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<document id="62659841F29ECF7496033493BD9BF5C8" ID-CLB-Dataset="6085" ID-DOI="10.11646/phytotaxa.31.1.1" ID-GBIF-Dataset="2185a8cf-c15c-448b-b8c0-6055976408ed" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894648" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622649568931" checkinUser="felipe" docAuthor="Gonzales, Jasivia &amp; Kessler, Michael" docDate="2011" docId="0380F37CFFEA7F0399A6E7FED6A7F921" docLanguage="en" docName="Phytotaxa.31.1-54.pdf.imf" docOrigin="Phytotaxa 31" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Sticherus tomentosus Smith 1990" docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="FFB98B04FFC57F2C9931E37FD712FFAA" masterDocTitle="A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="48" updateTime="1698998516251" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="72CA29305362AEEE3D05BDA542A51707">A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species</mods:title>
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<mods:namePart id="5B0AC5701F3747942B92E29C0A57A761">Gonzales, Jasivia</mods:namePart>
<mods:affiliation id="000B5A0019ABB1F6A34FFC506EEF7A15">Herbario Nacional de Bolivia, Casilla 10077, La Paz, Bolivia. Systematic Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland. E-mail: michael. kessler @ systbot. uzh. ch</mods:affiliation>
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<mods:title id="E393012243320EB6B44A1E242D2FC110">Phytotaxa</mods:title>
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<heading id="D0DEF506FFEA7F0399A6E7FED408FB31" box="[151,794,1153,1179]" fontSize="11" level="2" pageId="47" pageNumber="48" reason="3">
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<treatmentCitation id="0A88647BFFEA7F0399A6E7FED405FB31" author="Smith, A. R." box="[151,791,1153,1179]" page=")" pageId="47" pageNumber="48" year="1990">
<taxonomicName id="4C2939E9FFEA7F0399A6E7FED405FB31" ID-CoL="52HY3" authority="(Cav. ex Sw.) Smith (1990: 255)" authorityName="Smith" authorityPageNumber="255" authorityYear="1990" baseAuthorityName="Cav. ex Sw." box="[151,791,1153,1179]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="B95D9E78FFEA7F0399A6E7FED687FB31" bold="true" box="[151,405,1153,1179]" italics="true" pageId="47" pageNumber="48">Sticherus tomentosus</emphasis>
(Cav. ex Sw.)
<bibRefCitation id="EFB83F9BFFEA7F039B71E7FED405FB31" author="Smith, A. R." box="[576,791,1153,1179]" pageId="47" pageNumber="48" pagination="249 - 273" refId="ref33868" refString="Smith, A. R. (1990) Pteridophytes of the Venezuelan Guayana: new species. Annals of the Missouri Botanical Garden 77: 249 - 273." type="journal article" year="1990">Smith (1990: 255)</bibRefCitation>
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.
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</paragraph>
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<treatmentCitation id="0A88647BFFEA7F0399A6E7B0D5B7FB4F" author="Swartz, O." box="[151,677,1230,1254]" page=")" pageId="47" pageNumber="48" year="1804">
<taxonomicName id="4C2939E9FFEA7F0399A6E7B0D5B7FB4F" authority="Cav. ex Swartz (1804: 177)" authorityName="Swartz" authorityPageNumber="177" authorityYear="1804" box="[151,677,1230,1254]" class="Tentaculata" family="Mertensiidae" genus="Mertensia" kingdom="Animalia" order="Cydippida" pageId="47" pageNumber="48" phylum="Ctenophora" rank="species" species="tomentosa">
<emphasis id="B95D9E78FFEA7F0399A6E7B0D66AFB4C" box="[151,376,1231,1254]" italics="true" pageId="47" pageNumber="48">Mertensia tomentosa</emphasis>
Cav. ex
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</taxonomicName>
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.
<treatmentCitation id="0A88647BFFEA7F039B9EE7B0D21FFB4F" author="Sprengel, C. P. J." box="[687,1293,1230,1254]" page=")" pageId="47" pageNumber="48" year="1827">
<taxonomicName id="4C2939E9FFEA7F039B9EE7B0D21FFB4F" ID-CoL="6KGT7" authority="(Cav. ex Sw.) Sprengel (1827: 27)" authorityName="Sprengel" authorityPageNumber="27" authorityYear="1827" baseAuthorityName="Sw." box="[687,1293,1230,1254]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B95D9E78FFEA7F039B9EE7B0D48BFB4C" box="[687,921,1231,1254]" italics="true" pageId="47" pageNumber="48">Gleichenia tomentosa</emphasis>
(Cav. ex Sw.)
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.
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</paragraph>
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<emphasis id="B95D9E78FFEA7F0399F7E791D639FAA2" bold="true" box="[198,299,1262,1288]" pageId="47" pageNumber="48">Type:—</emphasis>
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.
<emphasis id="B95D9E78FFEA7F0398B5E790D576FAA2" box="[388,612,1263,1288]" italics="true" pageId="47" pageNumber="48">
<location id="8EF614B1FFEA7F0398B5E790D6E1FAA2" LSID="urn:lsid:plazi:treatment:0380F37CFFEA7F0399A6E7FED6A7F921:8EF614B1FFEA7F0398B5E790D6E1FAA2" box="[388,499,1263,1288]" country="Peru" name="Collector" pageId="47" pageNumber="48">Collector</location>
unknown
</emphasis>
(
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<collectionCode id="ED38DAAFFFEA7F039BD2E790D5E7FAA3" box="[739,757,1263,1289]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="47" pageNumber="48" type="Herbarium">S</collectionCode>
,
<emphasis id="B95D9E78FFEA7F039A33E790D4D1FAA2" box="[770,963,1263,1288]" italics="true" pageId="47" pageNumber="48">herb. Cavanilles</emphasis>
,
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<collectingCountry id="F33E02FAFFEA7F039AE3E790D4E5FAA3" box="[978,1015,1263,1289]" name="United States of America" pageId="47" pageNumber="48">US</collectingCountry>
photo).
</paragraph>
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<subSubSection id="C33311E1FFEA7F0399F7E66AD6A7F921" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8B96426AFFEA7F0399F7E66AD208FAD6" blockId="47.[151,1436,1230,1675]" pageId="47" pageNumber="48">
<emphasis id="B95D9E78FFEA7F0399F7E66AD53DFA85" bold="true" box="[198,559,1301,1327]" pageId="47" pageNumber="48">Distribution and ecology:—</emphasis>
The Guayana Highlands in southeastern
<collectingCountry id="F33E02FAFFEA7F039D1EE66AD3B5FA85" box="[1071,1191,1301,1327]" name="Venezuela" pageId="47" pageNumber="48">Venezuela</collectingCountry>
(to be expected in adjacent
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) and the Andes in northwestern
<collectingCountry id="F33E02FAFFEA7F039BCFE643D46DFAFC" box="[766,895,1340,1366]" name="Venezuela" pageId="47" pageNumber="48">Venezuela</collectingCountry>
,
<collectingCountry id="F33E02FAFFEA7F039ABCE643D31BFAFC" box="[909,1033,1340,1366]" name="Colombia" pageId="47" pageNumber="48">Colombia</collectingCountry>
,
<collectingCountry id="F33E02FAFFEA7F039D26E643D36DFAFC" box="[1047,1151,1340,1366]" name="Ecuador" pageId="47" pageNumber="48">Ecuador</collectingCountry>
, and
<collectingCountry id="F33E02FAFFEA7F039DF3E643D3ECFAFC" box="[1218,1278,1340,1366]" name="Peru" pageId="47" pageNumber="48">Peru</collectingCountry>
. Is is locally common along humid forest edges, along roads, on landslides, and along streams at
<quantity id="4CD1EF8FFFEA7F039D40E61DD204FAD6" box="[1137,1302,1378,1404]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="3.0" metricValueMin="1.7" pageId="47" pageNumber="48" unit="m" value="2350.0" valueMax="3000.0" valueMin="1700.0">17003000 m</quantity>
.
</paragraph>
<paragraph id="8B96426AFFEA7F0399F7E6F6D6A7F921" blockId="47.[151,1436,1230,1675]" pageId="47" pageNumber="48">
<emphasis id="B95D9E78FFEA7F0399F7E6F6D621FA09" bold="true" box="[198,307,1417,1443]" pageId="47" pageNumber="48">Notes:—</emphasis>
Morphologically, this species is at the center of a complex of seven closely related species which include:
<taxonomicName id="4C2939E9FFEA7F0399CEE6CFD6D3FA63" authorityName="Ching" authorityYear="1940" baseAuthorityName="Sodiro" box="[255,449,1456,1481]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="blepharolepis">
<emphasis id="B95D9E78FFEA7F0399CEE6CFD6D3FA63" box="[255,449,1456,1481]" italics="true" pageId="47" pageNumber="48">S. blepharolepis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2939E9FFEA7F0398E1E6CFD574FA63" authorityName="Copeland" authorityYear="1947" baseAuthorityName="Christ &amp; Rosenst." box="[464,614,1456,1481]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="buchtienii">
<emphasis id="B95D9E78FFEA7F0398E1E6CFD574FA63" box="[464,614,1456,1481]" italics="true" pageId="47" pageNumber="48">S. buchtienii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2939E9FFEA7F039B47E6CFD5C5FA63" authorityName="J. Gonzales" authorityYear="2011" box="[630,727,1456,1481]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="jacha">
<emphasis id="B95D9E78FFEA7F039B47E6CFD5C5FA63" box="[630,727,1456,1481]" italics="true" pageId="47" pageNumber="48">S. jacha</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2939E9FFEA7F039BD6E6CFD473FA63" authorityName="J. Gonzales" authorityYear="2011" baseAuthorityName="J. Gonzales" baseAuthorityYear="2011" box="[743,865,1456,1481]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="lanosus">
<emphasis id="B95D9E78FFEA7F039BD6E6CFD473FA63" box="[743,865,1456,1481]" italics="true" pageId="47" pageNumber="48">S. lanosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2939E9FFEA7F039A41E6CFD307FA63" authorityName="Smith" authorityYear="1990" baseAuthorityName="Cav. ex Sw." box="[880,1045,1456,1481]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="B95D9E78FFEA7F039A41E6CFD307FA63" box="[880,1045,1456,1481]" italics="true" pageId="47" pageNumber="48">S. tomentosus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2939E9FFEA7F039D69E6CFD3D9FA63" authorityName="Copeland" authorityYear="1947" baseAuthorityName="Kunze" box="[1112,1227,1456,1481]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="velatus">
<emphasis id="B95D9E78FFEA7F039D69E6CFD3D9FA63" box="[1112,1227,1456,1481]" italics="true" pageId="47" pageNumber="48">S. velatus</emphasis>
</taxonomicName>
. Traditionally all seven taxa have been considered to belong to a single, variable species (
<bibRefCitation id="EFB83F9BFFEA7F039AC1E6A8D3E2FA5B" author="Tryon, R. M. &amp; Stolze, R. G." box="[1008,1264,1495,1521]" pageId="47" pageNumber="48" pagination="1 - 122" refId="ref34267" refString="Tryon, R. M. &amp; Stolze, R. G. (1989) Pteridophyta of Peru. Part II. 13. Pteridaceae - 15. Dennstaedtiaceae. Fieldiana Botany, n. s. 22: 1 - 122." type="book chapter" year="1989">Tryon &amp; Stolze 1989</bibRefCitation>
,
<bibRefCitation id="EFB83F9BFFEA7F039DCDE6A8D641F9BD" author="Ostergaard A. &amp; Ollgaard, E." pageId="47" pageNumber="48" pagination="107 - 169" refId="ref33047" refString="Ostergaard A., Ollgaard, E. &amp; B. (2001) Gleicheniaceae. In Harling, G. &amp; Andersson, L. (eds.), Flora of Ecuador 66: 107 - 169." type="journal article" year="2001">Østergaard &amp; Øllgaard 2001</bibRefCitation>
). In their study of the genus in
<collectingCountry id="F33E02FAFFEA7F039BDBE682D447F9BD" box="[746,853,1533,1559]" name="Ecuador" pageId="47" pageNumber="48">Ecuador</collectingCountry>
,
<bibRefCitation id="EFB83F9BFFEA7F039A55E682D3F7F9BD" author="Ostergaard A. &amp; Ollgaard, E." box="[868,1253,1533,1559]" pageId="47" pageNumber="48" pagination="107 - 169" refId="ref33047" refString="Ostergaard A., Ollgaard, E. &amp; B. (2001) Gleicheniaceae. In Harling, G. &amp; Andersson, L. (eds.), Flora of Ecuador 66: 107 - 169." type="journal article" year="2001">Østergaard &amp; Øllgaard (2001)</bibRefCitation>
recognized
<taxonomicName id="4C2939E9FFEA7F039CB4E681D625F997" authorityName="Ching" authorityYear="1940" baseAuthorityName="Sodiro" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="blepharolepis">
<emphasis id="B95D9E78FFEA7F039CB4E681D625F997" italics="true" pageId="47" pageNumber="48">S. blepharolepis</emphasis>
</taxonomicName>
as a distinct species. A closer examination of the morphological variability of the complex over its entire geographical distribution has convinced us that there are seven distinct entities, which are treated here at the species level.
</paragraph>
</subSubSection>
</treatment>
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