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<mods:title id="CECA4C441C43AE71753D98BC69583DD9">Tasmanian Phoridae (Diptera) and some additional Australasian species</mods:title>
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<heading id="D0EC81D84224FFEDFDFAFE06FC65FD86" box="[619,963,500,525]" centered="true" fontSize="10" level="2" pageId="112" pageNumber="616" reason="3">
<emphasis id="B96FEAA64224FFEDFDFAFE06FD16FD86" bold="true" box="[619,688,500,524]" pageId="112" pageNumber="616">Genus</emphasis>
<taxonomicName id="4C1B4D374224FFEDFD28FE06FC65FD86" ID-CoL="6G76" authority="Beyer, 1966" authorityName="Beyer" authorityYear="1966" box="[697,963,500,525]" class="Insecta" family="Phoridae" genus="Paraphiura" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="616" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAA64224FFEDFD28FE06FC93FD86" bold="true" box="[697,821,500,524]" italics="true" pageId="112" pageNumber="616">Paraphiura</emphasis>
Beyer, 1966
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species.
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<taxonomicName id="4C1B4D374224FFEDFE57FDE4FCE0FDA1" authority="Beyer, 1966: 171" authorityName="Beyer" authorityPageNumber="171" authorityYear="1966" box="[454,838,534,556]" class="Insecta" family="Phoridae" genus="Paraphiura" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="616" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B96FEAA64224FFEDFE57FDE4FD36FDA1" box="[454,656,534,555]" italics="true" pageId="112" pageNumber="616">Paraphiura australis</emphasis>
Beyer, 1966: 171
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.
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<paragraph id="8BA436B44224FFEDFEF0FDBFFB91FCAE" blockId="112.[313,1269,589,1283]" pageId="112" pageNumber="616">
Beyer (1966) assigned this genus to the tribe
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, which has traditionally been distinguished from the
<taxonomicName id="4C1B4D374224FFEDFD1AFD9FFCACFD0F" authorityName="Peterson" authorityYear="1987" box="[651,778,621,645]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="616" phylum="Arthropoda" rank="tribe" tribe="Metopinini">Metopinini</taxonomicName>
by the different inclination of the supraantennal bristles. In
<taxonomicName id="4C1B4D374224FFEDFDB0FD7FFD07FD2F" box="[545,673,653,677]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="616" phylum="Arthropoda" rank="tribe" tribe="Beckerinini">Beckerinini</taxonomicName>
the SAs are porrect (inlined forward horizontally), often divergently, or reclinate (inclined upwards). Those of the
<taxonomicName id="4C1B4D374224FFEDFBD7FD5FFB63FD4F" authorityName="Peterson" authorityYear="1987" box="[1094,1221,685,709]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="616" phylum="Arthropoda" rank="tribe" tribe="Metopinini">Metopinini</taxonomicName>
are proclinate (inclined forwards and downwards). Genera lacking SAs have been somewhat arbitrarily assigned to one of these tribes on the basis of a general resemblance to genera previously assigned to the tribe in question.
</paragraph>
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In the case of
<taxonomicName id="4C1B4D374224FFEDFD98FCDEFD6BFCCF" authority="Beyer" authorityName="Beyer" authorityYear="1966" box="[521,717,812,837]" class="Insecta" family="Phoridae" genus="Paraphiura" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="616" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAA64224FFEDFD98FCDEFD22FCCE" box="[521,644,812,836]" italics="true" pageId="112" pageNumber="616">Paraphiura</emphasis>
Beyer
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assigned it to the
<taxonomicName id="4C1B4D374224FFEDFC36FCDEFB81FCCE" box="[935,1063,812,836]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="616" phylum="Arthropoda" rank="tribe" tribe="Beckerinini">Beckerinini</taxonomicName>
on the basis of a single pair of divergent SAs; having given the frontal bristle formula as 4-4-4 plus two divergent
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bristles. However, the head of his only specimen (see below) was damaged, with its palps being lost altogether. It seems that this damage also resulted in the loss of one of the two pairs of
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bristles, and that Beyer failed to observe their sockets. There are not only four (not two)
<collectionCode id="ED0AAE714224FFEDFCE5FC3EFC31FC6E" box="[884,919,972,996]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="112" pageNumber="616">SA</collectionCode>
bristles in the
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species of this genus but the upper pair is proclinate and the lower pair is divergent and reclinate (figure 60A). This situation indicates that the precise inclination of the
<collectionCode id="ED0AAE714224FFEDFB40FBF9FB52FBA9" box="[1233,1268,1035,1059]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="112" pageNumber="616">SA</collectionCode>
bristles is not a reliable diagnostic character for distinguishing the two tribes. Indeed the shift from proclinate SAs to these bristles being more porrect or reclinate seems to be largely a secondary consequence of a shortening of the frons (e.g. figure 60A). Such a shortening appears to have occurred independently in a number of phorid genera and cannot be regarded as a unique synapomorphy. I therefore formally propose the transfer of this genus to the
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and the abolition of the tribe
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as it is presently conceived.
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