465 lines
68 KiB
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465 lines
68 KiB
XML
<document id="468E4E491FD56CC6E220F6BDE4E15915" ID-CLB-Dataset="22100" ID-DOI="10.5852/ejt.2017.321" ID-GBIF-Dataset="caeead37-b6ce-4f03-96d2-ae8c8beb8573" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829282" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589562339121" checkinUser="carolina" docAuthor="Maxwell, Erin E. & Stumpf, Sebastian" docDate="2017" docId="03B22572FF99FF8DFDFFFA26BC1BCA21" docLanguage="en" docName="ejt-2017-321.pdf.imf" docOrigin="European Journal of Taxonomy 321 (321)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Saurorhynchus anningae Maxwell & Stumpf 2017, sp. nov." docType="treatment" docUuid="F16C4AFF-4A3B-4C35-8902-F95446F9B1EB" docUuidSource="ZooBank" docVersion="9" lastPageNumber="13" masterDocId="FF8B5D0AFF91FF81FFB1FFF9BF47CD01" masterDocTitle="Revision of Saurorhynchus (Actinopterygii: Saurichthyidae) from the Early Jurassic of England and Germany" masterLastPageNumber="29" masterPageNumber="1" pageNumber="9" updateTime="1698839856834" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B28264910A62DBCB831999523B2DDC33">Revision of Saurorhynchus (Actinopterygii: Saurichthyidae) from the Early Jurassic of England and Germany</mods:title>
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<mods:namePart id="164D94C6038BF982D56CCFB20DFD6A4E">Maxwell, Erin E.</mods:namePart>
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<mods:affiliation id="4352FBFC01D425C6C68CD9B119D9F9D6">Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany. Institut für Geographie und Geologie, Ernst-Moritz-Arndt-Universität Greifswald, Friedrich-Ludwig-Jahn Strasse 17 a, 17489 Greifswald, Germany.</mods:affiliation>
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<mods:nameIdentifier id="8BA6B40E718BF6EDA2A57493FD75BB6D" type="email">erin.maxwell@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="3F1604E958405635DA5AFC1DB5F499D5">Stumpf, Sebastian</mods:namePart>
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<mods:affiliation id="C91B9C32D549A349D986CCE9D01466E6">Email: sebastian. stumpf @ stud. uni-greifswald. de & urn: lsid: zoobank. org: author: 8 DADF 851 - CFA 5 - 469 E- 9628 - AD 020803 B 424</mods:affiliation>
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<mods:nameIdentifier id="7D822A5C5D7F68C78E400E5EDCF73125" type="email">sebastian.stumpf@stud.uni-greifswald.de</mods:nameIdentifier>
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<mods:title id="1A02BB229B7B69A593004630C0D5DDD4">European Journal of Taxonomy</mods:title>
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<treatment id="03B22572FF99FF8DFDFFFA26BC1BCA21" ID-DOI="http://doi.org/10.5281/zenodo.3848070" ID-GBIF-Taxon="164159995" ID-Zenodo-Dep="3848070" LSID="urn:lsid:zoobank.org:act:F16C4AFF-4A3B-4C35-8902-F95446F9B1EB" httpUri="http://treatment.plazi.org/id/03B22572FF99FF8DFDFFFA26BC1BCA21" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
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<subSubSection id="C301C7EFFF99FF89FDFFFA26BCA2C8FB" box="[590,997,1503,1530]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BA49464FF99FF89FDFFFA26BCA2C8FB" blockId="8.[368,1219,1503,1600]" box="[590,997,1503,1530]" pageId="8" pageNumber="9">
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<heading id="D0EC2308FF99FF89FDFFFA26BCA2C8FB" box="[590,997,1503,1530]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
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<taxonomicName id="4C1BEFE7FF99FF89FDFFFA26BCC4C8FB" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[590,899,1503,1530]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
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<emphasis id="B96F4876FF99FF89FDFFFA26BCC4C8FB" bold="true" box="[590,899,1503,1530]" italics="true" pageId="8" pageNumber="9">Saurorhynchus anningae</emphasis>
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<taxonomicNameLabel id="A25CF50DFF99FF89FC3AFA19BCA2C8FB" box="[907,997,1504,1530]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C301C7EFFF99FF89FEC1F9FABC82CB41" pageId="8" pageNumber="9" type="description">
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<uri id="FF8A9866FF99FF89FEC1F9FABB84CB1F" box="[368,1219,1539,1566]" pageId="8" pageNumber="9">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBDAEB1FF99FF89FD23F9FABB84CB1F" box="[658,1219,1539,1566]" pageId="8" pageNumber="9">F16C4AFF-4A3B-4C35-8902-F95446F9B1EB</uuid>
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<paragraph id="8BA49464FF99FF89FDC1F9DFBC82CB41" blockId="8.[368,1219,1503,1600]" box="[624,965,1574,1600]" pageId="8" pageNumber="9">
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<figureCitation id="132088E1FF99FF89FDC1F9DFBD8ACB41" box="[624,717,1574,1600]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="8" pageNumber="9">Figs 1C</figureCitation>
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,
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<figureCitation id="132088E1FF99FF89FD6AF9DFBDB8CB41" box="[731,767,1574,1600]" captionStart="Fig" captionStartId="6.[189,232,1647,1673]" captionTargetBox="[205,1385,332,1597]" captionTargetId="figure@6.[204,1385,329,1597]" captionTargetPageId="6" captionText="Fig. 2. Reconstruction of the dermal skull in Early Jurassic saurichthyids. A. Saurorhynchus brevirostris (Woodward, 1895). B. Saurorhynchus anningae sp. nov. C. Saurorhynchus hauffi sp. nov. D. Saurorhynchus acutus (Agassiz, 1844). Not to scale. Dashed lines indicate sensory canals, grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: ang = angular; bs = basisphenoid; cl = cleithrum; d = dentary; dh = dermohyal; dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l = lacrimal; l.ex = lateral extrascapular; m = maxilla; md.c = mandibular sensory canal; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; op = opercle; p = parietal; pm = rostropremaxilla; pop = preopercle; ps = parasphenoid; sa = supraangular; scl = supracleithrum; so = supraorbital; so.c = supraorbital sensory canal." figureDoi="http://doi.org/10.5281/zenodo.3829286" httpUri="https://zenodo.org/record/3829286/files/figure.png" pageId="8" pageNumber="9">2B</figureCitation>
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,
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<figureCitation id="132088E1FF99FF89FCBAF9DFBC76CB41" box="[779,817,1574,1600]" captionStart="Fig" captionStartId="9.[189,232,1718,1744]" captionTargetBox="[195,1396,311,1622]" captionTargetId="graphics@9.[317,1359,731,1036]" captionTargetPageId="9" captionText="Fig. 3. Dorsal skull roof. A. Saurorhynchus anningae sp. nov. (NHMUK PV P 964); note that this skull has been dorsoventrally compressed. B. Saurorhynchus hauffi sp. nov., three-dimensionally preserved skull GG 20001. Fine dashed lines indicate sensory canals, large dashes indicate broken or incomplete areas; grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l.ex = lateral extrascapular; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; p = parietal; pm = rostropremaxilla; so.c = supraorbital sensory canal. Scale bars = 10 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829288" httpUri="https://zenodo.org/record/3829288/files/figure.png" pageId="8" pageNumber="9">3A</figureCitation>
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,
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||
<figureCitation id="132088E1FF99FF89FC8FF9DFBC23CB41" box="[830,868,1574,1600]" captionStart="Fig" captionStartId="10.[189,232,1683,1709]" captionTargetBox="[189,1399,933,1642]" captionTargetId="figure@10.[189,1399,929,1642]" captionTargetPageId="10" captionText="Fig. 4. Orbitotemporal region, illustrating variation in the braincase of Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov. (based on NHMUK PV P 36227). B, E. Saurorhynchus brevirostris (Woodward, 1895) (photo pertains to NHMUK PV P 4878, mirrored). C, F. Saurorhynchus hauffi sp. nov. (photo is of SMNS 51888, mirrored). D. Saurorhynchus acutus (Agassiz, 1844) (based on SMNS 87737). Abbreviations: asc = ascending process of the parasphenoid; bs = basisphenoid; dpt = dermopterotic; dsp = dermosphenotic; f.ic+ep = foramen for the internal carotid artery and efferent pseudobranchial artery; f.oma = f.om = foramen for the great ophthalmic artery; my = posterior myodome; n = neomorph; nc = neurocranium; orb = orbit; psr = parasphenoid rostrum; rot = foramen for the lateral otic ramus; spi = ventral opening of the spiracular canal; tf.c = trigeminofacial chamber. Scale bars: E–F = 2 mm. E. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829291" httpUri="https://zenodo.org/record/3829291/files/figure.png" pageId="8" pageNumber="9">4A</figureCitation>
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,
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<figureCitation id="132088E1FF99FF89FCC1F9DFBCD2CB41" box="[880,917,1574,1600]" captionStart="Fig" captionStartId="14.[189,232,1828,1854]" captionTargetBox="[198,1392,811,1788]" captionTargetId="figure@14.[195,1392,808,1788]" captionTargetPageId="14" captionText="Fig. 6. Posteriormost laniary dentition. A. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV P 4878), note the lingually curved crowns. B. Saurorhynchus anningae sp. nov. (NHMUK PV P 27569), note the incisivlücke associated with the premaxillary laniary tooth and the absence of lateral crypts associated with the two flanking mandibular laniaries. Photos © The Trustees of the Natural History Museum, London. C. S. hauffi sp. nov. (SMNS 51007), mid-rostral dentition illustrating the relationship between the incisivlücken, laniaries, and flanking smaller teeth. Scale bars: 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3829295" httpUri="https://zenodo.org/record/3829295/files/figure.png" pageId="8" pageNumber="9">6B</figureCitation>
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,
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<figureCitation id="132088E1FF99FF89FC10F9DFBC82CB41" box="[929,965,1574,1600]" captionStart="Fig" captionStartId="16.[189,232,1753,1779]" captionTargetBox="[189,1399,836,1707]" captionTargetId="figure@16.[189,1399,832,1707]" captionTargetPageId="16" captionText="Fig. 7. Postcranium, Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov., neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790). B–D. Saurorhynchus hauffi sp. nov., SMNS 55057. B. Lepidotrichia of the anal fin. C. Relationship between the neural and haemal arches and the axonosts. D. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="8" pageNumber="9">7A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C301C7EFFF99FF89FF0CF99EBE15CBED" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="8BA49464FF99FF89FF0CF99EBC54CBA4" blockId="8.[189,1398,1639,1772]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="AB0BB34AFF99FF89FF0CF99EBC54CBA4" pageId="8" pageNumber="9">
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<taxonomicName id="4C1BEFE7FF99FF89FF0CF99EBE88CB80" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[189,463,1639,1665]" class="Actinopterygii" family="Saurichthyidae" genus="Belonorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="acutus">
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<emphasis id="B96F4876FF99FF89FF0CF99EBE88CB80" box="[189,463,1639,1665]" italics="true" pageId="8" pageNumber="9">Belonorhynchus acutus</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0ABAB275FF99FF89FE5BF99EBDF4CB83" box="[490,691,1639,1666]" pageId="8" pageNumber="9">
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<bibRefCitation id="EF8AE995FF99FF89FE5BF99EBDF4CB83" author="Woodward A. S." box="[490,691,1639,1666]" pageId="8" pageNumber="9" pagination="1 - 55" refId="ref15223" refString="Woodward A. S. 1890. The fossil fishes of the Hawkesbury Series at Gosford. Memoirs of the Geological Survey of New South Wales, Palaeontology 4: 1 - 55." type="journal article" year="1890">Woodward, 1890</bibRefCitation>
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||
</treatmentCitation>
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||
: pl. 8, fig. 7. —
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<treatmentCitation id="0ABAB275FF99FF89FCC1F99EBB75CB83" box="[880,1074,1639,1666]" pageId="8" pageNumber="9">
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<bibRefCitation id="EF8AE995FF99FF89FCC1F99EBB75CB83" author="Woodward A. S." box="[880,1074,1639,1666]" pageId="8" pageNumber="9" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward 1895</bibRefCitation>
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</treatmentCitation>
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||
, partim: 14–15, pl. 2, fig. 1.
|
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<taxonomicName id="4C1BEFE7FF99FF89FF0CF972BE86CBA5" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[189,449,1675,1701]" class="Actinopterygii" family="Saurichthyidae" genus="Acidorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="acutus">
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<emphasis id="B96F4876FF99FF89FF0CF972BE86CBA5" box="[189,449,1675,1701]" italics="true" pageId="8" pageNumber="9">Acidorhynchus acutus</emphasis>
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</taxonomicName>
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||
–
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||
<treatmentCitation id="0ABAB275FF99FF89FE6FF972BDF3CBA4" author="Stensio E." box="[478,692,1674,1701]" page="178" pageId="8" pageNumber="9" year="1925">
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<bibRefCitation id="EF8AE995FF99FF89FE6FF972BDF3CBA4" author="Stensio E." box="[478,692,1674,1701]" pageId="8" pageNumber="9" pagination="1 - 126" refId="ref14959" refString="Stensio E. 1925. Triassic fishes from Spitzbergen, Part II. Kungliga Svenska Vetenskapsakademiens Handlingar Tredje Serien 2 (1): 1 - 126." type="journal article" year="1925">Stensiö 1925: 178</bibRefCitation>
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||
</treatmentCitation>
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||
, fig. 58.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BA49464FF99FF89FF0CF957BE15CBED" blockId="8.[189,1398,1639,1772]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="AB0BB34AFF99FF89FF0CF957BE15CBED" pageId="8" pageNumber="9">
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<taxonomicName id="4C1BEFE7FF99FF89FF0CF957BE84CBC9" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[189,451,1710,1736]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="acutus">
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<emphasis id="B96F4876FF99FF89FF0CF957BE84CBC9" box="[189,451,1710,1736]" italics="true" pageId="8" pageNumber="9">Saurorhynchus acutus</emphasis>
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</taxonomicName>
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||
–
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||
<treatmentCitation id="0ABAB275FF99FF89FE55F957BDD7CBC9" box="[484,656,1710,1736]" pageId="8" pageNumber="9">
|
||
<bibRefCitation id="EF8AE995FF99FF89FE55F957BDD7CBC9" author="Gardiner B. G." box="[484,656,1710,1736]" pageId="8" pageNumber="9" pagination="241 - 384" refId="ref13301" refString="Gardiner B. G. 1960. A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias. Bulletin of the British Museum (Natural History): Geology 4 (7): 241 - 384." type="journal article" year="1960">Gardiner 1960</bibRefCitation>
|
||
</treatmentCitation>
|
||
, partim: 272–280, figs 19–21. —
|
||
<treatmentCitation id="0ABAB275FF99FF89FB98F957BA35CBC8" author="Forey P. L. & Longbottom A. & Mulley J." box="[1065,1394,1710,1737]" page="350" pageId="8" pageNumber="9" year="2010">
|
||
<bibRefCitation id="EF8AE995FF99FF89FB98F957BA35CBC8" author="Forey P. L. & Longbottom A. & Mulley J." box="[1065,1394,1710,1737]" pageId="8" pageNumber="9" pagination="341 - 371" refId="ref13236" refString="Forey P. L., Longbottom A. & Mulley J. 2010. Fishes - bony fishes. In: Lord A. R. & Davis P. G. (eds) Palaeontological Association Field Guides to Fossils, Number 13: Fossils from the Lower Lias of the Dorset Coast: 341 - 371. Palaeontological Association, London." type="book chapter" year="2010">
|
||
Forey
|
||
<emphasis id="B96F4876FF99FF89FBC6F956BBF4CBC9" box="[1143,1203,1710,1736]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
2010: 350–352
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 65, fig. 5.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF99FF89FF0CF8EABD47CA7F" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="8BA49464FF99FF89FF0CF8EABE03CA2C" blockId="8.[189,324,1811,1837]" box="[189,324,1811,1837]" pageId="8" pageNumber="9">
|
||
<heading id="D0EC2308FF99FF89FF0CF8EABE03CA2C" bold="true" box="[189,324,1811,1837]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="B96F4876FF99FF89FF0CF8EABE03CA2C" bold="true" box="[189,324,1811,1837]" pageId="8" pageNumber="9">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF99FF89FF0CF8BBBD47CA7F" blockId="8.[189,1399,1857,1919]" pageId="8" pageNumber="9">
|
||
Name modified from that derived by
|
||
<bibRefCitation id="EF8AE995FF99FF89FD33F8B8BC70CA5A" author="Agassiz L." box="[642,823,1857,1883]" pageId="8" pageNumber="9" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz (1844)</bibRefCitation>
|
||
in honour of Mary Anning, an important fossil collector from Lyme Regis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF64C4ECFF98FF88FF0CF94FBC4ECAE8" ID-DOI="http://doi.org/10.5281/zenodo.3829288" ID-Zenodo-Dep="3829288" httpUri="https://zenodo.org/record/3829288/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1718,1744]" targetBox="[195,1396,311,1622]" targetPageId="9">
|
||
<paragraph id="8BA49464FF98FF88FF0CF94FBC4ECAE8" blockId="9.[189,1399,1718,2026]" pageId="9" pageNumber="10">
|
||
<emphasis id="B96F4876FF98FF88FF0CF94FBE4DCBD1" bold="true" box="[189,266,1718,1744]" pageId="9" pageNumber="10">Fig. 3.</emphasis>
|
||
Dorsal skull roof.
|
||
<emphasis id="B96F4876FF98FF88FE56F94FBEBBCBD1" bold="true" box="[487,508,1718,1744]" pageId="9" pageNumber="10">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF98FF88FDBBF94FBC68CBD1" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[522,815,1718,1744]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF98FF88FDBBF94FBC68CBD1" box="[522,815,1718,1744]" italics="true" pageId="9" pageNumber="10">Saurorhynchus anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF98FF88FC87F94FBCD7CBD1" box="[822,912,1718,1744]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NHMUK PV P 964); note that this skull has been dorsoventrally compressed.
|
||
<emphasis id="B96F4876FF98FF88FDCAF920BDC8CBF2" bold="true" box="[635,655,1753,1779]" pageId="9" pageNumber="10">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF98FF88FD2EF920BCDCCBF2" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[671,923,1753,1779]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF98FF88FD2EF920BCDCCBF2" box="[671,923,1753,1779]" italics="true" pageId="9" pageNumber="10">Saurorhynchus hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF98FF88FC15F923BB44CBF5" box="[932,1027,1754,1780]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, three-dimensionally preserved skull GG 20001. Fine dashed lines indicate sensory canals, large dashes indicate broken or incomplete areas; grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l.ex = lateral extrascapular; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; p = parietal; pm = rostropremaxilla; so.c = supraorbital sensory canal. Scale bars = 10 mm. A. Photo © The Trustees of the Natural History Museum, London.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C301C7EFFF9BFF8BFF0CFEF2BD87CF9C" pageId="10" pageNumber="11" type="materials_examined">
|
||
<paragraph id="8BA49464FF9BFF8BFF0CFEF2BECBCC24" blockId="10.[189,396,267,293]" box="[189,396,267,293]" pageId="10" pageNumber="11">
|
||
<heading id="D0EC2308FF9BFF8BFF0CFEF2BECBCC24" bold="true" box="[189,396,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="B96F4876FF9BFF8BFF0CFEF2BECBCC24" bold="true" box="[189,396,267,293]" pageId="10" pageNumber="11">Material studied</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B739E39FF9BFF8BFF5BFEC3BE60CC9B" ID-GBIF-Occurrence="2620280304" collectionCode="NHMUK" country="United Kingdom" location="UNITED KINGDOM" pageId="10" pageNumber="11" specimenCode="NHMUK PV P 3791" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BA49464FF9BFF8BFF5BFEC3BE1BCC55" blockId="10.[189,1399,314,410]" box="[234,348,314,340]" pageId="10" pageNumber="11">
|
||
<emphasis id="B96F4876FF9BFF8BFF5BFEC3BE1BCC55" bold="true" box="[234,348,314,340]" pageId="10" pageNumber="11">
|
||
<typeStatus id="54A02AC6FF9BFF8BFF5BFEC3BE1BCC55" box="[234,348,314,340]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9BFF8BFF0CFEA4BE60CC9B" blockId="10.[189,1399,314,410]" pageId="10" pageNumber="11">
|
||
<collectingCountry id="F30CD4F4FF9BFF8BFF0CFEA4BE8FCC76" box="[189,456,349,375]" name="United Kingdom" pageId="10" pageNumber="11">UNITED KINGDOM</collectingCountry>
|
||
: a skull and pectoral girdle preserved in lateral view (
|
||
<collectionCode id="ED0A0CA1FF9BFF8BFBDAFEA4BB9BCC76" box="[1131,1244,349,375]" collectionName="NHMUK" pageId="10" pageNumber="11">NHMUK</collectionCode>
|
||
PV P 3791,
|
||
<figureCitation id="132088E1FF9BFF8BFF0CFE79BE50CC9B" box="[189,279,384,410]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="10" pageNumber="11">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BA49464FF9BFF8BFF0CFE38BED3CCDA" blockId="10.[189,404,449,475]" box="[189,404,449,475]" pageId="10" pageNumber="11">
|
||
<heading id="D0EC2308FF9BFF8BFF0CFE38BED3CCDA" bold="true" box="[189,404,449,475]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="B96F4876FF9BFF8BFF0CFE38BED3CCDA" bold="true" box="[189,404,449,475]" pageId="10" pageNumber="11">Stratum typicum</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9BFF8BFF0CFE09BCA7CF2C" blockId="10.[189,1399,496,557]" pageId="10" pageNumber="11">
|
||
<collectingCountry id="F30CD4F4FF9BFF8BFF0CFE09BE8ECF0B" box="[189,457,496,522]" name="United Kingdom" pageId="10" pageNumber="11">UNITED KINGDOM</collectingCountry>
|
||
: Lower Lias. Based on lithological comparisons, most likely the Black Ven Member of the Charmouth Mudstone Formation (
|
||
<bibRefCitation id="EF8AE995FF9BFF8BFCB5FDEABC93CF2C" author="Forey P. L. & Longbottom A. & Mulley J." box="[772,980,531,557]" pageId="10" pageNumber="11" pagination="341 - 371" refId="ref13236" refString="Forey P. L., Longbottom A. & Mulley J. 2010. Fishes - bony fishes. In: Lord A. R. & Davis P. G. (eds) Palaeontological Association Field Guides to Fossils, Number 13: Fossils from the Lower Lias of the Dorset Coast: 341 - 371. Palaeontological Association, London." type="book chapter" year="2010">
|
||
Forey
|
||
<emphasis id="B96F4876FF9BFF8BFCE3FDEDBCC2CF2C" box="[850,901,531,557]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9BFF8BFF0CFDACBE2ECF6F" blockId="10.[189,361,596,623]" box="[189,361,596,623]" pageId="10" pageNumber="11">
|
||
<heading id="D0EC2308FF9BFF8BFF0CFDACBE2ECF6F" bold="true" box="[189,361,596,623]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="B96F4876FF9BFF8BFF0CFDACBE2ECF6F" bold="true" box="[189,361,596,623]" pageId="10" pageNumber="11">Locus typicus</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9BFF8BFF0CFD7ABD87CF9C" blockId="10.[189,704,643,669]" box="[189,704,643,669]" pageId="10" pageNumber="11">
|
||
<collectingCountry id="F30CD4F4FF9BFF8BFF0CFD7ABE84CF9C" box="[189,451,643,669]" name="United Kingdom" pageId="10" pageNumber="11">UNITED KINGDOM</collectingCountry>
|
||
: Lyme Regis,
|
||
<collectingRegion id="49DF5A86FF9BFF8BFDDBFD7ABDFBCF9C" box="[618,700,643,669]" country="United Kingdom" name="Dorset" pageId="10" pageNumber="11">Dorset</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF9BFF8AFF0CFD3DBEA3CF26" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
|
||
<paragraph id="8BA49464FF9BFF8BFF0CFD3DBE8ECFDF" blockId="10.[189,457,708,734]" box="[189,457,708,734]" pageId="10" pageNumber="11">
|
||
<heading id="D0EC2308FF9BFF8BFF0CFD3DBE8ECFDF" bold="true" box="[189,457,708,734]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="B96F4876FF9BFF8BFF0CFD3DBE8ECFDF" bold="true" box="[189,457,708,734]" pageId="10" pageNumber="11">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9BFF8AFF0CFD0ABEA3CF26" blockId="10.[189,1399,755,852]" lastBlockId="11.[189,1399,267,551]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
This species can be differentiated from all other species of
|
||
<taxonomicName id="4C1BEFE7FF9BFF8BFCD6FD0ABB50CE0C" authorityName="Reis" authorityYear="1892" box="[871,1047,755,781]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF9BFF8BFCD6FD0ABB50CE0C" box="[871,1047,755,781]" italics="true" pageId="10" pageNumber="11">Saurorhynchus</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: slender, elongate skull (length to depth greater than in
|
||
<taxonomicName id="4C1BEFE7FF9BFF8BFC4AFCEEBBE5CE31" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[1019,1186,790,817]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9BFF8BFC4AFCEEBBE5CE31" box="[1019,1186,790,817]" italics="true" pageId="10" pageNumber="11">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
); posterior dorsal skull roof essentially parallel to the long axis of the skull; postorbital segment much longer than the depth of the lower jaw (almost double) and double the length of the orbit (unlike in
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFB87FEF5BBD8CC24" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[1078,1183,268,294]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9AFF8AFB87FEF5BBD8CC24" box="[1078,1183,268,294]" italics="true" pageId="11" pageNumber="12">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
); anterior external narial opening teardrop shaped, widening ventrally; parasphenoid rostrum bearing small denticles ventral to the orbit; foramen for the internal carotid and efferent pseudobranchial arteries displaced posteriorly relative to the basisphenoid and almost entirely enclosed by the parasphenoid (unlike in
|
||
<emphasis id="B96F4876FF9AFF8AFB7AFE83BE6FCCB8" italics="true" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFB7AFE83BA35CC95" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[1227,1394,378,404]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="brevirostris">S. brevirostris</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFF0CFE66BE6FCCB8" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[189,296,415,441]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acutus">S. acutus</taxonomicName>
|
||
</emphasis>
|
||
); posterior edge of mandible straight or only weakly sinusoidal, becoming gradually rounded ventrally; angle between the posterior and ventral edges of the mandible greater than 90 degrees; acrodin caps of posterior laniaries not directed lingually (unlike in
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFCDBFE10BB49CF03" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[874,1038,488,515]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9AFF8AFCDBFE10BB49CF03" box="[874,1038,488,515]" italics="true" pageId="11" pageNumber="12">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
); opercle roughly triangular in shape, tapering ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF64C4ECFF9BFF8BFF0CF96ABC80CAE8" ID-DOI="http://doi.org/10.5281/zenodo.3829291" ID-Zenodo-Dep="3829291" httpUri="https://zenodo.org/record/3829291/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1683,1709]" targetBox="[189,1399,933,1642]" targetPageId="10">
|
||
<paragraph id="8BA49464FF9BFF8BFF0CF96ABC80CAE8" blockId="10.[189,1399,1683,2026]" pageId="10" pageNumber="11">
|
||
<emphasis id="B96F4876FF9BFF8BFF0CF96ABE49CBAC" bold="true" box="[189,270,1683,1709]" pageId="10" pageNumber="11">Fig. 4.</emphasis>
|
||
Orbitotemporal region, illustrating variation in the braincase of Early Jurassic saurichthyids.
|
||
<emphasis id="B96F4876FF9BFF8BFF0CF94FBF95CBD1" bold="true" box="[189,210,1718,1744]" pageId="10" pageNumber="11">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9BFF8BFF51F94FBD44CBD1" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[224,515,1718,1744]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9BFF8BFF51F94FBD44CBD1" box="[224,515,1718,1744]" italics="true" pageId="10" pageNumber="11">Saurorhynchus anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9BFF8BFDBBF94FBD24CBD1" box="[522,611,1718,1744]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(based on NHMUK PV P 36227).
|
||
<emphasis id="B96F4876FF9BFF8BFC46F94FBB6BCBD1" bold="true" box="[1015,1068,1718,1744]" pageId="10" pageNumber="11">B, E</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9BFF8BFB88F94FBED0CBF5" authority="(Woodward, 1895)" baseAuthorityName="Woodward" baseAuthorityYear="1895" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9BFF8BFB88F94FBA30CBD1" box="[1081,1399,1718,1744]" italics="true" pageId="10" pageNumber="11">Saurorhynchus brevirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF9BFF8BFF74F920BEC8CBF5" author="Woodward A. S." box="[197,399,1753,1780]" pageId="10" pageNumber="11" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(photo pertains to NHMUK PV P 4878, mirrored).
|
||
<emphasis id="B96F4876FF9BFF8BFC53F920BB52CBF2" bold="true" box="[994,1045,1753,1779]" pageId="10" pageNumber="11">C, F</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9BFF8BFB93F920BA5DCBF2" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1058,1306,1753,1779]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9BFF8BFB93F920BA5DCBF2" box="[1058,1306,1753,1779]" italics="true" pageId="10" pageNumber="11">Saurorhynchus hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9BFF8BFAAEF923BA30CBF5" box="[1311,1399,1754,1780]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(photo is of SMNS 51888, mirrored).
|
||
<emphasis id="B96F4876FF9BFF8BFDDCF904BDC5CA16" bold="true" box="[621,642,1789,1815]" pageId="10" pageNumber="11">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9BFF8BFD3EF905BB0ACA17" authority="(Agassiz, 1844)" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[655,1101,1788,1814]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9BFF8BFD3EF905BCD6CA17" box="[655,913,1788,1814]" italics="true" pageId="10" pageNumber="11">Saurorhynchus acutus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF9BFF8BFC2FF905BB02CA17" author="Agassiz L." box="[926,1093,1788,1814]" pageId="10" pageNumber="11" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz, 1844</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(based on SMNS 87737). Abbreviations: asc = ascending process of the parasphenoid; bs = basisphenoid; dpt = dermopterotic; dsp = dermosphenotic; f.ic+ep = foramen for the internal carotid artery and efferent pseudobranchial artery; f.oma = f.om = foramen for the great ophthalmic artery; my = posterior myodome; n = neomorph; nc = neurocranium; orb = orbit; psr = parasphenoid rostrum; rot = foramen for the lateral otic ramus; spi = ventral opening of the spiracular canal; tf.c = trigeminofacial chamber. Scale bars: E–F = 2 mm. E. Photo © The Trustees of the Natural History Museum, London.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C301C7EFFF9AFF8DFF0CFDA9BA70C9B9" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8BA49464FF9AFF8AFF0CFDA9BE09CF6B" blockId="11.[189,334,592,618]" box="[189,334,592,618]" pageId="11" pageNumber="12">
|
||
<heading id="D0EC2308FF9AFF8AFF0CFDA9BE09CF6B" bold="true" box="[189,334,592,618]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B96F4876FF9AFF8AFF0CFDA9BE09CF6B" bold="true" box="[189,334,592,618]" pageId="11" pageNumber="12">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9AFF8AFF0CFD79BCB1CFE5" blockId="11.[189,1398,640,741]" pageId="11" pageNumber="12">
|
||
The material referred herein to
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFD8CFD78BD8CCF9B" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[573,715,641,667]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9AFF8AFD8CFD78BD8CCF9B" box="[573,715,641,667]" italics="true" pageId="11" pageNumber="12">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9AFF8AFD67FD78BC73CF9A" box="[726,820,641,667]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been well-described by
|
||
<bibRefCitation id="EF8AE995FF9AFF8AFB23FD79BA14CF9A" author="Gardiner B. G." box="[1170,1363,640,667]" pageId="11" pageNumber="12" pagination="241 - 384" refId="ref13301" refString="Gardiner B. G. 1960. A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias. Bulletin of the British Museum (Natural History): Geology 4 (7): 241 - 384." type="journal article" year="1960">Gardiner (1960)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFF0CFD5FBE60CFBE" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[189,295,678,704]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9AFF8AFF0CFD5FBE60CFBE" box="[189,295,678,704]" italics="true" pageId="11" pageNumber="12">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
. The following description focuses on details in which our interpretations differ from those of Gardiner, or those structures for which additional details can be noted.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9AFF8AFF0CFCF4BD8DCE70" blockId="11.[189,1399,781,881]" pageId="11" pageNumber="12">
|
||
The
|
||
<typeStatus id="54A02AC6FF9AFF8AFF45FCF4BE1CCE26" box="[244,347,781,807]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
skull is only
|
||
<quantity id="4CE33981FF9AFF8AFDB1FCF4BD12CE29" box="[512,597,781,808]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="11" pageNumber="12" unit="mm" value="89.0">89 mm</quantity>
|
||
long, however the largest skull referable to
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFBD9FCF7BBB3CE26" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1128,1268,782,808]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9AFF8AFBD9FCF7BBB3CE26" box="[1128,1268,782,808]" italics="true" pageId="11" pageNumber="12">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9AFF8AFB4FFCF7BA1DCE29" box="[1278,1370,782,808]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is
|
||
<quantity id="4CE33981FF9AFF8AFF0CFCCBBE66CE4C" box="[189,289,818,845]" metricMagnitude="-1" metricUnit="m" metricValue="1.3699999999999999" pageId="11" pageNumber="12" unit="mm" value="137.0">137 mm</quantity>
|
||
in length (Suppl. Table 1). Assuming similar proportions to
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFC4DFCCBBB19CE4D" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1020,1118,818,844]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9AFF8AFC4DFCCBBB19CE4D" box="[1020,1118,818,844]" italics="true" pageId="11" pageNumber="12">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9AFF8AFBD9FCCABB80CE4C" box="[1128,1223,819,845]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, fork length is estimated at
|
||
<quantity id="4CE33981FF9AFF8AFEE0FCAEBEDDCE70" box="[337,410,855,881]" metricMagnitude="-1" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="12" unit="cm" value="48.0">48 cm</quantity>
|
||
in the largest individuals.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9AFF8AFF0CFC63BBF0C9DA" blockId="11.[189,1399,922,1243]" pageId="11" pageNumber="12">The antorbital rostrum is composed of the rostropremaxilla, nasaloantorbitals, and frontals. There are two external narial openings, the anterior of which is teardrop shaped and somewhat narrower than reconstructed by Gardiner. The posterior narial opening is small and circular in outline. The suture between the maxilla and premaxilla occurs ventral to the nares. Between the maxilla and the nasaloantorbital is an extremely weakly ossified area that would have borne the infraorbital canal. This space corresponds to the lacrimal, and is more anteriorly extensive than in Gardiner’s reconstruction. Gardiner noted interrostral elements in some of the specimens he examined; we could not confirm the presence of these separate ossifications during restudy of the material. He also noted separate nasal and antorbital ossifications; this observation appears to be based on post-mortem breakage.</paragraph>
|
||
<paragraph id="8BA49464FF9AFF8AFF0CFAFDBB03C8D7" blockId="11.[189,1399,1284,1494]" pageId="11" pageNumber="12">
|
||
The frontal plays a small role in the dorsal edge of the orbit (
|
||
<figureCitation id="132088E1FF9AFF8AFC2EFAFDBCB8C81F" box="[927,1023,1284,1310]" captionStart="Fig" captionStartId="9.[189,232,1718,1744]" captionTargetBox="[195,1396,311,1622]" captionTargetId="graphics@9.[317,1359,731,1036]" captionTargetPageId="9" captionText="Fig. 3. Dorsal skull roof. A. Saurorhynchus anningae sp. nov. (NHMUK PV P 964); note that this skull has been dorsoventrally compressed. B. Saurorhynchus hauffi sp. nov., three-dimensionally preserved skull GG 20001. Fine dashed lines indicate sensory canals, large dashes indicate broken or incomplete areas; grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l.ex = lateral extrascapular; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; p = parietal; pm = rostropremaxilla; so.c = supraorbital sensory canal. Scale bars = 10 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829288" httpUri="https://zenodo.org/record/3829288/files/figure.png" pageId="11" pageNumber="12">Fig. 3A</figureCitation>
|
||
). Posterior to the lateral frontal is a small, triangular dermosphenotic, which carries the infraorbital sensory canal and excludes the dermopterotic from the orbit. Ventral to the dermosphenotic is a curved element forming the posterior edge of the orbit and carrying the infraorbital sensory canal; this was identified by Gardiner as the dermosphenotic but is here considered to be the penultimate infraorbital. In some of the most heavily ossified specimens, this element can be quite robust and heavily ornamented.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9AFF8AFF0CFA06BE8FCB86" blockId="11.[189,1399,1535,1671]" pageId="11" pageNumber="12">
|
||
The lateral extrascapular extends as far anteriorly as the hyomandibula-dermopterotic articulation, unlike in
|
||
<taxonomicName id="4C1BEFE7FF9AFF8AFE9BF9DDBED3CB3F" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[298,404,1572,1598]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9AFF8AFE9BF9DDBED3CB3F" box="[298,404,1572,1598]" italics="true" pageId="11" pageNumber="12">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
, and more anteriorly than in Gardiner’s reconstruction. A dermohyal is present, fused to the lateral surface of the dorsal hyomandibula (as noted by
|
||
<bibRefCitation id="EF8AE995FF9AFF8AFC1AF9B0BB1CCB65" author="Gardiner B. G." box="[939,1115,1609,1636]" pageId="11" pageNumber="12" pagination="241 - 384" refId="ref13301" refString="Gardiner B. G. 1960. A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias. Bulletin of the British Museum (Natural History): Geology 4 (7): 241 - 384." type="journal article" year="1960">Gardiner 1960</bibRefCitation>
|
||
). The opercle is also as described by Gardiner.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9AFF8AFF0CF948BD0DCA82" blockId="11.[189,1399,1712,1924]" pageId="11" pageNumber="12">
|
||
None of the specimens examined in the present study had a well-preserved ascending process of the parasphenoid, so the posterior position of the foramen for the orbital artery could not be confirmed. Anterior to the ascending process, the parasphenoid bears denticles along its ventral surface. In lateral view, the foramen for the internal carotid and efferent pseudobranchial arteries lies ventral to the opening for the posterior myodome (
|
||
<figureCitation id="132088E1FF9AFF8AFDA2F8BDBD32CA5F" box="[531,629,1860,1886]" captionStart="Fig" captionStartId="10.[189,232,1683,1709]" captionTargetBox="[189,1399,933,1642]" captionTargetId="figure@10.[189,1399,929,1642]" captionTargetPageId="10" captionText="Fig. 4. Orbitotemporal region, illustrating variation in the braincase of Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov. (based on NHMUK PV P 36227). B, E. Saurorhynchus brevirostris (Woodward, 1895) (photo pertains to NHMUK PV P 4878, mirrored). C, F. Saurorhynchus hauffi sp. nov. (photo is of SMNS 51888, mirrored). D. Saurorhynchus acutus (Agassiz, 1844) (based on SMNS 87737). Abbreviations: asc = ascending process of the parasphenoid; bs = basisphenoid; dpt = dermopterotic; dsp = dermosphenotic; f.ic+ep = foramen for the internal carotid artery and efferent pseudobranchial artery; f.oma = f.om = foramen for the great ophthalmic artery; my = posterior myodome; n = neomorph; nc = neurocranium; orb = orbit; psr = parasphenoid rostrum; rot = foramen for the lateral otic ramus; spi = ventral opening of the spiracular canal; tf.c = trigeminofacial chamber. Scale bars: E–F = 2 mm. E. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829291" httpUri="https://zenodo.org/record/3829291/files/figure.png" pageId="11" pageNumber="12">Fig. 4A</figureCitation>
|
||
), rather than ventral to the anterior basisphenoid as in Toarcian species (
|
||
<bibRefCitation id="EF8AE995FF9AFF8AFE90F890BEEDCA82" author="Wenz S." box="[289,426,1897,1923]" pageId="11" pageNumber="12" pagination="1 - 276" refId="ref15053" refString="Wenz S. 1967. Complements a l'etude des poissons actinopterygiens du Jurassique Francais. Cahiers de Paleontologie 1967: 1 - 276." type="journal article" year="1967">Wenz 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8AE995FF9AFF8AFE07F890BD7DCA85" author="Thies D." box="[438,570,1897,1924]" pageId="11" pageNumber="12" pagination="183 - 203" refId="ref14987" refString="Thies D. 1985. Funde von Acidorhynchus brevirostris (Woodward 1895) aus dem Posidonienschiefer (Unter-Toarcium) NW-Deutschlands. Palaeontographica Abt. A 187 (4 - 6): 183 - 203." type="journal article" year="1985">Thies 1985</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9AFF8AFF0CF855BA31CAEA" blockId="11.[189,1399,1964,2028]" pageId="11" pageNumber="12">
|
||
The mandible consists of three elements in lateral view: the dentary, angular and supraangular. The configuration of the angular-dentary suture reconstructed by
|
||
<bibRefCitation id="EF8AE995FF9AFF8AFC2FF828BB18CAED" author="Gardiner B. G." box="[926,1119,2001,2028]" pageId="11" pageNumber="12" pagination="241 - 384" refId="ref13301" refString="Gardiner B. G. 1960. A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias. Bulletin of the British Museum (Natural History): Geology 4 (7): 241 - 384." type="journal article" year="1960">Gardiner (1960)</bibRefCitation>
|
||
is incorrect, as argued
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9DFF8DFF0CFEF2BB6ECC48" blockId="12.[189,1399,267,329]" pageId="12" pageNumber="13">
|
||
elsewhere (
|
||
<bibRefCitation id="EF8AE995FF9DFF8DFEF3FEF2BEA6CC27" author="Griffith J." box="[322,481,267,294]" pageId="12" pageNumber="13" pagination="344 - 354" refId="ref13417" refString="Griffith J. 1962. The Triassic fish Saurichthys krambergeri Schlosser. Palaeontology 5 (2): 344 - 354." type="journal article" year="1962">Griffith 1962</bibRefCitation>
|
||
). The angular is extensively exposed along the lateral lower jaw, and extends anteriorly approximately as far as the anterior edge of the nasaloantorbital.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9DFF8DFF0CFE88BE91CE13" blockId="12.[189,1399,369,786]" pageId="12" pageNumber="13">
|
||
The dentary and rostropremaxilla bear large (laniary) teeth, each flanked by a pair of smaller teeth. Laniary teeth are all positioned well anterior to the external narial openings. The posteriormost laniary teeth have relatively straight crowns with acrodin caps, which fit into corresponding pits in the opposite jaw (incisivlücken). Among the posteriormost laniary teeth, the upper teeth overlap the dentary, resulting in incisivlücken on the lateral lower jaw, whereas the lower jaw is slightly narrower and so the laniaries do not project outside the mouth, resulting in an absence of incisivlücken on the lateral surface of the posterior premaxilla (
|
||
<figureCitation id="132088E1FF9DFF8DFDD1FDBFBDFBCF60" box="[608,700,582,609]" captionStart="Fig" captionStartId="14.[189,232,1828,1854]" captionTargetBox="[198,1392,811,1788]" captionTargetId="figure@14.[195,1392,808,1788]" captionTargetPageId="14" captionText="Fig. 6. Posteriormost laniary dentition. A. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV P 4878), note the lingually curved crowns. B. Saurorhynchus anningae sp. nov. (NHMUK PV P 27569), note the incisivlücke associated with the premaxillary laniary tooth and the absence of lateral crypts associated with the two flanking mandibular laniaries. Photos © The Trustees of the Natural History Museum, London. C. S. hauffi sp. nov. (SMNS 51007), mid-rostral dentition illustrating the relationship between the incisivlücken, laniaries, and flanking smaller teeth. Scale bars: 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3829295" httpUri="https://zenodo.org/record/3829295/files/figure.png" pageId="12" pageNumber="13">Fig. 6B</figureCitation>
|
||
). The largest teeth are found in the middle of the tooth row; the anteriormost laniary teeth are quite small, similar in size to the flanking teeth. All teeth consist of an acrodin cap, a lightly corrugated region of collar enamel, and an uncorrugated base. Plicidentine appears to be developed around the very base of the tooth, but does not form external ridges above the level of the jawbone. The difference in tooth shape observed between posterior and anterior teeth in
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFF0CFD01BE26CE13" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[189,353,760,786]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9DFF8DFF0CFD01BE26CE13" box="[189,353,760,786]" italics="true" pageId="12" pageNumber="13">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
is absent.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9DFF8DFF0CFCC3BA70C9B9" blockId="12.[189,1399,826,1208]" pageId="12" pageNumber="13">
|
||
Few specimens are available with well-preserved postcrania. The
|
||
<typeStatus id="54A02AC6FF9DFF8DFC4BFCC3BB26CE55" box="[1018,1121,826,852]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
specimen preserves a discrete supracleithrum bearing ornamentation on its external surface (
|
||
<figureCitation id="132088E1FF9DFF8DFBB5FCA4BB26CE76" box="[1028,1121,861,887]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="12" pageNumber="13">Fig. 1C</figureCitation>
|
||
). A triradiate cleithrum is also present. NHMUK PV P 3790 preserves the most complete postcranium. The neural arch-like elements are small and blocky, similar to those of many saurichthyids. Anteriorly, neural spines are small or absent, and elongate anterior and posterior zygapophyses overlap to form a lattice. In the region around the median fins, neural spines appear to be present, but poor preservation makes this homology interpretation questionable. Haemal spines are preserved in a 1:1 relationship with the neural arch-like elements along the posterior part of the block, dorsal to some poorly preserved anal axonosts. The haemal spines are elongate and bifurcate ventrally. Only a single scale row is present anterior to the median fins, the mid-dorsal scale row (
|
||
<figureCitation id="132088E1FF9DFF8DFD33FB83BDA7C995" box="[642,736,1146,1172]" captionStart="Fig" captionStartId="16.[189,232,1753,1779]" captionTargetBox="[189,1399,836,1707]" captionTargetId="figure@16.[189,1399,832,1707]" captionTargetPageId="16" captionText="Fig. 7. Postcranium, Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov., neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790). B–D. Saurorhynchus hauffi sp. nov., SMNS 55057. B. Lepidotrichia of the anal fin. C. Relationship between the neural and haemal arches and the axonosts. D. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="12" pageNumber="13">Fig. 7A</figureCitation>
|
||
). The scales are smooth and needle-like, tapering at their anterior and posterior ends. Posterior to the anal axonosts, a mid-ventral scale row is also present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF9DFF8DFF0CFB26BD55C8FC" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="8BA49464FF9DFF8DFF0CFB26BE68C9F8" blockId="12.[189,303,1247,1273]" box="[189,303,1247,1273]" pageId="12" pageNumber="13">
|
||
<heading id="D0EC2308FF9DFF8DFF0CFB26BE68C9F8" bold="true" box="[189,303,1247,1273]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B96F4876FF9DFF8DFF0CFB26BE68C9F8" bold="true" box="[189,303,1247,1273]" pageId="12" pageNumber="13">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9DFF8DFF0CFAF7BD55C8FC" blockId="12.[189,1399,1294,1533]" pageId="12" pageNumber="13">
|
||
<bibRefCitation id="EF8AE995FF9DFF8DFF0CFAF7BEE1C829" author="Agassiz L." box="[189,422,1294,1320]" pageId="12" pageNumber="13" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">
|
||
Agassiz (1844:
|
||
<quantity id="4CE33981FF9DFF8DFECBFAF7BEE1C829" box="[378,422,1294,1320]" metricMagnitude="0" metricUnit="m" metricValue="3.6322" pageId="12" pageNumber="13" unit="in" value="143.0">143</quantity>
|
||
</bibRefCitation>
|
||
in pt. 2) originally coined the name “
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFCC2FAF7BBD0C829" box="[883,1175,1294,1320]" class="Actinopterygii" family="Aspidorhynchidae" genus="Belonostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidorhychiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="anningiae">
|
||
<emphasis id="B96F4876FF9DFF8DFCC2FAF7BBD0C829" box="[883,1175,1294,1320]" italics="true" pageId="12" pageNumber="13">Belonostomus anningiae</emphasis>
|
||
</taxonomicName>
|
||
” for specimens of
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFF0CFACBBE2AC84D" authorityName="Reis" authorityYear="1892" box="[189,365,1330,1356]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF9DFF8DFF0CFACBBE2AC84D" box="[189,365,1330,1356]" italics="true" pageId="12" pageNumber="13">Saurorhynchus</emphasis>
|
||
</taxonomicName>
|
||
from the Lower Lias of the
|
||
<collectingCountry id="F30CD4F4FF9DFF8DFD03FACBBD98C84D" box="[690,735,1330,1356]" name="United Kingdom" pageId="12" pageNumber="13">UK</collectingCountry>
|
||
, but failed to describe the material or identify a
|
||
<typeStatus id="54A02AC6FF9DFF8DFAB9FACBBA34C84D" box="[1288,1395,1330,1356]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
. Subsequently,
|
||
<bibRefCitation id="EF8AE995FF9DFF8DFED7FAACBD7CC86E" author="Woodward A. S." box="[358,571,1365,1391]" pageId="12" pageNumber="13" pagination="144 - 148" refId="ref15136" refString="Woodward A. S. 1888. On some remains of Squatina cranei, sp. nov., and the mandible of Belonostomus cinctus, from the chalk of Sussex, preserved in the collection of Henry Willett, Esq., F. G. S., Brighton Museum. Quarterly Journal of the Geological Society of London 44: 144 - 148. Available from http: // biodiversitylibrary. org / page / 37059663 [accessed 28 Mar. 2017]." type="journal article" year="1888">Woodward (1888)</bibRefCitation>
|
||
figured a non-diagnostic specimen as
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFC46FAAFBBCCC86E" box="[1015,1163,1366,1391]" class="Actinopterygii" family="Aspidorhynchidae" genus="Belonostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidorhychiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="anningiae">
|
||
<emphasis id="B96F4876FF9DFF8DFC46FAAFBBCCC86E" box="[1015,1163,1366,1391]" italics="true" pageId="12" pageNumber="13">B. anningiae</emphasis>
|
||
</taxonomicName>
|
||
, but did not provide a description either and subsequently synonymized
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFCA0FA83BCE2C892" box="[785,933,1402,1427]" class="Actinopterygii" family="Aspidorhynchidae" genus="Belonostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidorhychiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="anningiae">
|
||
<emphasis id="B96F4876FF9DFF8DFCA0FA83BCE2C892" box="[785,933,1402,1427]" italics="true" pageId="12" pageNumber="13">B. anningiae</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFC54FA80BE46C8B6" authority="(Woodward 1895)" baseAuthorityName="Woodward" baseAuthorityYear="1895" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9DFF8DFC54FA80BBA0C892" box="[997,1255,1401,1427]" italics="true" pageId="12" pageNumber="13">Saurorhynchus acutus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF9DFF8DFB45FA80BFBEC8B6" author="Woodward A. S." pageId="12" pageNumber="13" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Thus, the name
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFE78FA65BDB2C8B7" box="[457,757,1436,1462]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="anningiae">
|
||
<emphasis id="B96F4876FF9DFF8DFE78FA65BDB2C8B7" box="[457,757,1436,1462]" italics="true" pageId="12" pageNumber="13">Saurorhynchus anningiae</emphasis>
|
||
</taxonomicName>
|
||
is considered a
|
||
<emphasis id="B96F4876FF9DFF8DFC00FA64BB11C8B7" box="[945,1110,1436,1462]" italics="true" pageId="12" pageNumber="13">nomen nudum</emphasis>
|
||
, and is still available for the longirostrine material from the Charmouth Mudstone Formation (Sinemurian) of the
|
||
<collectingCountry id="F30CD4F4FF9DFF8DFB62FA39BBBAC8DB" box="[1235,1277,1472,1498]" name="United Kingdom" pageId="12" pageNumber="13">UK</collectingCountry>
|
||
as per the original intention of Agassiz.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF9DFF8DFF0CF9DCBC1BCA21" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="8BA49464FF9DFF8DFF0CF9DCBE08CB3E" blockId="12.[189,335,1573,1599]" box="[189,335,1573,1599]" pageId="12" pageNumber="13">
|
||
<heading id="D0EC2308FF9DFF8DFF0CF9DCBE08CB3E" bold="true" box="[189,335,1573,1599]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B96F4876FF9DFF8DFF0CF9DCBE08CB3E" bold="true" box="[189,335,1573,1599]" pageId="12" pageNumber="13">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9DFF8DFF0CF9ADBC1BCA21" blockId="12.[189,1399,1620,1825]" pageId="12" pageNumber="13">
|
||
This species is currently known from the “Lower Lias” of Lyme Regis and Charmouth (
|
||
<collectingCountry id="F30CD4F4FF9DFF8DFB02F9ADBBA7CB6F" box="[1203,1248,1620,1646]" name="United Kingdom" pageId="12" pageNumber="13">UK</collectingCountry>
|
||
) only. Based on lithological comparisons,
|
||
<bibRefCitation id="EF8AE995FF9DFF8DFDA9F981BDBDCB93" author="Forey P. L. & Longbottom A. & Mulley J." box="[536,762,1656,1682]" pageId="12" pageNumber="13" pagination="341 - 371" refId="ref13236" refString="Forey P. L., Longbottom A. & Mulley J. 2010. Fishes - bony fishes. In: Lord A. R. & Davis P. G. (eds) Palaeontological Association Field Guides to Fossils, Number 13: Fossils from the Lower Lias of the Dorset Coast: 341 - 371. Palaeontological Association, London." type="book chapter" year="2010">
|
||
Forey
|
||
<emphasis id="B96F4876FF9DFF8DFDD7F980BDE4CB93" box="[614,675,1656,1682]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
surmised that the material of
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFBD4F981BBB5CB93" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1125,1266,1656,1682]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9DFF8DFBD4F981BBB5CB93" box="[1125,1266,1656,1682]" italics="true" pageId="12" pageNumber="13">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9DFF8DFB4AF981BA1FCB93" box="[1275,1368,1656,1682]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
at the NHMUK originated from the Black Ven Mudstone Member of the Charmouth Mudstone Formation (?
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFF65F939BE62CBD8" baseAuthorityName="Sowerby" baseAuthorityYear="1824" box="[212,293,1728,1753]" class="Cephalopoda" family="Asteroceratidae" genus="Caenisites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="turneri">
|
||
<emphasis id="B96F4876FF9DFF8DFF65F939BE62CBD8" box="[212,293,1728,1753]" italics="true" pageId="12" pageNumber="13">turneri</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFEFFF939BD7DCBD8" authority="Zones" authorityName="Zones" box="[334,570,1727,1753]" class="Cephalopoda" family="Echioceratidae" genus="Echioceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="raricostatum">
|
||
<emphasis id="B96F4876FF9DFF8DFEFFF939BEA1CBD8" box="[334,486,1728,1753]" italics="true" pageId="12" pageNumber="13">raricostatum</emphasis>
|
||
Zones
|
||
</taxonomicName>
|
||
; latest early Sinemurian to late Sinemurian). The single specimen for which more specific stratigraphic provenance is available, NHMUK PV P 27569 from the
|
||
<taxonomicName id="4C1BEFE7FF9DFF8DFB6BF91ABE35CA21" authority="Zone" authorityName="Zone" class="Cephalopoda" family="Oxynoticeratidae" genus="Oxynoticeras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="oxynotum">
|
||
<emphasis id="B96F4876FF9DFF8DFB6BF91ABE68CA21" italics="true" pageId="12" pageNumber="13">Oxynoticeras oxynotum</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
(late Sinemurian), falls within this range.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |