treatments-xml/data/E0/65/87/E06587DF5C67FFEEFCA2637CFC263695.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.5658661" ID-GBIF-Dataset="02f2e34f-46b2-40e9-a287-850d7cdf7300" ID-GBIF-Taxon="158486009" ID-Zenodo-Dep="5658661" ID-ZooBank="urn:lsid:zoobank.org:pub:936DFF93-AAA3-460F-AD5D-8EC342CB3132" checkinTime="1562016359349" checkinUser="plazi" docAuthor="Li, Xuankun &amp; Yeates, David K" docDate="2019" docId="E06587DF5C67FFEEFCA2637CFC263695" docLanguage="en" docName="australEntomology.58.192-203.pdf" docOrigin="Austral Entomology 58" docStyle="DocumentStyle{}" docTitle="Paramonovius nightking Li &amp; Yeates 2019, sp. nov" docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="99" masterDocId="1C5CFFA75C63FFE9FF8C6466FFF03011" masterDocTitle="A new genus and species of an unusual Australian winter bee fly (Diptera: Bombyliidae) with discussion on its phylogenetic position" masterLastPageNumber="203" masterPageNumber="92" pageId="4" pageNumber="96" updateTime="1636473863582" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new genus and species of an unusual Australian winter bee fly (Diptera: Bombyliidae) with discussion on its phylogenetic position</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Xuankun</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yeates, David K</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Austral Entomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-02-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>58</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>92</mods:start>
<mods:end>203</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1111/aen.12361</mods:identifier>
<mods:identifier type="GBIF-Dataset">02f2e34f-46b2-40e9-a287-850d7cdf7300</mods:identifier>
<mods:identifier type="Zenodo-Dep">3265414</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:936DFF93-AAA3-460F-AD5D-8EC342CB3132</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5658661" ID-GBIF-Taxon="158486009" ID-Zenodo-Dep="5658661" LSID="urn:lsid:plazi:treatment:E06587DF5C67FFEEFCA2637CFC263695" httpUri="http://treatment.plazi.org/id/E06587DF5C67FFEEFCA2637CFC263695" lastPageId="7" lastPageNumber="99" pageId="4" pageNumber="96">
<subSubSection box="[814,1198,1818,1842]" pageId="4" pageNumber="96" type="nomenclature">
<paragraph blockId="4.[814,1198,1818,1842]" box="[814,1198,1818,1842]" pageId="4" pageNumber="96">
<emphasis bold="true" box="[814,1198,1818,1842]" pageId="4" pageNumber="96">
<taxonomicName authority="Li &amp; Yeates, 2019" authorityName="Li &amp; Yeates" authorityYear="2019" box="[814,1106,1818,1842]" class="Insecta" family="Bombyliidae" genus="Paramonovius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="96" phylum="Arthropoda" rank="species" species="nightking" status="sp. nov">Paramonovius nightking</taxonomicName>
<taxonomicNameLabel box="[1113,1198,1818,1842]" pageId="4" pageNumber="96" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="96" type="description">
<paragraph blockId="4.[814,1372,1867,1953]" pageId="4" pageNumber="96">
(
<figureCitation box="[821,909,1867,1890]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="4.[145,181,1556,1577]" captionStartId-1="5.[144,180,1166,1187]" captionStartId-2="6.[145,181,1787,1808]" captionTargetBox-0="[341,1247,131,1524]" captionTargetBox-1="[340,1246,113,1133]" captionTargetBox-2="[393,1205,132,1745]" captionTargetId-0="figure@4.[340,1248,129,1524]" captionTargetId-1="figure@5.[339,1247,127,1134]" captionTargetId-2="figure@6.[382,1207,127,1756]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Fig. 2. Paramonovius nightking sp. nov. ♂: (a) dorsal; (b) lateral; (c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral; (g) head, frontal; (h) head, profile; (i) hind tarsi and claw. Scale bars = 1 mm (a, b, di); = 0.1 mm (c)." captionText-1="Fig. 3. Paramonovius nightking sp. nov. ♀: (a) dorsal; (b) lateral; (c) head, dorsal; (d) head, lateral; (e) head, frontal; (f) head, profile; (g) hind tarsi and claw. Scale bars = 1 mm." captionText-2="Fig. 4. Paramonovius nightking sp. nov. Genitalia (ae (♂); f, g (♀)): (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral; (d) epandrium, dorsal; (e) sternite 8, ventral; (f) genitalia and spermathecae. Scale bars = 0.1 mm. Abbreviations: acant spi = acanthophorite spine; aed sh = aedeagal sheath; b spmth dt = basal spermathecal duct; cerc = cercus; d brg = dorsal bridge; d spmth dt = distal spermathecal duct; ej apod = ejaculatory apodeme; epand = epandrium; gen fk = genital fork; goncx = gonocoxite; goncx apod = gonocoxalapodeme; gonst = gonostylus; lat ej proc = lateral ejaculatory process; ph = phallus; spmth = spermatheca; spm pmp = sperm pump; tg 9+10 = tergites 9+10." httpUri-0="https://zenodo.org/record/3265418/files/figure.png" httpUri-1="https://zenodo.org/record/3265420/files/figure.png" httpUri-2="https://zenodo.org/record/3265422/files/figure.png" pageId="4" pageNumber="96">Figs 24</figureCitation>
) http://zoobank.org/
<uri pageId="4" pageNumber="96">
urn:lsid:zoobank.org:pub:
<uuid pageId="4" pageNumber="96">936DFF93- AAA3-460F-AD5D-8EC342CB3132</uuid>
</uri>
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/3265420/files/figure.png" pageId="5" pageNumber="97" startId="5.[144,180,1166,1187]" targetBox="[340,1246,113,1133]" targetPageId="5">
<paragraph blockId="5.[144,1443,1165,1216]" pageId="5" pageNumber="97">
<emphasis box="[144,466,1166,1187]" italics="true" pageId="5" pageNumber="97">
<emphasis bold="true" box="[144,207,1166,1187]" italics="true" pageId="5" pageNumber="97">Fig. 3.</emphasis>
<taxonomicName authority="Li &amp; Yeates, 2019" authorityName="Li &amp; Yeates" authorityYear="2019" box="[236,466,1166,1187]" class="Insecta" family="Bombyliidae" genus="Paramonovius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="nightking" status="sp. nov.">Paramonovius nightking</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[472,547,1166,1187]" pageId="5" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
♀: (a) dorsal; (b) lateral; (c) head, dorsal; (d) head, lateral; (e) head, frontal; (f) head, profile; (g) hind tarsi and claw. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="97" type="materials_examined">
<paragraph blockId="5.[144,370,1273,1297]" box="[144,370,1273,1297]" pageId="5" pageNumber="97">
<heading bold="true" box="[144,370,1273,1297]" fontSize="10" level="3" pageId="5" pageNumber="97" reason="2">
<emphasis bold="true" box="[144,370,1273,1297]" pageId="5" pageNumber="97">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2273363824" collectedFrom="Scattered Banksia with low open shrubs on white sand. On Leucopogon oxycedrus" collectingDate="2012-05-19" collectionCode="WAM" collectorName="J. &amp; F. Hort" country="Australia" latitude="-31.96443" location="Kent Road" longLatPrecision="1" longitude="116.51224" municipality="Wandoo National Park" pageId="5" pageNumber="97" specimenCode="E97796" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="holotype">
<paragraph blockId="5.[144,231,1339,1360]" box="[144,231,1339,1360]" pageId="5" pageNumber="97">
<heading bold="true" box="[144,231,1339,1360]" fontSize="9" level="5" pageId="5" pageNumber="97" reason="2">
<emphasis bold="true" box="[144,231,1339,1360]" italics="true" pageId="5" pageNumber="97">
<typeStatus box="[144,231,1339,1360]" pageId="5" pageNumber="97">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[144,774,1386,1504]" pageId="5" pageNumber="97">
<specimenCount box="[144,163,1386,1409]" pageId="5" pageNumber="97" type="male"></specimenCount>
<collectingCountry box="[175,315,1386,1408]" name="Australia" pageId="5" pageNumber="97">AUSTRALIA</collectingCountry>
<collectingRegion box="[328,513,1386,1408]" country="Australia" name="Western Australia" pageId="5" pageNumber="97">Western Australia</collectingRegion>
,
<collectingMunicipality box="[529,770,1386,1408]" pageId="5" pageNumber="97">Wandoo National Park</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E06587DF5C67FFEEFCA2637CFC263695:6D1360125C66FFECFF1C61ECFF0D35B1" box="[144,253,1418,1440]" country="Australia" latitude="-31.96443" longLatPrecision="1" longitude="116.51224" municipality="Wandoo National Park" name="Kent Road" pageId="5" pageNumber="97" stateProvince="Western Australia">Kent Road</location>
,
<location LSID="urn:lsid:plazi:treatment:E06587DF5C67FFEEFCA2637CFC263695:6D1360125C66FFECFE8A61ECFECC35B1" box="[262,316,1418,1440]" country="Australia" latitude="-31.96443" longLatPrecision="1" longitude="116.51224" municipality="Wandoo National Park" name="York" pageId="5" pageNumber="97" stateProvince="Western Australia">York</location>
,
<geoCoordinate box="[324,426,1418,1440]" degrees="31.96443" direction="south" orientation="latitude" pageId="5" pageNumber="97" precision="1" value="-31.96443">-31.96443</geoCoordinate>
<geoCoordinate box="[431,539,1418,1440]" degrees="116.51224" direction="east" orientation="longitude" pageId="5" pageNumber="97" precision="1" value="116.51224">116.51224</geoCoordinate>
,
<collectedFrom pageId="5" pageNumber="97">
Scattered
<taxonomicName box="[644,724,1418,1440]" class="Magnoliopsida" family="Proteaceae" genus="Banksia" kingdom="Plantae" order="Proteales" pageId="5" pageNumber="97" phylum="Tracheophyta" rank="genus">Banksia</taxonomicName>
with low open shrubs on white sand. On
<taxonomicName box="[506,735,1450,1472]" class="Magnoliopsida" family="Ericaceae" genus="Leucopogon" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="97" phylum="Tracheophyta" rank="species" species="oxycedrus">
<emphasis box="[506,735,1450,1472]" italics="true" pageId="5" pageNumber="97">Leucopogon oxycedrus</emphasis>
</taxonomicName>
</collectedFrom>
,
<date pageId="5" pageNumber="97" value="2012-05-19">
<collectingDate pageId="5" pageNumber="97" value="2012-05-19">19 May 2012</collectingDate>
</date>
,
<collectorName box="[259,275,1482,1504]" pageId="5" pageNumber="97">J.</collectorName>
and
<collectorName box="[324,395,1482,1504]" pageId="5" pageNumber="97">F. Hort</collectorName>
(
<collectionCode box="[409,477,1482,1504]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="97" type="Museum">WAM</collectionCode>
:
<accessionNumber box="[487,569,1482,1504]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E97796" pageId="5" pageNumber="97">
<specimenCode box="[487,569,1482,1504]" pageId="5" pageNumber="97">E97796</specimenCode>
</accessionNumber>
).
</paragraph>
</materialsCitation>
<paragraph blockId="5.[144,238,1549,1570]" box="[144,238,1549,1570]" pageId="5" pageNumber="97">
<heading bold="true" box="[144,238,1549,1570]" fontSize="9" level="5" pageId="5" pageNumber="97" reason="2">
<emphasis bold="true" box="[144,238,1549,1570]" italics="true" pageId="5" pageNumber="97">
<typeStatus box="[144,238,1549,1570]" pageId="5" pageNumber="97">Paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[144,774,1596,1747]" pageId="5" pageNumber="97">
<materialsCitation ID-GBIF-Occurrence="2273363825" collectingDate="2012-05-19" collectionCode="WAM" collectorName="J. and F. Hort" country="Australia" county="Wandoo National Park" latitude="-32.118" location="York" longLatPrecision="5" longitude="116.60557" municipality="Kent Road" pageId="5" pageNumber="97" specimenCount="9" specimenCount-female="2" specimenCount-male="7" stateProvince="Western Australia" typeStatus="paratype">
<specimenCount box="[144,175,1596,1619]" pageId="5" pageNumber="97" type="male">7♂</specimenCount>
,
<specimenCount box="[193,224,1596,1619]" pageId="5" pageNumber="97" type="female">2♀</specimenCount>
:
<collectingCountry box="[242,383,1596,1618]" name="Australia" pageId="5" pageNumber="97">AUSTRALIA</collectingCountry>
<collectingRegion box="[394,579,1596,1618]" country="Australia" name="Western Australia" pageId="5" pageNumber="97">Western Australia</collectingRegion>
,
<collectingCounty pageId="5" pageNumber="97">Wandoo National Park</collectingCounty>
,
<collectingMunicipality box="[200,309,1628,1650]" pageId="5" pageNumber="97">Kent Road</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E06587DF5C67FFEEFCA2637CFC263695:6D1360125C66FFECFEB2623AFE833663" box="[318,371,1628,1650]" country="Australia" county="Wandoo National Park" latitude="-32.118" longLatPrecision="5" longitude="116.60557" municipality="Kent Road" name="York" pageId="5" pageNumber="97" stateProvince="Western Australia">York</location>
,
<geoCoordinate box="[380,469,1628,1650]" degrees="32.1180" direction="south" orientation="latitude" pageId="5" pageNumber="97" precision="5" value="-32.118">-32.1180</geoCoordinate>
<geoCoordinate box="[474,582,1628,1650]" degrees="116.60557" direction="east" orientation="longitude" pageId="5" pageNumber="97" precision="1" value="116.60557">116.60557</geoCoordinate>
, Jarrah open woodland with low open mixed shrubs. Feeding
<collectedFrom pageId="5" pageNumber="97">
on
<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Leucopogon" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="97" phylum="Tracheophyta" rank="species" species="oxycedrus">
<emphasis italics="true" pageId="5" pageNumber="97">Leucopogon oxycedrus</emphasis>
</taxonomicName>
</collectedFrom>
,
<date box="[262,410,1692,1714]" pageId="5" pageNumber="97" value="2012-05-19">
<collectingDate box="[262,410,1692,1714]" pageId="5" pageNumber="97" value="2012-05-19">19 May 2012</collectingDate>
</date>
,
<collectorName box="[426,580,1692,1714]" pageId="5" pageNumber="97">J. and F. Hort</collectorName>
(
<collectionCode box="[600,669,1692,1714]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="97" type="Museum">WAM</collectionCode>
:
<accessionNumber box="[683,761,1692,1714]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E97797" pageId="5" pageNumber="97">E97797</accessionNumber>
<accessionNumber box="[144,311,1724,1746]" isEnumeration="true" pageId="5" pageNumber="97">E97798, E97801</accessionNumber>
<accessionNumber box="[324,404,1724,1746]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E97807" pageId="5" pageNumber="97">E97807</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="5.[144,286,1791,1812]" box="[144,286,1791,1812]" pageId="5" pageNumber="97">
<heading bold="true" box="[144,286,1791,1812]" fontSize="9" level="5" pageId="5" pageNumber="97" reason="2">
<emphasis bold="true" box="[144,286,1791,1812]" italics="true" pageId="5" pageNumber="97">Other material</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[144,773,1837,1956]" pageId="5" pageNumber="97">
<materialsCitation ID-GBIF-Occurrence="2273363826" collectedFrom="Scattered Banksia with low open shrubs on white sand. On Leucopogon oxycedrus" collectingDate="2012-05-19" collectionCode="WAM" collectorName="J. &amp; F. Hort" country="Australia" latitude="-31.96443" location="Kent Road" longLatPrecision="1" longitude="116.51224" municipality="Flynn State Forest" pageId="5" pageNumber="97" specimenCode="E97799-E97800" specimenCount="2" specimenCount-male="2" stateProvince="Western Australia">
<specimenCount box="[144,175,1837,1860]" pageId="5" pageNumber="97" type="male">2♂</specimenCount>
<collectingCountry box="[184,324,1838,1860]" name="Australia" pageId="5" pageNumber="97">AUSTRALIA</collectingCountry>
<collectingRegion box="[332,515,1838,1860]" country="Australia" name="Western Australia" pageId="5" pageNumber="97">Western Australia</collectingRegion>
,
<collectingMunicipality box="[526,714,1838,1860]" pageId="5" pageNumber="97">Flynn State Forest</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E06587DF5C67FFEEFCA2637CFC263695:6D1360125C66FFECFD596348FF373775" country="Australia" latitude="-31.96443" longLatPrecision="1" longitude="116.51224" municipality="Flynn State Forest" name="Kent Road" pageId="5" pageNumber="97" stateProvince="Western Australia">Kent Road</location>
,
<location LSID="urn:lsid:plazi:treatment:E06587DF5C67FFEEFCA2637CFC263695:6D1360125C66FFECFF5E6328FEF83775" box="[210,264,1870,1892]" country="Australia" latitude="-31.96443" longLatPrecision="1" longitude="116.51224" municipality="Flynn State Forest" name="York" pageId="5" pageNumber="97" stateProvince="Western Australia">York</location>
,
<geoCoordinate box="[274,375,1870,1892]" degrees="31.96443" direction="south" orientation="latitude" pageId="5" pageNumber="97" precision="1" value="-31.96443">-31.96443</geoCoordinate>
<geoCoordinate box="[382,490,1870,1892]" degrees="116.51224" direction="east" orientation="longitude" pageId="5" pageNumber="97" precision="1" value="116.51224">116.51224</geoCoordinate>
,
<collectedFrom pageId="5" pageNumber="97">
Scattered
<taxonomicName box="[599,678,1870,1892]" class="Magnoliopsida" family="Proteaceae" genus="Banksia" kingdom="Plantae" order="Proteales" pageId="5" pageNumber="97" phylum="Tracheophyta" rank="genus">
<emphasis box="[599,678,1870,1892]" italics="true" pageId="5" pageNumber="97">Banksia</emphasis>
</taxonomicName>
with low open shrubs on white sand. On
<taxonomicName box="[494,730,1902,1924]" class="Magnoliopsida" family="Ericaceae" genus="Leucopogon" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="97" phylum="Tracheophyta" rank="species" species="oxycedrus">
<emphasis box="[494,730,1902,1924]" italics="true" pageId="5" pageNumber="97">Leucopogon oxycedrus</emphasis>
</taxonomicName>
</collectedFrom>
,
<date pageId="5" pageNumber="97" value="2012-05-19">
<collectingDate pageId="5" pageNumber="97" value="2012-05-19">19 May 2012</collectingDate>
</date>
,
<collectorName box="[259,275,1934,1956]" pageId="5" pageNumber="97">J.</collectorName>
and
<collectorName box="[324,395,1934,1956]" pageId="5" pageNumber="97">F. Hort</collectorName>
(
<collectionCode box="[409,477,1934,1956]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="97" type="Museum">WAM</collectionCode>
:
<specimenCode box="[487,659,1933,1956]" pageId="5" pageNumber="97">
<accessionNumber box="[487,564,1934,1956]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E97799" pageId="5" pageNumber="97">E97799</accessionNumber>
<accessionNumber box="[577,659,1934,1956]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E97800" pageId="5" pageNumber="97">E97800</accessionNumber>
</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="97" type="diagnosis">
<paragraph blockId="5.[813,928,1273,1297]" box="[813,928,1273,1297]" pageId="5" pageNumber="97">
<heading bold="true" box="[813,928,1273,1297]" fontSize="10" level="3" pageId="5" pageNumber="97" reason="2">
<emphasis bold="true" box="[813,928,1273,1297]" pageId="5" pageNumber="97">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[813,1443,1322,1472]" pageId="5" pageNumber="97">Body mostly covered with thick pale pruinescence. Scape and pedicel brownish yellow, flagellum black. Scutum with large black square markings in each corner and one thin black stripe in the midline. Wing membrane infuscated on anterior half, with inconspicuous lighter markings.</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="99" pageId="5" pageNumber="97" type="description">
<paragraph blockId="5.[813,954,1524,1548]" box="[813,954,1524,1548]" pageId="5" pageNumber="97">
<heading bold="true" box="[813,954,1524,1548]" fontSize="10" level="3" pageId="5" pageNumber="97" reason="2">
<emphasis bold="true" box="[813,954,1524,1548]" pageId="5" pageNumber="97">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[813,863,1590,1611]" box="[813,863,1590,1611]" pageId="5" pageNumber="97">
<heading bold="true" box="[813,863,1590,1611]" fontSize="9" level="5" pageId="5" pageNumber="97" reason="2">
<emphasis bold="true" box="[813,863,1590,1611]" italics="true" pageId="5" pageNumber="97">Male</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[813,1351,1637,1660]" box="[813,1351,1637,1660]" pageId="5" pageNumber="97">
Body length
<quantity box="[942,1077,1637,1660]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="10.9" metricValueMin="8.7" pageId="5" pageNumber="97" unit="mm" value="9.8" valueMax="10.9" valueMin="8.7">8.710.9 mm</quantity>
, wing length
<quantity box="[1212,1347,1637,1660]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.15" metricValueMin="0.9099999999999999" pageId="5" pageNumber="97" unit="mm" value="10.3" valueMax="11.5" valueMin="9.1">9.111.5 mm</quantity>
.
</paragraph>
<paragraph blockId="5.[813,1444,1710,1956]" lastBlockId="7.[144,774,142,740]" lastPageId="7" lastPageNumber="99" pageId="5" pageNumber="97">
<emphasis bold="true" box="[813,869,1710,1732]" pageId="5" pageNumber="97">Head</emphasis>
. Head about 2.2× wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to black hairs and scales. Eye narrowly separate by 0.5× width of ocellus. Frons short, 2.0× length of ocellar tubercle, with thick pale pruinescence and few golden scales. Ocellar tubercle slightly raised, brown to blackish brown with grey pruinescence, with long black hairs turning dark yellow at base. Face with thick pale pruinescence and long black hairs except turning dark yellow at base, parafacial area with long white scales admixed with a few long hairs. Gena with thick pale pruinescence and long white scales admixed with few long hairs. Clypeus with sparse pale pruinescence and otherwise bare. Occiput with dense white hairs and scales, dorsal central area admixed with fine black hairs and strong yellow hairs. Posterior eye margin slightly convex. Antennal scape and pedicel brownish yellow with thick pale pruinescence, scape with long black hairs turning dark yellow at base, pedicel with short white to pale yellow hairs; flagellum black without pruinescence, subapex with 35 long hairs. Scape 3.4× as long as wide, and 2.8× as long as pedicel, uniform from base to apex. Pedicel 1.6× as long as wide. Flagellum 12.0× as long as wide, 1.9× as long as scape + pedicel, 2.6× as long as scape, conical and slightly laterally compressed, one-segmented with apical stylus (
<figureCitation box="[332,404,590,612]" captionStart="Fig" captionStartId="4.[145,181,1556,1577]" captionTargetBox="[341,1247,131,1524]" captionTargetId="figure@4.[340,1248,129,1524]" captionTargetPageId="4" captionText="Fig. 2. Paramonovius nightking sp. nov. ♂: (a) dorsal; (b) lateral; (c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral; (g) head, frontal; (h) head, profile; (i) hind tarsi and claw. Scale bars = 1 mm (a, b, di); = 0.1 mm (c)." httpUri="https://zenodo.org/record/3265418/files/figure.png" pageId="7" pageNumber="99">Fig. 2c</figureCitation>
). Palpus short, not extending beyond oral cavity, black with blackish brown hairs, one-segmented, without palpal pit. Mouthparts slender, 3.2× as long as eye length, 4.8× as long as head length, labellum thin and filiform (
<figureCitation box="[151,222,718,740]" captionStart="Fig" captionStartId="4.[145,181,1556,1577]" captionTargetBox="[341,1247,131,1524]" captionTargetId="figure@4.[340,1248,129,1524]" captionTargetPageId="4" captionText="Fig. 2. Paramonovius nightking sp. nov. ♂: (a) dorsal; (b) lateral; (c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral; (g) head, frontal; (h) head, profile; (i) hind tarsi and claw. Scale bars = 1 mm (a, b, di); = 0.1 mm (c)." httpUri="https://zenodo.org/record/3265418/files/figure.png" pageId="7" pageNumber="99">Fig. 2f</figureCitation>
).
</paragraph>
<caption httpUri="https://zenodo.org/record/3265422/files/figure.png" pageId="6" pageNumber="98" startId="6.[145,181,1787,1808]" targetBox="[393,1205,132,1745]" targetPageId="6">
<paragraph blockId="6.[145,1444,1787,1955]" pageId="6" pageNumber="98">
<emphasis box="[145,469,1787,1808]" italics="true" pageId="6" pageNumber="98">
<emphasis bold="true" box="[145,209,1787,1808]" italics="true" pageId="6" pageNumber="98">Fig. 4.</emphasis>
<taxonomicName authority="Li &amp; Yeates, 2019" authorityName="Li &amp; Yeates" authorityYear="2019" box="[238,469,1787,1808]" class="Insecta" family="Bombyliidae" genus="Paramonovius" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="98" phylum="Arthropoda" rank="species" species="nightking" status="sp. nov.">
Paramonovius
<taxonomicName box="[381,469,1787,1808]" pageId="6" pageNumber="98" rank="species" species="nightking" status="sp. nov.">nightking</taxonomicName>
</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[475,551,1787,1808]" pageId="6" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
Genitalia (ae (♂); f, g (♀)): (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral; (d) epandrium, dorsal; (e) sternite 8, ventral; (f) genitalia and spermathecae. Scale bars = 0.1 mm. Abbreviations: acant spi = acanthophorite spine; aed sh = aedeagal sheath; b spmth dt = basal spermathecal duct; cerc = cercus; d brg = dorsal bridge; d spmth dt = distal spermathecal duct; ej apod = ejaculatory apodeme; epand = epandrium; gen fk = genital fork; goncx = gonocoxite; goncx apod = gonocoxal apodeme; gonst = gonostylus; lat ej proc = lateral ejaculatory process; ph = phallus; spmth = spermatheca; spm pmp = sperm pump; tg 9+10 = tergites 9+10.
</paragraph>
</caption>
<paragraph blockId="7.[144,774,782,1252]" pageId="7" pageNumber="99">
<emphasis bold="true" box="[144,222,782,804]" pageId="7" pageNumber="99">Thorax</emphasis>
. Integumental colour of scutum mostly black with dense pale pruinescence except postalar callus brownish yellow, scutum with large black square markings in each corner and one thin black stripe in the midline. Scutum covered with short white and golden scales, admixed with some long black hairs. Five brownish yellow notopleural setae present. Postalar callus with long black hairs. Scutellum brownish yellow with sparse pale pruinescence, with short white scales on anterior margin, dense short black scales and long black hairs on posterior margin, admixed with short golden scales. Pleura black with thick pale pruinescence, anepisternum with long white scales except anterior half with strong yellow hairs, katepisternum, katepimeron and dorsal margin of metepisternum with long white scales, coxae with dense long white scales, anepimeron, meron, laterotergite and mediotergite bare.
</paragraph>
<paragraph blockId="7.[144,775,1294,1540]" pageId="7" pageNumber="99">
<emphasis bold="true" box="[144,193,1294,1316]" pageId="7" pageNumber="99">Legs</emphasis>
. Legs mostly brownish yellow. Fore and mid femora with black scales on anterior face, hind femur with black scales on dorsal half, ventral half of femora with white to pale yellow hairs. Femora strong, with anterior blackish brown bristles. Other hairs and bristles on legs brown. Fore tibia 2.2× longer than fore basitarsus, mid tibia 2.5× longer than mid basitarsus, hind tibia 2.0× longer than hind basitarsus. Claw and pulvillus enlarged, as long as distitarsus (
<figureCitation box="[459,528,1518,1540]" captionStart="Fig" captionStartId="4.[145,181,1556,1577]" captionTargetBox="[341,1247,131,1524]" captionTargetId="figure@4.[340,1248,129,1524]" captionTargetPageId="4" captionText="Fig. 2. Paramonovius nightking sp. nov. ♂: (a) dorsal; (b) lateral; (c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral; (g) head, frontal; (h) head, profile; (i) hind tarsi and claw. Scale bars = 1 mm (a, b, di); = 0.1 mm (c)." httpUri="https://zenodo.org/record/3265418/files/figure.png" pageId="7" pageNumber="99">
Figs
<date box="[509,528,1518,1540]" pageId="7" pageNumber="99">2i</date>
</figureCitation>
and
<figureCitation box="[577,604,1518,1540]" captionStart="Fig" captionStartId="5.[144,180,1166,1187]" captionTargetBox="[340,1246,113,1133]" captionTargetId="figure@5.[339,1247,127,1134]" captionTargetPageId="5" captionText="Fig. 3. Paramonovius nightking sp. nov. ♀: (a) dorsal; (b) lateral; (c) head, dorsal; (d) head, lateral; (e) head, frontal; (f) head, profile; (g) hind tarsi and claw. Scale bars = 1 mm." httpUri="https://zenodo.org/record/3265420/files/figure.png" pageId="7" pageNumber="99">
<quantity box="[577,604,1518,1540]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="7" pageNumber="99" unit="g" value="3.0">3g</quantity>
</figureCitation>
).
</paragraph>
<paragraph blockId="7.[144,773,1582,1796]" pageId="7" pageNumber="99">
<emphasis bold="true" box="[144,211,1582,1604]" pageId="7" pageNumber="99">Wings</emphasis>
. Wing membrane infuscated on anterior half, with inconspicuous light markings on the area around base of vein
<emphasis box="[743,758,1614,1636]" italics="true" pageId="7" pageNumber="99">R</emphasis>
<emphasis box="[758,767,1624,1639]" italics="true" pageId="7" pageNumber="99">
<subScript attach="left" box="[758,767,1624,1639]" fontSize="6" pageId="7" pageNumber="99">4</subScript>
</emphasis>
, crossvein
<emphasis box="[247,282,1646,1668]" italics="true" pageId="7" pageNumber="99">r-m</emphasis>
, crossvein
<emphasis box="[403,465,1646,1668]" italics="true" pageId="7" pageNumber="99">dm-cu</emphasis>
and apex of cell
<emphasis box="[656,686,1646,1668]" italics="true" pageId="7" pageNumber="99">bm</emphasis>
. Cell
<emphasis box="[754,764,1646,1668]" italics="true" pageId="7" pageNumber="99">r</emphasis>
<emphasis box="[764,773,1656,1671]" italics="true" pageId="7" pageNumber="99">
<subScript attach="left" box="[764,773,1656,1671]" fontSize="6" pageId="7" pageNumber="99">5</subScript>
</emphasis>
widely open; cell
<emphasis box="[316,338,1678,1700]" italics="true" pageId="7" pageNumber="99">br</emphasis>
nearly as long as cell
<emphasis box="[550,580,1678,1700]" italics="true" pageId="7" pageNumber="99">bm</emphasis>
, crossvein
<emphasis box="[690,725,1678,1700]" italics="true" pageId="7" pageNumber="99">r-m</emphasis>
arising from base of cell
<emphasis box="[363,393,1710,1732]" italics="true" pageId="7" pageNumber="99">dm</emphasis>
; crossvein
<emphasis box="[509,553,1710,1732]" italics="true" pageId="7" pageNumber="99">m-m</emphasis>
long, 2.0× as long as crossvein
<emphasis box="[241,276,1742,1764]" italics="true" pageId="7" pageNumber="99">r-m</emphasis>
; cell
<emphasis box="[328,364,1742,1764]" italics="true" pageId="7" pageNumber="99">cup</emphasis>
open (
<figureCitation box="[429,501,1742,1764]" captionStart="Fig" captionStartId="4.[145,181,1556,1577]" captionTargetBox="[341,1247,131,1524]" captionTargetId="figure@4.[340,1248,129,1524]" captionTargetPageId="4" captionText="Fig. 2. Paramonovius nightking sp. nov. ♂: (a) dorsal; (b) lateral; (c) flagellum; (d) wing; (e) head, dorsal; (f) head, lateral; (g) head, frontal; (h) head, profile; (i) hind tarsi and claw. Scale bars = 1 mm (a, b, di); = 0.1 mm (c)." httpUri="https://zenodo.org/record/3265418/files/figure.png" pageId="7" pageNumber="99">Fig. 2d</figureCitation>
). Haltere stem yellow, knob dark yellow.
</paragraph>
<paragraph blockId="7.[144,773,1838,1956]" lastBlockId="7.[813,1443,142,676]" pageId="7" pageNumber="99">
<emphasis bold="true" box="[144,247,1838,1860]" pageId="7" pageNumber="99">Abdomen</emphasis>
. Integumental colour of tergites black with thick grey pruinescence. Tergite 1 with pale yellow hairs admixed with long black hairs on posterior margin; tergite 2 covered with white scales admixed with brown to black hairs on posterior half and short golden scales on posterior margin; tergites 3 to 7 compact, with golden scales admixed with black hairs, and with thick short white scales posterolaterally; tergites 26 with median stripe consisting of dense, decumbent short white scales. Tergites 79 rotated, with epandrium and gonocoxite rotated 180°.
<emphasis box="[1075,1168,302,324]" italics="true" pageId="7" pageNumber="99">Genitalia</emphasis>
. Epandrium nearly rectangular, anterior margin deeply concave, posterolateral corner with thick long hairs (
<figureCitation box="[1073,1158,366,388]" captionStart="Fig" captionStartId="6.[145,181,1787,1808]" captionTargetBox="[393,1205,132,1745]" captionTargetId="figure@6.[382,1207,127,1756]" captionTargetPageId="6" captionText="Fig. 4. Paramonovius nightking sp. nov. Genitalia (ae (♂); f, g (♀)): (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral; (d) epandrium, dorsal; (e) sternite 8, ventral; (f) genitalia and spermathecae. Scale bars = 0.1 mm. Abbreviations: acant spi = acanthophorite spine; aed sh = aedeagal sheath; b spmth dt = basal spermathecal duct; cerc = cercus; d brg = dorsal bridge; d spmth dt = distal spermathecal duct; ej apod = ejaculatory apodeme; epand = epandrium; gen fk = genital fork; goncx = gonocoxite; goncx apod = gonocoxalapodeme; gonst = gonostylus; lat ej proc = lateral ejaculatory process; ph = phallus; spmth = spermatheca; spm pmp = sperm pump; tg 9+10 = tergites 9+10." httpUri="https://zenodo.org/record/3265422/files/figure.png" pageId="7" pageNumber="99">Fig. 4d</figureCitation>
). Hypandrium fused with gonocoxite. Gonocoxite elongate, basal half wide and apical half abrupt narrowed. Ejaculatory apodeme large (
<figureCitation box="[1336,1431,430,452]" captionStart="Fig" captionStartId="6.[145,181,1787,1808]" captionTargetBox="[393,1205,132,1745]" captionTargetId="figure@6.[382,1207,127,1756]" captionTargetPageId="6" captionText="Fig. 4. Paramonovius nightking sp. nov. Genitalia (ae (♂); f, g (♀)): (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral; (d) epandrium, dorsal; (e) sternite 8, ventral; (f) genitalia and spermathecae. Scale bars = 0.1 mm. Abbreviations: acant spi = acanthophorite spine; aed sh = aedeagal sheath; b spmth dt = basal spermathecal duct; cerc = cercus; d brg = dorsal bridge; d spmth dt = distal spermathecal duct; ej apod = ejaculatory apodeme; epand = epandrium; gen fk = genital fork; goncx = gonocoxite; goncx apod = gonocoxalapodeme; gonst = gonostylus; lat ej proc = lateral ejaculatory process; ph = phallus; spmth = spermatheca; spm pmp = sperm pump; tg 9+10 = tergites 9+10." httpUri="https://zenodo.org/record/3265422/files/figure.png" pageId="7" pageNumber="99">Fig. 4a,b</figureCitation>
); gonocoxal apodeme strong and pointed anteriorly; anterior arm of aedeagal sheath long and wide, extending beyond gonocoxal margin; lateral ejaculatory process strong; inner apex of gonocoxite sharp; outer apex of gonocoxite sharp; dorsal bridge without lateral hollow; aedeagal sheath with acute projection on subapex; gonostylus small and strongly incurved; phallus long and strong (
<figureCitation box="[1068,1142,654,676]" captionStart="Fig" captionStartId="6.[145,181,1787,1808]" captionTargetBox="[393,1205,132,1745]" captionTargetId="figure@6.[382,1207,127,1756]" captionTargetPageId="6" captionText="Fig. 4. Paramonovius nightking sp. nov. Genitalia (ae (♂); f, g (♀)): (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral; (d) epandrium, dorsal; (e) sternite 8, ventral; (f) genitalia and spermathecae. Scale bars = 0.1 mm. Abbreviations: acant spi = acanthophorite spine; aed sh = aedeagal sheath; b spmth dt = basal spermathecal duct; cerc = cercus; d brg = dorsal bridge; d spmth dt = distal spermathecal duct; ej apod = ejaculatory apodeme; epand = epandrium; gen fk = genital fork; goncx = gonocoxite; goncx apod = gonocoxalapodeme; gonst = gonostylus; lat ej proc = lateral ejaculatory process; ph = phallus; spmth = spermatheca; spm pmp = sperm pump; tg 9+10 = tergites 9+10." httpUri="https://zenodo.org/record/3265422/files/figure.png" pageId="7" pageNumber="99">Fig. 4c</figureCitation>
).
</paragraph>
<paragraph blockId="7.[813,886,719,740]" box="[813,886,719,740]" pageId="7" pageNumber="99">
<heading bold="true" box="[813,886,719,740]" fontSize="9" level="5" pageId="7" pageNumber="99" reason="2">
<emphasis bold="true" box="[813,886,719,740]" italics="true" pageId="7" pageNumber="99">Female</emphasis>
</heading>
</paragraph>
<paragraph blockId="7.[813,1443,765,1172]" pageId="7" pageNumber="99">
Body length 10.811.0 mm, wing length
<quantity box="[1233,1382,765,788]" metricMagnitude="-2" metricUnit="m" metricValue="1.155" metricValueMax="1.16" metricValueMin="1.15" pageId="7" pageNumber="99" unit="mm" value="11.55" valueMax="11.6" valueMin="11.5">11.511.6 mm</quantity>
. Very similar to male, except frons black with thick pale pruinescence, 3.0× as wide as ocellar tubercle, frons with long brown to black hairs admixed with short golden scales (
<figureCitation box="[1246,1323,862,884]" captionStart="Fig" captionStartId="5.[144,180,1166,1187]" captionTargetBox="[340,1246,113,1133]" captionTargetId="figure@5.[339,1247,127,1134]" captionTargetPageId="5" captionText="Fig. 3. Paramonovius nightking sp. nov. ♀: (a) dorsal; (b) lateral; (c) head, dorsal; (d) head, lateral; (e) head, frontal; (f) head, profile; (g) hind tarsi and claw. Scale bars = 1 mm." httpUri="https://zenodo.org/record/3265420/files/figure.png" pageId="7" pageNumber="99">Fig. 3c</figureCitation>
). Tergite 8 fused and sclerotized, with some sparse hairs. Around 30 acanthophorite spines present on each side, slightly curved apically. Genital fork strong and broad, enlarged basally. Sperm pump strong and about as long as basal spermathecal duct, clothed in longitudinal muscle, with lateral papillae; sperm pump basal collars present and apical collars absent; common and basal spermathecal duct not elongate; apical spermathecal duct shorter than spermatheca; spermatheca elongate, cylindrical with apex spherical (
<figureCitation box="[915,986,1150,1172]" captionStart="Fig" captionStartId="6.[145,181,1787,1808]" captionTargetBox="[393,1205,132,1745]" captionTargetId="figure@6.[382,1207,127,1756]" captionTargetPageId="6" captionText="Fig. 4. Paramonovius nightking sp. nov. Genitalia (ae (♂); f, g (♀)): (a) genital capsule, dorsal; (b) genital capsule, ventral; (c) genital capsule, lateral; (d) epandrium, dorsal; (e) sternite 8, ventral; (f) genitalia and spermathecae. Scale bars = 0.1 mm. Abbreviations: acant spi = acanthophorite spine; aed sh = aedeagal sheath; b spmth dt = basal spermathecal duct; cerc = cercus; d brg = dorsal bridge; d spmth dt = distal spermathecal duct; ej apod = ejaculatory apodeme; epand = epandrium; gen fk = genital fork; goncx = gonocoxite; goncx apod = gonocoxalapodeme; gonst = gonostylus; lat ej proc = lateral ejaculatory process; ph = phallus; spmth = spermatheca; spm pmp = sperm pump; tg 9+10 = tergites 9+10." httpUri="https://zenodo.org/record/3265422/files/figure.png" pageId="7" pageNumber="99">Fig. 4f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="99" type="discussion">
<paragraph blockId="7.[813,921,1212,1236]" box="[813,921,1212,1236]" pageId="7" pageNumber="99">
<heading bold="true" box="[813,921,1212,1236]" fontSize="10" level="3" pageId="7" pageNumber="99" reason="2">
<emphasis bold="true" box="[813,921,1212,1236]" pageId="7" pageNumber="99">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="7.[813,1443,1262,1316]" pageId="7" pageNumber="99">
Wing with short stump vein sometimes present on crossvein
<emphasis box="[813,875,1294,1316]" italics="true" pageId="7" pageNumber="99">m-cua</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="99" type="etymology">
<paragraph blockId="7.[813,943,1357,1381]" box="[813,943,1357,1381]" pageId="7" pageNumber="99">
<heading bold="true" box="[813,943,1357,1381]" fontSize="10" level="3" pageId="7" pageNumber="99" reason="2">
<emphasis bold="true" box="[813,943,1357,1381]" pageId="7" pageNumber="99">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="7.[813,1443,1406,1556]" pageId="7" pageNumber="99">
This species is named after the Night King in the American fantasy drama
<emphasis box="[957,1126,1438,1460]" italics="true" pageId="7" pageNumber="99">Game of Thrones</emphasis>
, because all the specimens were collected in winter and the fly is mostly covered in thick pale pruinescence. The specific name is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection box="[813,982,1596,1668]" pageId="7" pageNumber="99" type="distribution">
<paragraph blockId="7.[813,959,1596,1620]" box="[813,959,1596,1620]" pageId="7" pageNumber="99">
<heading bold="true" box="[813,959,1596,1620]" fontSize="10" level="3" pageId="7" pageNumber="99" reason="2">
<emphasis bold="true" box="[813,959,1596,1620]" pageId="7" pageNumber="99">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="7.[813,982,1646,1668]" box="[813,982,1646,1668]" pageId="7" pageNumber="99">Endemic to WA.</paragraph>
</subSubSection>
</treatment>
</document>