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<document id="EFF472B79C0601E67914584175C2DC94" ID-CLB-Dataset="57600" ID-DOI="10.11646/zootaxa.5278.2.2" ID-GBIF-Dataset="e1a99a41-0d7a-4bd1-b5e2-f009b582403b" ID-ISSN="1175-5326" ID-Zenodo-Dep="7906073" ID-ZooBank="0D60B8EA-2637-422D-96A7-F48ACE362CCC" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683529066879" checkinUser="plazi" docAuthor="Mukherjee, Bindarika &amp; Hazra, Niladri" docDate="2023" docId="038EF23FFFE6FFF1FF22F738FBB1FB88" docLanguage="en" docName="zootaxa.5278.2.2.pdf" docOrigin="Zootaxa 5278 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Saetheria circinata Mukherjee &amp; Hazra 2023, sp. n." docType="treatment" docUuid="60F40CC2-CCA8-44E2-B256-7CB9E40BC76F" docUuidSource="ZooBank" docVersion="7" lastPageNumber="253" masterDocId="FFB78A47FFEAFFFFFFB5F373FFDAFFD0" masterDocTitle="Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae)" masterLastPageNumber="263" masterPageNumber="239" pageNumber="251" updateTime="1699515339190" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2496B3471857D06E2C2A8434001D1E86" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1B4548F8B9B5EC5EA8ECA0E94F0615BF">
<mods:title id="B685C53FFA5E62559898CE1ED8989074">Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae)</mods:title>
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<mods:name id="BAFF745E9ADDB9454F0846A65B9819AB" type="personal">
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<mods:namePart id="AF675115DDC7627BEB0D6DCD36C67361">Mukherjee, Bindarika</mods:namePart>
<mods:nameIdentifier id="6EB86F100E4BD9223ED0DB629CFD7FB5" type="ORCID">0000-0002-9792-7931</mods:nameIdentifier>
<mods:affiliation id="8BC7DFFDB6C1556F838333FE1C9E6B9A">Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, India &amp; bindabdn @ gmail. com; https: // orcid. org / 0000 - 0002 - 9792 - 7931</mods:affiliation>
<mods:nameIdentifier id="1A73EACF6670EEA750A2C9269994BB56" type="email">bindabdn@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="C7B6A98D3E648F19A4863CD0516E04B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2893F0FB65EC4B6DC6E6A256F975C792">Hazra, Niladri</mods:namePart>
<mods:affiliation id="ACAADFF0FCC6B6ADACE29B1E15EEB2E4">Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, India</mods:affiliation>
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<mods:title id="C0FF86B763FC0693CB51D9F17FA7B612">Zootaxa</mods:title>
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<mods:part id="5A4CFA0D019B5A7C61954E4412D79C5C">
<mods:date id="D5E8077DFA6BE852C07394223B8ED5B3">2023</mods:date>
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<mods:number id="C60FED4D1B60634C94F619CD25E6DF06">2023-05-05</mods:number>
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<mods:number id="E8474EAF37084E88747CBAA1195E3A21">5278</mods:number>
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<mods:classification id="998C607E078D9FCA2BABE637DFF14F5C">journal article</mods:classification>
<mods:identifier id="A4AEA7D7031E66C5215B7AB3CB0A36F9" type="CLB-Dataset">57600</mods:identifier>
<mods:identifier id="921344BAF5F17741AE11283A1394B139" type="DOI">10.11646/zootaxa.5278.2.2</mods:identifier>
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<mods:identifier id="0985262E885C81E8F813DCE2BA9FF3DF" type="ZooBank">0D60B8EA-2637-422D-96A7-F48ACE362CCC</mods:identifier>
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<treatment id="038EF23FFFE6FFF1FF22F738FBB1FB88" ID-DOI="http://doi.org/10.5281/zenodo.7906109" ID-GBIF-Taxon="209588533" ID-Zenodo-Dep="7906109" LSID="urn:lsid:zoobank.org:act:60F40CC2-CCA8-44E2-B256-7CB9E40BC76F" httpUri="http://treatment.plazi.org/id/038EF23FFFE6FFF1FF22F738FBB1FB88" lastPageId="14" lastPageNumber="253" pageId="12" pageNumber="251">
<subSubSection id="C33D10A2FFE6FFF3FF22F738FDF2FB5A" pageId="12" pageNumber="251" type="nomenclature">
<paragraph id="8B984329FFE6FFF3FF22F738FE1DFBB6" blockId="12.[151,974,1099,1199]" box="[151,455,1099,1126]" pageId="12" pageNumber="251">
<heading id="D0D0F445FFE6FFF3FF22F738FE1DFBB6" bold="true" box="[151,455,1099,1126]" fontSize="11" level="1" pageId="12" pageNumber="251" reason="1">
<emphasis id="B9539F3BFFE6FFF3FF22F738FE1DFBB6" bold="true" box="[151,455,1099,1126]" pageId="12" pageNumber="251">
<taxonomicName id="4C2738AAFFE6FFF3FF22F738FEA4FBB6" ID-CoL="BDNMG" authority="Mukherjee &amp; Hazra, 2023" authorityName="Mukherjee &amp; Hazra" authorityYear="2023" box="[151,382,1099,1126]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="circinata" status="sp. nov.">
<emphasis id="B9539F3BFFE6FFF3FF22F738FEA4FBB6" bold="true" box="[151,382,1099,1126]" italics="true" pageId="12" pageNumber="251">Saetheria circinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2602240FFE6FFF3FE30F73FFE1DFBB6" box="[389,455,1100,1126]" pageId="12" pageNumber="251" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B984329FFE6FFF3FF22F703FDF2FB5A" blockId="12.[151,974,1099,1199]" box="[151,552,1136,1162]" pageId="12" pageNumber="251">GenBank Accession No. OP683428</paragraph>
</subSubSection>
<subSubSection id="C33D10A2FFE6FFF3FF22F7E7FC14FB7F" box="[151,974,1172,1199]" pageId="12" pageNumber="251" type="description">
<paragraph id="8B984329FFE6FFF3FF22F7E7FC14FB7F" blockId="12.[151,974,1099,1199]" box="[151,974,1172,1199]" pageId="12" pageNumber="251">
<uri id="FFB64F2BFFE6FFF3FF22F7E7FC14FB7F" box="[151,974,1172,1199]" pageId="12" pageNumber="251">
urn:lsid:zoobank.org:act:
<uuid id="FF8179FCFFE6FFF3FE1FF7E7FC14FB7F" box="[426,974,1172,1199]" pageId="12" pageNumber="251">60F40CC2-CCA8-44E2-B256-7CB9E40BC76F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33D10A2FFE6FFF3FF22F7AFFE74FA1E" pageId="12" pageNumber="251" type="materials_examined">
<paragraph id="8B984329FFE6FFF3FF22F7AFFACDFAB2" blockId="12.[151,1437,1244,2026]" pageId="12" pageNumber="251">
<emphasis id="B9539F3BFFE6FFF3FF22F7AFFE58FB26" bold="true" box="[151,386,1244,1270]" pageId="12" pageNumber="251">Material examined.</emphasis>
<typeStatus id="549CFD8BFFE6FFF3FE3AF7AFFE2FFB26" box="[399,501,1244,1270]" pageId="12" pageNumber="251" type="holotype">Holotype</typeStatus>
male, labelled ‘
<materialsCitation id="3B4F4974FFE6FFF3FD01F7AFFCB0FACA" ID-GBIF-Occurrence="4101311307" collectingDate="2020-02-04" collectorName="Coll. N. Hazra'" country="India" latitude="23.2393" location="Purba Barddhaman" longLatPrecision="7" longitude="87.8512" pageId="12" pageNumber="251" specimenCount="1" stateProvince="West Bengal" typeStatus="holotype">
<typeStatus id="549CFD8BFFE6FFF3FD01F7AFFCC5FB26" box="[692,799,1244,1270]" pageId="12" pageNumber="251" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C2738AAFFE6FFF3FC98F7AFFBDFFB26" authority="Mukherjee &amp; Hazra, 2023" authorityName="Mukherjee &amp; Hazra" authorityYear="2023" box="[813,1029,1244,1270]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="circinata" status="sp. nov.">
<emphasis id="B9539F3BFFE6FFF3FC98F7AFFBDFFB26" box="[813,1029,1244,1270]" italics="true" pageId="12" pageNumber="251">Saetheria circinata</emphasis>
</taxonomicName>
<emphasis id="B9539F3BFFE6FFF3FBA6F7AFFB8CFB26" bold="true" box="[1043,1110,1244,1270]" pageId="12" pageNumber="251">
<taxonomicNameLabel id="A2602240FFE6FFF3FBA6F7AFFB8CFB26" box="[1043,1110,1244,1270]" pageId="12" pageNumber="251" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<collectingCountry id="F33003B9FFE6FFF3FBDEF7AFFB7CFB26" box="[1131,1190,1244,1270]" name="India" pageId="12" pageNumber="251">India</collectingCountry>
,
<collectingRegion id="49E38DCBFFE6FFF3FB02F7AFFA90FB26" box="[1207,1354,1244,1270]" country="India" name="West Bengal" pageId="12" pageNumber="251">West Bengal</collectingRegion>
,
<location id="8EF815F2FFE6FFF3FAE9F7AFFEF9FACA" LSID="urn:lsid:plazi:treatment:038EF23FFFE6FFF1FF22F738FBB1FB88:8EF815F2FFE6FFF3FAE9F7AFFEF9FACA" country="India" latitude="23.2393" longLatPrecision="7" longitude="87.8512" name="Purba Barddhaman" pageId="12" pageNumber="251" stateProvince="West Bengal">Purba Barddhaman</location>
[
<geoCoordinate id="EE1325EEFFE6FFF3FE9BF673FE6AFACA" box="[302,432,1280,1306]" degrees="23.2393" direction="north" orientation="latitude" pageId="12" pageNumber="251" precision="5" value="23.2393">23.2393° N</geoCoordinate>
,
<geoCoordinate id="EE1325EEFFE6FFF3FE0CF673FDECFACA" box="[441,566,1280,1307]" degrees="87.8512" direction="east" orientation="longitude" pageId="12" pageNumber="251" precision="5" value="87.8512">87.8512° E</geoCoordinate>
],
<date id="FF9965E9FFE6FFF3FDF2F673FD63FACA" box="[583,697,1280,1306]" pageId="12" pageNumber="251" value="2020-02-04">
<collectingDate id="EFDD9C01FFE6FFF3FDF2F673FD63FACA" box="[583,697,1280,1306]" pageId="12" pageNumber="251" value="2020-02-04">04.ii.2020</collectingDate>
</date>
,
<collectorName id="26D226FFFFE6FFF3FD77F673FCB0FACA" box="[706,874,1280,1306]" pageId="12" pageNumber="251">Coll. N. Hazra’</collectorName>
</materialsCitation>
.
<materialsCitation id="3B4F4974FFE6FFF3FCC7F673FE9AFAEE" ID-GBIF-Occurrence="4101311318" collectingDate="2020-06-25" collectorName="Coll. N. Hazra." country="India" latitude="23.2393" location="Purba Barddhaman" longLatPrecision="7" longitude="87.8512" pageId="12" pageNumber="251" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="paratype">
<typeStatus id="549CFD8BFFE6FFF3FCC7F673FC08FACA" box="[882,978,1280,1306]" pageId="12" pageNumber="251" type="paratype">Paratype</typeStatus>
<specimenCount id="9D2188A0FFE6FFF3FC62F673FBFBFACA" box="[983,1057,1280,1306]" pageId="12" pageNumber="251" type="male">1 male</specimenCount>
, same data as holotype,
<date id="FF9965E9FFE6FFF3FA95F673FA42FACB" box="[1312,1432,1280,1307]" pageId="12" pageNumber="251" value="2020-06-25">
<collectingDate id="EFDD9C01FFE6FFF3FA95F673FA42FACB" box="[1312,1432,1280,1307]" pageId="12" pageNumber="251" value="2020-06-25">25.vi.2020</collectingDate>
</date>
,
<collectorName id="26D226FFFFE6FFF3FF22F657FE9AFAEE" box="[151,320,1316,1342]" pageId="12" pageNumber="251">Coll. N. Hazra.</collectorName>
</materialsCitation>
<materialsCitation id="3B4F4974FFE6FFF3FEF2F657FB92FAEE" ID-GBIF-Occurrence="4101311309" box="[327,1096,1316,1343]" collectingDate="2021-02-15" collectorName="Coll. N. Hazra." country="India" latitude="23.2393" location="Purba Barddhaman" longLatPrecision="7" longitude="87.8512" pageId="12" pageNumber="251" specimenCount="2" specimenCount-male="2" stateProvince="West Bengal" typeStatus="paratype">
<typeStatus id="549CFD8BFFE6FFF3FEF2F657FE68FAEE" box="[327,434,1316,1342]" pageId="12" pageNumber="251" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D2188A0FFE6FFF3FE0FF657FDC8FAEE" box="[442,530,1316,1342]" pageId="12" pageNumber="251" type="male">2 males</specimenCount>
, same data as holotype,
<date id="FF9965E9FFE6FFF3FC94F657FC48FAEF" box="[801,914,1316,1343]" pageId="12" pageNumber="251" value="2021-02-15">
<collectingDate id="EFDD9C01FFE6FFF3FC94F657FC48FAEF" box="[801,914,1316,1343]" pageId="12" pageNumber="251" value="2021-02-15">15.ii.2021</collectingDate>
</date>
,
<collectorName id="26D226FFFFE6FFF3FC2BF657FB92FAEE" box="[926,1096,1316,1342]" pageId="12" pageNumber="251">Coll. N. Hazra.</collectorName>
</materialsCitation>
<materialsCitation id="3B4F4974FFE6FFF3FBFAF657FDFFFAB2" ID-GBIF-Occurrence="4101311302" collectingDate="2021-06-14" collectorName="Coll N. Hazra" country="India" latitude="23.2393" location="Purba Barddhaman" longLatPrecision="7" longitude="87.8512" pageId="12" pageNumber="251" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="paratype">
<typeStatus id="549CFD8BFFE6FFF3FBFAF657FB75FAEE" box="[1103,1199,1316,1342]" pageId="12" pageNumber="251" type="paratype">Paratype</typeStatus>
<specimenCount id="9D2188A0FFE6FFF3FB03F657FADEFAEE" box="[1206,1284,1316,1342]" pageId="12" pageNumber="251" type="male">1 male</specimenCount>
, same data as holotype,
<date id="FF9965E9FFE6FFF3FEB0F63BFEA7FAB2" box="[261,381,1352,1378]" pageId="12" pageNumber="251" value="2021-06-14">
<collectingDate id="EFDD9C01FFE6FFF3FEB0F63BFEA7FAB2" box="[261,381,1352,1378]" pageId="12" pageNumber="251" value="2021-06-14">14.vi.2021</collectingDate>
</date>
,
<collectorName id="26D226FFFFE6FFF3FE3DF63BFDFFFAB2" box="[392,549,1352,1378]" pageId="12" pageNumber="251">Coll N. Hazra</collectorName>
</materialsCitation>
.
<materialsCitation id="3B4F4974FFE6FFF3FD9AF63BFACDFAB2" ID-GBIF-Occurrence="4101311326" box="[559,1303,1352,1378]" collectingDate="2021-11-19" collectorName="Coll. N. Hazra." country="India" latitude="23.2393" location="Purba Barddhaman" longLatPrecision="7" longitude="87.8512" pageId="12" pageNumber="251" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="paratype">
<typeStatus id="549CFD8BFFE6FFF3FD9AF63BFD55FAB2" box="[559,655,1352,1378]" pageId="12" pageNumber="251" type="paratype">Paratype</typeStatus>
<specimenCount id="9D2188A0FFE6FFF3FD23F63BFD38FAB2" box="[662,738,1352,1378]" pageId="12" pageNumber="251" type="male">1 male</specimenCount>
, same data as holotype,
<date id="FF9965E9FFE6FFF3FC58F63BFBBEFAB2" box="[1005,1124,1352,1378]" pageId="12" pageNumber="251" value="2021-11-19">
<collectingDate id="EFDD9C01FFE6FFF3FC58F63BFBBEFAB2" box="[1005,1124,1352,1378]" pageId="12" pageNumber="251" value="2021-11-19">19.xi.2021</collectingDate>
</date>
,
<collectorName id="26D226FFFFE6FFF3FBC5F63BFACDFAB2" box="[1136,1303,1352,1378]" pageId="12" pageNumber="251">Coll. N. Hazra.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B984329FFE6FFF3FF72F61FFE74FA1E" blockId="12.[151,1437,1244,2026]" pageId="12" pageNumber="251">
<emphasis id="B9539F3BFFE6FFF3FF72F61FFE08FA56" bold="true" box="[199,466,1388,1414]" pageId="12" pageNumber="251">Diagnostic characters.</emphasis>
The distinguished features of new species are as follows: apically slightly broadened anal point with a median ridge, Y shaped anal tergite band, anal tergite band and nearly triangular shaped ventral lobe of superior volsella.
</paragraph>
</subSubSection>
<subSubSection id="C33D10A2FFE6FFF1FF72F6ABFE48FC51" lastPageId="14" lastPageNumber="253" pageId="12" pageNumber="251" type="etymology">
<paragraph id="8B984329FFE6FFF3FF72F6ABFE83F9C6" blockId="12.[151,1437,1244,2026]" pageId="12" pageNumber="251">
<emphasis id="B9539F3BFFE6FFF3FF72F6ABFE94FA22" bold="true" box="[199,334,1496,1522]" pageId="12" pageNumber="251">Etymology.</emphasis>
The name ‘
<taxonomicName id="4C2738AAFFE6FFF3FE6CF6AAFDE7FA22" authorityName="Mukherjee &amp; Hazra" authorityYear="2023" box="[473,573,1497,1522]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="circinata">
<emphasis id="B9539F3BFFE6FFF3FE6CF6AAFDE7FA22" box="[473,573,1497,1522]" italics="true" pageId="12" pageNumber="251">circinata</emphasis>
</taxonomicName>
’, originated from a Latin word ‘
<emphasis id="B9539F3BFFE6FFF3FC06F6AAFBF8FA22" box="[947,1058,1497,1522]" italics="true" pageId="12" pageNumber="251">circinatus</emphasis>
’, refers to circular dorsal lobe of superior volsella.
</paragraph>
<paragraph id="8B984329FFE6FFF3FF72F553FE91F9E9" blockId="12.[151,1437,1244,2026]" box="[199,331,1567,1594]" pageId="12" pageNumber="251">
<emphasis id="B9539F3BFFE6FFF3FF72F553FE91F9E9" bold="true" box="[199,331,1567,1594]" pageId="12" pageNumber="251">Male (n=6)</emphasis>
</paragraph>
<paragraph id="8B984329FFE6FFF3FF72F537FA58F952" blockId="12.[151,1437,1244,2026]" pageId="12" pageNumber="251">
Total length 2.75–2.8,
<quantity id="4CDFEECCFFE6FFF3FE70F537FDE0F98E" box="[453,570,1604,1631]" metricMagnitude="-3" metricUnit="m" metricValue="2.775" pageId="12" pageNumber="251" unit="mm" value="2.775">2.775 mm</quantity>
. Wing length 1.7–1.85,
<quantity id="4CDFEECCFFE6FFF3FCF2F537FC74F98E" box="[839,942,1604,1630]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" pageId="12" pageNumber="251" unit="mm" value="1.77">1.77 mm</quantity>
. Costal length 1.65–1.7,
<quantity id="4CDFEECCFFE6FFF3FB70F537FAF6F98E" box="[1221,1324,1604,1630]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="12" pageNumber="251" unit="mm" value="1.67">1.67 mm</quantity>
. Antennal length 0.82–0.85,
<quantity id="4CDFEECCFFE6FFF3FED7F51BFE0DF952" box="[354,471,1640,1667]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" pageId="12" pageNumber="251" unit="mm" value="0.835">0.835 mm</quantity>
. Colouration. Thorax with dark brown marking; leg and abdomen yellowish brown.
</paragraph>
<paragraph id="8B984329FFE6FFF3FF72F5FFFE22F8C3" blockId="12.[151,1437,1244,2026]" pageId="12" pageNumber="251">Head. Head width 450–460, 455 µm. Temporal setae 8–9 (IV 4–5, OV 3–4, Po 0–1). Clypeal setae 12. Frontal tubercles present. Eyes bare with dorsomedial extension of 130–140, 135 µm. AR 1.2–1.41, 1.30; ultimate flagellomere 460–480, 470 µm long. Palpomere lengths (I–V) (µm): 23–25, 24: 30: 80.5–82, 81.25: 103.5: 149.5– 152, 150. CA 0.54–0.55, 0.545.</paragraph>
<paragraph id="8B984329FFE6FFF3FF72F46FFC8FF8E6" blockId="12.[151,1437,1244,2026]" box="[199,853,1820,1846]" pageId="12" pageNumber="251">Thorax. Scutal tubercle absent. Ac 8, Dc 7, Pa 3, Scts 2–3.</paragraph>
<paragraph id="8B984329FFE6FFF3FF72F433FC91F8AE" blockId="12.[151,1437,1244,2026]" pageId="12" pageNumber="251">
Wing (
<figureCitation id="131C5FACFFE6FFF3FEA0F433FEABF88A" box="[277,369,1856,1882]" captionStart="FIGURE 4" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[266,1330,181,1910]" captionTargetId="figure-17@13.[254,1334,181,1920]" captionTargetPageId="13" captionText="FIGURE 4. Adult male Saetheria circinata sp. n. A. Wing, B. Hypopygium (dorsal), C. Hypopygium, scale 0.01 mm, D. Superior volsella, scale 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.7906087" httpUri="https://zenodo.org/record/7906087/files/figure.png" pageId="12" pageNumber="251">Fig. 4A</figureCitation>
). VR 1.14–1.16, 1.15. R
<subScript id="17A3416CFFE6FFF3FD32F421FD55F8B0" attach="left" box="[647,655,1874,1888]" fontSize="6" pageId="12" pageNumber="251">1</subScript>
and R
<subScript id="17A3416CFFE6FFF3FD6EF421FD2FF8B0" attach="left" box="[731,757,1874,1888]" fontSize="6" pageId="12" pageNumber="251">4+5</subScript>
bare. Brachiolum with 1 seta. Squama with 7 fringed setae. FCu distinctly distal to
<collectingRegion id="49E38DCBFFE6FFF3FE2BF417FE17F8AE" box="[414,461,1892,1918]" country="Kenya" name="Lamu" pageId="12" pageNumber="251">RM</collectingRegion>
. Anal lobe moderately developed.
</paragraph>
<paragraph id="8B984329FFE6FFF3FF72F4FBFEB9F83A" blockId="12.[151,1437,1244,2026]" pageId="12" pageNumber="251">Legs. Fore tibia with 2 setae. Mid legs with 2 tibial spurs; 16 µm and 18.4 µm long with 22–26 lateral teeth. Hind leg with 2 tibial spurs; 18.4 µm and 20.7 µm long with 29–32 teeth. Lengths and proportions of leg segments shown in table IV.</paragraph>
<caption id="DF5813A1FFE7FFF2FF2DF4E4FE31F803" ID-DOI="http://doi.org/10.5281/zenodo.7906087" ID-Zenodo-Dep="7906087" httpUri="https://zenodo.org/record/7906087/files/figure.png" pageId="13" pageNumber="252" startId="13.[152,255,1943,1967]" targetBox="[266,1330,181,1910]" targetPageId="13" targetType="figure">
<paragraph id="8B984329FFE7FFF2FF2DF4E4FE31F803" blockId="13.[151,1437,1943,2004]" pageId="13" pageNumber="252">
<emphasis id="B9539F3BFFE7FFF2FF2DF4E4FEC1F87F" bold="true" box="[152,283,1943,1967]" pageId="13" pageNumber="252">FIGURE 4.</emphasis>
Adult male
<taxonomicName id="4C2738AAFFE7FFF2FE14F4EBFDBCF87F" authority="Mukherjee &amp; Hazra, 2023" authorityName="Mukherjee &amp; Hazra" authorityYear="2023" box="[417,614,1944,1968]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="252" phylum="Arthropoda" rank="species" species="circinata" status="sp. nov.">
<emphasis id="B9539F3BFFE7FFF2FE14F4EBFDBCF87F" box="[417,614,1944,1968]" italics="true" pageId="13" pageNumber="252">Saetheria circinata</emphasis>
</taxonomicName>
<emphasis id="B9539F3BFFE7FFF2FDDBF4EBFD70F860" bold="true" box="[622,682,1944,1968]" pageId="13" pageNumber="252">
<taxonomicNameLabel id="A2602240FFE7FFF2FDDBF4EBFD70F860" box="[622,682,1944,1968]" pageId="13" pageNumber="252" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A. Wing, B. Hypopygium (dorsal), C. Hypopygium, scale 0.01 mm, D. Superior volsella, scale 0.01 mm.
</paragraph>
</caption>
<caption id="DF5813A1FFE4FFF1FF2DF3E4FC39FF7F" box="[152,995,151,175]" pageId="14" pageNumber="253" startId="14.[152,244,151,175]" targetBox="[152,1395,191,576]" targetIsTable="true" targetPageId="14">
<paragraph id="8B984329FFE4FFF1FF2DF3E4FC39FF7F" blockId="14.[152,995,151,175]" box="[152,995,151,175]" pageId="14" pageNumber="253">
<emphasis id="B9539F3BFFE4FFF1FF2DF3E4FEC1FF7F" bold="true" box="[152,283,151,175]" pageId="14" pageNumber="253">TABLE IV.</emphasis>
Lengths (µm) and proportions of legs in
<taxonomicName id="4C2738AAFFE4FFF1FD61F3E4FC7AFF7F" authority="Mukherjee &amp; Hazra, 2023" authorityName="Mukherjee &amp; Hazra" authorityYear="2023" box="[724,928,151,175]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="253" phylum="Arthropoda" rank="species" species="circinata" status="sp. nov.">
<emphasis id="B9539F3BFFE4FFF1FD61F3E4FC7AFF7F" box="[724,928,151,175]" italics="true" pageId="14" pageNumber="253">Saetheria circinata</emphasis>
</taxonomicName>
<emphasis id="B9539F3BFFE4FFF1FC12F3E4FC39FF7F" bold="true" box="[935,995,151,175]" pageId="14" pageNumber="253">
<taxonomicNameLabel id="A2602240FFE4FFF1FC12F3E4FC39FF7F" box="[935,995,151,175]" pageId="14" pageNumber="253" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</caption>
<paragraph id="8B984329FFE4FFF1FEC2F3B3FAA9FDEA" pageId="14" pageNumber="253">
<table id="F927B189FFE40000FF2DF3CCFAA9FD90" box="[152,1395,191,576]" colsContinueFrom="7.[152,1408,1301,1685]" gridcols="6" gridrows="9" pageId="14" pageNumber="253">
<tr id="3517416BFFE40000FF2DF3CCFAA9FF0D" box="[152,1395,191,221]" gridrow="0" pageId="14" pageNumber="253" rowspan-0="1">
<th id="76C62817FFE40000FEC2F3CCFDD9FF0D" box="[375,515,191,221]" gridcol="1" gridrow="0" pageId="14" pageNumber="253">Fe</th>
<th id="76C62817FFE40000FDFEF3CCFD0CFF0D" box="[587,726,191,221]" gridcol="2" gridrow="0" pageId="14" pageNumber="253">Ti</th>
<th id="76C62817FFE40000FCA8F3CCFC14FF0D" box="[797,974,191,221]" gridcol="3" gridrow="0" pageId="14" pageNumber="253">Ta1</th>
<th id="76C62817FFE40000FC42F3CCFB7CFF0D" box="[1015,1190,191,221]" gridcol="4" gridrow="0" pageId="14" pageNumber="253">Ta2</th>
<th id="76C62817FFE40000FB67F3CCFAA9FF0D" box="[1234,1395,191,221]" gridcol="5" gridrow="0" pageId="14" pageNumber="253">Ta3</th>
</tr>
<tr id="3517416BFFE40000FF2DF394FAA9FED5" box="[152,1395,231,261]" gridrow="1" pageId="14" pageNumber="253">
<th id="76C62817FFE40000FF2DF394FF61FED5" box="[152,187,231,261]" gridcol="0" gridrow="1" pageId="14" pageNumber="253">P1</th>
<td id="76C62817FFE40000FEC2F394FDD9FED5" box="[375,515,231,261]" gridcol="1" gridrow="1" pageId="14" pageNumber="253">600–610, 605</td>
<td id="76C62817FFE40000FDFEF394FD0CFED5" box="[587,726,231,261]" gridcol="2" gridrow="1" pageId="14" pageNumber="253">400–410, 405</td>
<td id="76C62817FFE40000FCA8F394FC14FED5" box="[797,974,231,261]" gridcol="3" gridrow="1" pageId="14" pageNumber="253">–</td>
<td id="76C62817FFE40000FC42F394FB7CFED5" box="[1015,1190,231,261]" gridcol="4" gridrow="1" pageId="14" pageNumber="253">–</td>
<td id="76C62817FFE40000FB67F394FAA9FED5" box="[1234,1395,231,261]" gridcol="5" gridrow="1" pageId="14" pageNumber="253">–</td>
</tr>
<tr id="3517416BFFE40000FF2DF263FAA9FEFE" box="[152,1395,272,302]" gridrow="2" pageId="14" pageNumber="253">
<th id="76C62817FFE40000FF2DF263FF61FEFE" box="[152,187,272,302]" gridcol="0" gridrow="2" pageId="14" pageNumber="253">P 2</th>
<td id="76C62817FFE40000FEC2F263FDD9FEFE" box="[375,515,272,302]" gridcol="1" gridrow="2" pageId="14" pageNumber="253">520–550, 535</td>
<td id="76C62817FFE40000FDFEF263FD0CFEFE" box="[587,726,272,302]" gridcol="2" gridrow="2" pageId="14" pageNumber="253">450–460, 455</td>
<td id="76C62817FFE40000FCA8F263FC14FEFE" box="[797,974,272,302]" gridcol="3" gridrow="2" pageId="14" pageNumber="253">280</td>
<td id="76C62817FFE40000FC42F263FB7CFEFE" box="[1015,1190,272,302]" gridcol="4" gridrow="2" pageId="14" pageNumber="253">150</td>
<td id="76C62817FFE40000FB67F263FAA9FEFE" box="[1234,1395,272,302]" gridcol="5" gridrow="2" pageId="14" pageNumber="253">90</td>
</tr>
<tr id="3517416BFFE40000FF2DF24BFAA9FE86" box="[152,1395,312,342]" gridrow="3" pageId="14" pageNumber="253">
<th id="76C62817FFE40000FF2DF24BFF61FE86" box="[152,187,312,342]" gridcol="0" gridrow="3" pageId="14" pageNumber="253">P3</th>
<td id="76C62817FFE40000FEC2F24BFDD9FE86" box="[375,515,312,342]" gridcol="1" gridrow="3" pageId="14" pageNumber="253">610–630, 620</td>
<td id="76C62817FFE40000FDFEF24BFD0CFE86" box="[587,726,312,342]" gridcol="2" gridrow="3" pageId="14" pageNumber="253">600–610, 605</td>
<td id="76C62817FFE40000FCA8F24BFC14FE86" box="[797,974,312,342]" gridcol="3" gridrow="3" pageId="14" pageNumber="253">400</td>
<td id="76C62817FFE40000FC42F24BFB7CFE86" box="[1015,1190,312,342]" gridcol="4" gridrow="3" pageId="14" pageNumber="253">240</td>
<td id="76C62817FFE40000FB67F24BFAA9FE86" box="[1234,1395,312,342]" gridcol="5" gridrow="3" pageId="14" pageNumber="253">200</td>
</tr>
<tr id="3517416BFFE40000FF2DF2F2FAA9FE49" box="[152,1395,385,409]" gridrow="4" pageId="14" pageNumber="253">
<th id="76C62817FFE40000FF2DF2F2FAA9FE49" box="[152,1395,385,409]" colspan="6" colspanRight="5" gridcol="0" gridrow="4" pageId="14" pageNumber="253">
<emphasis id="B9539F3BFFE4FFF1FF2DF2F2FECDFE49" bold="true" box="[152,279,385,409]" pageId="14" pageNumber="253">Continued.</emphasis>
</th>
</tr>
<tr id="3517416BFFE40000FF2DF2D9FAA9FE18" box="[152,1395,426,456]" gridrow="5" pageId="14" pageNumber="253" rowspan-0="1">
<td id="76C62817FFE40000FEC2F2D9FDD9FE18" box="[375,515,426,456]" gridcol="1" gridrow="5" pageId="14" pageNumber="253">Ta4</td>
<td id="76C62817FFE40000FDFEF2D9FD0CFE18" box="[587,726,426,456]" gridcol="2" gridrow="5" pageId="14" pageNumber="253">Ta 5</td>
<td id="76C62817FFE40000FCA8F2D9FC14FE18" box="[797,974,426,456]" gridcol="3" gridrow="5" pageId="14" pageNumber="253">LR</td>
<td id="76C62817FFE40000FC42F2D9FB7CFE18" box="[1015,1190,426,456]" gridcol="4" gridrow="5" pageId="14" pageNumber="253">BV</td>
<td id="76C62817FFE40000FB67F2D9FAA9FE18" box="[1234,1395,426,456]" gridcol="5" gridrow="5" pageId="14" pageNumber="253">SV</td>
</tr>
<tr id="3517416BFFE40000FF2DF2A1FAA9FE20" box="[152,1395,466,496]" gridrow="6" pageId="14" pageNumber="253">
<th id="76C62817FFE40000FF2DF2A1FF61FE20" box="[152,187,466,496]" gridcol="0" gridrow="6" pageId="14" pageNumber="253">P 1</th>
<td id="76C62817FFE40000FEC2F2A1FDD9FE20" box="[375,515,466,496]" gridcol="1" gridrow="6" pageId="14" pageNumber="253">–</td>
<td id="76C62817FFE40000FDFEF2A1FD0CFE20" box="[587,726,466,496]" gridcol="2" gridrow="6" pageId="14" pageNumber="253">–</td>
<td id="76C62817FFE40000FCA8F2A1FC14FE20" box="[797,974,466,496]" gridcol="3" gridrow="6" pageId="14" pageNumber="253">–</td>
<td id="76C62817FFE40000FC42F2A1FB7CFE20" box="[1015,1190,466,496]" gridcol="4" gridrow="6" pageId="14" pageNumber="253">–</td>
<td id="76C62817FFE40000FB67F2A1FAA9FE20" box="[1234,1395,466,496]" gridcol="5" gridrow="6" pageId="14" pageNumber="253">–</td>
</tr>
<tr id="3517416BFFE40000FF2DF289FAA9FDC8" box="[152,1395,506,536]" gridrow="7" pageId="14" pageNumber="253">
<th id="76C62817FFE40000FF2DF289FF61FDC8" box="[152,187,506,536]" gridcol="0" gridrow="7" pageId="14" pageNumber="253">P2</th>
<td id="76C62817FFE40000FEC2F289FDD9FDC8" box="[375,515,506,536]" gridcol="1" gridrow="7" pageId="14" pageNumber="253">50</td>
<td id="76C62817FFE40000FDFEF289FD0CFDC8" box="[587,726,506,536]" gridcol="2" gridrow="7" pageId="14" pageNumber="253">50</td>
<td id="76C62817FFE40000FCA8F289FC14FDC8" box="[797,974,506,536]" gridcol="3" gridrow="7" pageId="14" pageNumber="253">0.57–0.62, 0.59</td>
<td id="76C62817FFE40000FC42F289FB7CFDC8" box="[1015,1190,506,536]" gridcol="4" gridrow="7" pageId="14" pageNumber="253">3.6–3.7, 3.65</td>
<td id="76C62817FFE40000FB67F289FAA9FDC8" box="[1234,1395,506,536]" gridcol="5" gridrow="7" pageId="14" pageNumber="253">3.57–3.7, 3.63</td>
</tr>
<tr id="3517416BFFE40000FF2DF151FAA9FD90" box="[152,1395,546,576]" gridrow="8" pageId="14" pageNumber="253">
<th id="76C62817FFE40000FF2DF151FF61FD90" box="[152,187,546,576]" gridcol="0" gridrow="8" pageId="14" pageNumber="253">P3</th>
<td id="76C62817FFE40000FEC2F151FDD9FD90" box="[375,515,546,576]" gridcol="1" gridrow="8" pageId="14" pageNumber="253">120</td>
<td id="76C62817FFE40000FDFEF151FD0CFD90" box="[587,726,546,576]" gridcol="2" gridrow="8" pageId="14" pageNumber="253">70</td>
<td id="76C62817FFE40000FCA8F151FC14FD90" box="[797,974,546,576]" gridcol="3" gridrow="8" pageId="14" pageNumber="253">0.63–0.65, 0.64</td>
<td id="76C62817FFE40000FC42F151FB7CFD90" box="[1015,1190,546,576]" gridcol="4" gridrow="8" pageId="14" pageNumber="253">2.55–2.60, 2.57</td>
<td id="76C62817FFE40000FB67F151FAA9FD90" box="[1234,1395,546,576]" gridcol="5" gridrow="8" pageId="14" pageNumber="253">3.02–3.1, 3.06</td>
</tr>
</table>
</paragraph>
<paragraph id="8B984329FFE4FFF1FF72F119FE48FC51" blockId="14.[151,1437,618,1112]" pageId="14" pageNumber="253">
Hypopygium (
<figureCitation id="131C5FACFFE4FFF1FEDFF119FE2BFD54" box="[362,497,618,644]" captionStart="FIGURE 4" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[266,1330,181,1910]" captionTargetId="figure-17@13.[254,1334,181,1920]" captionTargetPageId="13" captionText="FIGURE 4. Adult male Saetheria circinata sp. n. A. Wing, B. Hypopygium (dorsal), C. Hypopygium, scale 0.01 mm, D. Superior volsella, scale 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.7906087" httpUri="https://zenodo.org/record/7906087/files/figure.png" pageId="14" pageNumber="253">Figs. 4B–D</figureCitation>
). Anal tergite band Y shaped. Anal point slightly expanded apically, 37–40, 38.5 µm long with a median ridge like appearance. Laterosternite IX with 3 setae on each side of anal point. Longitudinal and transverse sternapodeme 75–78, 76.5 µm and 73.5 µm long respectively. Superior volsella 55–58, 56.5 µm long and 25.3 µm wide, nearly triangular in shape, consisting of dorsal and ventral lobe, dorsal lobe having roughly circular base (23 µm long) with digitiform projection (27–28 µm long) while ventral lobe nearly triangular shaped bearing 2 setae (1 seta on heel and another one placed subapically). Inferior volsella with a rounded apical part covered with microtrichia. Gonocoxite 122–126, 124 µm long. Gonostylus 126.5–131, 128.75 µm long. HR 0.96–0.97, 0.965. HV 2.18– 2.29, 2.235.
</paragraph>
</subSubSection>
<subSubSection id="C33D10A2FFE4FFF1FF72F0F9FD9EFC74" box="[199,580,906,932]" pageId="14" pageNumber="253" type="distribution">
<paragraph id="8B984329FFE4FFF1FF72F0F9FD9EFC74" blockId="14.[151,1437,618,1112]" box="[199,580,906,932]" pageId="14" pageNumber="253">
<emphasis id="B9539F3BFFE4FFF1FF72F0F9FEBBFC74" bold="true" box="[199,353,906,932]" pageId="14" pageNumber="253">Distribution.</emphasis>
<collectingCountry id="F33003B9FFE4FFF1FED2F0F9FE79FC74" box="[359,419,906,932]" name="India" pageId="14" pageNumber="253">India</collectingCountry>
,
<collectingRegion id="49E38DCBFFE4FFF1FE1BF0F9FD9AFC74" box="[430,576,906,932]" country="India" name="West Bengal" pageId="14" pageNumber="253">West Bengal</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D10A2FFE4FFF1FF72F0DDFBB1FB88" pageId="14" pageNumber="253" type="discussion">
<paragraph id="8B984329FFE4FFF1FF72F0DDFBB1FB88" blockId="14.[151,1437,618,1112]" pageId="14" pageNumber="253">
<emphasis id="B9539F3BFFE4FFF1FF72F0DDFEE1FC18" bold="true" box="[199,315,942,968]" pageId="14" pageNumber="253">Remarks.</emphasis>
Presence of roughly bilobed superior volsella having digitiform projection on dorsal lobe, Y shaped anal tergite band and spatulated anal point confirm the positioning of new species into genus
<taxonomicName id="4C2738AAFFE4FFF1FB2FF0A1FAB6FC3C" authority="Jackson." authorityName="Jackson." box="[1178,1388,978,1004]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B9539F3BFFE4FFF1FB2FF0A1FADFFC3C" box="[1178,1285,978,1004]" italics="true" pageId="14" pageNumber="253">Saetheria</emphasis>
Jackson.
</taxonomicName>
The new species closely related with
<taxonomicName id="4C2738AAFFE4FFF1FDBBF085FCE5FBC0" authority="Yan et al., 2011" authorityName="Yan" authorityYear="2011" box="[526,831,1014,1040]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="253" phylum="Arthropoda" rank="species" species="digitata">
<emphasis id="B9539F3BFFE4FFF1FDBBF085FD59FBC0" box="[526,643,1014,1040]" italics="true" pageId="14" pageNumber="253">S. digitata</emphasis>
<bibRefCitation id="EFB63ED8FFE4FFF1FD3EF085FCE5FBC0" author="Yan, C. &amp; Saether, O. A. &amp; Wang, X." box="[651,831,1014,1040]" pageId="14" pageNumber="253" pagination="34 - 42" refId="ref27197" refString="Yan, C., Saether, O. A. &amp; Wang, X. (2011) Saetheria Jackson from the Sino-Indian Region (Diptera: Chironomidae). Zootaxa, 3040 (1), 34 - 42. https: // doi. org / 10.11646 / zootaxa. 3040.1.5" type="journal article" year="2011">
Yan
<emphasis id="B9539F3BFFE4FFF1FD0AF084FD28FBC0" box="[703,754,1014,1040]" italics="true" pageId="14" pageNumber="253">et al</emphasis>
.,
<quantity id="4CDFEECCFFE4FFF1FCBDF085FCE5FBC0" box="[776,831,1014,1040]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="14" pageNumber="253" unit="in" value="2011.0">2011</quantity>
</bibRefCitation>
</taxonomicName>
in having more or less similar gonostylus and dorsal lobe of superior volsella but presence of triangular like ventral lobe of superior volsella with 1 seta at apical end and somewhat apically spatulated anal point can segregate the new species from
<taxonomicName id="4C2738AAFFE4FFF1FC44F74DFBBEFB88" box="[1009,1124,1086,1112]" class="Insecta" family="Chironomidae" genus="Saetheria" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="253" phylum="Arthropoda" rank="species" species="digitata">
<emphasis id="B9539F3BFFE4FFF1FC44F74DFBBEFB88" box="[1009,1124,1086,1112]" italics="true" pageId="14" pageNumber="253">S. digitata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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