16" type="Museum">MRAC 64722</specimenCode>
(Supplementary file SM.01: Fiche-210)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA436F8F841F30FFF24D6255F2FCD3F" blockId="16.[189,1399,267,435]" box="[234,550,267,293]" pageId="16" pageNumber="17">
<emphasis id="B96FEAEAF841F30FFF24D6255F2FCD3F" bold="true" box="[234,550,267,293]" pageId="16" pageNumber="17">Other material examined</emphasis>
</paragraph>
<paragraph id="8BA436F8F841F30FFF73D6005C4CCDA9" blockId="16.[189,1399,267,435]" pageId="16" pageNumber="17">
<materialsCitation id="3B733CA5F841F30FFF73D6005F4BCD52" ID-GBIF-Occurrence="2830976306" box="[189,578,302,329]" country="Kenya" location="Lake Victoria" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Lake Victoria">
<collectingCountry id="F30C7668F841F30FFF73D6005C16CD52" box="[189,287,302,328]" name="Kenya" pageId="16" pageNumber="17">KENYA</collectingCountry>
<emphasis id="B96FEAEAF841F30FFE8AD6005F4BCD52" bold="true" box="[324,578,302,328]" pageId="16" pageNumber="17">
<location id="8EC46023F841F30FFE8AD6005CFDCD52" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFE8AD6005CFDCD52" box="[324,500,302,328]" country="Kenya" name="Lake Victoria" pageId="16" pageNumber="17" stateProvince="Lake Victoria">Lake Victoria</location>
basin
</emphasis>
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFDACD6005F67CD71" ID-GBIF-Occurrence="2830976316" collectionCode="BMNH" country="Kenya" county="Luambwa" location="Nyanza Province" municipality="Nzoia River" pageId="16" pageNumber="17" specimenCode="BMNH 1960.6.7.37" specimenCount="1" stateProvince="Lake Victoria" typeStatus="holotype">
<specimenCount id="9D1DFD71F841F30FFDACD6005FB3CD53" box="[610,698,302,329]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
,
<typeStatus id="54A0885AF841F30FFD04D6005E38CD52" box="[714,817,302,328]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1B4D7BF841F30FFC91D6015ED7CD52" baseAuthorityName="Whitehead" baseAuthorityYear="1960" box="[863,990,303,328]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="cercops">
<emphasis id="B96FEAEAF841F30FFC91D6015ED7CD52" box="[863,990,303,328]" italics="true" pageId="16" pageNumber="17">E. cercops</emphasis>
</taxonomicName>
;
<collectingCounty id="62C54E74F841F30FFC3FD6005960CD52" box="[1009,1129,302,328]" pageId="16" pageNumber="17">Luambwa</collectingCounty>
,
<collectingMunicipality id="6BC0AC82F841F30FFBB7D6005804CD52" box="[1145,1293,302,328]" pageId="16" pageNumber="17">Nzoia River</collectingMunicipality>
,
<location id="8EC46023F841F30FFAD3D6015C20CD71" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFAD3D6015C20CD71" country="Kenya" county="Luambwa" municipality="Nzoia River" name="Nyanza Province" pageId="16" pageNumber="17" stateProvince="Lake Victoria">Nyanza Province</location>
; 1960;
<specimenCode id="DBBD9E83F841F30FFE4CD67C5F67CD71" box="[386,622,337,364]" pageId="16" pageNumber="17">BMNH 1960.6.7.37</specimenCode>
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFD45D67C5C48CDA9" ID-GBIF-Occurrence="2830976319" collectionCode="SM" country="Kenya" county="Luambwa" location="Nyanza Province" municipality="Nzoia River" pageId="16" pageNumber="17" specimenCode="BMNH 1960.6.7.38, 1960.6.7.50" specimenCount="7" stateProvince="Lake Victoria" typeStatus="paratype">
<specimenCount id="9D1DFD71F841F30FFD45D67C5FEBCD76" box="[651,738,338,364]" pageId="16" pageNumber="17">7 specs</specimenCount>
[of
<quantity id="4CE39B1DF841F30FFCD8D67F5E5ACD71" box="[790,851,337,363]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="16" pageNumber="17" unit="in" value="13.0">13 in</quantity>
lot],
<typeStatus id="54A0885AF841F30FFC5BD67C590CCD76" box="[917,1029,338,364]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C1B4D7BF841F30FFBFED67C59A5CD71" baseAuthorityName="Whitehead" baseAuthorityYear="1960" box="[1072,1196,338,363]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="cercops">
<emphasis id="B96FEAEAF841F30FFBFED67C59A5CD71" box="[1072,1196,338,363]" italics="true" pageId="16" pageNumber="17">E. cercops</emphasis>
</taxonomicName>
; same collection data as for holotype;
<specimenCode id="DBBD9E83F841F30FFE07D65B5FB1CD94" box="[457,696,372,399]" pageId="16" pageNumber="17">BMNH 1960.6.7.38</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFD26D65A5E7ACD95" box="[744,883,372,399]" pageId="16" pageNumber="17">1960.6.7.50</specimenCode>
(Supplementary file SM.01: Fiche-177 to Fiche-183)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA436F8F841F30FFF73D6F7593FC9EB" blockId="16.[189,1399,472,1521]" pageId="16" pageNumber="17">
<materialsCitation id="3B733CA5F841F30FFF73D6F75E3DCE20" ID-GBIF-Occurrence="2830976317" collectingDate="2016-11-08" collectionCode="HIPE, MT, SM" collectorName="GenBank &amp; Supplementary" country="Uganda" latitude="-0.45855555" location="Nchwera River" longLatPrecision="1" longitude="29.801943" municipality="Nchwera River" pageId="16" pageNumber="17" specimenCode="RMCA 2016.035.P.0148, 2016.035.P.0149" specimenCount="1" stateProvince="Lake Edward">
<collectingCountry id="F30C7668F841F30FFF73D6F75C33CDE9" box="[189,314,473,499]" name="Uganda" pageId="16" pageNumber="17">UGANDA</collectingCountry>
<emphasis id="B96FEAEAF841F30FFEA7D6F75F3ECDE9" bold="true" box="[361,567,473,499]" pageId="16" pageNumber="17">
<collectingRegion id="49DFF81AF841F30FFEA7D6F75F14CDE9" box="[361,541,473,499]" pageId="16" pageNumber="17">Lake Edward</collectingRegion>
</emphasis>
<specimenCount id="9D1DFD71F841F30FFD89D6F75FA8CDEE" box="[583,673,473,500]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
[of
<quantity id="4CE39B1DF841F30FFD2DD6F75E11CDE9" box="[739,792,473,499]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
lot];
<location id="8EC46023F841F30FFCA6D6F7592CCDE9" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFCA6D6F7592CCDE9" box="[872,1061,473,499]" country="Uganda" latitude="-0.45855555" longLatPrecision="1" longitude="29.801943" municipality="Nchwera River" name="Nchwera River" pageId="16" pageNumber="17" stateProvince="Lake Edward">Nchwera River</location>
,
<location id="8EC46023F841F30FFBF7D6F75823CDE9" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFBF7D6F75823CDE9" box="[1081,1322,473,499]" country="Uganda" latitude="-0.45855555" longLatPrecision="1" longitude="29.801943" municipality="Nchwera River" name="Rwenshama-Ishasha" pageId="16" pageNumber="17" stateProvince="Lake Edward">Rwenshama-Ishasha</location>
road;
<geoCoordinate id="EE2F503FF841F30FFF73D6D45C46CE0C" box="[189,335,506,534]" degrees="0" direction="south" minutes="27" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="30.8" value="-0.45855555">0°27ʹ30.8ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFE95D6D45CF7CE0D" box="[347,510,506,535]" degrees="29" direction="east" minutes="48" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="07.0" value="29.801943">29°48ʹ07.0ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFDC4D6D25F92CE0C" box="[522,667,508,535]" pageId="16" pageNumber="17" value="2016-11-08">
<collectingDate id="EFE1E9D0F841F30FFDC4D6D25F92CE0C" box="[522,667,508,535]" pageId="16" pageNumber="17" value="2016-11-08">8 Nov. 2016</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFC09D6D25945CE0D" box="[967,1100,508,535]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251114" pageId="16" pageNumber="17">MT251114</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFB99D6D2587ECE0D" box="[1111,1399,508,535]" pageId="16" pageNumber="17">RMCA 2016.035.P.0148</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFF12D5315CA9CE20" box="[220,416,543,570]" pageId="16" pageNumber="17">2016.035.P.0149</specimenCode>
(Supplementary file SM.01: E31)
</materialsCitation>
.•
<materialsCitation id="3B733CA5F841F30FFC84D50E5803CE47" ID-GBIF-Occurrence="2830976321" collectingDate="2016-11-08" collectionCode="SM" collectorName="Supplementary" country="Uganda" latitude="-0.45855555" location="Rwenshama-Ishasha" longLatPrecision="1" longitude="29.801943" municipality="Nchwera River" pageId="16" pageNumber="17" specimenCode="RMCA 2016.035.P.0171, 2016.035.P.0172" specimenCount="1" stateProvince="Lake Edward">
<specimenCount id="9D1DFD71F841F30FFC84D50E5E95CE20" box="[842,924,544,570]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
[of
<quantity id="4CE39B1DF841F30FFC00D50E5EF2CE23" box="[974,1019,543,570]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
lot]; same collection data as for preceding;
<specimenCode id="DBBD9E83F841F30FFEF1D56D5F69CE47" box="[319,608,578,605]" pageId="16" pageNumber="17">RMCA 2016.035.P.0171</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFD4AD56C5E41CE47" box="[644,840,578,605]" pageId="16" pageNumber="17">2016.035.P.0172</specimenCode>
(Supplementary file SM.01: HP1028)
</materialsCitation>
.•
<materialsCitation id="3B733CA5F841F30FFAD3D56D5957CEBE" ID-GBIF-Occurrence="2830976313" collectingDate="2018-01-20" collectionCode="HIPE, MT, SM" collectorName="GenBank &amp; Supplementary" country="Uganda" latitude="-0.092" location="Lake Edward" longLatPrecision="1" longitude="29.758417" municipality="Kayanja" pageId="16" pageNumber="17" specimenCode="RMCA 2018.008.P.0247" specimenCount="1" stateProvince="Lake Edward">
<specimenCount id="9D1DFD71F841F30FFAD3D56D587BCE47" box="[1309,1394,579,605]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="6BC0AC82F841F30FFF73D5485C2BCE9A" box="[189,290,614,640]" pageId="16" pageNumber="17">Kayanja</collectingMunicipality>
, offshore,
<location id="8EC46023F841F30FFE6FD5485F4DCE9A" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFE6FD5485F4DCE9A" box="[417,580,614,640]" country="Uganda" latitude="-0.092" longLatPrecision="1" longitude="29.758417" municipality="Kayanja" name="Lake Edward" pageId="16" pageNumber="17" stateProvince="Lake Edward">Lake Edward</location>
;
<geoCoordinate id="EE2F503FF841F30FFD9CD54A5FEECE9A" box="[594,743,612,640]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="31.2" value="-0.092">0°05ʹ31.2ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFD38D54A5E94CE9A" box="[758,925,612,640]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="30.3" value="29.758417">29°45ʹ30.3ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFC65D5485941CE9A" box="[939,1096,614,640]" pageId="16" pageNumber="17" value="2018-01-20">
<collectingDate id="EFE1E9D0F841F30FFC65D5485941CE9A" box="[939,1096,614,640]" pageId="16" pageNumber="17" value="2018-01-20">20 Jan. 2018</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFF73D5A75C4DCEBE" box="[189,324,649,676]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251133" pageId="16" pageNumber="17">MT251133</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFE98D5A75F77CEB9" box="[342,638,649,675]" pageId="16" pageNumber="17">RMCA 2018.008.P.0247</specimenCode>
(Supplementary file SM.01: HP2599)
</materialsCitation>
.•
<materialsCitation id="3B733CA5F841F30FFBB4D5A75E9DCEF0" ID-GBIF-Occurrence="2830976301" collectingDate="2018-01-31" collectionCode="HIPE, SM" collectorName="Supplementary" country="Uganda" latitude="-0.31180558" location="Kisenji" longLatPrecision="1" longitude="29.86" municipality="Nyamweru River" pageId="16" pageNumber="17" specimenCode="RBINS 25585, 25586" specimenCount="2" stateProvince="Lake Edward">
<specimenCount id="9D1DFD71F841F30FFBB4D5A759D4CEBE" box="[1146,1245,649,676]" pageId="16" pageNumber="17">2 specs</specimenCount>
.;
<collectingMunicipality id="6BC0AC82F841F30FFB3AD5A45C09CEDC" pageId="16" pageNumber="17">Nyamweru River</collectingMunicipality>
, mouth,
<location id="8EC46023F841F30FFEBBD5825CC7CEDC" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFEBBD5825CC7CEDC" box="[373,462,684,710]" country="Uganda" latitude="-0.31180558" longLatPrecision="1" longitude="29.86" municipality="Nyamweru River" name="Kisenji" pageId="16" pageNumber="17" stateProvince="Lake Edward">Kisenji</location>
;
<geoCoordinate id="EE2F503FF841F30FFE2FD5845F77CEDC" box="[481,638,682,710]" degrees="0" direction="south" minutes="18" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="42.5" value="-0.31180558">0°1842.5″ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFD5DD5845E48CEDD" box="[659,833,682,711]" degrees="29" direction="east" minutes="51" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="36.0" value="29.86">29°5136.0″ E</geoCoordinate>
;
<date id="FFA51038F841F30FFC9BD5825EF7CEDC" box="[853,1022,684,711]" pageId="16" pageNumber="17" value="2018-01-31">
<collectingDate id="EFE1E9D0F841F30FFC9BD5825EF7CEDC" box="[853,1022,684,711]" pageId="16" pageNumber="17" value="2018-01-31">31 Jan. 2018</collectingDate>
</date>
; HIPE exped.;
<specimenCode id="DBBD9E83F841F30FFB09D582587FCEDD" box="[1223,1398,684,711]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">RBINS 25585</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFF11D5E15C21CEF0" box="[223,296,719,746]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">25586</specimenCode>
(Supplementary file SM.01: Fiche-118, Fiche-119)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFC77D5FE5928CF4E" ID-GBIF-Occurrence="2830976310" collectingDate="2016-10-31" collectionCode="HIPE, SM" collectorName="Supplementary" country="Uganda" county="River" latitude="-0.096" location="Lake George" longLatPrecision="1" longitude="30.681" municipality="Lake George" pageId="16" pageNumber="17" specimenCode="RMCA 2016.035.P.0159, 2016.035.P.0163" specimenCount="5" stateProvince="Lake George">
<emphasis id="B96FEAEAF841F30FFC77D5FE5957CEF3" bold="true" box="[953,1118,719,746]" pageId="16" pageNumber="17">
<collectingMunicipality id="6BC0AC82F841F30FFC77D5FE5957CEF3" box="[953,1118,719,746]" pageId="16" pageNumber="17">Lake George</collectingMunicipality>
</emphasis>
<specimenCount id="9D1DFD71F841F30FFBB1D5E159D2CEF0" box="[1151,1243,719,746]" pageId="16" pageNumber="17">5 specs</specimenCount>
;
<locationDeviation id="9A162FDAF841F30FFB25D5E15C9CCF17" pageId="16" pageNumber="17">
River
<quantity id="4CE39B1DF841F30FFAF9D5FE587FCEF3" box="[1335,1398,719,746]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="km" value="2.0">2 km</quantity>
east of Kitongore
</locationDeviation>
;
<geoCoordinate id="EE2F503FF841F30FFE6BD5DF5F35CF17" box="[421,572,753,781]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="45.6" value="-0.096">0°05ʹ45.6ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFD83D5DF5FFFCF17" box="[589,758,753,781]" degrees="30" direction="east" minutes="40" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="51.6" value="30.681">30°40ʹ51.6ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFCC9D5DD5EA2CF17" box="[775,939,755,781]" pageId="16" pageNumber="17" value="2016-10-31">
<collectingDate id="EFE1E9D0F841F30FFCC9D5DD5EA2CF17" box="[775,939,755,781]" pageId="16" pageNumber="17" value="2016-10-31">31 Oct. 2016</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFB22D5DC587BCF17" box="[1260,1394,754,781]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251105" pageId="16" pageNumber="17">MT251105</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFF73D4385C4ACF2B" box="[189,323,790,817]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251120" pageId="16" pageNumber="17">MT251120</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFE93D4385CEDCF2B" box="[349,484,790,817]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251121" pageId="16" pageNumber="17">MT251121</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFE30D4385F8DCF2B" box="[510,644,790,817]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251106" pageId="16" pageNumber="17">MT251106</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFD50D4385E2CCF2B" box="[670,805,790,817]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251107" pageId="16" pageNumber="17">MT251107</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFCF1D4385966CF2B" box="[831,1135,790,817]" pageId="16" pageNumber="17">RMCA 2016.035.P.0159</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFB7CD438587FCF2B" box="[1202,1398,790,817]" pageId="16" pageNumber="17">2016.035.P.0163</specimenCode>
(Supplementary file SM.01: HP592, HP595, HP596, HP600, HP602)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFB84D417586CCF80" ID-GBIF-Occurrence="2830976302" collectingDate="2016-10-31" collectionCode="HIPE, SM" collectorName="GenBank &amp; Supplementary" country="Uganda" latitude="0.039" location="Mpanga River" longLatPrecision="1" longitude="30.289999" municipality="Lake George" pageId="16" pageNumber="17" specimenCode="RMCA 2016.035.P.0164, 2016.035.P.0165" specimenCount="2" stateProvince="Lake George">
<specimenCount id="9D1DFD71F841F30FFB84D41759A2CF4E" box="[1098,1195,825,852]" pageId="16" pageNumber="17">2 specs</specimenCount>
;
<location id="8EC46023F841F30FFB71D414587BCF49" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFB71D414587BCF49" box="[1215,1394,825,852]" country="Uganda" latitude="0.039" longLatPrecision="1" longitude="30.289999" municipality="Lake George" name="Mpanga River" pageId="16" pageNumber="17" stateProvince="Lake George">Mpanga River</location>
, mouth,
<collectingMunicipality id="6BC0AC82F841F30FFED6D4725CBCCF6C" box="[280,437,860,886]" pageId="16" pageNumber="17">Lake George</collectingMunicipality>
;
<geoCoordinate id="EE2F503FF841F30FFE0FD4745F52CF6D" box="[449,603,858,887]" degrees="0" direction="north" minutes="02" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="20.4" value="0.039">0°02ʹ20.4ʺ N</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFDA6D4745E07CF6D" box="[616,782,858,887]" degrees="30" direction="east" minutes="17" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="24.0" value="30.289999">30°17ʹ24.0ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFCD5D4725EB0CF6C" box="[795,953,860,886]" pageId="16" pageNumber="17" value="2016-10-31">
<collectingDate id="EFE1E9D0F841F30FFCD5D4725EB0CF6C" box="[795,953,860,886]" pageId="16" pageNumber="17" value="2016-10-31">31 Oct. 2016</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFB23D472587BCF6D" box="[1261,1394,860,887]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251119" pageId="16" pageNumber="17">MT251119</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFF73D4515C4DCF80" box="[189,324,895,922]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251104" pageId="16" pageNumber="17">MT251104</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFE81D4AE5F67CF80" box="[335,622,895,922]" pageId="16" pageNumber="17">RMCA 2016.035.P.0164</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFD5DD4515E5ECF80" box="[659,855,895,922]" pageId="16" pageNumber="17">2016.035.P.0165</specimenCode>
(Supplementary file SM.01: HP575, HP579)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFF73D48D59B3CFFA" ID-GBIF-Occurrence="2830976311" collectingDate="2017-04-01" collectionCode="HIPE, MT, SM" collectorName="GenBank &amp; Supplementary" country="Uganda" latitude="0.35005555" location="Mahoma River" longLatPrecision="1" longitude="30.278055" municipality="Mahoma River" pageId="16" pageNumber="17" specimenCode="RMCA 2017.006.P.0244" specimenCount="1" stateProvince="Lake George">
<specimenCount id="9D1DFD71F841F30FFF73D48D5C1BCFA7" box="[189,274,931,957]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
;
<location id="8EC46023F841F30FFED1D48D5CDBCFA7" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFED1D48D5CDBCFA7" box="[287,466,931,957]" country="Uganda" latitude="0.35005555" longLatPrecision="1" longitude="30.278055" municipality="Mahoma River" name="Mahoma River" pageId="16" pageNumber="17" stateProvince="Lake George">Mahoma River</location>
,
<locationDeviation id="9A162FDAF841F30FFE10D48D5FD3CFA7" box="[478,730,931,957]" pageId="16" pageNumber="17">Rwimi-Nyabani road</locationDeviation>
;
<geoCoordinate id="EE2F503FF841F30FFD28D48F5E89CFA7" box="[742,896,929,957]" degrees="0" direction="north" minutes="21" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="00.2" value="0.35005555">0°21ʹ00.2ʺ N</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFC42D48F5938CFA7" box="[908,1073,929,957]" degrees="30" direction="east" minutes="16" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="41.0" value="30.278055">30°16ʹ41.0ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFBF0D48D59C2CFA7" box="[1086,1227,931,957]" pageId="16" pageNumber="17" value="2017-04-01">
<collectingDate id="EFE1E9D0F841F30FFBF0D48D59C2CFA7" box="[1086,1227,931,957]" pageId="16" pageNumber="17" value="2017-04-01">1 Apr. 2017</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFEF3D4E85CCBCFFB" box="[317,450,966,993]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251110" pageId="16" pageNumber="17">MT251110</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFE1ED4E85FFACFFA" box="[464,755,966,992]" pageId="16" pageNumber="17">RMCA 2017.006.P.0244</specimenCode>
(Supplementary file SM.01: HP1878)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFB19D4E8596DC83D" ID-GBIF-Occurrence="2830976315" collectingDate="2018-02-02" collectionCode="HIPE, MT, SM" collectorName="GenBank &amp; Supplementary" country="Uganda" latitude="-0.08461111" location="Kashaka Bay" longLatPrecision="1" longitude="30.179333" municipality="Lake George" pageId="16" pageNumber="17" specimenCode="RMCA 2018.008.P.0248" specimenCount="1" stateProvince="Lake George">
<specimenCount id="9D1DFD71F841F30FFB19D4E85827CFFA" box="[1239,1326,966,992]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="6BC0AC82F841F30FFAF2D4E85C1FC819" pageId="16" pageNumber="17">Lake George</collectingMunicipality>
,
<location id="8EC46023F841F30FFEEFD4C75CB4C81E" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFEEFD4C75CB4C81E" box="[289,445,1001,1028]" country="Uganda" latitude="-0.08461111" longLatPrecision="1" longitude="30.179333" municipality="Lake George" name="Kashaka Bay" pageId="16" pageNumber="17" stateProvince="Lake George">Kashaka Bay</location>
, south of inlet;
<geoCoordinate id="EE2F503FF841F30FFDBCD4C95E0DC819" box="[626,772,999,1027]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="04.6" value="-0.08461111">0°05ʹ04.6ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFCDFD4C95EBCC81E" box="[785,949,999,1028]" degrees="30" direction="east" minutes="10" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="45.6" value="30.179333">30°10ʹ45.6ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFC0FD4C75946C819" box="[961,1103,1001,1027]" pageId="16" pageNumber="17" value="2018-02-02">
<collectingDate id="EFE1E9D0F841F30FFC0FD4C75946C819" box="[961,1103,1001,1027]" pageId="16" pageNumber="17" value="2018-02-02">2 Feb. 2018</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFF73D3225C4BC83D" box="[189,322,1036,1063]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251111" pageId="16" pageNumber="17">MT251111</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFE98D3225F89C83C" box="[342,640,1036,1062]" pageId="16" pageNumber="17">RMCA 2018.008.P.0248</specimenCode>
(Supplementary file SM.01: HP3246)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFB5AD322599FC88B" ID-GBIF-Occurrence="2830976312" collectingDate="2016-10-30" collectionCode="HIPE, SM" collectorName="GenBank &amp; Supplementary" country="Uganda" latitude="-0.123833336" location="Kazinga Channel" longLatPrecision="1" longitude="30.052082" municipality="Kyambura River" pageId="16" pageNumber="17" specimenCode="RMCA 2016.035.P.0155, 2016.035.P.0158" specimenCount="4" stateProvince="Kazinga Channel">
<emphasis id="B96FEAEAF841F30FFB5AD322587FC83C" bold="true" box="[1172,1398,1036,1062]" pageId="16" pageNumber="17">
<location id="8EC46023F841F30FFB5AD322587FC83C" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFB5AD322587FC83C" box="[1172,1398,1036,1062]" country="Uganda" latitude="-0.123833336" longLatPrecision="1" longitude="30.052082" municipality="Kyambura River" name="Kazinga Channel" pageId="16" pageNumber="17" stateProvince="Kazinga Channel">Kazinga Channel</location>
</emphasis>
<specimenCount id="9D1DFD71F841F30FFF1AD31E5C3BC850" box="[212,306,1072,1098]" pageId="16" pageNumber="17">4 specs</specimenCount>
;
<location id="8EC46023F841F30FFE8AD3015F06C853" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFE8AD3015F06C853" box="[324,527,1071,1097]" country="Uganda" latitude="-0.123833336" longLatPrecision="1" longitude="30.052082" municipality="Kyambura River" name="Kyambura River" pageId="16" pageNumber="17" stateProvince="Kazinga Channel">Kyambura River</location>
, mouth,
<location id="8EC46023F841F30FFDB1D3015E5DC853" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFDB1D3015E5DC853" box="[639,852,1071,1097]" country="Uganda" latitude="-0.123833336" longLatPrecision="1" longitude="30.052082" municipality="Kyambura River" name="Kazinga Channel" pageId="16" pageNumber="17" stateProvince="Kazinga Channel">Kazinga Channel</location>
;
<geoCoordinate id="EE2F503FF841F30FFCABD3005EF4C850" box="[869,1021,1070,1098]" degrees="0" direction="south" minutes="07" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="25.8" value="-0.123833336">0°07ʹ25.8ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFBC1D30059B0C850" box="[1039,1209,1070,1098]" degrees="30" direction="east" minutes="03" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="07.5" value="30.052082">30°03ʹ07.5ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFB05D31E587BC853" box="[1227,1394,1071,1098]" pageId="16" pageNumber="17" value="2016-10-30">
<collectingDate id="EFE1E9D0F841F30FFB05D31E587BC853" box="[1227,1394,1071,1098]" pageId="16" pageNumber="17" value="2016-10-30">30 Oct. 2016</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFE3CD37C5F71C877" box="[498,632,1106,1133]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251100" pageId="16" pageNumber="17">MT251100</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFD47D37C5E06C877" box="[649,783,1106,1133]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251101" pageId="16" pageNumber="17">MT251101</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFCEFD37C5EAEC877" box="[801,935,1106,1133]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251102" pageId="16" pageNumber="17">MT251102</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFC76D37C5936C877" box="[952,1087,1106,1133]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251103" pageId="16" pageNumber="17">MT251103</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFB9ED37D587FC877" box="[1104,1398,1106,1133]" pageId="16" pageNumber="17">RMCA 2016.035.P.0155</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFF2BD3585CA0C88B" box="[229,425,1142,1169]" pageId="16" pageNumber="17">2016.035.P.0158</specimenCode>
(Supplementary file SM.01: HP562(2), HP563 to HP565)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFB0CD3585C68C8CD" ID-GBIF-Occurrence="2830976303" collectingDate="2018-02-04" collectionCode="HIPE, SM" collectorName="Supplementary" country="Uganda" latitude="-0.123833336" location="Kazinga Channel" longLatPrecision="1" longitude="30.052082" municipality="Kyambura River" pageId="16" pageNumber="17" specimenCode="RBINS 25588, 25594" specimenCount="7" stateProvince="Kazinga Channel">
<specimenCount id="9D1DFD71F841F30FFB0CD358582CC88A" box="[1218,1317,1142,1168]" pageId="16" pageNumber="17">7 specs</specimenCount>
; same locality;
<date id="FFA51038F841F30FFEE6D3B75CB7C8A9" box="[296,446,1177,1203]" pageId="16" pageNumber="17" value="2018-02-04">
<collectingDate id="EFE1E9D0F841F30FFEE6D3B75CB7C8A9" box="[296,446,1177,1203]" pageId="16" pageNumber="17" value="2018-02-04">4 Feb. 2018</collectingDate>
</date>
; HIPE exped.;
<specimenCode id="DBBD9E83F841F30FFDB5D3B75E2CC8AE" box="[635,805,1177,1204]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">RBINS 25588</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFC9CD3B75E92C8AE" box="[850,923,1177,1204]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">25594</specimenCode>
(Supplementary file SM.01: Fiche-143 to Fiche-149)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFE4DD3925C2EC907" ID-GBIF-Occurrence="2830976320" collectingDate="2018-02-04" collectionCode="SM" collectorName="Supplementary" country="Uganda" latitude="-0.123833336" location="GenBank" longLatPrecision="1" longitude="30.052082" municipality="Kyambura River" pageId="16" pageNumber="17" specimenCode="RMCA 2018.008.P.0249, 2018.008.P.0252" specimenCount="4" stateProvince="Kazinga Channel">
<specimenCount id="9D1DFD71F841F30FFE4DD3925CE8C8CD" box="[387,481,1212,1239]" pageId="16" pageNumber="17">4 specs</specimenCount>
; same collection data as for preceding; GenBank:
<accessionNumber id="9448AB1BF841F30FFB9BD39259D5C8CD" box="[1109,1244,1212,1239]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251099" pageId="16" pageNumber="17">MT251099</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFB22D392587BC8CD" box="[1260,1394,1212,1239]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251112" pageId="16" pageNumber="17">MT251112</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFF73D3CE5CE9C8E0" box="[189,480,1248,1274]" pageId="16" pageNumber="17">RMCA 2018.008.P.0249</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFDC4D3F15FC7C8E0" box="[522,718,1247,1274]" pageId="16" pageNumber="17">2018.008.P.0252</specimenCode>
(Supplementary file SM.01: HP3312, HP3314, HP3317, HP3319)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFEF0D22D5987C95B" ID-GBIF-Occurrence="2830976314" collectingDate="2018-02-02" collectionCode="HIPE, SM" collectorName="Supplementary" country="Uganda" latitude="-0.20658334" location="Kyambura River" longLatPrecision="1" longitude="30.10675" municipality="Kyambura River" pageId="16" pageNumber="17" specimenCode="RBINS 25587" specimenCount="1" stateProvince="Kazinga Channel">
<specimenCount id="9D1DFD71F841F30FFEF0D22D5C9BC907" box="[318,402,1283,1309]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
;
<location id="8EC46023F841F30FFE53D22D5F6AC907" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFE53D22D5F6AC907" box="[413,611,1283,1309]" country="Uganda" latitude="-0.20658334" longLatPrecision="1" longitude="30.10675" municipality="Kyambura River" name="Kyambura River" pageId="16" pageNumber="17" stateProvince="Kazinga Channel">Kyambura River</location>
,
<locationDeviation id="9A162FDAF841F30FFDA3D22D592CC907" box="[621,1061,1283,1309]" pageId="16" pageNumber="17">middle course, north of Lake Katinda</locationDeviation>
;
<geoCoordinate id="EE2F503FF841F30FFBE1D22F59CBC907" box="[1071,1218,1281,1309]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="23.7" value="-0.20658334">0°12ʹ23.7ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFB00D22F5878C907" box="[1230,1393,1281,1309]" degrees="30" direction="east" minutes="06" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="24.3" value="30.10675">30°06ʹ24.3ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFF73D2085C59C95A" box="[189,336,1318,1344]" pageId="16" pageNumber="17" value="2018-02-02">
<collectingDate id="EFE1E9D0F841F30FFF73D2085C59C95A" box="[189,336,1318,1344]" pageId="16" pageNumber="17" value="2018-02-02">2 Feb. 2018</collectingDate>
</date>
; HIPE exped.;
<specimenCode id="DBBD9E83F841F30FFDC9D2085FB9C95B" box="[519,688,1318,1345]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">RBINS 25587</specimenCode>
(Supplementary file SM.01: Fiche-128)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFB62D2085981C9B0" ID-GBIF-Occurrence="2830976309" collectingDate="2016-11-04" collectionCode="HIPE, SM" collectorName="Supplementary" country="Uganda" county="Kazinga Channel" latitude="-0.136" location="Kazinga Channel" longLatPrecision="1" longitude="30.041" municipality="Queen Elisabeth Bush Lodge" pageId="16" pageNumber="17" specimenCode="RMCA 2016.035.P.0166" specimenCount="6" stateProvince="Kazinga Channel">
<specimenCount id="9D1DFD71F841F30FFB62D208580EC95A" box="[1196,1287,1318,1345]" pageId="16" pageNumber="17">5 specs</specimenCount>
;
<location id="8EC46023F841F30FFADBD2085C2BC979" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFADBD2085C2BC979" country="Uganda" county="Kazinga Channel" latitude="-0.136" longLatPrecision="1" longitude="30.041" municipality="Queen Elisabeth Bush Lodge" name="Kazinga Channel" pageId="16" pageNumber="17" stateProvince="Kazinga Channel">Kazinga Channel</location>
,
<locationDeviation id="9A162FDAF841F30FFEE1D2645FC5C979" box="[303,716,1353,1380]" pageId="16" pageNumber="17">near Queen Elisabeth Bush Lodge</locationDeviation>
;
<geoCoordinate id="EE2F503FF841F30FFD17D2695E67C979" box="[729,878,1351,1379]" degrees="0" direction="south" minutes="08" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="09.6" value="-0.136">0°08ʹ09.6ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFCB3D269592AC97E" box="[893,1059,1351,1380]" degrees="30" direction="east" minutes="02" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="27.6" value="30.041">30°02ʹ27.6ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFBFFD26759C1C979" box="[1073,1224,1353,1380]" pageId="16" pageNumber="17" value="2016-11-04">
<collectingDate id="EFE1E9D0F841F30FFBFFD26759C1C979" box="[1073,1224,1353,1380]" pageId="16" pageNumber="17" value="2016-11-04">4 Nov. 2016</collectingDate>
</date>
; HIPE exped.; GenBank:
<accessionNumber id="9448AB1BF841F30FFE8ED2425CCCC99D" box="[320,453,1388,1415]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251123" pageId="16" pageNumber="17">MT251123</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFE18D2425F55C99D" box="[470,604,1388,1415]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251124" pageId="16" pageNumber="17">MT251124</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFDA2D2425FFBC99D" box="[620,754,1388,1415]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251125" pageId="16" pageNumber="17">MT251125</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFCCCD2425E81C99D" box="[770,904,1388,1415]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251108" pageId="16" pageNumber="17">MT251108</accessionNumber>
,
<accessionNumber id="9448AB1BF841F30FFC56D2425916C99D" box="[920,1055,1388,1415]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT251109" pageId="16" pageNumber="17">MT251109</accessionNumber>
;
<specimenCode id="DBBD9E83F841F30FFBE1D242585DC99D" box="[1071,1364,1388,1415]" pageId="16" pageNumber="17">RMCA 2016.035.P.0166</specimenCode>
to
<specimenCount id="9D1DFD71F841F30FFF73D2A15C88C9B0" box="[189,385,1423,1450]" pageId="16" pageNumber="17">2016.035.P.0170</specimenCount>
(Supplementary file SM.01: HP719 to HP721, HP723, HP724)
</materialsCitation>
<materialsCitation id="3B733CA5F841F30FFB66D2BE593BC9EB" ID-GBIF-Occurrence="2830976305" collectingDate="2018-02-04" collectionCode="HIPE, SM" collectorName="Supplementary" country="Uganda" latitude="-0.121527776" location="Kazinga Channel" longLatPrecision="1" longitude="30.04972" municipality="Kazinga Channel" pageId="16" pageNumber="17" specimenCode="RBINS 25595, 25596" specimenCount="2" stateProvince="Kazinga Channel">
<specimenCount id="9D1DFD71F841F30FFB66D2BE580CC9B0" box="[1192,1285,1424,1450]" pageId="16" pageNumber="17">2 specs</specimenCount>
;
<location id="8EC46023F841F30FFADBD2A15C2BC9D7" LSID="urn:lsid:plazi:treatment:03B287EEF85FF30EFD90D6845856C948:8EC46023F841F30FFADBD2A15C2BC9D7" country="Uganda" latitude="-0.121527776" longLatPrecision="1" longitude="30.04972" municipality="Kazinga Channel" name="Kazinga Channel" pageId="16" pageNumber="17" stateProvince="Kazinga Channel">Kazinga Channel</location>
,
<locationDeviation id="9A162FDAF841F30FFEE0D29D5FD1C9D7" box="[302,728,1459,1485]" pageId="16" pageNumber="17">opposite of mouth Kyambura River</locationDeviation>
;
<geoCoordinate id="EE2F503FF841F30FFD2AD29F5E71C9D7" box="[740,888,1457,1485]" degrees="0" direction="south" minutes="07" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="17.5" value="-0.121527776">0°07ʹ17.5ʺ S</geoCoordinate>
,
<geoCoordinate id="EE2F503FF841F30FFC48D29F5925C9D7" box="[902,1068,1457,1485]" degrees="30" direction="east" minutes="02" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="59.0" value="30.04972">30°02ʹ59.0ʺ E</geoCoordinate>
;
<date id="FFA51038F841F30FFBF7D29D59C3C9D7" box="[1081,1226,1459,1485]" pageId="16" pageNumber="17" value="2018-02-04">
<collectingDate id="EFE1E9D0F841F30FFBF7D29D59C3C9D7" box="[1081,1226,1459,1485]" pageId="16" pageNumber="17" value="2018-02-04">4 Feb. 2018</collectingDate>
</date>
; HIPE exped.;
<specimenCode id="DBBD9E83F841F30FFF73D2F85C6DC9EB" box="[189,356,1494,1521]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">RBINS 25595</specimenCode>
to
<specimenCode id="DBBD9E83F841F30FFE47D2F85CDBC9EB" box="[393,466,1494,1521]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="16" pageNumber="17" type="Museum">25596</specimenCode>
(Supplementary file SM.01: Fiche-121, Fiche-123)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016573F841F30EFF73D1395C02CFD3" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BA436F8F841F30FFF73D1395C47CA2B" blockId="16.[189,334,1559,1585]" box="[189,334,1559,1585]" pageId="16" pageNumber="17">
<heading id="D0EC8194F841F30FFF73D1395C47CA2B" bold="true" box="[189,334,1559,1585]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B96FEAEAF841F30FFF73D1395C47CA2B" bold="true" box="[189,334,1559,1585]" pageId="16" pageNumber="17">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436F8F841F30EFF73D1685F88CE78" blockId="16.[189,1399,1606,2020]" lastBlockId="17.[189,1399,267,610]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Based on the
<typeStatus id="54A0885AF841F30FFEA6D1685CDDCA7A" box="[360,468,1606,1632]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
and one
<typeStatus id="54A0885AF841F30FFD85D1685FE0CA7A" box="[587,745,1606,1632]" pageId="16" pageNumber="17" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C1B4D7BF841F30FFCD6D1695E82CA7A" box="[792,907,1606,1632]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF841F30FFCD6D1695E82CA7A" box="[792,907,1606,1632]" italics="true" pageId="16" pageNumber="17">E. alberti</emphasis>
</taxonomicName>
, the type series of
<taxonomicName id="4C1B4D7BF841F30FFBB0D16959F4CA7A" baseAuthorityName="Whitehead" baseAuthorityYear="1960" box="[1150,1277,1607,1632]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="cercops">
<emphasis id="B96FEAEAF841F30FFBB0D16959F4CA7A" box="[1150,1277,1607,1632]" italics="true" pageId="16" pageNumber="17">E. cercops</emphasis>
</taxonomicName>
, which is synonymised here as
<taxonomicName id="4C1B4D7BF841F30FFE7BD1445F2BCA99" box="[437,546,1641,1667]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF841F30FFE7BD1445F2BCA99" box="[437,546,1641,1667]" italics="true" pageId="16" pageNumber="17">E. alberti</emphasis>
</taxonomicName>
, and 37 additional specimens from the Lake Edward system, identified as
<taxonomicName id="4C1B4D7BF841F30FFF73D1A35C25CABC" box="[189,300,1676,1702]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF841F30FFF73D1A35C25CABC" box="[189,300,1676,1702]" italics="true" pageId="16" pageNumber="17">E. alberti</emphasis>
</taxonomicName>
. The
<typeStatus id="54A0885AF841F30FFEBED1A25CD5CABC" box="[368,476,1676,1702]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
is illustrated in
<figureCitation id="13202A7DF841F30FFD54D1A25FE6CABD" box="[666,751,1676,1703]" captionStart="Fig" captionStartId="14.[189,232,1956,1982]" captionTargetBox="[189,1398,1090,1915]" captionTargetId="figure@14.[189,1398,1090,1918]" captionTargetPageId="14" captionText="Fig. 7. (a) Lectotype of Enteromius alberti (Poll, 1939) (MRAC 64723) with 74.3 mm SL. (b) Fresh specimen of E. alberti (RMCA 2018.008.P.0248 HP 3246) with 62.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3981083" httpUri="https://zenodo.org/record/3981083/files/figure.png" pageId="16" pageNumber="17">Fig. 7a</figureCitation>
. Maximum observed size:
<quantity id="4CE39B1DF841F30FFBE5D1A2599DCABD" box="[1067,1172,1676,1703]" metricMagnitude="-2" metricUnit="m" metricValue="7.43" pageId="16" pageNumber="17" unit="mm" value="74.3">74.3 mm</quantity>
SL. Morphometric and meristic data are given in
<tableCitation id="C6990343F841F30FFDD2D1815F7FCAD0" box="[540,630,1711,1738]" captionStart="Table 2" captionStartId="12.[179,248,531,559]" captionText="Table 2 (continued on the next page). Measurements and meristics of Enteromius alberti (Poll, 1939) (n = 47, *= 46, ** = 45), the types of Enteromius mimus (Boulenger, 1912) (n = 9), and Enteromius cf. mimus (n = 76, * = 75, ** = 74, *** = 73)." pageId="16" pageNumber="17">Table 2</tableCitation>
. Body fusiform, largest depth anterior to dorsal fin. Dorsal profile from tip of snout to origin of dorsal fin slightly convex, slightly tapering to base of caudal fin. Ventral profile from operculum to origin of pelvic fin slightly convex, slightly tapering to posterior end of anal fin base, then slightly concave to caudal fin. Head small. Eye large and round, located dorso-laterally, closer to tip of snout than to distal margin of operculum, interorbital profile slightly convex. Snout rounded and mouth subterminal. Two pairs of barbels, anterior barbel usually slightly shorter than posterior with anterior one usually reaching up to vertical through posterior margin of eye, while posterior one usually up to vertical trough posterior margin of pre-operculum. Dorsal fin with 3 unbranched and 7 to 9 branched rays, distal margin straight to slightly concave, origin located slightly anterior [1 or 2 lateral line scale(s)] to vertical through pelvic fin insertion. First branched dorsal fin ray longest, posterior rays decreasing progressively in size. Pectoral fin with 1 unbranched and 13 to 16 branched rays, distal profile straight to slightly convex, not reaching anterior base of pelvic fin. Pelvic fin with 1 unbranched and 6 to 8 branched rays, distal margin slightly convex. Anal fin with 3 unbranched and 5 to 6 branched fin rays, distal margin slightly concave. Caudal fin forked with outer rays twice as long as median ones and both lobes rounded and of similar size. Anus and urogenital opening situated immediately in front of anal fin base. Scales cycloid, rounded and radially striate. Lateral line completely pored with many lateral line scales [2734 (median 31)], and gently curved downwards over abdomen but running straight along middle of caudal peduncle and ending at base of caudal fin. Lateral-line scales smaller on caudal peduncle than below dorsal fin.
</paragraph>
<paragraph id="8BA436F8F840F30EFF73D5A75C70CEBE" blockId="17.[189,377,649,676]" box="[189,377,649,676]" pageId="17" pageNumber="18">
<heading id="D0EC8194F840F30EFF73D5A75C70CEBE" bold="true" box="[189,377,649,676]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B96FEAEAF840F30EFF73D5A75C70CEBE" bold="true" box="[189,377,649,676]" pageId="17" pageNumber="18">Colour pattern</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436F8F840F30EFF73D5975C02CFD3" blockId="17.[189,1399,696,969]" pageId="17" pageNumber="18">
No sexual dimorphism observed. In life, overall background colour of body silvery, darker toward dorsum and lighter towards belly (
<figureCitation id="13202A7DF840F30EFD93D5F55FBECEEC" box="[605,695,731,758]" captionStart="Fig" captionStartId="14.[189,232,1956,1982]" captionTargetBox="[189,1398,1090,1915]" captionTargetId="figure@14.[189,1398,1090,1918]" captionTargetPageId="14" captionText="Fig. 7. (a) Lectotype of Enteromius alberti (Poll, 1939) (MRAC 64723) with 74.3 mm SL. (b) Fresh specimen of E. alberti (RMCA 2018.008.P.0248 HP 3246) with 62.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3981083" httpUri="https://zenodo.org/record/3981083/files/figure.png" pageId="17" pageNumber="18">Fig. 7b</figureCitation>
). Thick yellowish midlateral band, often with one to three darker spots. All fins translucent. In preserved specimens, overall background colour of body yellowish, greyish dorsally (
<figureCitation id="13202A7DF840F30EFE49D40C5CD5CF26" box="[391,476,802,828]" captionStart="Fig" captionStartId="14.[189,232,1956,1982]" captionTargetBox="[189,1398,1090,1915]" captionTargetId="figure@14.[189,1398,1090,1918]" captionTargetPageId="14" captionText="Fig. 7. (a) Lectotype of Enteromius alberti (Poll, 1939) (MRAC 64723) with 74.3 mm SL. (b) Fresh specimen of E. alberti (RMCA 2018.008.P.0248 HP 3246) with 62.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3981083" httpUri="https://zenodo.org/record/3981083/files/figure.png" pageId="17" pageNumber="18">Fig. 7a</figureCitation>
). One to three dark spots on flanks on thick silvery midlateral band; spots fused into mid-lateral line in some specimens, overlying silvery band and starting posterior to operculum. First spot situated anterior to dorsal fin origin, median one below last ray of dorsal fin and last one at caudal fin base. Specimens with more than three spots are rare (
<specimenCount id="9D1DFD71F840F30EFCA5D4A55EF8CFBF" box="[875,1009,907,933]" pageId="17" pageNumber="18" type="generic">1 specimen</specimenCount>
with 4 spots and another with 6 spots).
</paragraph>
</subSubSection>
<subSubSection id="C3016573F840F30EFF73D4DE5856C948" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BA436F8F840F30EFF73D4DE5CCDC810" blockId="17.[189,452,1008,1034]" box="[189,452,1008,1034]" pageId="17" pageNumber="18">
<emphasis id="B96FEAEAF840F30EFF73D4DE5C5FC810" bold="true" box="[189,342,1008,1034]" pageId="17" pageNumber="18">Distribution</emphasis>
(
<figureCitation id="13202A7DF840F30EFEA8D4DE5CB2C810" box="[358,443,1008,1034]" captionStart="Fig" captionStartId="21.[189,232,1851,1877]" captionTargetBox="[340,1248,797,1800]" captionTargetId="figure@21.[332,1255,789,1808]" captionTargetPageId="21" captionText="Fig. 10. Overview of the sample locations of E. alberti (Poll. 1939) (●) and E. cf. mimus (Boulenger, 1912) (■), including the specimens from Tshambi, in the Lake Edward system. The location of the lectotype and the paralectotypes of E. alberti (♦). The approximate location of the lectotype and the paralectotypes of E. mimus (Boulenger, 1912) (▼) and the holotype and paratypes of E. cercops (Whitehead, 1960) (▲) are indicated in the inset." figureDoi="http://doi.org/10.5281/zenodo.3981089" httpUri="https://zenodo.org/record/3981089/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
)
</paragraph>
<paragraph id="8BA436F8F840F30EFF73D3315856C948" blockId="17.[189,1399,1054,1363]" pageId="17" pageNumber="18">
<taxonomicName id="4C1B4D7BF840F30EFF73D3315C91C822" box="[189,408,1054,1080]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF840F30EFF73D3315C91C822" box="[189,408,1054,1080]" italics="true" pageId="17" pageNumber="18">Enteromius alberti</emphasis>
</taxonomicName>
occurs in river systems throughout the Lake Edward basin. For the Congolese part of the basin, it is known from its
<typeStatus id="54A0885AF840F30EFDD4D36C5F44C846" box="[538,589,1090,1116]" pageId="17" pageNumber="18">type</typeStatus>
locality, the Rutshuru River, May-Ya-Moto (Poll 1939). Although the Congolese part of the system was not sampled during the recent expeditions, we identified specimens from the collection at the RMCA (86-01-P-479 to 500 and 86-09-P-70 to 71) from Vitshumbi, located in the southern Congolese part of the Lake Edward system, as
<taxonomicName id="4C1B4D7BF840F30EFCBDD3825EE8C8DF" box="[883,993,1195,1221]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF840F30EFCBDD3825EE8C8DF" box="[883,993,1195,1221]" italics="true" pageId="17" pageNumber="18">E. alberti</emphasis>
</taxonomicName>
. In the Ugandan part, we collected the species in the Mahoma, Mpanga, Nchwera, Nyamweru and Kyambura (an affluent of the Rusangwe) Rivers, in Lake Edward offshore at Kayanja, and in Lake George at Kashaka Bay. For the Lake Victoria basin,
<taxonomicName id="4C1B4D7BF840F30EFEC3D2385C77C935" box="[269,382,1301,1327]" family="Cyprinidae" genus="Enteromius" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="alberti">
<emphasis id="B96FEAEAF840F30EFEC3D2385C77C935" box="[269,382,1301,1327]" italics="true" pageId="17" pageNumber="18">E. alberti</emphasis>
</taxonomicName>
is known from the Nzoia River, the Malawa River and the Middle Akagera system (
<bibRefCitation id="EF8A4B09F840F30EFF0BD2165C82C948" author="Whitehead P. J. P." box="[197,395,1336,1362]" firstAuthor="Whitehead" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="17" pageNumber="18" pagination="106 - 119" part="62" refId="ref17994" refString="Whitehead P. J. P. 1960. Three new cyprinid fishes of the genus Barbus from the Lake Victoria basin. Revue de Zoologie et de Botanique africaines 62 (1 - 2): 106 - 119." title="Three new cyprinid fishes of the genus Barbus from the Lake Victoria basin" type="journal article" year="1960">Whitehead 1960</bibRefCitation>
;
<bibRefCitation id="EF8A4B09F840F30EFE59D2165F6CC948" author="Greenwood P. H." box="[407,613,1336,1362]" firstAuthor="Greenwood" journalOrPublisher="The Uganda Society, Kampala" pageId="17" pageNumber="18" refId="ref15852" refString="Greenwood P. H. 1966. The Fishes of Uganda. The Uganda Society, Kampala." title="The Fishes of Uganda" type="book" year="1966">Greenwood 1966</bibRefCitation>
;
<bibRefCitation id="EF8A4B09F840F30EFDBCD217596BC948" author="De Vos L. &amp; Thys van den Audenaerde D. F. E." box="[626,1122,1336,1363]" firstAuthor="De Vos" journalOrPublisher="Revue d'Hydrobiologie tropicale" pageId="17" pageNumber="18" pagination="141 - 159" part="23" refId="ref15481" refString="De Vos L. &amp; Thys van den Audenaerde D. F. E. 1990. Petit Barbus (Pisces, Cyprinidae) du Rwanda. Revue d'Hydrobiologie tropicale 23 (2): 141 - 159." title="Petit Barbus (Pisces, Cyprinidae) du Rwanda" type="journal article" year="1990">De Vos &amp; Thys van den Audenaerde 1990</bibRefCitation>
;
<bibRefCitation id="EF8A4B09F840F30EFBA0D216585AC948" author="Seegers L. &amp; De Vos L. &amp; Okeyo D. O." box="[1134,1363,1336,1362]" etAl="et al." firstAuthor="Seegers" journalOrPublisher="Journal of East African Natural History" pageId="17" pageNumber="18" pagination="11 - 47" part="92" refId="ref17474" refString="Seegers L., De Vos L. &amp; Okeyo D. O. 2003. Annotated checklist of the freshwater fishes of Kenya (excluding the lacustrine haplochromines from Lake Victoria). Journal of East African Natural History 92: 11 - 47. https: // doi. org / 10.2982 / 0012 - 8317 (2003) 92 % 5 B 11: ACOTFF % 5 D 2.0. CO; 2" title="Annotated checklist of the freshwater fishes of Kenya (excluding the lacustrine haplochromines from Lake Victoria)" type="journal article" year="2003">
Seegers
<emphasis id="B96FEAEAF840F30EFB1FD2175802C948" box="[1233,1291,1336,1362]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2003
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>