310 lines
35 KiB
XML
310 lines
35 KiB
XML
<document id="1CC0E0F00FCA18A40DC8211BD22E7B45" ID-DOI="10.5281/zenodo.273398" ID-GBIF-Dataset="ae452b89-2798-4a2a-8bd9-963dead78ebc" ID-ISSN="11755326" ID-Zenodo-Dep="273398" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761708870" checkinUser="plazi" docAuthor="Howden, Henry F." docDate="2006" docId="0380EE1A1958FFEAFE94F8EA3AEB0691" docLanguage="en" docName="zt01137p052.pdf" docOrigin="Zootaxa 1137" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Neoathyreus julietae Howden, 2006, new species" docType="treatment" docVersion="7" lastPageNumber="41" masterDocId="FFB996621959FFEEFF9CFB0C3B2E025F" masterDocTitle="New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini)" masterLastPageNumber="52" masterPageNumber="37" pageNumber="38" updateTime="1698215286212" updateUser="plazi">
|
||
<mods:mods id="34E58A80D85657B630BDBD6254A6ABD0" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="82B40583A0B3426E534E14B49AE0C990">
|
||
<mods:title id="6BB862F4437E1F418EF8C7C3A11E2040">New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="B966FACCAA4D187C1CC6D795BEFF2462" type="personal">
|
||
<mods:role id="53836864808A1DF573920C2AEBD2BF60">
|
||
<mods:roleTerm id="AA9B329F090A40697E7D9CB24ED10B93">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E02B4675AE740C87F8E0D703CBF8422A">Howden, Henry F.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1357EAAB01F385AEAA2FC83BCD941B40">text</mods:typeOfResource>
|
||
<mods:relatedItem id="7A431BD91ED282D6518274D1C1CAE698" type="host">
|
||
<mods:titleInfo id="19734FF5D4F05A328DC60271A2A131CD">
|
||
<mods:title id="FBB879138F36FA84F4CA52686664E6E3">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C699BF19A67CCB30D8A7EF86654D6C99">
|
||
<mods:date id="6849D07AB47CEE79F036DEAB98E277DF">2006</mods:date>
|
||
<mods:detail id="6583DA5685D1CB8F3498D5791877EC4C" type="volume">
|
||
<mods:number id="FEE030827D111843CAFF96CB5DA45F82">1137</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FE9A6E549488F492BF38B8FAAACA91D0" unit="page">
|
||
<mods:start id="80B06146AF2A0C5DA8B6E60E64CF748C">37</mods:start>
|
||
<mods:end id="70E75EB594AD1DF51108BA5550BC24EA">52</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="CB2FBE5FC7F6B65C9802C2196CFC9851">journal article</mods:classification>
|
||
<mods:identifier id="E44E136662446E645F0EBB588D359575" type="DOI">10.5281/zenodo.273398</mods:identifier>
|
||
<mods:identifier id="21E465F040BB75FC3A612CBDF724FBAB" type="GBIF-Dataset">ae452b89-2798-4a2a-8bd9-963dead78ebc</mods:identifier>
|
||
<mods:identifier id="922D1FB51D47C56484079746FDEE484A" type="ISSN">11755326</mods:identifier>
|
||
<mods:identifier id="C50A96069EEEDE15320B044800B7C00D" type="Zenodo-Dep">273398</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0380EE1A1958FFEAFE94F8EA3AEB0691" ID-DOI="http://doi.org/10.5281/zenodo.6260964" ID-GBIF-Taxon="119345038" ID-Zenodo-Dep="6260964" LSID="urn:lsid:plazi:treatment:0380EE1A1958FFEAFE94F8EA3AEB0691" httpUri="http://treatment.plazi.org/id/0380EE1A1958FFEAFE94F8EA3AEB0691" lastPageId="4" lastPageNumber="41" pageId="1" pageNumber="38">
|
||
<subSubSection id="C3330C871958FFEFFE94F8EA3804065F" box="[264,810,998,1024]" pageId="1" pageNumber="38" type="nomenclature">
|
||
<paragraph id="8B965F0C1958FFEFFE94F8EA3804065F" blockId="1.[264,810,998,1024]" box="[264,810,998,1024]" pageId="1" pageNumber="38">
|
||
<heading id="D0DEE8601958FFEFFE94F8EA3804065F" bold="true" box="[264,810,998,1024]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
|
||
<emphasis id="B95D831E1958FFEFFE94F8EA3804065F" bold="true" box="[264,810,998,1024]" pageId="1" pageNumber="38">
|
||
<emphasis id="B95D831E1958FFEFFE94F8EA3927065F" bold="true" box="[264,521,998,1024]" italics="true" pageId="1" pageNumber="38">
|
||
<taxonomicName id="4C29248F1958FFEFFE94F8EA392D065F" box="[264,515,998,1024]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="julietae" status="sp. nov.">Neoathyreus julietae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A26E3E651958FFEFFD8CF8EA398E065F" box="[528,672,998,1024]" pageId="1" pageNumber="38" rank="species">new species</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="131243891958FFEFFD2CF8EA380F065F" box="[688,801,998,1024]" captionStart="FIGURES 1 – 6" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[294,1292,473,1717]" captionTargetId="figure@3.[292,1296,473,1722]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Neoathyreus spp. Figs. 1 – 3. N. julietae, n. sp.: 1, Head and pronotum; 2, Male genitalia, dorsal view; 3, Male genitalia, lateral view. Figs. 4 – 6, N. moraguesi, n. sp.: 4, Head and pronotum; 5, Male genitalia, dorsal view; 6, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273399/files/figure.png" pageId="1" pageNumber="38">Figs. 1–3</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3330C871958FFECFE94FF3D38C60446" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8B965F0C1958FFECFE94FF3D3AB50149" blockId="1.[264,1323,1073,1899]" lastBlockId="2.[264,1324,284,1881]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
|
||
<emphasis id="B95D831E1958FFEFFE94FF3D3AAF0614" bold="true" box="[264,385,1073,1099]" pageId="1" pageNumber="38">
|
||
<typeStatus id="5492E1AE1958FFEFFE94FF3D3A520614" box="[264,380,1073,1099]" pageId="1" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male, length
|
||
<quantity id="4CD1F2E91958FFEFFDB4FF3D39B80614" box="[552,662,1073,1099]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="1" pageNumber="38" unit="mm" value="10.9">10.9 mm</quantity>
|
||
, greatest width
|
||
<quantity id="4CD1F2E91958FFEFFCCEFF3D389E0614" box="[850,944,1073,1099]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="38" unit="mm" value="7.4">7.4 mm</quantity>
|
||
. Dorsally with labrum, clypeus, and lateral edges of pronotum near foveae reddish brown, remainder of pronotum black tinged with brown; elytra black; ventral surfaces varying from reddish brown to dark brown. Labrum coarsely rugose, particularly across basal third. Clypeus coarsely rugose, transverse carina defined by row of irregular transverse tubercles; anterior oblique carinae absent; posterior transverse clypeal carinae distinct on each side of median tubercle, tubercle only slightly higher and anterior to lateral tubercle on each side. Vertex very slightly depressed between eyes, surface coarsely, contiguously granulate; between granules surface with numerous erect, pale tan setae. Gena with outer margins elevated, outer anterior angle slightly acute, surface of gena granulate. Pronotum (
|
||
<figureCitation id="131243891958FFEFFBCBFE953FB507EC" box="[1111,1179,1433,1459]" captionStart="FIGURES 1 – 6" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[294,1292,473,1717]" captionTargetId="figure@3.[292,1296,473,1722]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Neoathyreus spp. Figs. 1 – 3. N. julietae, n. sp.: 1, Head and pronotum; 2, Male genitalia, dorsal view; 3, Male genitalia, lateral view. Figs. 4 – 6, N. moraguesi, n. sp.: 4, Head and pronotum; 5, Male genitalia, dorsal view; 6, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273399/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
|
||
) with small fossa on each side contiguous with anterior marginal bead posterior to eye; width of fossa approximately equal to width of adjacent bead; between fossae margin gradually elevated to midline, midline with indistinct tubercle on bead; tubercle extending posteriorly as low, narrow, glabrous ridge for approximately
|
||
<quantity id="4CD1F2E91958FFEFFD6AFD35387A040C" box="[758,852,1593,1619]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="38" unit="mm" value="0.7">0.7 mm</quantity>
|
||
. Pronotum with inner and outer carinae welldefined, inner carinae moderately thick with anterior ends separated by distance approximately equal to distance between outer edges of eyes; inner pronotal carinae from anterior ends almost straight, gradually converging, becoming parallel at posterior end of pronotal concavity, terminating
|
||
<quantity id="4CD1F2E91958FFEFFD0BFDD539D404AC" box="[663,762,1753,1779]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="38" unit="mm" value="0.5">0.5 mm</quantity>
|
||
before posterior margin. Pronotal concavity deepest medially between anterior ends of inner carinae, surface granulatepunctate anteriorly, becoming smooth in posterior fourth of concavity; setose in anterior third between granules and near inner edge of carinae. Outer pronotal carina on each side in posterior half
|
||
<quantity id="4CD1F2E9195BFFECFE31FA1039290369" box="[429,519,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="39" unit="mm" value="1.7">1.7 mm</quantity>
|
||
long, curved, slightly closer to pronotal margin than to inner carina. Lateral pronotal fovea on each side shallow, elongate, surface mostly smooth; pronotal marginal bead absent in indentation below fovea. Pronotal surface laterad to inner carinae closely, coarsely granulate, posteriorly granules contiguous; many long, erect, pale tan setae arising from bases of granules. Elytron with several vague strial indentations near base of disc, surface of disc coarsely, contiguously granulate, granules near base transversely oval, becoming elongate in apical third; numerous tan or black, almosterect setae arising between granules. Pygidium brown, apex broadly rounded. Metasternum between middle coxae slightly convex, anterior edge near midline obliquely angulate, midline represented by narrow, depressed line, surface on either side relatively uniformly setosepunctate. Foretibia (
|
||
<figureCitation id="13124389195BFFECFDDBF9A039AB0099" box="[583,645,684,710]" captionStart="FIGURES 1 – 6" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[294,1292,473,1717]" captionTargetId="figure@3.[292,1296,473,1722]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Neoathyreus spp. Figs. 1 – 3. N. julietae, n. sp.: 1, Head and pronotum; 2, Male genitalia, dorsal view; 3, Male genitalia, lateral view. Figs. 4 – 6, N. moraguesi, n. sp.: 4, Head and pronotum; 5, Male genitalia, dorsal view; 6, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273399/files/figure.png" pageId="2" pageNumber="39">Fig.1</figureCitation>
|
||
) with five teeth on outer edge. Genital capsule with apex of ventral lobe rounded, fringed with setae, flattened surface of lobe glabrous. Genitalia as in
|
||
<figureCitation id="13124389195BFFECFEBAF9F03ABB0149" box="[294,405,764,790]" captionStart="FIGURES 1 – 6" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[294,1292,473,1717]" captionTargetId="figure@3.[292,1296,473,1722]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Neoathyreus spp. Figs. 1 – 3. N. julietae, n. sp.: 1, Head and pronotum; 2, Male genitalia, dorsal view; 3, Male genitalia, lateral view. Figs. 4 – 6, N. moraguesi, n. sp.: 4, Head and pronotum; 5, Male genitalia, dorsal view; 6, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273399/files/figure.png" pageId="2" pageNumber="39">Figs. 2, 3</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B965F0C195BFFECFEA4F82839250659" blockId="2.[264,1324,284,1881]" pageId="2" pageNumber="39">
|
||
<emphasis id="B95D831E195BFFECFEA4F8283A870161" bold="true" box="[312,425,804,830]" pageId="2" pageNumber="39">Allotype.</emphasis>
|
||
Female, length
|
||
<quantity id="4CD1F2E9195BFFECFDF1F82839F40161" box="[621,730,804,830]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="39" unit="mm" value="10.5">10.5 mm</quantity>
|
||
, greatest width
|
||
<quantity id="4CD1F2E9195BFFECFC05F82838D70161" box="[921,1017,804,830]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="39" unit="mm" value="6.7">6.7 mm</quantity>
|
||
. Dorsally lacking reddish brown areas on head and pronotum, these areas black tinged with brown, elytra black. Similar to
|
||
<typeStatus id="5492E1AE195BFFECFE1AF8783AC201D1" box="[390,492,884,910]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
except as follows: clypeus with irregular transverse carina very close to apical margin; median tubercle on clypeus less developed, no higher than lateral tubercles; pronotal concavity reduced in depth, posterior smooth area no more than half length of that of male.
|
||
</paragraph>
|
||
<paragraph id="8B965F0C195BFFECFEA4FF1838DA07AE" blockId="2.[264,1324,284,1881]" pageId="2" pageNumber="39">
|
||
<emphasis id="B95D831E195BFFECFEA4FF183AE30671" bold="true" box="[312,461,1044,1070]" pageId="2" pageNumber="39">
|
||
<typeStatus id="5492E1AE195BFFECFEA4FF183A5B0671" box="[312,373,1044,1070]" pageId="2" pageNumber="39">Type</typeStatus>
|
||
Series.
|
||
</emphasis>
|
||
<typeStatus id="5492E1AE195BFFECFE48FF18396A0671" box="[468,580,1044,1070]" pageId="2" pageNumber="39" type="holotype">Holotype</typeStatus>
|
||
, male,
|
||
<collectingCountry id="F33E1F9C195BFFECFD0AFF1839C10671" box="[662,751,1044,1070]" name="Bolivia" pageId="2" pageNumber="39">Bolivia</collectingCountry>
|
||
, Dep. Sta Cruz, Buena Vista, 1922, R. C. Robert, Howden coll. ex A. Martínez coll. (
|
||
<collectionCode id="ED38C7C9195BFFECFD36FF3038290609" box="[682,775,1084,1110]" pageId="2" pageNumber="39">CMNC</collectionCode>
|
||
). Allotype, female,
|
||
<collectingCountry id="F33E1F9C195BFFECFC73FF303F660609" box="[1007,1096,1084,1110]" name="Bolivia" pageId="2" pageNumber="39">Bolivia</collectingCountry>
|
||
, D Sta. Cruz, Pica Ichilo, Buena Vista, Tacu, III.[19]51, Martínez, Howden coll. ex A. Martínez coll. (
|
||
<collectionCode id="ED38C7C9195BFFECFE8CFF803A4306F9" box="[272,365,1164,1190]" pageId="2" pageNumber="39">CMNC</collectionCode>
|
||
).
|
||
<typeStatus id="5492E1AE195BFFECFE1EFF803AD906F9" box="[386,503,1164,1190]" pageId="2" pageNumber="39" type="paratype">Paratypes</typeStatus>
|
||
,
|
||
<specimenCount id="9D2F9485195BFFECFD9FFF80394C06F9" box="[515,610,1164,1190]" pageId="2" pageNumber="39" type="male">8 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D2F9485195BFFECFDF3FF8039DA06F9" box="[623,756,1164,1190]" pageId="2" pageNumber="39" type="female">13 females</specimenCount>
|
||
.
|
||
<collectingCountry id="F33E1F9C195BFFECFC9EFF80387406F9" box="[770,858,1164,1190]" name="Bolivia" pageId="2" pageNumber="39">
|
||
<emphasis id="B95D831E195BFFECFC9EFF80387406F9" bold="true" box="[770,858,1164,1190]" pageId="2" pageNumber="39">Bolivia</emphasis>
|
||
</collectingCountry>
|
||
: Cochabamba,
|
||
<specimenCount id="9D2F9485195BFFECFB88FF803F5C06F9" box="[1044,1138,1164,1190]" pageId="2" pageNumber="39" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D2F9485195BFFECFB1CFF803FC406F9" box="[1152,1258,1164,1190]" pageId="2" pageNumber="39" type="female">1 female</specimenCount>
|
||
,
|
||
<quantity id="4CD1F2E9195BFFECFB64FF803A03068F" metricMagnitude="4" metricUnit="m" metricValue="6.75" pageId="2" pageNumber="39" unit="km" value="67.5">67.5 km</quantity>
|
||
E Villa Tunari, Est. Biol. Valle Sajta, Univ. San Simon,
|
||
<quantity id="4CD1F2E9195BFFECFC4DFFBA3F0E068F" box="[977,1056,1206,1232]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="39" unit="m" value="300.0">300 m</quantity>
|
||
, 17 0 6' 19"
|
||
<geoCoordinate id="EE1D39CB195BFFECFB21FFBA3E05068F" box="[1213,1323,1204,1232]" direction="south" orientation="latitude" pageId="2" pageNumber="39" precision="925" value="-64.76667">S 64 46'</geoCoordinate>
|
||
57" W,
|
||
<date id="FF9779CC195BFFECFEFFFFD33AD306A6" box="[355,509,1247,1273]" pageId="2" pageNumber="39" value="1999-02-09" valueMax="1999-02-13" valueMin="1999-02-09">9–13.II.1999</date>
|
||
, F. Génier, lowland rain forest, ex FIT, 99068,9; Dpto. Sta. Cruz:
|
||
<specimenCount id="9D2F9485195BFFECFA87FFD33A61077D" pageId="2" pageNumber="39" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D2F9485195BFFECFEC0FE043AFF077D" box="[348,465,1288,1314]" pageId="2" pageNumber="39" type="female">9 females</specimenCount>
|
||
,
|
||
<quantity id="4CD1F2E9195BFFECFE42FE043934077D" box="[478,538,1288,1314]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="39" unit="km" value="5.0">5 km</quantity>
|
||
SSE Buena Vista, Hotel Flora y Fauna,
|
||
<quantity id="4CD1F2E9195BFFECFC60FE043F65077D" box="[1020,1099,1288,1314]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="39" unit="m" value="440.0">440 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1D39CB195BFFECFBCBFE043FD7077D" box="[1111,1273,1286,1314]" direction="south" orientation="latitude" pageId="2" pageNumber="39" precision="1" value="-17.49875">17 29.925' S</geoCoordinate>
|
||
<geoCoordinate id="EE1D39CB195BFFECFA9EFE043AAD0712" direction="west" orientation="longitude" pageId="2" pageNumber="39" precision="1" value="-63.652134">63 39.128' W</geoCoordinate>
|
||
, 6–15, 15–24,
|
||
<date id="FF9779CC195BFFECFDACFE3F39C10712" box="[560,751,1331,1357]" pageId="2" pageNumber="39" value="2003-12-24" valueMax="2003-12-31" valueMin="2003-12-24">24–31.XII.2003</date>
|
||
, S. & J. Peck, forest FIT;
|
||
<specimenCount id="9D2F9485195BFFECFB83FE3F3FBD0712" box="[1055,1171,1331,1357]" pageId="2" pageNumber="39" type="female">2 females</specimenCount>
|
||
,
|
||
<quantity id="4CD1F2E9195BFFECFB03FE3F3FDF0712" box="[1183,1265,1331,1357]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="39" unit="km" value="3.7">3.7 km</quantity>
|
||
SSE Buena Vista, Hotel Flora y Fauna,
|
||
<geoCoordinate id="EE1D39CB195BFFECFD2CFE52380C0727" box="[688,802,1372,1400]" direction="south" orientation="latitude" pageId="2" pageNumber="39" precision="925" value="-17.483334">17 29' S</geoCoordinate>
|
||
<geoCoordinate id="EE1D39CB195BFFECFCB0FE5238860727" box="[812,936,1372,1400]" direction="west" orientation="longitude" pageId="2" pageNumber="39" precision="925" value="-63.55">63 33' W</geoCoordinate>
|
||
,
|
||
<date id="FF9779CC195BFFECFC2BFE523F1D0727" box="[951,1075,1374,1400]" pageId="2" pageNumber="39">28.IV–2.V</date>
|
||
,
|
||
<date id="FF9779CC195BFFECFBDEFE523FF50727" box="[1090,1243,1374,1400]" pageId="2" pageNumber="39" value="2004-05-07" valueMax="2004-05-12" valueMin="2004-05-07">7–12.V.2004</date>
|
||
. A. R. Cline, FIT;
|
||
<specimenCount id="9D2F9485195BFFECFE3BFE8B392B07FE" box="[423,517,1415,1441]" pageId="2" pageNumber="39" type="male">1 male</specimenCount>
|
||
, Ichilo, P[arque] N[acional] A[mboró] (Saguayo),
|
||
<quantity id="4CD1F2E9195BFFECFB01FE8B3FD807FE" box="[1181,1270,1415,1441]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="39" unit="m" value="500.0">500 m</quantity>
|
||
,
|
||
<date id="FF9779CC195BFFECFA91FE8B3AA10796" pageId="2" pageNumber="39" value="1991-03-08" valueMax="1991-03-12" valueMin="1991-03-08">8– 12.III.1991</date>
|
||
, Pablo Bettella;
|
||
<specimenCount id="9D2F9485195BFFECFDC9FEA339EE0796" box="[597,704,1455,1481]" pageId="2" pageNumber="39" type="female">1 female</specimenCount>
|
||
,
|
||
<collectingCountry id="F33E1F9C195BFFECFD51FEA338640796" box="[717,842,1455,1481]" name="Saint Pierre and Miquelon" pageId="2" pageNumber="39">San Pedro</collectingCountry>
|
||
,
|
||
<date id="FF9779CC195BFFECFCC4FEA338CE0796" box="[856,992,1455,1481]" pageId="2" pageNumber="39" value="1990-06-26">26.VI.1990</date>
|
||
, C. J. Pruett.
|
||
<collectingCountry id="F33E1F9C195BFFECFB18FEA23FEE0797" box="[1156,1216,1454,1480]" name="Peru" pageId="2" pageNumber="39">
|
||
<emphasis id="B95D831E195BFFECFB18FEA23FEE0797" bold="true" box="[1156,1216,1454,1480]" pageId="2" pageNumber="39">Peru</emphasis>
|
||
</collectingCountry>
|
||
:
|
||
<specimenCount id="9D2F9485195BFFECFB4EFEA33E080796" box="[1234,1318,1455,1481]" pageId="2" pageNumber="39" type="male">1 male</specimenCount>
|
||
, near Satipo, Junin,>
|
||
<quantity id="4CD1F2E9195BFFECFE65FEDB397D07AE" box="[505,595,1495,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="39" unit="m" value="1000.0">1000 m</quantity>
|
||
(Yungas),
|
||
<date id="FF9779CC195BFFECFD4EFEDB386E07AE" box="[722,832,1495,1521]" pageId="2" pageNumber="39" value="2004-12">XII.2004</date>
|
||
, flying at dusk.
|
||
</paragraph>
|
||
<paragraph id="8B965F0C195BFFECFEA4FEF338C60446" blockId="2.[264,1324,284,1881]" box="[312,1000,1535,1561]" pageId="2" pageNumber="39">
|
||
<typeStatus id="5492E1AE195BFFECFEA4FEF33A840446" box="[312,426,1535,1561]" pageId="2" pageNumber="39" type="paratype">Paratypes</typeStatus>
|
||
are in:
|
||
<collectionCode id="ED38C7C9195BFFECFD9EFEF3397E0446" box="[514,592,1535,1561]" pageId="2" pageNumber="39">ABTS</collectionCode>
|
||
,
|
||
<collectionCode id="ED38C7C9195BFFECFDC7FEF3399A0446" box="[603,692,1535,1561]" pageId="2" pageNumber="39">CMNC</collectionCode>
|
||
,
|
||
<collectionCode id="ED38C7C9195BFFECFD5CFEF338380446" box="[704,790,1535,1561]" pageId="2" pageNumber="39">HAHC</collectionCode>
|
||
,
|
||
<collectionCode id="ED38C7C9195BFFECFCBEFEF338540446" box="[802,890,1535,1561]" httpUri="http://grbio.org/cool/zsd4-j9gb" name="National Kweiyang Medical College" pageId="2" pageNumber="39">NKMC</collectionCode>
|
||
<collectionCode id="ED38C7C9195BFFECFC1EFEF338CA0446" box="[898,996,1535,1561]" pageId="2" pageNumber="39">WBWC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3330C87195BFFEAFEA4FD2A39380671" lastPageId="4" lastPageNumber="41" pageId="2" pageNumber="39" type="discussion">
|
||
<paragraph id="8B965F0C195BFFEDFEA4FD2A3F5403F1" blockId="2.[264,1324,284,1881]" lastBlockId="3.[264,1324,284,430]" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
|
||
<emphasis id="B95D831E195BFFECFEA4FD2A3A9F041F" bold="true" box="[312,433,1574,1600]" pageId="2" pageNumber="39">Remarks.</emphasis>
|
||
Variation in the
|
||
<typeStatus id="5492E1AE195BFFECFDEEFD2B398A041E" box="[626,676,1575,1601]" pageId="2" pageNumber="39">type</typeStatus>
|
||
series is moderate. Length varies from 9.5 to 11.0 mm, and greatest width from 5.5 to 7.0 mm. The transverse anterior clypeal carina may be vague or distinct; the posterior clypeal carina and associated tubercles are reduced in height in small specimens. The inner pronotal carina and concavity are both reduced in small specimens; the height and thickness of the carinae are less and the depth and posterior smooth area of the concavity are similarly reduced; in one specimen the smooth area is almost absent. In one specimen the lateral marginal bead of the pronotum is complete. The reddish brown color on the head and sides of the pronotum is lacking in more than half of the series, these being brownish black to black. The number of teeth, five, on the foretibia is quite constant, but there are six teeth in one female. There are also slight differences in the shape of the apical lobes of the male genitalia but this is, at least partly, because of distortion when dry or differences caused by positioning.
|
||
</paragraph>
|
||
<caption id="DF560F84195AFFEDFE94FDCC3A4F051F" httpUri="https://zenodo.org/record/273399/files/figure.png" pageId="3" pageNumber="40" targetBox="[294,1292,473,1717]" targetPageId="3">
|
||
<paragraph id="8B965F0C195AFFEDFE94FDCC3A4F051F" blockId="3.[264,1323,1728,1856]" pageId="3" pageNumber="40">
|
||
<emphasis id="B95D831E195AFFEDFE94FDCC3A980487" bold="true" box="[264,438,1728,1752]" pageId="3" pageNumber="40">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C29248F195AFFEDFE22FDCD396A0487" box="[446,580,1729,1752]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95D831E195AFFEDFE22FDCD396A0487" box="[446,580,1729,1752]" italics="true" pageId="3" pageNumber="40">Neoathyreus</emphasis>
|
||
</taxonomicName>
|
||
spp. Figs. 1–3.
|
||
<emphasis id="B95D831E195AFFEDFC9BFDCD38510487" box="[775,895,1729,1752]" italics="true" pageId="3" pageNumber="40">
|
||
<taxonomicName id="4C29248F195AFFEDFC9BFDCD38550487" box="[775,891,1729,1752]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="julietae" status="sp. nov.">N. julietae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A26E3E65195AFFEDFC14FDCD38E80487" box="[904,966,1729,1752]" pageId="3" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
: 1, Head and pronotum; 2, Male genitalia, dorsal view; 3, Male genitalia, lateral view. Figs. 4–6,
|
||
<emphasis id="B95D831E195AFFEDFC5FFDEF3F7704A5" box="[963,1113,1763,1786]" italics="true" pageId="3" pageNumber="40">
|
||
<taxonomicName id="4C29248F195AFFEDFC5FFDEF3F7B04A5" box="[963,1109,1763,1786]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="moraguesi" status="sp. nov.">N. moraguesi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A26E3E65195AFFEDFBFDFDEF3FB004A5" box="[1121,1182,1763,1786]" pageId="3" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
: 4, Head and pronotum; 5, Male genitalia, dorsal view; 6, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B965F0C195DFFEAFEA4FA103F6603A1" blockId="4.[264,1323,284,1230]" pageId="4" pageNumber="41">
|
||
<taxonomicName id="4C29248F195DFFEAFEA4FA103905036A" box="[312,555,284,309]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="julietae">
|
||
<emphasis id="B95D831E195DFFEAFEA4FA103905036A" box="[312,555,284,309]" italics="true" pageId="4" pageNumber="41">Neoathyreus julietae</emphasis>
|
||
</taxonomicName>
|
||
will usually key to couplet
|
||
<quantity id="4CD1F2E9195DFFEAFCE8FA10389E0369" box="[884,944,284,310]" metricMagnitude="0" metricUnit="m" metricValue="1.1176" pageId="4" pageNumber="41" unit="in" value="44.0">44 in</quantity>
|
||
<bibRefCitation id="EFB822FD195DFFEAFC24FA103F5B0369" author="Howden" box="[952,1141,284,310]" pageId="4" pageNumber="52" refString="Howden, H. F. (1985) A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae). Contributions of the American Entomological Institute, 21, 1 - 95." type="journal article" year="1985">Howden (1985)</bibRefCitation>
|
||
. In that couplet most specimens would key to
|
||
<taxonomicName id="4C29248F195DFFEAFDEEFA4838730301" authority="Howden" authorityName="Howden" box="[626,861,324,350]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="B95D831E195DFFEAFDEEFA4839C30302" box="[626,749,324,349]" italics="true" pageId="4" pageNumber="41">N. inermis</emphasis>
|
||
Howden
|
||
</taxonomicName>
|
||
, but differ from that species by having fewer teeth on the foretibia, 7 or 8 for
|
||
<taxonomicName id="4C29248F195DFFEAFD75FA60384803DA" box="[745,870,364,389]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="B95D831E195DFFEAFD75FA60384803DA" box="[745,870,364,389]" italics="true" pageId="4" pageNumber="41">N. inermis</emphasis>
|
||
</taxonomicName>
|
||
; also the male genitalia are radically different. The other species in couplet 44,
|
||
<taxonomicName id="4C29248F195DFFEAFD60FA983F2803F1" authority="Klug" authorityName="Klug" box="[764,1030,404,430]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="lanuginosus">
|
||
<emphasis id="B95D831E195DFFEAFD60FA98388303F2" box="[764,941,404,429]" italics="true" pageId="4" pageNumber="41">N. lanuginosus</emphasis>
|
||
(Klug)
|
||
</taxonomicName>
|
||
, is close to
|
||
<taxonomicName id="4C29248F195DFFEAFB0DFA983E2203F2" box="[1169,1292,404,429]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="julietae">
|
||
<emphasis id="B95D831E195DFFEAFB0DFA983E2203F2" box="[1169,1292,404,429]" italics="true" pageId="4" pageNumber="41">N. julietae</emphasis>
|
||
</taxonomicName>
|
||
in key characters, but differs in dorsal color and in the shape of the male genitalia. Both
|
||
<taxonomicName id="4C29248F195DFFEAFA8CFAB03AB803A2" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="lanuginosus">
|
||
<emphasis id="B95D831E195DFFEAFA8CFAB03AB803A2" italics="true" pageId="4" pageNumber="41">N. lanuginosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C29248F195DFFEAFE53FAE8396703A2" box="[463,585,484,509]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="julietae">
|
||
<emphasis id="B95D831E195DFFEAFE53FAE8396703A2" box="[463,585,484,509]" italics="true" pageId="4" pageNumber="41">N. julietae</emphasis>
|
||
</taxonomicName>
|
||
occur in the Buena Vista region of
|
||
<collectingCountry id="F33E1F9C195DFFEAFC75FAE83F6D03A1" box="[1001,1091,484,510]" name="Bolivia" pageId="4" pageNumber="41">Bolivia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B965F0C195DFFEAFEA4F90039380671" blockId="4.[264,1323,284,1230]" pageId="4" pageNumber="41">
|
||
While the two species that key out to couplet
|
||
<date id="FF9779CC195DFFEAFCE0F90038F60079" box="[892,984,524,550]" pageId="4" pageNumber="41" value="1944-05">44 may</date>
|
||
be somewhat related to
|
||
<emphasis id="B95D831E195DFFEAFA8CF9003A490012" italics="true" pageId="4" pageNumber="41">
|
||
<taxonomicName id="4C29248F195DFFEAFA8CF9003A4D0012" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="julietae">N. julietae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the most closely related species is probably
|
||
<taxonomicName id="4C29248F195DFFEAFCE8F9383F430011" authority="Howden" authorityName="Howden" box="[884,1133,564,590]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B95D831E195DFFEAFCE8F9383F2E0012" box="[884,1024,564,589]" italics="true" pageId="4" pageNumber="41">N. obscurus</emphasis>
|
||
Howden
|
||
</taxonomicName>
|
||
, described from four specimens from
|
||
<collectingCountry id="F33E1F9C195DFFEAFD92F95039770029" box="[526,601,604,630]" name="Brazil" pageId="4" pageNumber="41">Brazil</collectingCountry>
|
||
. In my 1985 key,
|
||
<taxonomicName id="4C29248F195DFFEAFCDCF95038FF002A" box="[832,977,604,629]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B95D831E195DFFEAFCDCF95038FF002A" box="[832,977,604,629]" italics="true" pageId="4" pageNumber="41">N. obscurus</emphasis>
|
||
</taxonomicName>
|
||
will key to couplet 30; this includes species with four teeth on the foretibia. This species should also have been keyed to species with five teeth on the foretibia as one
|
||
<typeStatus id="5492E1AE195DFFEAFCCCF9A0389B0099" box="[848,949,684,710]" pageId="4" pageNumber="41" type="paratype">paratype</typeStatus>
|
||
has a very small fifth tooth. A small series of
|
||
<emphasis id="B95D831E195DFFEAFE5EF9D8397700B2" box="[450,601,724,749]" italics="true" pageId="4" pageNumber="41">
|
||
<taxonomicName id="4C29248F195DFFEAFE5EF9D8397A00B2" box="[450,596,724,749]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="obscurus">N. obscurus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
recently collected in Mato Grosso,
|
||
<collectingCountry id="F33E1F9C195DFFEAFB90F9D83F7800B1" box="[1036,1110,724,750]" name="Brazil" pageId="4" pageNumber="41">Brazil</collectingCountry>
|
||
, by F. Génier, all have a small fifth tooth on the foretibia, as do most
|
||
<emphasis id="B95D831E195DFFEAFCF7F9F038C3014A" box="[875,1005,764,789]" italics="true" pageId="4" pageNumber="41">
|
||
<taxonomicName id="4C29248F195DFFEAFCF7F9F038C4014A" box="[875,1002,764,789]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="julietae">N. julietae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Externally the two species are not always separable because of variation in the teeth of the foretibia and in the marginal pronotal bead. The bead is usually complete in
|
||
<taxonomicName id="4C29248F195DFFEAFC5EF8403F7C013A" box="[962,1106,844,869]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B95D831E195DFFEAFC5EF8403F7C013A" box="[962,1106,844,869]" italics="true" pageId="4" pageNumber="41">N. obscurus</emphasis>
|
||
</taxonomicName>
|
||
and broken in
|
||
<taxonomicName id="4C29248F195DFFEAFA8CF8403A4E01D2" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="julietae">
|
||
<emphasis id="B95D831E195DFFEAFA8CF8403A4E01D2" italics="true" pageId="4" pageNumber="41">N. julietae</emphasis>
|
||
</taxonomicName>
|
||
; however, in the series at hand, the reverse is true for one specimen of each species. The only certain way to distinguish the two species is by the differences in the male genitalia and, as far as is known, by the different ranges of the two species. This complex is a good example of the desirability of having a good series when dealing with closely related species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3330C87195DFFEAFEA4FF303AEB0691" pageId="4" pageNumber="41" type="etymology">
|
||
<paragraph id="8B965F0C195DFFEAFEA4FF303AEB0691" blockId="4.[264,1323,284,1230]" pageId="4" pageNumber="41">
|
||
<emphasis id="B95D831E195DFFEAFEA4FF303AEA0609" bold="true" box="[312,452,1084,1110]" pageId="4" pageNumber="41">Etymology.</emphasis>
|
||
This species,
|
||
<emphasis id="B95D831E195DFFEAFD10FF303836060A" box="[652,792,1084,1109]" italics="true" pageId="4" pageNumber="41">
|
||
<taxonomicName id="4C29248F195DFFEAFD10FF30383A060A" box="[652,788,1084,1109]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="julietae">N. julietae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
is named for Julieta Ledezma, Curator, Entomology, NKMC, Santa Cruz,
|
||
<collectingCountry id="F33E1F9C195DFFEAFD03FF6839D60621" box="[671,760,1124,1150]" name="Bolivia" pageId="4" pageNumber="41">Bolivia</collectingCountry>
|
||
, who was extremely helpful to my wife and me during our visit to
|
||
<collectingCountry id="F33E1F9C195DFFEAFE7BFF80396F06F9" box="[487,577,1164,1190]" name="Bolivia" pageId="4" pageNumber="41">Bolivia</collectingCountry>
|
||
. She has shown the same consideration to other entomologists visiting
|
||
<collectingCountry id="F33E1F9C195DFFEAFEFBFFB83AEF0691" box="[359,449,1204,1230]" name="Bolivia" pageId="4" pageNumber="41">Bolivia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |