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<document id="12099F4E0446F6405034CD25F425B2FA" ID-DOI="10.1080/00222933.2011.642415" ID-ISSN="1464-5262" ID-Zenodo-Dep="5202591" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628967769572" checkinUser="felipe" docAuthor="Andrássy, István" docDate="2012" docId="038A2E0A691DCE5BFDC6FE3EFB90FAB5" docLanguage="en" docName="JNatHist.46.7-8.453-494.pdf" docOrigin="Journal of Natural History 46 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2011.642415" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Egtitus biformis Andrássy 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="482" masterDocId="FFB356726901CE44FFE8FFEEFFC0FFC0" masterDocTitle="On the “ Paractinolaimus ” genus group (Nematoda: Actinolaimidae), with description of five new and two rare species of Egtitus Thorne, 1967" masterLastPageNumber="494" masterPageNumber="453" pageNumber="479" updateTime="1699057353532" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0B6427A3C501C19384485B996419F510">On the “ Paractinolaimus ” genus group (Nematoda: Actinolaimidae), with description of five new and two rare species of Egtitus Thorne, 1967</mods:title>
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<mods:namePart id="C2758B8B6E5F14BE0010CD98306DC91E">Andrássy, István</mods:namePart>
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<mods:title id="E8C09E462B66B2ADAAA64F5EACA0CB37">Journal of Natural History</mods:title>
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<mods:date id="B91EE53FCB4FCA2876CBC48888BCF01D">2012</mods:date>
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<mods:number id="82F60F8A0C10A6CAA04D279C29FDF1BD">2012-02-29</mods:number>
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<mods:number id="35EC32B2B67242AB2CBF7E880D2F5DF7">46</mods:number>
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<mods:number id="33BAED26F0EDF8C7F2038B255444AA0D">7 - 8</mods:number>
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<heading id="D0D42870691DCE58FDC6FE3EFCF9FE28" box="[558,825,464,489]" centered="true" fontSize="10" level="2" pageId="28" pageNumber="479" reason="3">
<taxonomicName id="4C23E49F691DCE58FDC6FE3EFD1FFE29" authority="Andrássy, 2012" authorityName="Andrássy" authorityYear="2012" box="[558,735,464,489]" class="Adenophorea" family="Actinolaimidae" genus="Egtitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="28" pageNumber="479" phylum="Nematoda" rank="species" species="biformis" status="sp. nov.">
<emphasis id="B957430E691DCE58FDC6FE3EFD1FFE29" bold="true" box="[558,735,464,489]" italics="true" pageId="28" pageNumber="479">Egtitus biformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264FE75691DCE58FD0DFE3FFCF9FE28" box="[741,825,465,488]" pageId="28" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B9C9F1C691DCE58FDFAFE1EFC94FDC8" blockId="28.[530,852,464,520]" box="[530,852,496,520]" pageId="28" pageNumber="479">
(
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and
<figureCitation id="13188399691DCE58FD14FE1EFC8DFDC8" box="[764,845,496,520]" captionStart="Figure 11" captionStartId="30.[211,278,1570,1592]" captionTargetBox="[211,1170,161,1535]" captionTargetId="figure-5@30.[211,1171,161,1537]" captionTargetPageId="30" captionText="Figure 11. Egtitus biformis sp. nov. (A) Tip of the anterior testis; (B) tip of the posterior testis; (C) posterior end of male. Scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5202613" httpUri="https://zenodo.org/record/5202613/files/figure.png" pageId="28" pageNumber="479">11AC</figureCitation>
)
</paragraph>
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<materialsCitation id="3B4B9541691DCE58FF3BFDDAFB49FD8C" box="[211,1161,561,588]" collectionCode="V" pageId="28" pageNumber="479" specimenCount="1" typeStatus="holotype">
<typeStatus id="549821BE691DCE58FF3BFDDAFEFEFD8C" box="[211,318,564,588]" pageId="28" pageNumber="479" type="holotype">Holotype</typeStatus>
(smaller) female: L
<emphasis id="B957430E691DCE58FDF7FDDBFDF4FD8C" box="[543,564,565,588]" italics="true" pageId="28" pageNumber="479">=</emphasis>
<quantity id="4CDB32F9691DCE58FDD3FDDAFD5FFD8C" box="[571,671,564,588]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="28" pageNumber="479" unit="mm" value="1.72">1.72 mm</quantity>
; a
<emphasis id="B957430E691DCE58FD56FDDBFD13FD8C" box="[702,723,565,588]" italics="true" pageId="28" pageNumber="479">=</emphasis>
45; b
<emphasis id="B957430E691DCE58FCFFFDDBFCECFD8C" box="[791,812,565,588]" italics="true" pageId="28" pageNumber="479">=</emphasis>
3.2; c
<emphasis id="B957430E691DCE58FC9CFDDBFC49FD8C" box="[884,905,565,588]" italics="true" pageId="28" pageNumber="479">=</emphasis>
10;
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<superScript id="7C563254691DCE58FC50FDDBFC09FD82" attach="left" box="[952,969,561,588]" fontSize="7" pageId="28" pageNumber="479"></superScript>
=
</emphasis>
6.7;
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<emphasis id="B957430E691DCE58FBDDFDDBFB8AFD8C" box="[1077,1098,565,588]" italics="true" pageId="28" pageNumber="479">=</emphasis>
50%.
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</paragraph>
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<typeStatus id="549821BE691DCE58FF3BFD8CFEF7FDBA" box="[211,311,610,634]" pageId="28" pageNumber="479" type="paratype">Paratype</typeStatus>
(larger) females (
<emphasis id="B957430E691DCE58FDECFD8DFDF0FDBA" box="[516,560,611,634]" italics="true" pageId="28" pageNumber="479">n =</emphasis>
2): L
<emphasis id="B957430E691DCE58FD93FD8DFD50FDBA" box="[635,656,611,634]" italics="true" pageId="28" pageNumber="479">=</emphasis>
<quantity id="4CDB32F9691DCE58FD71FD8FFCFDFDB9" box="[665,829,609,633]" metricMagnitude="-3" metricUnit="m" metricValue="2.3449999999999998" metricValueMax="2.42" metricValueMin="2.27" pageId="28" pageNumber="479" unit="mm" value="2.3449999999999998" valueMax="2.42" valueMin="2.27">2.272.42 mm</quantity>
; a
<emphasis id="B957430E691DCE58FC89FD8DFCB6FDBA" box="[865,886,611,634]" italics="true" pageId="28" pageNumber="479">=</emphasis>
3638; b
<emphasis id="B957430E691DCE58FC02FD8DFC3FFDBA" box="[1002,1023,611,634]" italics="true" pageId="28" pageNumber="479">=</emphasis>
3.74.0; c
<emphasis id="B957430E691DCE58FB97FD8DFB54FDBA" box="[1151,1172,611,634]" italics="true" pageId="28" pageNumber="479">=</emphasis>
10.711.0;
<emphasis id="B957430E691DCE58FEA5FD6CFEBCFD5A" box="[333,380,638,666]" italics="true" pageId="28" pageNumber="479">
<superScript id="7C563254691DCE58FEA5FD6CFE9EFD4F" attach="left" box="[333,350,638,665]" fontSize="7" pageId="28" pageNumber="479"></superScript>
=
</emphasis>
6.56.8; V
<emphasis id="B957430E691DCE58FE15FD6DFDD2FD5A" box="[509,530,643,666]" italics="true" pageId="28" pageNumber="479">=</emphasis>
48%.
<typeStatus id="549821BE691DCE58FDB0FD6CFD7CFD5A" box="[600,700,642,666]" pageId="28" pageNumber="479" type="paratype">Paratype</typeStatus>
male: L
<emphasis id="B957430E691DCE58FCCAFD6DFCF7FD5A" box="[802,823,643,666]" italics="true" pageId="28" pageNumber="479">=</emphasis>
<quantity id="4CDB32F9691DCE58FCD6FD6FFC63FD59" box="[830,931,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="28" pageNumber="479" unit="mm" value="1.62">1.62 mm</quantity>
; a
<emphasis id="B957430E691DCE58FC2BFD6DFC18FD5A" box="[963,984,643,666]" italics="true" pageId="28" pageNumber="479">=</emphasis>
34; b
<emphasis id="B957430E691DCE58FBF7FD6DFBF4FD5A" box="[1055,1076,643,666]" italics="true" pageId="28" pageNumber="479">=</emphasis>
3.0; c
<emphasis id="B957430E691DCE58FB97FD6DFB54FD5A" box="[1151,1172,643,666]" italics="true" pageId="28" pageNumber="479">=</emphasis>
49;
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<superScript id="7C563254691DCE58FF14FD4CFECCFD6E" attach="left" box="[252,268,669,697]" fontSize="7" pageId="28" pageNumber="479"></superScript>
=
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1.0.
</paragraph>
<paragraph id="8B9C9F1C691DCE58FF3BFD15FE60FCD3" blockId="28.[211,416,763,787]" box="[211,416,763,787]" pageId="28" pageNumber="479">
<emphasis id="B957430E691DCE58FF3BFD15FE60FCD3" box="[211,416,763,787]" italics="true" pageId="28" pageNumber="479">General characters</emphasis>
</paragraph>
<paragraph id="8B9C9F1C691DCE58FF3BFCCDFDEDFC1A" blockId="28.[211,1172,803,1305]" pageId="28" pageNumber="479">Body mostly twisted upon fixation, strongly tapered toward anterior end, 4048 µm (smaller specimens) or 6264 µm (larger specimens) wide at mid-region. Cuticle practically smooth, 3.04.0 µm thick on most of body. Lip region slightly offset, rounded, 1617 µm wide, somewhat wider than adjoining neck. Body at posterior end of pharynx 2.83.3 times as wide as lip region. Amphids caliciform with aperture half the corresponding body width.</paragraph>
<paragraph id="8B9C9F1C691DCE58FF16FC0CFE6FFAD9" blockId="28.[211,1172,803,1305]" pageId="28" pageNumber="479">
Labial and dental chambers well separated, the former 10 µm, the latter 67 µm broad. No pharyngeal denticles. Stylet chamber unusually roomy and thick-walled, 9 µm broad, seen clearly under lower magnification. Guiding ring at 1718 µm from anterior end. Onchia as usual, simple. Odontostyle 2426 µm long, 1.51.6 times longer than labial diameter, 2.02.2 µm thick at middle, slightly thicker than cuticle at the same level. The aperture occupying about one-third stylet length. Pharynx 550610 µm long, at 4849% expanded. Because of the strong muscular structure, the pharyngeal gland nuclei are rather indistinct. D
<emphasis id="B957430E691DCE58FD10FB2CFCCDFB19" box="[760,781,1218,1241]" italics="true" pageId="28" pageNumber="479">=</emphasis>
5052%,
<collectingCountry id="F334DF8C691DCE58FC94FB2FFC5FFB19" box="[892,927,1217,1241]" name="American Samoa" pageId="28" pageNumber="479">AS</collectingCountry>
nuclei not visible, PS
<emphasis id="B957430E691DCE58FF3BFB0CFF28FB39" box="[211,232,1250,1273]" italics="true" pageId="28" pageNumber="479">=</emphasis>
6870%. Glandularium 260296 µm long. Posterior end of pharynx with a shallow disc; cardia conoid.
</paragraph>
<paragraph id="8B9C9F1C691DCE58FF3BFAB5FEE2FAB3" blockId="28.[211,290,1371,1395]" box="[211,290,1371,1395]" pageId="28" pageNumber="479">
<emphasis id="B957430E691DCE58FF3BFAB5FEE2FAB3" box="[211,290,1371,1395]" italics="true" pageId="28" pageNumber="479">Female</emphasis>
</paragraph>
<paragraph id="8B9C9F1C691DCE5BFF3BFA6DFEBFFF1F" blockId="28.[211,1172,1411,1722]" lastBlockId="31.[143,1103,167,223]" lastPageId="31" lastPageNumber="482" pageId="28" pageNumber="479">
Reproductive system didelphic, amphidelphic. Gonads long, anterior branch 5.56.2 body widths long or occupying 1516% of body length, posterior branch 6.66.8 body widths long or occupying 1718% of body length. Vulva a short longitudinal slit, with sclerotized inner labia. Vagina 22 µm (shorter female) or 3233 µm (longer females), occupying half or less of the body width. Uteri bipartite with proximal roomy and distal narrower sections. The roomy chambers of the uteri in
<specimenCount id="9D255495691DCE58FB82F9CDFEE8F99A" pageId="28" pageNumber="479" type="female">two females</specimenCount>
(the longer ones) were packed with sperm. Uterine eggs not found. Distance between posterior end of pharynx and vulva shorter (0.70.9 times) than the pharynx itself. Rectum 1.51.8 times and pre-rectum 3.24.0 times the anal body diameter long. Vulvaanus distance equal to 3.94.7 tail lengths. Tail 178220 µm long or 910% of entire length of body, first conoid then strongly tapered to its very fine filiform tip, slightly bent dorsally.
</paragraph>
<caption id="DF5CCF94691CCE59FF67F9D4FDC7F9AD" ID-DOI="http://doi.org/10.5281/zenodo.5202611" ID-Zenodo-Dep="5202611" httpUri="https://zenodo.org/record/5202611/files/figure.png" pageId="29" pageNumber="480" startId="29.[143,210,1594,1616]" targetBox="[143,1102,161,1558]" targetPageId="29">
<paragraph id="8B9C9F1C691CCE59FF67F9D4FDC7F9AD" blockId="29.[143,1104,1594,1646]" pageId="29" pageNumber="480">
Figure 10.
<taxonomicName id="4C23E49F691CCE59FEEEF9D5FE68F991" authority="Andrássy, 2012" authorityName="Andrássy" authorityYear="2012" box="[262,424,1595,1617]" class="Adenophorea" family="Actinolaimidae" genus="Egtitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="29" pageNumber="480" phylum="Nematoda" rank="species" species="biformis" status="sp. nov.">
<emphasis id="B957430E691CCE59FEEEF9D5FE68F991" box="[262,424,1595,1617]" italics="true" pageId="29" pageNumber="480">Egtitus biformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264FE75691CCE59FE5AF9D5FDC2F990" box="[434,514,1595,1616]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
(A) Anterior end; (B) cardial region; (C) vulval region; (D, E) female tails. Scale bars 20 µm.
</paragraph>
</caption>
<caption id="DF5CCF94691FCE5AFF3BF9CCFD53F995" ID-DOI="http://doi.org/10.5281/zenodo.5202613" ID-Zenodo-Dep="5202613" httpUri="https://zenodo.org/record/5202613/files/figure.png" pageId="30" pageNumber="481" startId="30.[211,278,1570,1592]" targetBox="[211,1170,161,1535]" targetPageId="30">
<paragraph id="8B9C9F1C691FCE5AFF3BF9CCFD53F995" blockId="30.[211,1172,1570,1622]" pageId="30" pageNumber="481">
Figure 11.
<taxonomicName id="4C23E49F691FCE5AFEAFF9CDFE26F9F9" authority="Andrássy, 2012" authorityName="Andrássy" authorityYear="2012" box="[327,486,1571,1593]" class="Adenophorea" family="Actinolaimidae" genus="Egtitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="30" pageNumber="481" phylum="Nematoda" rank="species" species="biformis" status="sp. nov.">
<emphasis id="B957430E691FCE5AFEAFF9CDFE26F9F9" box="[327,486,1571,1593]" italics="true" pageId="30" pageNumber="481">Egtitus biformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264FE75691FCE5AFE05F9CDFDFAF9F8" box="[493,570,1571,1592]" pageId="30" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
(A) Tip of the anterior testis; (B) tip of the posterior testis; (C) posterior end of male. Scale bars 20 µm.
</paragraph>
</caption>
<paragraph id="8B9C9F1C691ECE5BFF67FECAFF09FEFC" blockId="31.[143,201,292,316]" box="[143,201,292,316]" pageId="31" pageNumber="482">
<emphasis id="B957430E691ECE5BFF67FECAFF09FEFC" box="[143,201,292,316]" italics="true" pageId="31" pageNumber="482">Male</emphasis>
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FEA3FBE3FE03" blockId="31.[143,1104,332,452]" pageId="31" pageNumber="482">
Testes two, comparatively short, each three body widths long with reflexed distal end. Spermatozoa fusiform. Spicula dorylaimoid, 58 µm long in curvature. Ventromedial supplements distinct, well spaced,
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number. Pre-rectum beginning at the level of the second supplement. Tail 34 µm long or 2.0% of body length, conoid-rounded.
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FDE7FE7BFDE1" blockId="31.[143,443,521,545]" box="[143,443,521,545]" pageId="31" pageNumber="482">
<emphasis id="B957430E691ECE5BFF67FDE7FE7BFDE1" box="[143,443,521,545]" italics="true" pageId="31" pageNumber="482">Diagnosis and relationships</emphasis>
</paragraph>
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Body either under
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and slender, or over
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and more robust, lip region narrow, stylet chamber unusually roomy, odontostyle about one-and-a-half times the lip region width long, female gonads long, male gonads short and distally ventrally reflexed, supplements 10, female tail medium long, finely pointed.
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF52FD5EFD44FC47" blockId="31.[143,1104,560,904]" pageId="31" pageNumber="482">
This interesting species shows the usual structures typical for
<taxonomicName id="4C23E49F691ECE5BFC97FD5FFC0EFD08" authorityName=", Thorne" authorityYear="1967" box="[895,974,689,712]" class="Adenophorea" family="Actinolaimidae" genus="Egtitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="31" pageNumber="482" phylum="Nematoda" rank="genus">
<emphasis id="B957430E691ECE5BFC97FD5FFC0EFD08" box="[895,974,689,712]" italics="true" pageId="31" pageNumber="482">Egtitus</emphasis>
</taxonomicName>
, but at the same time it has some special characters. These latter are: the stylet chamber is roomy, the testes are reflexed on their distal ends, and the body size is different among specimens of the same population (the shorter female amounts to 7175% the length of the longer females). As for the shape of the stylet chamber, the representatives of
<taxonomicName id="4C23E49F691ECE5BFF67FCA1FEDBFCA7" authorityName="Kahn, Ahmad and Jairajpuri" authorityYear="1994" box="[143,283,847,871]" class="Adenophorea" family="Actinolaimidae" genus="Stopractinca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="31" pageNumber="482" phylum="Nematoda" rank="genus">
<emphasis id="B957430E691ECE5BFF67FCA1FEDBFCA7" box="[143,283,847,871]" italics="true" pageId="31" pageNumber="482">Stopractinca</emphasis>
</taxonomicName>
are similar in this respect, but their onchia are bi-tipped and the males have a great number (1825) of supplements.
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FC23FEFAFC24" blockId="31.[143,314,973,996]" box="[143,314,973,996]" pageId="31" pageNumber="482">
<emphasis id="B957430E691ECE5BFF67FC23FEFAFC24" box="[143,314,973,996]" italics="true" pageId="31" pageNumber="482">Type specimens</emphasis>
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FC1AFEC0FB8B" blockId="31.[143,1103,1012,1099]" pageId="31" pageNumber="482">
<typeStatus id="549821BE691ECE5BFF67FC1AFF3AFBCC" box="[143,250,1012,1036]" pageId="31" pageNumber="482" type="holotype">Holotype</typeStatus>
female on slide no. 13711.
<materialsCitation id="3B4B9541691ECE5BFDDBFC1AFF3CFB8B" collectionCode="ELTE" county="Deposited" location="University" municipality="Department of Systematic Zoology" pageId="31" pageNumber="482" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Budapest" typeStatus="paratype">
<typeStatus id="549821BE691ECE5BFDDBFC1AFD65FBCC" box="[563,677,1012,1036]" pageId="31" pageNumber="482" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D255495691ECE5BFD58FC1AFCF9FBCC" box="[688,825,1012,1036]" pageId="31" pageNumber="482" type="female">two females</specimenCount>
,
<specimenCount id="9D255495691ECE5BFCADFC1AFC6CFBCC" box="[837,940,1012,1036]" pageId="31" pageNumber="482" type="male">one male</specimenCount>
and
<specimenCount id="9D255495691ECE5BFC0FFC1AFF7AFBEB" pageId="31" pageNumber="482" type="juvenile">one juvenile</specimenCount>
.
<collectingCounty id="62FDE790691ECE5BFF20FBFDFEFBFBEB" box="[200,315,1043,1067]" pageId="31" pageNumber="482">Deposited</collectingCounty>
at the
<collectingMunicipality id="6BF80566691ECE5BFE7CFBFAFCECFBEB" box="[404,812,1043,1067]" pageId="31" pageNumber="482">Department of Systematic Zoology</collectingMunicipality>
of the
<collectionCode id="ED3207D9691ECE5BFC6EFBFAFC0AFBEB" box="[902,970,1044,1067]" pageId="31" pageNumber="482">ELTE</collectionCode>
<location id="8EFCC9C7691ECE5BFC3DFBFDFB8CFBEB" LSID="urn:lsid:plazi:treatment:038A2E0A691DCE5BFDC6FE3EFB90FAB5:8EFCC9C7691ECE5BFC3DFBFDFB8CFBEB" box="[981,1100,1043,1067]" county="Deposited" municipality="Department of Systematic Zoology" name="University" pageId="31" pageNumber="482" stateProvince="Budapest">University</location>
,
<collectingRegion id="49E751FE691ECE5BFF67FBDDFF3CFB8B" box="[143,252,1075,1099]" country="Hungary" name="Budapest" pageId="31" pageNumber="482">Budapest</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FB7FFE61FB68" blockId="31.[143,417,1168,1192]" box="[143,417,1168,1192]" pageId="31" pageNumber="482">
<emphasis id="B957430E691ECE5BFF67FB7FFE61FB68" box="[143,417,1168,1192]" italics="true" pageId="31" pageNumber="482">Type habitat and locality</emphasis>
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FB57FD3CFB30" blockId="31.[143,1103,1208,1264]" pageId="31" pageNumber="482">
<materialsCitation id="3B4B9541691ECE5BFF67FB57FD38FB30" collectingDate="1969-09" collectorName="J. Balogh" country="Papua New Guinea" county="Fly River" location="Wet" municipality="Kiunga" pageId="31" pageNumber="482" specimenCount="1" stateProvince="Western">
<location id="8EFCC9C7691ECE5BFF67FB57FF7BFB10" LSID="urn:lsid:plazi:treatment:038A2E0A691DCE5BFDC6FE3EFB90FAB5:8EFCC9C7691ECE5BFF67FB57FF7BFB10" box="[143,187,1209,1232]" country="Papua New Guinea" county="Fly River" municipality="Kiunga" name="Wet" pageId="31" pageNumber="482" stateProvince="Western">Wet</location>
fallen leaves from a rain forest,
<collectingMunicipality id="6BF80566691ECE5BFDF6FB56FDB3FB10" box="[542,627,1208,1232]" pageId="31" pageNumber="482">Kiunga</collectingMunicipality>
by the
<collectingCounty id="62FDE790691ECE5BFD2CFB56FCEFFB10" box="[708,815,1208,1232]" pageId="31" pageNumber="482">Fly River</collectingCounty>
,
<collectingRegion id="49E751FE691ECE5BFCD1FB57FC3EFB10" box="[825,1022,1208,1232]" country="Papua New Guinea" name="Western" pageId="31" pageNumber="482">Western Province</collectingRegion>
,
<collectingCountry id="F334DF8C691ECE5BFBE0FB57FEE0FB30" name="Papua New Guinea" pageId="31" pageNumber="482">Papua New Guinea</collectingCountry>
; collected in
<date id="FF9DB9DC691ECE5BFE59FB36FDA5FB30" box="[433,613,1240,1264]" pageId="31" pageNumber="482" value="1969-09">
<collectingDate id="EFD94034691ECE5BFE59FB36FDA5FB30" box="[433,613,1240,1264]" pageId="31" pageNumber="482" value="1969-09">September 1969</collectingDate>
</date>
by
<collectorName id="26D6FACA691ECE5BFD66FB37FD38FB30" box="[654,760,1240,1264]" pageId="31" pageNumber="482">J. Balogh</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FADBFEC7FA8D" blockId="31.[143,263,1333,1357]" box="[143,263,1333,1357]" pageId="31" pageNumber="482">
<emphasis id="B957430E691ECE5BFF67FADBFEC7FA8D" box="[143,263,1333,1357]" italics="true" pageId="31" pageNumber="482">Etymology</emphasis>
</paragraph>
<paragraph id="8B9C9F1C691ECE5BFF67FAB3FB90FAB5" blockId="31.[143,1104,1373,1397]" box="[143,1104,1373,1397]" pageId="31" pageNumber="482">
The epithet
<taxonomicName id="4C23E49F691ECE5BFEFEFAB3FEB0FAB5" authorityName="Andrássy" authorityYear="2012" box="[278,368,1373,1397]" class="Adenophorea" family="Actinolaimidae" genus="Egtitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="31" pageNumber="482" phylum="Nematoda" rank="species" species="biformis">
<emphasis id="B957430E691ECE5BFEFEFAB3FEB0FAB5" box="[278,368,1373,1397]" italics="true" pageId="31" pageNumber="482">biformis</emphasis>
</taxonomicName>
comes from the Latin, and means: dimorphous, or, of dual shape.
</paragraph>
</subSubSection>
</treatment>
</document>