351 lines
42 KiB
XML
351 lines
42 KiB
XML
<document id="C193BA9F58FFFB49522A1C4852DEDD4D" ID-DOI="10.11646/zootaxa.4020.3.5" ID-GBIF-Dataset="9b91a64c-e23b-416a-b1e0-0eceb046ca45" ID-ISSN="1175-5326" ID-Zenodo-Dep="240877" ID-ZooBank="00EB426F-43F8-4701-BAA6-8C54C9B6B44E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461270911891" checkinUser="plazi" docAuthor="Harvey, Michael B., O’Connell, Kyle A., Barraza, Gabriel, Riyanto, Awal, Kurniawan, Nia & Smith, Eric N." docDate="2015" docId="039C296EFFE5FFA014F69BC160ECF98B" docLanguage="en" docName="zt04020p516.pdf" docOrigin="Zootaxa 4020 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus semicinctus Harvey, O’Connell, Barraza, Riyanto, Kurniawan & Smith, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="507" masterDocId="FFA55116FFEDFFAC14619F0E620CFFA7" masterDocTitle="Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the Southern Bukit Barisan Range of Sumatra and an estimation of their phylogeny" masterLastPageNumber="516" masterPageNumber="495" pageNumber="503" updateTime="1698611758727" updateUser="plazi">
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<mods:title id="6FBA7C148079CA6792BD98B6976D6A5F">Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the Southern Bukit Barisan Range of Sumatra and an estimation of their phylogeny</mods:title>
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<mods:namePart id="54D92A03CD9A41393F46377F8B0E9922">Harvey, Michael B.</mods:namePart>
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<mods:namePart id="7B0170D7AE21B305E795C63CC128F1C1">O’Connell, Kyle A.</mods:namePart>
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</mods:role>
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<mods:namePart id="9A7E182625081DE30F77D4ABA69649C5">Barraza, Gabriel</mods:namePart>
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<mods:namePart id="7E1B19F939143AF13A67959CCCDF912B">Riyanto, Awal</mods:namePart>
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<mods:namePart id="424A43B281104F0FB3556BF6C011BC34">Kurniawan, Nia</mods:namePart>
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<mods:namePart id="7150EFE855063C88017520B2D19CFE39">Smith, Eric N.</mods:namePart>
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<mods:title id="7C705950247955CA8A0641E55828D57B">Zootaxa</mods:title>
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<mods:date id="D3B276CEE8A6BE75ED1AF12783FD9B22">2015</mods:date>
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<mods:number id="1210069771A07115DF1A2ABDE47467F7">4020</mods:number>
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<treatment id="039C296EFFE5FFA014F69BC160ECF98B" ID-DOI="http://doi.org/10.5281/zenodo.5625844" ID-GBIF-Taxon="119638815" ID-Zenodo-Dep="5625844" LSID="urn:lsid:plazi:treatment:039C296EFFE5FFA014F69BC160ECF98B" httpUri="http://treatment.plazi.org/id/039C296EFFE5FFA014F69BC160ECF98B" lastPageId="12" lastPageNumber="507" pageId="8" pageNumber="503">
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<emphasis id="B941446AFFE5FFA414F69BC1672DFB4D" bold="true" box="[151,1313,1231,1258]" pageId="8" pageNumber="503">
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<taxonomicName id="4C35E3FBFFE5FFA414F69BC163DEFB4E" box="[151,466,1231,1257]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="503" phylum="Chordata" rank="species" species="semicinctus" status="sp. nov.">
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<emphasis id="B941446AFFE5FFA414F69BC163DEFB4E" bold="true" box="[151,466,1231,1257]" italics="true" pageId="8" pageNumber="503">Cyrtodactylus semicinctus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A272F911FFE5FFA415B89BDE603BFB4D" box="[473,567,1232,1258]" pageId="8" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
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Harvey, O’Connell, Barraza, Riyanto, Kurniawan & Smith
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C32FCBF3FFE5FFA414F69A186602FA44" pageId="8" pageNumber="503" type="materials_examined">
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<paragraph id="8B8A9878FFE5FFA414F69A1866A8FAD0" blockId="8.[151,1436,1302,2011]" pageId="8" pageNumber="503">
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<emphasis id="B941446AFFE5FFA414F69A186307FA88" bold="true" box="[151,267,1302,1327]" pageId="8" pageNumber="503">
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<typeStatus id="548E26DAFFE5FFA414F69A18630AFA88" box="[151,262,1302,1327]" pageId="8" pageNumber="503" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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An adult male (
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<collectionCode id="ED2400BDFFE5FFA415AB9A196004FA88" box="[458,520,1303,1327]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="503">MZB</collectionCode>
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9703, collector’s tag
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<collectionCode id="ED2400BDFFE5FFA417679A196137FA88" LSID="urn:lsid:biocol.org:col:13248" box="[774,827,1303,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13248" name="Hubei College for Nationalities" pageId="8" pageNumber="503">ENS</collectionCode>
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14958,
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<figureCitation id="130E84FDFFE5FFA417FD9A1861ECFA88" box="[924,992,1302,1327]" captionStart="FIGURE 5" captionStartId="9.[151,249,1239,1261]" captionTargetBox="[151,1436,181,1226]" captionTargetId="figure@9.[151,1436,181,1226]" captionTargetPageId="9" captionText="FIGURE 5. Adult male holotype (A, MZB 9703, head length 20.5 mm) and adult female paratypes (B, UTA R- 61588, SVL 47 mm; C – D, MZB 9694, SVL 83 mm) of Cyrtodactylus semicinctus from montane forest on trail above Kayu Aro, leading to top of Mt. Kerinci, Jambi, Sumatra, Indonesia." httpUri="https://zenodo.org/record/240882/files/figure.png" pageId="8" pageNumber="503">Fig. 5</figureCitation>
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) from montane forest on a trail above Kayu Aro, leading to the top of Mt. Kerinci, Kabupaten Kerinci, Jambi Province, Sumatra,
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<collectingCountry id="F322D8E8FFE5FFA410A19A34673CFAF4" box="[1216,1328,1338,1363]" name="Indonesia" pageId="8" pageNumber="503">Indonesia</collectingCountry>
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,
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<quantity id="4CCD359DFFE5FFA4115C9A34679BFAF5" box="[1341,1431,1338,1363]" metricMagnitude="3" metricUnit="m" metricValue="1.905" pageId="8" pageNumber="503" unit="m" value="1905.0">1905 m</quantity>
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,
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<geoCoordinate id="EE01FEBFFFE5FFA414F69A506319FAD0" box="[151,277,1374,1399]" direction="south" orientation="latitude" pageId="8" pageNumber="503" precision="1" value="-1.74159">1.74159° S</geoCoordinate>
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,
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<geoCoordinate id="EE01FEBFFFE5FFA415439A5063B4FAD0" box="[290,440,1374,1399]" direction="east" orientation="longitude" pageId="8" pageNumber="503" precision="1" value="101.26015">101.26015°E</geoCoordinate>
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, collected on
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<date id="FF8BBEB8FFE5FFA416339A5160EBFAD0" box="[594,743,1375,1399]" pageId="8" pageNumber="503" value="2013-06-24">24 June 2013</date>
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by M. B. Harvey and K. A. O’Connell.
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||
</paragraph>
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||
<paragraph id="8B8A9878FFE5FFA414A69A8C6602FA44" blockId="8.[151,1436,1302,2011]" pageId="8" pageNumber="503">
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<emphasis id="B941446AFFE5FFA414A69A8C6363FA3C" bold="true" box="[199,367,1410,1435]" pageId="8" pageNumber="503">
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<typeStatus id="548E26DAFFE5FFA414A69A8C6333FA3C" box="[199,319,1410,1435]" pageId="8" pageNumber="503" type="paratype">Paratypes</typeStatus>
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(7).
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</emphasis>
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Two males (
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<collectionCode id="ED2400BDFFE5FFA416629A8D604DFA3C" box="[515,577,1411,1435]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="503">MZB</collectionCode>
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9695, 9696), four females (
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<collectionCode id="ED2400BDFFE5FFA417E19A8D61B2FA3C" box="[896,958,1411,1435]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="503">MZB</collectionCode>
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9694,
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<collectionCode id="ED2400BDFFE5FFA4106F9A8D6649FA3C" LSID="urn:lsid:biocol.org:col:34873" box="[1038,1093,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="8" pageNumber="503">UTA</collectionCode>
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<collectionCode id="ED2400BDFFE5FFA4102F9A8C6666FA3C" box="[1102,1130,1410,1435]" pageId="8" pageNumber="503">R-</collectionCode>
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61576, 61577, 61588), and one unsexed juvenile (
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<collectionCode id="ED2400BDFFE5FFA415F09AA963C3FA18" box="[401,463,1447,1471]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="503">MZB</collectionCode>
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9701) collected from the same trail as the
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<typeStatus id="548E26DAFFE5FFA417CA9AA8661CFA18" box="[939,1040,1446,1471]" pageId="8" pageNumber="503" type="holotype">holotype</typeStatus>
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,
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<quantity id="4CCD359DFFE5FFA4107B9AA866BBFA19" box="[1050,1207,1446,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.879" metricValueMax="2.006" metricValueMin="1.752" pageId="8" pageNumber="503" unit="m" value="1879.0" valueMax="2006.0" valueMin="1752.0">1752–2006 m</quantity>
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, on
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<date id="FF8BBEB8FFE5FFA410849AA96776FA18" box="[1253,1402,1447,1471]" pageId="8" pageNumber="503" value="2014-06-21">21 June 2014</date>
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by M. B. Harvey, E. N. Smith, and other members of the summer 2013 field party.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32FCBF3FFE5FFA414A69AE06736F87C" pageId="8" pageNumber="503" type="diagnosis">
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<paragraph id="8B8A9878FFE5FFA414A69AE06736F87C" blockId="8.[151,1436,1302,2011]" pageId="8" pageNumber="503">
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<emphasis id="B941446AFFE5FFA414A69AE0634DF9A0" bold="true" box="[199,321,1518,1543]" pageId="8" pageNumber="503">Diagnosis.</emphasis>
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A medium size species of
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<taxonomicName id="4C35E3FBFFE5FFA416FD9AE06135F9A0" box="[668,825,1518,1543]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="503" phylum="Chordata" rank="genus">
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<emphasis id="B941446AFFE5FFA416FD9AE06135F9A0" box="[668,825,1518,1543]" italics="true" pageId="8" pageNumber="503">Cyrtodactylus</emphasis>
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</taxonomicName>
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reaching
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<quantity id="4CCD359DFFE5FFA417D69AE16602F9A1" box="[951,1038,1519,1543]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="8" pageNumber="503" unit="mm" value="89.0">89 mm</quantity>
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SVL and distinguished from all congeners by the following combination of characters: (1) body moderately robust; limbs and digits relatively long; (2) tail 1.05–1.19 times longer than body; (3) subpyramidal, weakly keeled tubercles extending from parietal region and supraorbital skin to base of tail, 24–27 irregular longitudinal rows of tubercles at midbody, 29–35 irregular transverse rows between limbs; (4) dorsal antebrachium tuberculate, brachium lacking tubercles; (5) tubercles on tail restricted to basal third; (6) 33–44 ventrals in a transverse row at midbody; (7) subcaudals not transversely enlarged; (8) deep, subtriangular precloacal depression in males; (9) 36–38 femoral and precloacal pores in males, arrayed in a continuous series (fewer pore primordia usually present in females), sunken into precloacal depression; (10) scale at anterior apex of pore-bearing scales greatly enlarged, sunken into precloacal depression in males, superficial in females; (11) pore-bearing series sharply transitioning to tiny granular scales of postaxial surface of thigh; (12) 19–22 lamellae under fourth toe; (13) cloacal tubercles 2/2 or 3/3 contacting one another; (14) dorsal pattern of brown blotches widely edged in black; tubercles on flanks yellow to pale cream; (15) labials mostly yellow; wide U-shaped band around occiput to loreal region; (16) 5–7 transverse, irregular or broken bands on body; 9–11 regular bands on tail, incomplete ventrally, except near distal tip of some specimens.
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF4AC8F0FFE4FFA514F69BD96045FA8C" httpUri="https://zenodo.org/record/240882/files/figure.png" pageId="9" pageNumber="504" targetBox="[151,1436,181,1226]" targetPageId="9">
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||
<paragraph id="8B8A9878FFE4FFA514F69BD96045FA8C" blockId="9.[151,1435,1239,1323]" pageId="9" pageNumber="504">
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||
<emphasis id="B941446AFFE4FFA514F69BD9631EFB4A" bold="true" box="[151,274,1239,1261]" pageId="9" pageNumber="504">FIGURE 5.</emphasis>
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Adult male holotype (A, MZB 9703, head length 20.5 mm) and adult female paratypes (B, UTA R-61588, SVL 47 mm; C–D, MZB 9694, SVL 83 mm) of
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<taxonomicName id="4C35E3FBFFE4FFA5164B9BF86138FAAB" box="[554,820,1270,1292]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="504" phylum="Chordata" rank="species" species="semicinctus">
|
||
<emphasis id="B941446AFFE4FFA5164B9BF86138FAAB" box="[554,820,1270,1292]" italics="true" pageId="9" pageNumber="504">Cyrtodactylus semicinctus</emphasis>
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||
</taxonomicName>
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from montane forest on trail above Kayu Aro, leading to top of Mt. Kerinci, Jambi, Sumatra, Indonesia.
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="C32FCBF3FFE4FFA014A69A5C63D7FB07" lastPageId="12" lastPageNumber="507" pageId="9" pageNumber="504" type="description">
|
||
<paragraph id="8B8A9878FFE4FFA514A69A5C60A5F9B8" blockId="9.[151,1436,1362,2035]" pageId="9" pageNumber="504">
|
||
<emphasis id="B941446AFFE4FFA514A69A5C6355FACC" bold="true" box="[199,345,1362,1387]" pageId="9" pageNumber="504">Description.</emphasis>
|
||
Medium-size
|
||
<taxonomicName id="4C35E3FBFFE4FFA5159D9A5C6095FACC" box="[508,665,1362,1387]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="504" phylum="Chordata" rank="genus">
|
||
<emphasis id="B941446AFFE4FFA5159D9A5C6095FACC" box="[508,665,1362,1387]" italics="true" pageId="9" pageNumber="504">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
reaching at least
|
||
<quantity id="4CCD359DFFE4FFA517009A5D61BDFACD" box="[865,945,1363,1387]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="9" pageNumber="504" unit="mm" value="89.0">89 mm</quantity>
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SVL in females,
|
||
<quantity id="4CCD359DFFE4FFA5101B9A5C66C6FACD" box="[1146,1226,1362,1387]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="504" unit="mm" value="75.0">75 mm</quantity>
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in males; smallest juvenile
|
||
<quantity id="4CCD359DFFE4FFA514999A78634BFA29" box="[248,327,1398,1423]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="504" unit="mm" value="36.0">36 mm</quantity>
|
||
SVL; body moderately robust, accounting for 69.9–72.9% (
|
||
<emphasis id="B941446AFFE4FFA517899A7961FAFA29" box="[1000,1014,1399,1422]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 3) of SVL in males, 70.1–74.3% (
|
||
<emphasis id="B941446AFFE4FFA511EF9A796790FA29" box="[1422,1436,1399,1422]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 4) of SVL in females; tail 1.05–1.19 (1.10 ± 0.05,
|
||
<emphasis id="B941446AFFE4FFA516849A9560FFFA15" box="[741,755,1435,1458]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 5) times longer than body and accounting for 51.4–54.4% (52.4 ± 1.2,
|
||
<emphasis id="B941446AFFE4FFA5157E9AB16321FA71" box="[287,301,1471,1494]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 5) of total length; head triangular in dorsal view; head 27.1–30.1% of SVL, 62.4–71.1% as wide as long, and 37.6–40.3% as deep as wide in males (
|
||
<emphasis id="B941446AFFE4FFA516DC9AED60C7FA5D" box="[701,715,1507,1530]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 3) and 25.7–29.9% of SVL, 58.7–72.3% as wide as long, and 32.7–37.1% as deep as wide in females (
|
||
<emphasis id="B941446AFFE4FFA5163E99096061F9B9" box="[607,621,1543,1566]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 4).
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE4FFA514A69924674DF854" blockId="9.[151,1436,1362,2035]" pageId="9" pageNumber="504">
|
||
Rostral narrower than mental (76.7–99.5%, 89.5 ± 8.6,
|
||
<emphasis id="B941446AFFE4FFA5175B99256144F9E5" box="[826,840,1579,1602]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8, of width of mental), posteriorly forming border of nostril; dorsal margin of rostral deeply notched medially by shallow groove; rostral bordered posteriorly by three (50%,
|
||
<emphasis id="B941446AFFE4FFA51482997D62FDF92D" box="[227,241,1651,1674]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8), four (38%,
|
||
<emphasis id="B941446AFFE4FFA515DD997D63C6F92D" box="[444,458,1651,1674]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8), or five (12%,
|
||
<emphasis id="B941446AFFE4FFA516D0997D60B3F92D" box="[689,703,1651,1674]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8) postrostrals; postrostral at midline projecting into posterior margin of rostral; internarial region narrow, 17.2–21.1% (19.5 ± 1.4,
|
||
<emphasis id="B941446AFFE4FFA517DA999961C5F909" box="[955,969,1687,1710]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8) as wide as head; distance between anterior margins of orbit 29.2–39.6% (34.8 ± 3.9,
|
||
<emphasis id="B941446AFFE4FFA516BF99B560E0F975" box="[734,748,1723,1746]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8) as wide as head; shallow, key-hole shaped depression occupying most of prefrontal, frontal, and parietal areas (depression bordered laterally by slightly raised canthal, circumorbital, and supratemporal ridges and thickened skin at occiput); scales on dorsal snout granular, subpyrimidal, smooth, subequal in size, and slightly larger than granules of frontal and parietal regions; tubercles extending to parietal region and on supraorbital skin but absent from snout and depressed areas between eyes; tubercles on dorsal surface of head subpyrimidal and smooth, much smaller than those on body, each surrounded by seven or eight granular scales; dorsal body tubercles unicarinate, oval, slightly projecting from dorsum, about five times as large as dorsal granules, 24–27 (25 ± 1,
|
||
<emphasis id="B941446AFFE4FFA5177098B96113F869" box="[785,799,1975,1998]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8) longitudinal rows at midbody including scales in ventrolateral folds, 29–35 (32 ± 2,
|
||
<emphasis id="B941446AFFE4FFA5167C98D56027F855" box="[541,555,2011,2034]" italics="true" pageId="9" pageNumber="504">n</emphasis>
|
||
= 8) irregular transverse rows between limbs; dorsal granules smooth.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE7FFA614A69F996613FD0F" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="505">
|
||
Nostril oval, directed postero-laterally and somewhat dorsally; bordered by rostral, postrostral, supranasal, nasal, and narrow antero-dorsal process of first supralabial; supranasal rounded, about twice as large as adjacent postnasals; postnasals 4–6 (5 ± 0,
|
||
<emphasis id="B941446AFFE7FFA616749FEF602FFF5F" box="[533,547,225,248]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 16); snout length 37.3–44.7% (40.8 ± 2.1,
|
||
<emphasis id="B941446AFFE7FFA6107F9FEF6620FF5F" box="[1054,1068,225,248]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 8) of head length and distance from orbit to nostril accounting for 68.1–77.6% (72.7 ± 2.8,
|
||
<emphasis id="B941446AFFE7FFA6173F9E0B6160FEBB" box="[862,876,261,284]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 8) of snout length; loreal region sloping; lips flared; 16–23 (18 ± 2,
|
||
<emphasis id="B941446AFFE7FFA615FF9E2763A0FEE7" box="[414,428,297,320]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 8) loreals between nasal and anterior border of orbit; lorilabials in front of orbit flat, plate-like, enlarged relative to other loreals (1.5–2 times as large as loreals above them, but much smaller than supralabials); eye separated from supralabials by one or two longitudinal rows of tiny granules; diameter of orbit 20.8–26.6% (23.4 ± 2.0,
|
||
<emphasis id="B941446AFFE7FFA615CD9E9B63B6FE0B" box="[428,442,405,428]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 8) of head length; temporal region with granular scales separating large, subpyrimidal, and keeled tubercles; distance from posterior border of orbit to anterior border of auditory meatus 27.0–33.3% (29.7 ± 2.3,
|
||
<emphasis id="B941446AFFE7FFA615419ED36322FE53" box="[288,302,477,500]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 8) of head length; auditory meatus deep, narrowly elliptical, directed postero-dorsally at about 30° to vertical, positioned at posterior end of head and with its ventral margin at same level as rictus; smallest diameter of auditory meatus 60–70% smaller than greatest diameter; enlarged supralabials 8–11 (10 ± 1,
|
||
<emphasis id="B941446AFFE7FFA611689D2B671BFD9B" box="[1289,1303,549,572]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 15), last separated by 2–4 small granular scales from rictus (10–15 total supralabials; 7–9 from center of orbit to rostral); first supralabial with antero-dorsal extension reaching nostril and separating lorilabials from rostral; remaining supralabials subrectangular with rounded dorsal edges, reducing in size to rictus.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE7FFA614A69DBA673DFC03" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="505">
|
||
Mental triangular; infralabials 8–10 (9 ± 1,
|
||
<emphasis id="B941446AFFE7FFA616DD9DBB60C6FD6B" box="[700,714,693,716]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 15; 6–7 from center of orbit); pair of postmentals in contact medially; second pair of postmentals squarish, about half as wide as first pair, separated from one another medially by 6–8 (7 ± 1,
|
||
<emphasis id="B941446AFFE7FFA615299DF3635AFCB3" box="[328,342,765,788]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 8) small granular scales bordering first pair of postmentals; sublabial row decreasing in size posteriorly; gulars small, smooth, granular, nonimbricate, much smaller than ventrals; gulars grading into progressively larger, imbricate scales just in front of pectoral girdle; scale of chest like ventrals; ventrals smooth, imbricate, flat, circular, slightly smaller than tubercles on dorsum, but much smaller than dorsal granules; ventrals in transverse row at midbody 33–44 (38 ± 5,
|
||
<emphasis id="B941446AFFE7FFA616F19C836092FC03" box="[656,670,909,932]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 6); ventrolateral skin fold distinct, containing tubercles.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE7FFA614A69CA166AAFB4F" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="505">
|
||
Tail subcylindrical, though somewhat flattened dorsally and ventrally; tubercles restricted to 17.2–23.3% of intact tails (
|
||
<emphasis id="B941446AFFE7FFA615419CDB6322FC4B" box="[288,302,981,1004]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 2); distribution of tubercles at base of tail like dorsal body, distally tubercles not projecting and arrayed in 1–5 verticils; verticiles separated by 5–7 small scales along dorsal midline; tubercles on tail not extending to ventral side, although some scales of verticils may be slightly enlarged relative to other subcaudals; subcaudals flat, homogenous, two or three times as large as scales on sides of tail; each subcaudal about as long as wide; transversely enlarged subcaudal plates absent; in specimens with regenerated tails, subcaudals just proximal to regenerated portion enlarged: about half again as long as scale row proximal to them; cloacal tubercles rounded, smooth, two (88%,
|
||
<emphasis id="B941446AFFE7FFA615129BA3638DFB63" box="[371,385,1197,1220]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 16) or three (12%,
|
||
<emphasis id="B941446AFFE7FFA616159BA3608EFB63" box="[628,642,1197,1220]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 16, both counts of three in one specimen: UTA R-61577), in contact with one another, half again as large as dorsal tubercles; tip of tail capped by granular scales.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE7FFA614A69BFA667DFA42" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="505">
|
||
Brachium covered in small, homogenous, weakly keeled scales; brachial tubercles absent; antebrachium tuberculate; antebrachial tubercles subpyrimidal, unicarinate, about four times as large as granular scales between them, slightly smaller than those on dorsum; ventral surfaces of brachium and antebrachium covered in smooth, slightly imbricate, homogenous scales; scales on palm and dorsal surface of hand granular, smooth, weakly (ventrally) to moderately (dorsal surface) imbricate; scales under finger
|
||
<date id="FF8BBEB8FFE7FFA617DE9A8B6604FA3B" box="[959,1032,1412,1436]" pageId="10" pageNumber="505" value="2010-01-08">I 8–10</date>
|
||
(
|
||
<emphasis id="B941446AFFE7FFA610769A8B6629FA3B" box="[1047,1061,1413,1436]" italics="true" pageId="10" pageNumber="505">n</emphasis>
|
||
= 8 for all digits), under finger
|
||
<date id="FF8BBEB8FFE7FFA611E89A8B62D3FA67" pageId="10" pageNumber="505" value="2012-02-15">II 12–15</date>
|
||
, under finger
|
||
<date id="FF8BBEB8FFE7FFA615E19AA763E1FA67" box="[384,493,1447,1472]" pageId="10" pageNumber="505" value="1915-03-17">III 15–17</date>
|
||
, under finger
|
||
<date id="FF8BBEB8FFE7FFA616EC9AA760F7FA67" box="[653,763,1447,1472]" pageId="10" pageNumber="505" value="1916-04-18">IV 16–18</date>
|
||
, under finger
|
||
<date id="FF8BBEB8FFE7FFA617FC9AA761F3FA67" box="[925,1023,1447,1472]" pageId="10" pageNumber="505" value="2012-05-15">V 12–15</date>
|
||
(all counts excluding ungual scale); one or two scales widened and similar to lamellae extending onto palm at base of digits.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE7FFA614A69AE1676DF8A2" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="505">
|
||
Thigh, shank, and postaxial half of dorsal foot (at base of toes IV–V) tuberculate; tubercles of leg subpyrimidal, unicarinate, about same size as those on dorsum, about five times as large as small granular scales between them; preaxial and ventral surface of thigh covered in rounded, flat, imbricate scales; ventral thigh with enlarged row of pore-bearing scales and similarly enlarged row of scales anterior to pore-bearing row; this doubled row of scales arranged in an inverted V with its apex at midline of belly; abrupt transition to tiny granules on postaxial surface of thigh; postfemoral granules only one-fifth to one-sixth size of pore-bearing scales; sole covered in flat, imbricate, rounded scales; scales under toe
|
||
<date id="FF8BBEB8FFE7FFA616A999C7611CF947" box="[712,784,1736,1760]" pageId="10" pageNumber="505" value="2011-01-08">I 8–11</date>
|
||
, under toe
|
||
<date id="FF8BBEB8FFE7FFA617ED99C761E1F947" box="[908,1005,1735,1760]" pageId="10" pageNumber="505" value="2012-02-15">II 12–15</date>
|
||
, under toe
|
||
<date id="FF8BBEB8FFE7FFA6100999C766DFF947" box="[1128,1235,1736,1760]" pageId="10" pageNumber="505" value="1917-03-19">III 17–19</date>
|
||
; under toe
|
||
<date id="FF8BBEB8FFE7FFA6112F99C762B9F8A3" pageId="10" pageNumber="505" value="1919-04-22">IV 19– 22</date>
|
||
, under toe
|
||
<date id="FF8BBEB8FFE7FFA6154E99E3639EF8A3" box="[303,402,1772,1796]" pageId="10" pageNumber="505" value="1917-05-20">V 17–20</date>
|
||
; one or two scales widened and similar to lamellae extending onto sole at base of digits.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE7FFA614A6981E66AEF87A" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="505">
|
||
Arrangement of pore-bearing scales sexually dimorphic; both sexes with single, greatly enlarged scale at apex of pore-bearing series; apical scale twice as long as wide, with single pore or pore primordium at its posterior end; apical scale superficial in females and juveniles, sunken into precloacal depression in males; total pores 36 and
|
||
<quantity id="4CCD359DFFE7FFA611E1985662A1F832" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="10" pageNumber="505" unit="in" value="38.0">38 in</quantity>
|
||
two adult males (SVL
|
||
<quantity id="4CCD359DFFE7FFA615DA98726000F833" box="[443,524,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="505" unit="mm" value="75.0">75 mm</quantity>
|
||
and
|
||
<quantity id="4CCD359DFFE7FFA6162798726095F833" box="[582,665,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="505" unit="mm" value="68.0">68 mm</quantity>
|
||
, respectively), only
|
||
<quantity id="4CCD359DFFE7FFA617E3987261B1F832" box="[898,957,1916,1941]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="10" pageNumber="505" unit="in" value="14.0">14 in</quantity>
|
||
small male (SVL
|
||
<quantity id="4CCD359DFFE7FFA610F3987266E8F833" box="[1170,1252,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="505" unit="mm" value="55.0">55 mm</quantity>
|
||
) perhaps due to ontogenetic differences; two females (SVL
|
||
<quantity id="4CCD359DFFE7FFA616E798AE60DBF81F" box="[646,727,1952,1977]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="505" unit="mm" value="51.0">51 mm</quantity>
|
||
and
|
||
<quantity id="4CCD359DFFE7FFA6176E98AE616EF81F" box="[783,866,1952,1976]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="10" pageNumber="505" unit="mm" value="89.0">89 mm</quantity>
|
||
, respectively) with 17 and 19 pore primordia, one lacking primordia (SVL
|
||
<quantity id="4CCD359DFFE7FFA615CA98CA63F2F87B" box="[427,510,1988,2012]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="10" pageNumber="505" unit="mm" value="83.0">83 mm</quantity>
|
||
); in both sexes pores and/or primordia extending onto thigh.
|
||
</paragraph>
|
||
<caption id="DF4AC8F0FFE6FFA714F6983161B6F8D4" httpUri="https://zenodo.org/record/240883/files/figure.png" pageId="11" pageNumber="506" targetBox="[176,1419,208,1820]" targetPageId="11">
|
||
<paragraph id="8B8A9878FFE6FFA714F6983161B6F8D4" blockId="11.[151,1436,1855,1907]" pageId="11" pageNumber="506">
|
||
<emphasis id="B941446AFFE6FFA714F698316319F8F3" bold="true" box="[151,277,1855,1877]" pageId="11" pageNumber="506">FIGURE 6.</emphasis>
|
||
Dorsal and ventral color variation within the type series of
|
||
<taxonomicName id="4C35E3FBFFE6FFA717EA9831669BF8F2" box="[907,1175,1855,1877]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="506" phylum="Chordata" rank="species" species="semicinctus">
|
||
<emphasis id="B941446AFFE6FFA717EA9831669BF8F2" box="[907,1175,1855,1877]" italics="true" pageId="11" pageNumber="506">Cyrtodactylus semicinctus</emphasis>
|
||
</taxonomicName>
|
||
(from left to right, same specimens in bottom row: MZB 9694, 9696, 9703 (holotype), 9695, and 9701).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8A9878FFE6FFA014A698AE6647FE52" blockId="11.[151,1437,1952,2013]" lastBlockId="12.[151,1437,151,1581]" lastPageId="12" lastPageNumber="507" pageId="11" pageNumber="506">
|
||
<emphasis id="B941446AFFE6FFA714A698AE6343F81E" bold="true" box="[199,335,1952,1977]" pageId="11" pageNumber="506">Coloration.</emphasis>
|
||
In life,
|
||
<taxonomicName id="4C35E3FBFFE6FFA715CB98AE60DEF81F" box="[426,722,1952,1977]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="506" phylum="Chordata" rank="species" species="semicinctus">
|
||
<emphasis id="B941446AFFE6FFA715CB98AE60DEF81F" box="[426,722,1952,1977]" italics="true" pageId="11" pageNumber="506">Cyrtodactylus semicinctus</emphasis>
|
||
</taxonomicName>
|
||
has a light gray, greenish tan, or dull orange ground coloration with brown to reddish brown markings widely edged in black (MZB 9695 was in the process of shedding and its ground coloration was gray rather than orange or greenish tan). Narrow markings are entirely black. The dorsal tubercles are a paler shade than the dorsal ground color. On the flanks, even some of the tubercles within dark markings are this pale color, so that the flanks appear to be densely speckled. A yellow wash covers the labials, rostral, and supraciliary scales. These yellowish scales are mostly immaculate, although small, irregular black blotches mark the narial and internarial region and sometimes extend ventrally onto the rostral and or first supralabial. Dorsally, the head usually has one to three poorly defined blotches in the parietal region and a sharp black line edging the gray to yellow supraciliaries medially. This species has a distinctive U-shaped marking extending from the loreal regions around the occiput. The marking begins a short distance behind the nasal just above the enlarged series of lorilabials and widens just in front of the eye. Behind the eye and around the occiput, a slightly more narrow pale band edges the U-shaped band dorsally and accentuates it.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE1FFA014A69EF1609BFD9A" blockId="12.[151,1437,151,1581]" pageId="12" pageNumber="507">The iris is reddish brown to greenish bronze, edged in black and with a black reticulum. The ciliary body is only slightly lighter than the iris and indistict.</paragraph>
|
||
<paragraph id="8B8A9878FFE1FFA014A69D49634BFC02" blockId="12.[151,1437,151,1581]" pageId="12" pageNumber="507">
|
||
<taxonomicName id="4C35E3FBFFE1FFA014A69D4963E3FDC7" box="[199,495,583,608]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="507" phylum="Chordata" rank="species" species="semicinctus">
|
||
<emphasis id="B941446AFFE1FFA014A69D4963E3FDC7" box="[199,495,583,608]" italics="true" pageId="12" pageNumber="507">Cyrtodactylus semicinctus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130E84FDFFE1FFA0159E9D49604FFDC7" box="[511,579,583,608]" captionStart="FIGURE 6" captionStartId="11.[151,250,1855,1877]" captionTargetBox="[176,1419,208,1820]" captionTargetId="figure@11.[154,1433,193,1833]" captionTargetPageId="11" captionText="FIGURE 6. Dorsal and ventral color variation within the type series of Cyrtodactylus semicinctus (from left to right, same specimens in bottom row: MZB 9694, 9696, 9703 (holotype), 9695, and 9701)." httpUri="https://zenodo.org/record/240883/files/figure.png" pageId="12" pageNumber="507">Fig. 6</figureCitation>
|
||
) has one band on the neck, 5–7 (6 ± 1,
|
||
<emphasis id="B941446AFFE1FFA0106B9D476614FDC7" box="[1034,1048,585,608]" italics="true" pageId="12" pageNumber="507">n</emphasis>
|
||
= 8) bands on the body, and 9–11 (10 ± 1,
|
||
<emphasis id="B941446AFFE1FFA0149B9D636304FD23" box="[250,264,621,644]" italics="true" pageId="12" pageNumber="507">n</emphasis>
|
||
= 5) bands on the tail. Bands on the neck and body are jagged, irregular, and frequently broken on the flanks. When broken, blotches on the flanks may fuse with one another, sometimes to produce long sinuous markings. In particular, the band on the neck usually breaks up into blotches. However, the band on the neck does not fuse to the U-shaped nuchal band. Bands on the tail are slightly longer than the pale interspaces. Except near the distal tip of the tail of some specimens, the bands are not continuous ventrally (i.e., the tail is banded, not ringed; all specimens have at least the basal one-third of the tail not regenerated and none have ringed tails). Distally, the caudal interspaces become increasingly more pale. Regenerated tails are entirely black or mostly black with few scattered cream scales. The brachium, antebrachium, thigh, and shank each bears two or three jagged, irregular bands.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE1FFA014A69CBE63D7FB07" blockId="12.[151,1437,151,1581]" pageId="12" pageNumber="507">Except for the tail, the venter is mostly rose pink to grayish purple. Palms and soles are a darker shade of the same ground color, whereas the pore-bearing series are distinctly paler than scales around them. Some scales at the edges of the venter are the same color as the dorsal ground coloration. They form a diffuse reticulum against the rose pink to grayish purple ground color. This diffuse, greenish tan to orange reticulum is restricted to the lateral edges of the belly and pre- and postaxial margins of the ventral limbs, but extends across the entire gular region. The cloacal tubercles are greenish tan or orange. Ventrally, intact tails are tessellated with charcoal and elements of the dorsal ground coloration.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32FCBF3FFE1FFA014A69BA2616FFAF2" pageId="12" pageNumber="507" type="etymology">
|
||
<paragraph id="8B8A9878FFE1FFA014A69BA26370FAAA" blockId="12.[151,1437,151,1581]" pageId="12" pageNumber="507">
|
||
<emphasis id="B941446AFFE1FFA014A69BA26342FB62" bold="true" box="[199,334,1196,1221]" pageId="12" pageNumber="507">Etymology.</emphasis>
|
||
This species’ name
|
||
<taxonomicName id="4C35E3FBFFE1FFA0165F9BA360CDFB63" box="[574,705,1197,1220]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="507" phylum="Chordata" rank="species" species="semicinctus">
|
||
<emphasis id="B941446AFFE1FFA0165F9BA360CDFB63" box="[574,705,1197,1220]" italics="true" pageId="12" pageNumber="507">semicinctus</emphasis>
|
||
</taxonomicName>
|
||
is a masculine adjective derived from the Latin noun
|
||
<emphasis id="B941446AFFE1FFA0115D9BA36790FB63" box="[1340,1436,1197,1220]" italics="true" pageId="12" pageNumber="507">cinctum,</emphasis>
|
||
meaning girdle or belt, and prefix
|
||
<emphasis id="B941446AFFE1FFA016799BDF6047FB4F" box="[536,587,1233,1256]" italics="true" pageId="12" pageNumber="507">semi</emphasis>
|
||
, meaning partial or incomplete. The name alludes to the incomplete caudal rings of this species.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE1FFA014A69A196115FA97" blockId="12.[151,1437,151,1581]" box="[199,793,1303,1328]" pageId="12" pageNumber="507">
|
||
<emphasis id="B941446AFFE1FFA014A69A1963EAFA97" bold="true" box="[199,486,1303,1328]" pageId="12" pageNumber="507">Standard English name.</emphasis>
|
||
Kerinci Bent-toed Geckos.
|
||
</paragraph>
|
||
<paragraph id="8B8A9878FFE1FFA014A69A32616FFAF2" blockId="12.[151,1437,151,1581]" box="[199,867,1340,1365]" pageId="12" pageNumber="507">
|
||
<emphasis id="B941446AFFE1FFA014A69A32601CFAF2" bold="true" box="[199,528,1340,1365]" pageId="12" pageNumber="507">Standard Indonesian name.</emphasis>
|
||
Cicak Jari Lengkung Kerinci.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32FCBF3FFE1FFA014A69A5160ECF98B" pageId="12" pageNumber="507" type="distribution">
|
||
<paragraph id="8B8A9878FFE1FFA014A69A5160ECF98B" blockId="12.[151,1437,151,1581]" pageId="12" pageNumber="507">
|
||
<emphasis id="B941446AFFE1FFA014A69A51605DFADF" bold="true" box="[199,593,1375,1400]" pageId="12" pageNumber="507">Distribution and natural history.</emphasis>
|
||
This species is known only from the
|
||
<typeStatus id="548E26DAFFE1FFA0179F9A6F6622FADE" box="[1022,1070,1377,1401]" pageId="12" pageNumber="507">type</typeStatus>
|
||
locality. We collected the entire
|
||
<typeStatus id="548E26DAFFE1FFA014F69A8B62CBFA3A" box="[151,199,1413,1437]" pageId="12" pageNumber="507">type</typeStatus>
|
||
series between
|
||
<quantity id="4CCD359DFFE1FFA015E49A8A6028FA3B" box="[389,548,1412,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.879" metricValueMax="2.006" metricValueMin="1.752" pageId="12" pageNumber="507" unit="m" value="1879.0" valueMax="2006.0" valueMin="1752.0">1752–2006 m</quantity>
|
||
along a trail above the tiny hamlet of Kayu Aro. The trail extends through apparently pristine forest to the top of Mt. Kerinci and is frequented by hikers. Field parties collected
|
||
<taxonomicName id="4C35E3FBFFE1FFA0109E9AA96316FA43" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="507" phylum="Chordata" rank="species" species="semicinctus">
|
||
<emphasis id="B941446AFFE1FFA0109E9AA96316FA43" italics="true" pageId="12" pageNumber="507">Cyrtodactylus semicinctus</emphasis>
|
||
</taxonomicName>
|
||
on the forest floor at night (1950–2340 hrs) and we found most specimens around tree trunks. In field notes, Harvey noted that UTA R-61576 emitted a faint “baaap” when first grasped and a louder “raaank” when struggling as the senior author collected color notes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |