267 lines
52 KiB
XML
267 lines
52 KiB
XML
<document id="97A63CFA9DB696CAD4377EE277A35B4C" ID-DOI="10.1093/zoolinnean/zlad026" ID-ISSN="0024-4082" ID-Zenodo-Dep="8432514" ID-ZooBank="5F8CB822-E5A6-4E68-8DB6-A5E07104F898" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696940055617" checkinUser="plazi" docAuthor="Ferreira, Vinicius S., Barbosa, Felipe Francisco, Bocakova, Milada & Solodovnikov, Alexey" docDate="2023" docId="0388D85FFFEF64247460FC3EFE57FF46" docLanguage="en" docName="zlad026.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad026" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Xenomorphon Ferreira & Barbosa & Bocakova & Solodovnikov 2023, gen. nov." docType="treatment" docVersion="3" lastPageNumber="560" masterDocId="FFB1A027FFE964237548FF9AFFEEFFA2" masterDocTitle="An extraordinary case of elytra loss in Coleoptera (Elateroidea: Lycidae): discovery and placement of the first anelytrous adult male beetle" masterLastPageNumber="566" masterPageNumber="553" pageNumber="559" updateTime="1697046424687" updateUser="ExternalLinkService">
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<mods:title id="CF7BF07B9D5440735614F99C1F8232FE">An extraordinary case of elytra loss in Coleoptera (Elateroidea: Lycidae): discovery and placement of the first anelytrous adult male beetle</mods:title>
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<mods:namePart id="3F010613FB1A2CD47A5ED9F24A18DB54">Ferreira, Vinicius S.</mods:namePart>
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<mods:affiliation id="020CEBB6D9173FCCECAE808221F30DBC">Natural History Museum of Denmark, Zoological Museum, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="C2C5C775BD3D00B45F9C31A07E022A98" type="email">vinicius.sffl@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F62DD8276C9F8BBE946638587C8588A1">Barbosa, Felipe Francisco</mods:namePart>
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<mods:affiliation id="AEEAC699C9CF222A18BE093B94FCB524">Laboratório de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Avenida Carlos Chagas Filho, 373, Cidade Universitária, Ilha do Fundão, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:namePart id="821B9EA9B7D2B37C28A7527DCC56DFF4">Bocakova, Milada</mods:namePart>
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<mods:affiliation id="48723A5467ECB1D22F801AA6FE6D0F09">Department of Biology, Faculty of Education, Palacky University, Purkrabska 2, CZ- 77140, Olomouc, Czech Republic</mods:affiliation>
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<mods:namePart id="8DA106221593DFC5E39C857419A695E1">Solodovnikov, Alexey</mods:namePart>
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<mods:affiliation id="3CEF341DDBB7FC1AF082CA561253003D">Natural History Museum of Denmark, Zoological Museum, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:title id="51341D2F5F0FDA5873491B80C667E4DC">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5B6F5F8FA03731F1BD010E8DFF92DE0D">2023</mods:date>
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<mods:number id="832F7633CA21E31C7A4DA8556994847E">2023-07-01</mods:number>
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<treatment id="0388D85FFFEF64247460FC3EFE57FF46" ID-DOI="http://doi.org/10.5281/zenodo.8432516" ID-Zenodo-Dep="8432516" LSID="urn:lsid:plazi:treatment:0388D85FFFEF64247460FC3EFE57FF46" httpUri="http://treatment.plazi.org/id/0388D85FFFEF64247460FC3EFE57FF46" lastPageId="7" lastPageNumber="560" pageId="6" pageNumber="559">
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<subSubSection id="C33B3AC2FFEF64257460FC3EFD8CFC1D" box="[296,610,932,959]" pageId="6" pageNumber="559" type="nomenclature">
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<paragraph id="8B9E6949FFEF64257460FC3EFD8CFC1D" blockId="6.[296,610,932,959]" box="[296,610,932,959]" pageId="6" pageNumber="559">
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<heading id="D0D6DE25FFEF64257460FC3EFD8CFC1D" bold="true" box="[296,610,932,959]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="559" reason="3">
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<emphasis id="B955B55BFFEF64257460FC3EFD8CFC1D" bold="true" box="[296,610,932,959]" pageId="6" pageNumber="559">
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Genus
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<taxonomicName id="4C2112CAFFEF6425743AFC3EFDECFC1D" authority="Ferreira & Barbosa & Bocakova & Solodovnikov, 2023" authorityName="Ferreira & Barbosa & Bocakova & Solodovnikov" authorityYear="2023" box="[370,514,932,959]" class="Insecta" family="Lycidae" genus="Xenomorphon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B955B55BFFEF6425743AFC3EFDECFC1D" bold="true" box="[370,514,932,959]" italics="true" pageId="6" pageNumber="559">Xenomorphon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2660820FFEF6425774EFC3FFD8CFC1D" box="[518,610,933,959]" pageId="6" pageNumber="559" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C33B3AC2FFEF642474C5FC4FFE57FF46" lastPageId="7" lastPageNumber="560" pageId="6" pageNumber="559" type="description">
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<paragraph id="8B9E6949FFEF642574C5FC4FFE13FC4D" blockId="6.[397,509,981,1007]" box="[397,509,981,1007]" pageId="6" pageNumber="559">
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<emphasis id="B955B55BFFEF642574C5FC4FFE13FC4D" bold="true" box="[397,509,981,1007]" pageId="6" pageNumber="559">
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(
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<figureCitation id="131A75CCFFEF642574D0FC4FFE1CFC4D" box="[408,498,981,1007]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="3.[113,178,1650,1674]" captionStartId-1="4.[130,195,810,834]" captionStartId-2="5.[113,178,1772,1796]" captionTargetBox-0="[309,1263,147,1619]" captionTargetBox-2="[269,1303,147,1741]" captionTargetId-0="figure-155@3.[306,1266,144,1622]" captionTargetId-1="figure-644@4.[321,1281,144,782]" captionTargetId-2="figure-7@5.[266,1306,144,1744]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." captionText-1="Figure 3. Distribution of Xenomorphon baranowskii in Mexico." captionText-2="Figure 4. A, distribution of elytral morphology and habitats in paedomorphic Lycidae. Adapted from the tree topology presented by Kusy et al. (2019) and data from Bocakova (2014). B, Preliminary phylogenetic hypothesis for the placement of Xenomorphon baranowskii in the Calopterini resulting from the morphology-based maximum likelihood (ML) analysis; k-states Markov (Mkv+G) model; Extended majority-rule consensus (EMRC) tree.Green circles in the nodes represent branches with support levels ≥70% from at least one of the used reconstruction methods." figureDoi-0="http://doi.org/10.5281/zenodo.8432520" figureDoi-1="http://doi.org/10.5281/zenodo.8432524" figureDoi-2="http://doi.org/10.5281/zenodo.8432526" httpUri-0="https://zenodo.org/record/8432520/files/figure.png" httpUri-1="https://zenodo.org/record/8432524/files/figure.png" httpUri-2="https://zenodo.org/record/8432526/files/figure.png" pageId="6" pageNumber="559">Figs 2–4</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="8B9E6949FFEF642575C9FB8CFD3BFBEF" blockId="6.[129,778,1046,1101]" pageId="6" pageNumber="559">
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<emphasis id="B955B55BFFEF642575C9FB8CFE8BFB8F" box="[129,357,1046,1070]" italics="true" pageId="6" pageNumber="559">Zoobank registration.</emphasis>
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hưp://zoobank.org/
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<uri id="FFB0654BFFEF64257718FB8CFD3BFBEF" pageId="6" pageNumber="559">
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urn:lsid:zoobank. org:act:
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<uuid id="FF87539CFFEF64257581FBAFFD3BFBEF" box="[201,725,1077,1101]" pageId="6" pageNumber="559">D958AFA5-4ABA-4AD4-B3DD-7B40E67BCC35</uuid>
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</uri>
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</paragraph>
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<paragraph id="8B9E6949FFEF642575C9FBF4FD38FA23" blockId="6.[128,778,1134,1409]" pageId="6" pageNumber="559">
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<emphasis id="B955B55BFFEF642575C9FBF4FEBDFB24" box="[129,339,1134,1158]" italics="true" pageId="6" pageNumber="559">Differential diagnosis.</emphasis>
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Body dark brown, compressed, elytra and membranous wings absent (
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<figureCitation id="131A75CCFFEF642574E5FB14FE17FB04" box="[429,505,1166,1190]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A</figureCitation>
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). Pronotum subquadrate, with a median longitudinal carina developed, scutellar shield absent (
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<figureCitation id="131A75CCFFEF642575F4FB56FEDAFB46" box="[188,308,1228,1252]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, 2D</figureCitation>
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). Antennae inserted on gibbous prominence, distinctly serrate from antennomeres III–X, with pedicel ca. 3× longer than antennomere III (
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<figureCitation id="131A75CCFFEF64257480FA91FDF8FA81" box="[456,534,1291,1315]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A</figureCitation>
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,
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<figureCitation id="131A75CCFFEF64257760FA91FDA9FA81" box="[552,583,1291,1315]" captionStart="Figure 3" captionStartId="4.[130,195,810,834]" captionTargetId="figure-644@4.[321,1281,144,782]" captionTargetPageId="4" captionText="Figure 3. Distribution of Xenomorphon baranowskii in Mexico." figureDoi="http://doi.org/10.5281/zenodo.8432524" httpUri="https://zenodo.org/record/8432524/files/figure.png" pageId="6" pageNumber="559">3C</figureCitation>
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). Maxillary, labial palps and mandibles fully developed (
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<figureCitation id="131A75CCFFEF64257745FAB0FDB8FAE0" box="[525,598,1322,1346]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2E</figureCitation>
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). Legs short and stubby, tibiae tapered basally, gradually expanding towards the apex (
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<figureCitation id="131A75CCFFEF642575F7FAF3FEEBFA23" box="[191,261,1385,1409]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2B</figureCitation>
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). Male genitalia distinctly trilobate (
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<figureCitation id="131A75CCFFEF64257733FAF3FD2BFA23" box="[635,709,1385,1409]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2G</figureCitation>
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).
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</paragraph>
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<paragraph id="8B9E6949FFEF642575C8FA3CFF00F9F8" blockId="6.[128,778,1446,1626]" pageId="6" pageNumber="559">
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<emphasis id="B955B55BFFEF642575C8FA3CFF03FA1C" box="[128,237,1446,1470]" italics="true" pageId="6" pageNumber="559">Etymology.</emphasis>
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The name of this genus is an allusion to the unique appearance of the studied specimen, which at first thought was not even considered to be a beetle.
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<emphasis id="B955B55BFFEF64257755FA7FFDB8FA5E" box="[541,598,1509,1532]" italics="true" pageId="6" pageNumber="559">Xeno‐</emphasis>
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, from the Greek, ξένος, meaning strange, alien, exotic or different, and
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<emphasis id="B955B55BFFEF642577E3F999FCEAF9B9" box="[683,772,1539,1563]" italics="true" pageId="6" pageNumber="559">morphon</emphasis>
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, from the Greek μορφῶν, meaning of shape or form. Gender masculine.
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</paragraph>
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<paragraph id="8B9E6949FFEF642575C8F90AFD98F90A" blockId="6.[128,630,1679,1704]" box="[128,630,1679,1704]" pageId="6" pageNumber="559">
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<emphasis id="B955B55BFFEF642575C8F90AFDF1F905" box="[128,543,1679,1703]" italics="true" pageId="6" pageNumber="559">
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<typeStatus id="549AD7EBFFEF642575C8F90AFF5EF905" box="[128,176,1680,1703]" pageId="6" pageNumber="559">Type</typeStatus>
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species.
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<taxonomicName id="4C2112CAFFEF6425744CF915FDF1F905" authority="Ferreira & Barbosa & Bocakova & Solodovnikov, 2023" authorityName="Ferreira & Barbosa & Bocakova & Solodovnikov" authorityYear="2023" box="[260,543,1679,1703]" class="Insecta" family="Lycidae" genus="Xenomorphon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="baranowskii" status="sp. nov.">
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<emphasis id="B955B55BFFEF6425744CF915FDF1F905" bold="true" box="[260,543,1679,1703]" pageId="6" pageNumber="559">Xenomorphon baranowskii</emphasis>
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</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A2660820FFEF6425776CF90AFD98F90A" box="[548,630,1680,1704]" pageId="6" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B9E6949FFEF642575C8F950FF11F89D" blockId="6.[128,778,1737,2011]" pageId="6" pageNumber="559">
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<emphasis id="B955B55BFFEF642575C8F950FE92F943" box="[128,380,1737,1761]" italics="true" pageId="6" pageNumber="559">Description of adult male.</emphasis>
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General dorsal coloration dark brown, pedicel slightly lighter, yellowish (
|
||
<figureCitation id="131A75CCFFEF642574A5F972FDD5F8A2" box="[493,571,1768,1792]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A</figureCitation>
|
||
). Body moderately setose, with short, decumbent yellow setation throughout (
|
||
<figureCitation id="131A75CCFFEF642575C3F8BDFF00F89C" box="[139,238,1831,1855]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF642575D4F8DCFC4EFF45" blockId="6.[128,778,1737,2011]" lastBlockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF642575D4F8DCFF31F8FC" bold="true" box="[156,223,1862,1886]" pageId="6" pageNumber="559">Head:</emphasis>
|
||
as long as wide, widest at eyes, posteriorly covered, in part, by pronotum, hypognathous, inclined, frons strongly bulgy, bearing pits behind antennal insertion (
|
||
<figureCitation id="131A75CCFFEF64257726F81FFD17F83E" box="[622,761,1925,1949]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A–C, E</figureCitation>
|
||
). Posterior margin of epistoma clearly emarginate; labrum distinct from clypeus, transverse and small, apparently moveable and connected to clypeus with membrane. Eyes hemispherical, projecting anterolaterally in dorsal view, coarsely granulate (
|
||
<figureCitation id="131A75CCFFEF642570D5FF35FC61FF44" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A–C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF6425761DFF74FA58FE43" blockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF6425761DFF74FC2AFEA4" bold="true" box="[853,964,238,262]" pageId="6" pageNumber="559">Antennae:</emphasis>
|
||
distinctly serrate from antennomere III–IX, densely setose, dorsoventrally flaưened from III–XI, inserted in gibbous prominence at anterior distal portion of head, 11-segmented; in resting position reaching to basal half of body length (
|
||
<figureCitation id="131A75CCFFEF64257006FED6FA5FFEC6" box="[1358,1457,332,356]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, C</figureCitation>
|
||
). Scape pyriform, approximated at base by half distance of scape width, subconical (
|
||
<figureCitation id="131A75CCFFEF642576B4FE10FBABFE01" box="[1020,1093,394,419]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2C</figureCitation>
|
||
); pedicel minute, ca. 4× shorter than scape; antennomeres III-IX slightly increasing in length towards apex (
|
||
<figureCitation id="131A75CCFFEF6425763EFE53FC38FE43" box="[886,982,457,481]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
); antennomere XI round apically (
|
||
<figureCitation id="131A75CCFFEF6425707AFE53FA90FE43" box="[1330,1406,457,481]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="131A75CCFFEF642570C0FE53FA48FE43" box="[1416,1446,457,481]" captionStart="Figure 3" captionStartId="4.[130,195,810,834]" captionTargetId="figure-644@4.[321,1281,144,782]" captionTargetPageId="4" captionText="Figure 3. Distribution of Xenomorphon baranowskii in Mexico." figureDoi="http://doi.org/10.5281/zenodo.8432524" httpUri="https://zenodo.org/record/8432524/files/figure.png" pageId="6" pageNumber="559">3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF6425761DFE72FA81FDDC" blockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF6425761DFE72FC0EFDA2" bold="true" box="[853,992,488,512]" pageId="6" pageNumber="559">Mouthparts:</emphasis>
|
||
mandibles small, hook-shaped, opposable (
|
||
<figureCitation id="131A75CCFFEF642570D4FE72FCBAFD82" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2E</figureCitation>
|
||
). Maxillary palp four-segmented (
|
||
<figureCitation id="131A75CCFFEF64257185FD92FAF7FD82" box="[1229,1305,520,544]" captionStart="Figure 3" captionStartId="4.[130,195,810,834]" captionTargetId="figure-644@4.[321,1281,144,782]" captionTargetPageId="4" captionText="Figure 3. Distribution of Xenomorphon baranowskii in Mexico." figureDoi="http://doi.org/10.5281/zenodo.8432524" httpUri="https://zenodo.org/record/8432524/files/figure.png" pageId="6" pageNumber="559">Fig. 3B</figureCitation>
|
||
); palpomere II elongate, as long as the sum of I + III + IV (
|
||
<figureCitation id="131A75CCFFEF64257041FDBDFABFFD9D" box="[1289,1361,551,575]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2E</figureCitation>
|
||
); terminal palpomeres abruptly acuminate (
|
||
<figureCitation id="131A75CCFFEF642571D4FDDCFB06FDFC" box="[1180,1256,582,606]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2E</figureCitation>
|
||
). Labial palp threesegmented; terminal palpomere slightly blunt (
|
||
<figureCitation id="131A75CCFFEF64257050FDFCFAB0FDDC" box="[1304,1374,614,638]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF6425761DFD1FFBB5FCFA" blockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF6425761DFD1FFBDEFD3F" bold="true" box="[853,1072,645,669]" pageId="6" pageNumber="559">Thorax, prothorax:</emphasis>
|
||
pronotum nearly as long as wide, subrectangular (
|
||
<figureCitation id="131A75CCFFEF642576ACFD3EFBBBFD1E" box="[996,1109,676,700]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, D</figureCitation>
|
||
); margins distinct and developed, angles round, bearing a median longitudinal carina developed throughout (
|
||
<figureCitation id="131A75CCFFEF64257149FD79FB9DFD58" box="[1025,1139,739,763]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, D</figureCitation>
|
||
). Hypomeron slightly concave. Prosternum Y-shaped, posteriorly bifurcated apically, divergent towards procoxae (
|
||
<figureCitation id="131A75CCFFEF64257143FCB8FBB6FC98" box="[1035,1112,802,826]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2B</figureCitation>
|
||
). Protrochantins slender and exposed. Procoxal cavity open.
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF6425761CFCFAFB2CFBF1" blockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF6425761CFCFAFBD4FCDA" bold="true" box="[852,1082,864,888]" pageId="6" pageNumber="559">Thorax, mesothorax:</emphasis>
|
||
mesonotum plate-like, apically round, with angles slightly narrowed (
|
||
<figureCitation id="131A75CCFFEF6425713DFC1AFB30FC35" box="[1141,1246,896,920]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, D</figureCitation>
|
||
). Mesoscutum anteriorly divided by a tiny median suture (Supporting Information, Supplementary Material 4,
|
||
<figureCitation id="131A75CCFFEF6425712BFC24FB5EFC74" box="[1123,1200,958,982]" captionStart="Figure 1" captionStartId="2.[128,193,1449,1473]" captionTargetBox="[212,1390,147,1419]" captionTargetId="figure-208@2.[209,1393,144,1421]" captionTargetPageId="2" captionText="Figure 1. Diversity of reduction or complete loss of flight in different beetle families. A, Atractocerus procerus Schenkling, 1914 (Lymexylidae), Roura, French Guyana (image: Bernard Dupont). B, Thylodrias contractus Motschulsky, 1839 (Dermestidae) (female), North Carolina, USA (image: Maư Bertone).C, Carabus hungaricus Fabricius, 1792 (Carabidae), Hungary (image: Nikola Rahmé). D, Ripiphorus vierecki (Fall, 1907) (Ripiphoridae) (image: Glenn Seplak).E, Phosphaenus hemipterus (Goeze, 1777) (Lampyridae), France (image: Pierre Bornand). F, Berberomeloe majalis (Linnaeus, 1758) (Meloidae), Jimena de la Frontera, Andalucía, Spain (image:Gail Hampshire).Photographs were used with permission from the authors." figureDoi="http://doi.org/10.5281/zenodo.8432518" httpUri="https://zenodo.org/record/8432518/files/figure.png" pageId="6" pageNumber="559">Fig. S1</figureCitation>
|
||
). Elytra absent, with no vestiges of wing-base sclerites. Mesospiracles pill-shaped (
|
||
<figureCitation id="131A75CCFFEF642570D4FC44FCBAFBB7" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2B</figureCitation>
|
||
). Mesoventrite subtrapezoidal, with distinct visible suture connecting mesoventrite and mesanepisternum (
|
||
<figureCitation id="131A75CCFFEF64257020FB86FA5FFB96" box="[1384,1457,1052,1076]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2B</figureCitation>
|
||
). Mesotrochantins slender and exposed.
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF6425761CFBC1FC11FB52" blockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF6425761CFBC1FBD5FBD1" bold="true" box="[852,1083,1115,1139]" pageId="6" pageNumber="559">Thorax, metathorax:</emphasis>
|
||
metanotum plate-like, having round angles; scutellar shield absent (
|
||
<figureCitation id="131A75CCFFEF6425713FFBE0FB0FFB30" box="[1143,1249,1146,1170]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, D</figureCitation>
|
||
). Membranous wings absent, with no vestiges of wing-base sclerites (
|
||
<figureCitation id="131A75CCFFEF64257077FB03FA5CFB13" box="[1343,1458,1177,1202]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, D</figureCitation>
|
||
). Metaventrite convex, posterolateral angles acute; metadiscrimen complete (
|
||
<figureCitation id="131A75CCFFEF642576E0FB42FC00FB52" box="[936,1006,1240,1264]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF6425761DFB62FA53FA49" blockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF6425761DFB62FC7EFAB2" bold="true" box="[853,912,1272,1296]" pageId="6" pageNumber="559">Legs:</emphasis>
|
||
short and stubby (
|
||
<figureCitation id="131A75CCFFEF64257119FB6DFB5AFAAD" box="[1105,1204,1271,1296]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
). Coxae conical, obliquely positioned, inner margin nearly contiguous but not touching (
|
||
<figureCitation id="131A75CCFFEF6425760CFAACFC40FAEC" box="[836,942,1334,1358]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
); posterior face of metacoxae shallowly excavated. Trochanters shortened; subrectangular, not tubular (
|
||
<figureCitation id="131A75CCFFEF64257006FACFFA5FFACF" box="[1358,1457,1365,1390]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
). Femora and tibiae subequal in length (
|
||
<figureCitation id="131A75CCFFEF642571FFFAEFFAF9FA2E" box="[1207,1303,1397,1421]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
); femora parallel sided; tibiae tapered basally, gradually expanding towards the apex (
|
||
<figureCitation id="131A75CCFFEF6425763FFA29FC37FA69" box="[887,985,1459,1484]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
). Tarsal formula 5-5-5; all tarsomeres narrowed; terminal tarsomere bearing two simple claws at end (
|
||
<figureCitation id="131A75CCFFEF64257003FA49FA42FA48" box="[1355,1452,1491,1515]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF6425761DFA68FAF3F8A7" blockId="6.[825,1476,144,1797]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF6425761DFA68FC24F9A8" bold="true" box="[853,970,1522,1546]" pageId="6" pageNumber="559">Abdomen:</emphasis>
|
||
with eight ventrites (sternites II–IX); ventrite seven medially apically notched; ventrite eight lanceolate, elongate, apically acuminate with apex blunt (
|
||
<figureCitation id="131A75CCFFEF64257060F9ABFA9DF9EB" box="[1320,1395,1585,1609]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2B</figureCitation>
|
||
). Male genitalia typical elateriformia: trilobate, symmetrical (
|
||
<figureCitation id="131A75CCFFEF6425702CF9CAFA5EF9CA" box="[1380,1456,1616,1640]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2G</figureCitation>
|
||
); phallobase posteriorly round, median suture absent (
|
||
<figureCitation id="131A75CCFFEF6425702AF9F5FA5EF925" box="[1378,1456,1647,1671]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2G</figureCitation>
|
||
); parameres approximately half the length of median lobe, apices round (
|
||
<figureCitation id="131A75CCFFEF642576CFF934FC3FF964" box="[903,977,1710,1734]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2G</figureCitation>
|
||
); median lobe elongate, nearly parallel-sided, apically blunt, exposed portion approximately one-quarter longer the length of parameres + phallobase (
|
||
<figureCitation id="131A75CCFFEF6425718AF977FAE3F8A7" box="[1218,1293,1773,1797]" captionStart="Figure 2" captionStartId="3.[113,178,1650,1674]" captionTargetBox="[309,1263,147,1619]" captionTargetId="figure-155@3.[306,1266,144,1622]" captionTargetPageId="3" captionText="Figure 2. Xenomorphon baranowskii. A, C, D, dry specimen. B, E–G, photographed in glycerin.A, dorsal view.B, ventral view. C, detail of head in dorsal view.D, thorax in dorsal view.E, detail of mouthparts. F, detail of male terminalia in ventral view. G, male genitalia in dorsal, lateral and ventral view." figureDoi="http://doi.org/10.5281/zenodo.8432520" httpUri="https://zenodo.org/record/8432520/files/figure.png" pageId="6" pageNumber="559">Fig. 2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF64257671F8B3FA49F8C3" blockId="6.[825,1474,1833,1889]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF64257671F8B3FC58F8E3" box="[825,950,1833,1857]" italics="true" pageId="6" pageNumber="559">Distribution.</emphasis>
|
||
<collectingCountry id="F33629D9FFEF64257682F8B3FBF6F8E3" box="[970,1048,1833,1857]" name="Mexico" pageId="6" pageNumber="559">Mexico</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E5A7ABFFEF6425716EF8B3FB9AF8E3" box="[1062,1140,1833,1857]" country="Mexico" name="Oaxaca" pageId="6" pageNumber="559">Oaxaca</collectingRegion>
|
||
, near the municipality of Valle Nacional (estimated
|
||
<geoCoordinate id="EE150F8EFFEF64257144F8D3FB47F8C2" box="[1036,1193,1863,1889]" degrees="17" direction="north" minutes="14" orientation="latitude" pageId="6" pageNumber="559" precision="1" seconds="48.85" value="17.246902">17°14ʹ48.85″N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE150F8EFFEF642571FAF8D3FABBF8C2" box="[1202,1365,1863,1889]" degrees="96" direction="west" minutes="18" orientation="longitude" pageId="6" pageNumber="559" precision="1" seconds="15.31" value="-96.30425">96°18ʹ15.31″W</geoCoordinate>
|
||
;
|
||
<figureCitation id="131A75CCFFEF64257016F8D3FA78F8C3" box="[1374,1430,1865,1889]" captionStart="Figure 3" captionStartId="4.[130,195,810,834]" captionTargetId="figure-644@4.[321,1281,144,782]" captionTargetPageId="4" captionText="Figure 3. Distribution of Xenomorphon baranowskii in Mexico." figureDoi="http://doi.org/10.5281/zenodo.8432524" httpUri="https://zenodo.org/record/8432524/files/figure.png" pageId="6" pageNumber="559">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEF64257671F81FFA42F879" blockId="6.[825,1474,1925,2011]" pageId="6" pageNumber="559">
|
||
<emphasis id="B955B55BFFEF64257671F81FFB9FF83F" box="[825,1137,1925,1949]" italics="true" pageId="6" pageNumber="559">Biology, immatures and females.</emphasis>
|
||
Unknown. The only known specimen was collected in the leaf liưer of a pine–oak forest floor, and the female is presumed to be paedomorphic (see Discussion).
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEE64247539FF0AFDAAFF05" blockId="7.[113,580,144,168]" box="[113,580,144,168]" pageId="7" pageNumber="560">
|
||
<emphasis id="B955B55BFFEE64247539FF0AFE05FF0A" box="[113,491,144,168]" italics="true" pageId="7" pageNumber="560">Length (head through tip of abdomen).</emphasis>
|
||
<quantity id="4CD9C4ACFFEE642474A7FF0AFDAEFF05" box="[495,576,144,168]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="560" unit="mm" value="2.9">2.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E6949FFEE64247539FF57FE57FF46" blockId="7.[113,441,205,229]" box="[113,441,205,229]" pageId="7" pageNumber="560">
|
||
<emphasis id="B955B55BFFEE64247539FF57FE8FFF47" box="[113,353,205,229]" italics="true" pageId="7" pageNumber="560">Width (across humerus).</emphasis>
|
||
<quantity id="4CD9C4ACFFEE6424742DFF57FE58FF46" box="[357,438,205,229]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="560" unit="mm" value="0.5">0.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |