treatments-xml/data/BF/FA/EF/BFFAEFD58F8374B5423CC80E8485EC3F.xml
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<document id="B97BC5D32C4DE7317B1C651B45A8D946" ENCODING="UTF-8" ID-GBIF-Dataset="68b0d7de-1651-4945-8576-b7328a26175b" ModsDocID="z00249p001" checkinTime="1246893442886" checkinUser="thomas" docAuthor="Jonathan W. Armbruster" docDate="2003" docId="BFFAEFD58F8374B5423CC80E8485EC3F" docLanguage="en" docName="2003_Armbruster_2_gg1.xml" docOrigin="Zootaxa 249" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:DDFAA9D6-E4FA-4C3C-9749-CF0313D30F3C" docTitle="Hypostomus hemicochliodon Armbruster, 2003, new species" docType="treatment" docUuid="4125CE0B-B466-47CA-B90F-CF9073FEE6F8" docUuidSource="ZooBank" docVersion="14" lastPageNumber="31" masterDocId="A29EE709B7587AF1AD73A093A03CF987" masterDocTitle="The species of the Hypostomus cochliodon group (Siluriformes: Loricariidae)." masterLastPageNumber="60" masterPageNumber="1" pageNumber="29" updateTime="1701313868015" updateUser="plazi">
<mods:mods id="24EA38EC4B54F9CA02FD743B03F2DDEA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5E7B38C110A02314362E9734E3F23EFC">
<mods:title id="8CE43EEC8F16C15835B9998DC7DBE27C">The species of the Hypostomus cochliodon group (Siluriformes: Loricariidae).</mods:title>
</mods:titleInfo>
<mods:name id="002CEBF158F3FC06AD65018B6481C3EB" type="personal">
<mods:role id="8A19AA94D5430F06C4DEEAC487440CD7">
<mods:roleTerm id="43F78CE4F3F17B047FCF0B1F08F931D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ACF37352ABD8ACFFB9326D36CF8531E9">Jonathan W. Armbruster</mods:namePart>
</mods:name>
<mods:typeOfResource id="806E8D4CDCBDE0B65E701E4412071B94">text</mods:typeOfResource>
<mods:relatedItem id="5FFDEA34E50F1AD5CF2AE905EDA86803" type="host">
<mods:titleInfo id="711620B579C718F54EAA3A373B8A9218">
<mods:title id="5A8409C514E6FECB4460769A82608C47">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A22D2BD99D445922A0F35DAFD09F5D81">
<mods:date id="47C5BA1FC0CDC8791DBF8F8646683AAA">2003</mods:date>
<mods:detail id="9D75FAB3D84F465834C6EF5C188CE749" type="volume">
<mods:number id="CB987E35127581C59F7A67E1E0450584">249</mods:number>
</mods:detail>
<mods:extent id="2105D23611A64064E2DC895E1D97427F" unit="page">
<mods:start id="4DA13F491EC30B29EFCC9519EC67F9BB">1</mods:start>
<mods:end id="E919B2CA9117AEA326A9CCAD0EBBFF19">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="34082B9C11E940571D71473598B7BC7E">
<mods:url id="D5EEE06FF0856BA5601D2179BD45B55D">http://www.zoobank.org/urn:lsid:zoobank.org:pub:DDFAA9D6-E4FA-4C3C-9749-CF0313D30F3C</mods:url>
</mods:location>
<mods:classification id="76B2DD52D1A4C22A02E8998B7480CF92">journal article</mods:classification>
<mods:identifier id="769C059DA397ACAA46A88976F74F7658" type="Plazi-Custom">z00249p001</mods:identifier>
<mods:identifier id="B3FD3E85E69662E9BF7FBFED55C3C93C" type="ZooBank">DDFAA9D6-E4FA-4C3C-9749-CF0313D30F3C</mods:identifier>
</mods:mods>
<treatment id="BFFAEFD58F8374B5423CC80E8485EC3F" ID-DOI="http://doi.org/10.5281/zenodo.6274707" ID-GBIF-Taxon="100125132" ID-Zenodo-Dep="6274707" LSID="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" httpUri="http://treatment.plazi.org/id/BFFAEFD58F8374B5423CC80E8485EC3F" lastPageNumber="31" pageNumber="29">
<subSubSection id="674CB8BA8FF8B3C5703A1328A71A833F" type="nomenclature">
<paragraph id="8E4A536377D9539D8EA4A2E59A0CC27E" pageNumber="29">
<taxonomicName id="E932DAFA5E2C5E2C53F33EF9ECF94102" ID-CoL="3NZ7Y" LSID-ZBK="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus hemicochliodon Armbruster 2003:29" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="species" species="hemicochliodon">Hypostomus hemicochliodon</taxonomicName>
<taxonomicNameLabel id="22395697B5AC57AF65A448E57A91EA07" rank="species">new species</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FCCFD8D4A63B825B79F2AC48F59D1B7F" type="description">
<paragraph id="1209D714EDC091FF932DBB6C05EAB11F" pageNumber="29">(Fig. 12)</paragraph>
</subSubSection>
<subSubSection id="B72E319BE95FD8B305B7911EB61824E9" type="nomenclature">
<paragraph id="DF64A561A81B358639ABD0C4334C2E42" pageNumber="29">
<materialsCitation id="7C882A4AB11839083102EEF1216D674F" ID-GBIF-Occurrence="1058481078" country="PERU" stateProvince="Loreto">
<typeStatus id="69EA9452B0FC2F37BEC14EF4E7359AA4">Holotype</typeStatus>
:
<collectingCountry id="4104F25B59A41A4C8BD5CBA59E9D4A31">PERU</collectingCountry>
.
<normalizedToken id="0984A0AC27DE4E33591EDDBB1FC98883" originalValue="Río">Rio</normalizedToken>
Nanay,
<normalizedToken id="8B16B3741DDFDCD65C0BE2F6F96BCA17" originalValue="Río">Rio</normalizedToken>
Amazonas dr. at Pampa Chica, 4.54 km from Iquitos center, 269° bearing,
<geoCoordinate id="690FA06AAE6732E48F05E276D95A4062" direction="south" orientation="latitude" value="-3.7519445">03°4507”S</geoCoordinate>
,
<geoCoordinate id="5291DB5915AFBD6F493B4CCE4BE1C0C8" direction="west" orientation="longitude" value="-73.28306">73°1659”W</geoCoordinate>
,
<collectionCode id="BF82C57874DB1FB197B0B380D16B2784" collectionName="Peru, Lima, Universidad Nacional Mayor de San Marcos, Museo de Historia Natural">MUSM</collectionCode>
17442, 1, (126.2 mm SL).
</materialsCitation>
</paragraph>
<paragraph id="C96A147FEB711AA922E143E655037516" pageNumber="29">
<materialsCitation id="9DB1110C3E9D966A9CD3609BAEFA8960" ID-GBIF-Occurrence="1058481092" country="PERU" stateProvince="Amazonas">
<typeStatus id="8C7491B91CDFDC1CF8F850A6DE59C58C">Paratypes</typeStatus>
:
<collectingCountry id="669635CE1F3251F5CBCB443E6A30B9F6">PERU</collectingCountry>
.
<collectingRegion id="698E287196B27BE23789B314240F8B51">Amazonas</collectingRegion>
,
<normalizedToken id="31D7F44BA03390C690B84F9DEF9FAA65" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="022296B92AA5D7E6D6204F41D6AD285F" originalValue="Marañon">Maranon</normalizedToken>
dr. 500 m upstream from Caterpiza,
<collectionCode id="D2F71DB20883E0C25EF5C1543D8231BA" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
42013-4, 1, (87.8)
</materialsCitation>
.
<materialsCitation id="0D4B0987DFBC9FBE171B986D8D29BB01" ID-GBIF-Occurrence="1058481093" country="PERU" stateProvince="Amazonas">
<normalizedToken id="32046C791822102B5DA148B728067DFE" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="0ED9A53F853B19ED0AD90E9A2AE29B86" originalValue="Marañon">Maranon</normalizedToken>
dr. Caterpiza,
<collectionCode id="C4E5C2A2C7E84BC3C5D2C43F88D4F398" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
41886-2, 1, (100.3)
</materialsCitation>
<materialsCitation id="7E445231D901DE28260275AD3DA26724" ID-GBIF-Occurrence="1058481176" country="PERU" stateProvince="Amazonas">
and
<collectionCode id="8696F183FDF1D16BEAADCB8FCF25E9B3" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
41893-4
</materialsCitation>
,
<materialsCitation id="5784F68E3E96B51D79C1830F46881A98" ID-GBIF-Occurrence="1058481139" country="Peru" stateProvince="Amazonas">
<collectingRegion id="C73F3058D650CBEF37A6B061BABF91E8">Amazonas</collectingRegion>
, Quebrada,
<normalizedToken id="5A6BF3B91F7BD885BD397675C2A329BA" originalValue="Río">Rio</normalizedToken>
Santiago -
<normalizedToken id="FDEA7C47598A08284772CE3D7C94508C" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="A204278032B76FB44C79EA11B981D65F" originalValue="Marañon">Maranon</normalizedToken>
dr. quebrada enters
<normalizedToken id="BD1ED16D32E57798B3396344B433C0BE" originalValue="Río">Rio</normalizedToken>
Santiago from east, 2/3 length of island downstream of La Poza,
<collectionCode id="9DDA7449400C84DD1C9B60E91AB0A99B" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
39862-5, 1, (80.1)
</materialsCitation>
.
<materialsCitation id="68DF5D5FA0D6E0898CC847BF226B1354" ID-GBIF-Occurrence="1058481095" country="Peru" stateProvince="Amazonas">
Quebrada Kayamasa,
<normalizedToken id="E8B90E937BD96EA8A1E2E19E65D7DF9E" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="8EEF7FBE10F7060931E487EF8945EC26" originalValue="Marañon">Maranon</normalizedToken>
dr. 40 km upstream from Caterpiza,
<collectionCode id="A62D7E4139EB13ADAC25D5954462ABC9" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
41853-3, 1, (206.9)
</materialsCitation>
.
<materialsCitation id="9F16873BCB5AF7BBA7185D068FA13CD3" ID-GBIF-Occurrence="1058481067" country="Peru" stateProvince="Amazonas">
Quebrada Pastazilla,
<normalizedToken id="4124ED90B650CD144D81DCDA3E2823FA" originalValue="Río">Rio</normalizedToken>
Santiago -
<normalizedToken id="48CAA4D3082AB6426815C8EE0D118A02" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="4355CC8BABD9EE1E01EE82FD4D6635C3" originalValue="Marañon">Maranon</normalizedToken>
dr.
<collectionCode id="8CC92122101B1A9BA8DB244ADC121F1E" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
39947-5, 2, (109.7-118.9)
</materialsCitation>
<materialsCitation id="DEF37E10BCB57E8F78A98547C5BCEC81" ID-GBIF-Occurrence="1058481203" country="Peru" stateProvince="Amazonas">
and
<collectionCode id="BDB5E10FA95E5B3DF6AC7D09400DB3F7" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
39937-6, 1, (162.6)
</materialsCitation>
.
<materialsCitation id="1FCB9DA410AB70DAF1AE4BA5CA28603D" ID-GBIF-Occurrence="1058481021" country="Peru" stateProvince="Amazonas">
<normalizedToken id="DA50AEDBBB078D3728DB83B92E3D5A2F" originalValue="Río">Rio</normalizedToken>
Santiago,
<normalizedToken id="E8912D98CA60EE166650D993E313ED02" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="9D9ADBB32C98B03DCC3675E1B570179F" originalValue="Marañon">Maranon</normalizedToken>
dr. at La Poza,
<collectionCode id="0E05503F0BBBBE780E9205748D2E4217" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
97010, 1, (115.1)
</materialsCitation>
<materialsCitation id="ED045089B4E5AC7EBA8B36BCED99EF2F" ID-GBIF-Occurrence="1058481179" country="Peru" stateProvince="Amazonas">
and
<collectionCode id="E62C9EA199F5FC3C6120FAB8588DE6A7" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
41729-34, 1, (91.3)
</materialsCitation>
.
<materialsCitation id="C8EECC3566DD0DAC5EE92703563AAC42" ID-GBIF-Occurrence="1058481201" country="Peru" stateProvince="Loreto">
<collectingRegion id="FD49A27EF1FB4C4B6A186C6C2A354B38">Loreto</collectingRegion>
,
<normalizedToken id="7015745E52E5B123115D4C46FC65DA70" originalValue="Caño">Cano</normalizedToken>
Sacarito,
<normalizedToken id="F07BA8F56BCB843BEE945D5E3AF68BCB" originalValue="Río">Rio</normalizedToken>
Orosa -
<normalizedToken id="CCD87EADA15B32F72D6005B2E6651B61" originalValue="Río">Rio</normalizedToken>
Amazonas dr. about 35 min by boat upstream from mouth of Tonche
<normalizedToken id="96BDB50997D04DDA195B29E895604324" originalValue="Caño">Cano</normalizedToken>
, 67.1 mi ENE Iquitos,
<geoCoordinate id="6091862A12D3D42569C48E298AE171AC" direction="south" orientation="latitude" value="-3.6138887">03°3650”S</geoCoordinate>
,
<geoCoordinate id="2580DA1BEC5A9FF7F4F7107238D395C7" direction="west" orientation="longitude" value="-72.281944">72°1655”W</geoCoordinate>
,
<collectionCode id="1082E21793BED17FF280FF7B7F3ED571" collectionName="USA, Illinois, Champaign, Illinois Natural History Survey">INHS</collectionCode>
39690, 1, (84.6)
</materialsCitation>
,
<materialsCitation id="A94AE6583CB1E4E8128DAFDAE66681BA" ID-GBIF-Occurrence="1058481100" country="Peru" stateProvince="Loreto">
<collectionCode id="3AAF98FB543EE0621AFE9946A11086DE" collectionName="AUM">AUM</collectionCode>
28907, 1 (82.8)
</materialsCitation>
.
<materialsCitation id="B9C3FE59331ED0D93F8BACC67DF921EA" ID-GBIF-Occurrence="1058481055" country="Peru" stateProvince="Loreto">
<normalizedToken id="1C4BEE0C0D8D97814536193C99140EBE" originalValue="Río">Rio</normalizedToken>
Itaya,
<normalizedToken id="656B913ABD82D2BAD4623F4141256285" originalValue="Río">Rio</normalizedToken>
Amazonas dr. approx. 10 km S Santa Clara,
<collectionCode id="ED385943508F24D0FEA5C73866549511" collectionName="USA, Illinois, Champaign, Illinois Natural History Survey">INHS</collectionCode>
36878, 1, 1 cs, (99.8)
</materialsCitation>
<materialsCitation id="6E01DC201EB5B6613C192149B24ACB9C" ID-GBIF-Occurrence="1058481116" country="Peru" stateProvince="Loreto">
and
<collectionCode id="5A3E1FF0F97A9561927FDDC2DC7C1E7C" collectionName="USA, Illinois, Carbondale, Southern Illinois University, Research Museum of Zoology">SIUC</collectionCode>
36690, 1, (93.8)
</materialsCitation>
.
<materialsCitation id="D364A233D0F2E2AC146317495D88AA8A" ID-GBIF-Occurrence="1058481115" country="Peru" stateProvince="Loreto">
<normalizedToken id="A621A24FE4FB7C9DBA54FDDC4B8A6AF2" originalValue="Río">Rio</normalizedToken>
Itaya,
<normalizedToken id="B70D04C60DC47BA3A079488D41491DA4" originalValue="Río">Rio</normalizedToken>
Amazonas dr. ca. 4-5 km upstream from Iquitos (
<normalizedToken id="E508E522FDC10CCA63C6D9F4C11C34DD" originalValue="Belém">Belem</normalizedToken>
) above and below mouth of Quebrada Manzana,
<geoCoordinate id="A0B3CAB778474D8AF6400E79B89AE961" direction="south" orientation="latitude" value="-3.8030555">03°4771”S</geoCoordinate>
,
<geoCoordinate id="8EE63A41C2260446DA2F596401F97DE2" direction="west" orientation="longitude" value="-73.29139">73°1729”W</geoCoordinate>
,
<collectionCode id="66306325D3B028DCB663357A68996558" collectionName="USA, Illinois, Champaign, Illinois Natural History Survey">INHS</collectionCode>
39962, 1, (100.5)
</materialsCitation>
.
<materialsCitation id="8F73ACBC56EF1AD78F6B5CD08BB4FE47" ID-GBIF-Occurrence="1058481186" country="Peru" stateProvince="Loreto">
<normalizedToken id="28EB40F2ECE177B65AEB1CA2F8F84E89" originalValue="Río">Rio</normalizedToken>
Itaya,
<normalizedToken id="1874C6A4627852D9D6332024CF7917C7" originalValue="Río">Rio</normalizedToken>
Amazonas dr. near Iquitos,
<collectionCode id="C1C5A2BE75D127477F32FFCDA56995D3" collectionName="USA, New York, New York, American Museum of Natural History">AMNH</collectionCode>
218019, 1, (82.1)
</materialsCitation>
.
<materialsCitation id="5133F183C6A56FE6C1C5C43CD3536CD6" ID-GBIF-Occurrence="1058481191" country="Peru" stateProvince="Loreto">
<normalizedToken id="215386185AF760A089FD335B7E2EADF8" originalValue="Río">Rio</normalizedToken>
Nanay,
<normalizedToken id="DF5B2AF297154EA1D4E0FD149DD09E50" originalValue="Río">Rio</normalizedToken>
Amazonas dr. at Pampa Chica, 4.54 km from Iquitos center, 269°bearing,
<geoCoordinate id="F7C93022931C1A5BC0DFEAC106366599" direction="south" orientation="latitude" value="-3.7519445">03°4507”S</geoCoordinate>
,
<geoCoordinate id="31E1B97F22D0189F5738FBB361A0CB27" direction="west" orientation="longitude" value="-73.28306">73°1659”W</geoCoordinate>
,
<collectionCode id="86B0152DF24B1B8DC4949C561497E350" collectionName="USA, Illinois, Champaign, Illinois Natural History Survey">INHS</collectionCode>
50131, 2, 1 cs, (80.6- 107.8)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9ED5B9BC354EDFCD12712BE6E5572CEA" type="materials_examined">
<paragraph id="5BF808BDF6CE4AAC6BAD164F1CF2CC64" pageNumber="29">
<materialsCitation id="561EB147FF97808F0B9E6D80F4493C4C" ID-GBIF-Occurrence="1058481212" country="Brazil" stateProvince="Mato Grosso">
Nontype material:
<collectingCountry id="04DBEE1A95504B75B282149A61C0E6FB">BRAZIL</collectingCountry>
.
<collectingRegion id="9FBFBCC23D357D6A1F47D00C43960ED7">Mato Grosso</collectingRegion>
, Rio Juruena, Rio Tapajos dr. upper Rio Juruena,
<collectionCode id="4F6851A4C3485A1308F7380B1CB6A07A" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
199225, 1, (119.3)
</materialsCitation>
.
<materialsCitation id="3522ABC972676BCCB6840CC48916EFD9" ID-GBIF-Occurrence="1058481045" country="Brazil" stateProvince="Mato Grosso">
Rio Xingu, Rio Amazonas dr.
<normalizedToken id="D0886D9FD0C1D496D608E41C05EEEEFB" originalValue="São">Sao</normalizedToken>
Felix,
<collectionCode id="8A09F228F370580F046F32FAA97F8F12" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
207509, 1, (82.7)
</materialsCitation>
.
<materialsCitation id="197EE7C777E3D4FA6A3E3FF84751C6E8" ID-GBIF-Occurrence="1058481154" country="Ecuador" stateProvince="Napo">
<collectingCountry id="6FD0F7CA191050FC78A635F1B95461E7">ECUADOR</collectingCountry>
:
<collectingRegion id="56379A5E94C31DA5898751BED280BB1C">Napo</collectingRegion>
,
<normalizedToken id="609058D8AAFDF2F2321D1A1038709A2D" originalValue="Río">Rio</normalizedToken>
Jivino,
<normalizedToken id="F74DD7D06D1F7B79BB06A0CF382465DE" originalValue="Río">Rio</normalizedToken>
Napo dr. lower 4 km (mostly) to 0.6 km upstream from mouth (most between the two ports of Limoncocha),
<geoCoordinate id="75061392D0752E9B9AA60B0B63D64186" direction="south" orientation="latitude" value="-0.41">00°2436”S</geoCoordinate>
,
<geoCoordinate id="352FCD08CC01BDAE8C2BA7D61A37E115" direction="west" orientation="longitude" value="-76.65">76°3900”W</geoCoordinate>
,
<collectionCode id="39FBCB93ECDE651B3BE10461C13C0262" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
106021, 1, (105.3)
</materialsCitation>
.
<materialsCitation id="0C1701B7CADE80E436427086341C11FA" ID-GBIF-Occurrence="1058481104" country="Ecuador">
<normalizedToken id="253136D0BAFDAC813755A01340C1D003" originalValue="Río">Rio</normalizedToken>
Shushufindi,
<normalizedToken id="10113BBA018A16459CB2C71D8B7F0DCD" originalValue="Río">Rio</normalizedToken>
Aguarico dr. lower reaches, about 2 km from mouth in
<normalizedToken id="0675B278B512AE36171EDB6D0E63C360" originalValue="Río">Rio</normalizedToken>
Aquarico,
<geoCoordinate id="CED78009E76FE85BF63CC3668588C057" direction="south" orientation="latitude" value="-0.29166666">00°1730”S</geoCoordinate>
,
<geoCoordinate id="18C4C47AF85D50C7ECC4068527986B7C" direction="west" orientation="longitude" value="-76.42667">76°2536”W</geoCoordinate>
,
<collectionCode id="486DA97EAF22EA2FC92459481CB6632E" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
106025, 2, (141.5-157.9)
</materialsCitation>
.
<materialsCitation id="EE7CB61E25AF074DBB17EDD57A89F8DB" ID-GBIF-Occurrence="1058481120" country="Ecuador">
Tributary,
<normalizedToken id="C70B443DE1DF9550A4D8A6C1F67C4AFF" originalValue="Río">Rio</normalizedToken>
Payamino dr. few km upstream from San Jose de Payamino,
<geoCoordinate id="260ECFE959BDE19FA7A04057C7F08C1C" direction="south" orientation="latitude" value="-0.33666667">00°2012”S</geoCoordinate>
,
<geoCoordinate id="F44E8FCD5FA8CAA189A7E9AAFEF404E3" direction="west" orientation="longitude" value="-77.3">77°18W</geoCoordinate>
,
<collectionCode id="0859BAD235C05B297761AE633BC36C45" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
106024, 1, (158.0)
</materialsCitation>
.
<materialsCitation id="74609FB6613071E50571AB4861B494F1" ID-GBIF-Occurrence="1058481090" country="Ecuador">
Tributary,
<normalizedToken id="CD42D27E70455A6EB7E89D565DFC74D1" originalValue="Río">Rio</normalizedToken>
San Miguel dr. km 50, 4.5 km S of Tipischa,
<collectionCode id="390F5A75BD58120400D7C01F3F490440" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
106022, 1, (208.7)
</materialsCitation>
.
<materialsCitation id="BC403EAA26B672F7B89B99E9FA338CF5" ID-GBIF-Occurrence="1058481123" country="Ecuador">
Tributary,
<normalizedToken id="3403CEBABD0FC1A98E4BD568FF34E814" originalValue="Río">Rio</normalizedToken>
Churuyacu -
<normalizedToken id="FF9147FD674C59B531C92901E57C8EE8" originalValue="Río">Rio</normalizedToken>
Payamino dr. Tiuyacu, first tributary of
<normalizedToken id="DF90A72B748A684650689B005CF8B58D" originalValue="Río">Rio</normalizedToken>
Churuyacu upstream from mouth in
<normalizedToken id="BA2B530F2C69F6795E85586CA241C630" originalValue="Río">Rio</normalizedToken>
Payamino and near mouth of
<normalizedToken id="915FB52E4C9C063BB396EF0912F9CA39" originalValue="Río">Rio</normalizedToken>
Churuyacu,
<geoCoordinate id="83E4E9E1EA51D2645042FD82CC634C96" direction="south" orientation="latitude" value="-0.49166664">00°2930”S</geoCoordinate>
,
<geoCoordinate id="CE3CBEC022A1D930D402339D5B15FC73" direction="west" orientation="longitude" value="-77.3">77°18W</geoCoordinate>
,
<collectionCode id="E050CA1D6B08A4B707C1B72D30E7C746" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
106023, 3, (112.6-197.2)
</materialsCitation>
.
<materialsCitation id="23AD52F592C2C52429CF872B23ED1484" ID-GBIF-Occurrence="1058481105" country="Ecuador">
<normalizedToken id="936E585C764939BE928A9231E0371A1A" originalValue="Río">Rio</normalizedToken>
Corrientes,
<normalizedToken id="676F1A1BF5CF75E94ECC86EE83E7E4AA" originalValue="Río">Rio</normalizedToken>
Napo dr. eastern Ecuador,
<collectionCode id="6C6296483860C1EC226ABCC7B0D2FA46" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
92633, 3, 1 cs, (78.6- 89.8)
</materialsCitation>
.
<materialsCitation id="004C6FD376C6F56CD90646F593F046DD" ID-GBIF-Occurrence="1058481065" country="Venezuela" stateProvince="Amazonas">
<collectingCountry id="3894F0BB0CADD0B1A856EC65297C5306">VENEZUELA</collectingCountry>
.
<collectingRegion id="D686250DA7A9D85F2A1AF4DA5DA50DC9">Amazonas</collectingRegion>
,
<normalizedToken id="070CA7B245ED48D0FEF4105C79FC76F6" originalValue="Caño">Cano</normalizedToken>
Jenita,
<normalizedToken id="B657C018EF511D316E0F754DFE946DA7" originalValue="Río">Rio</normalizedToken>
Ocamo dr. Oramo department, in an old meander 1 km river km above the
<normalizedToken id="EAD1F392A58683F13C93F835D1FD38DE" originalValue="Río">Rio</normalizedToken>
Ocamo,
<geoCoordinate id="6499892D3BAEEBBAD3B51AE735C7F2F1" direction="north" orientation="latitude" value="2.7666667">02°46N</geoCoordinate>
,
<geoCoordinate id="3CCAF0249026A6DA1C7FA7143B97DD33" direction="west" orientation="longitude" value="-64.9">64°54W</geoCoordinate>
,
<collectionCode id="0D31DBD36B7AF0FC26E3D55D5356E4F0" collectionName="MCNG">MCNG</collectionCode>
24332, 1, (142.3)
</materialsCitation>
.
<materialsCitation id="F3DE8809062C07FE253BE620C609BCA2" ID-GBIF-Occurrence="1058481077" country="Venezuela">
Laguna
<normalizedToken id="3288DA25C24F1CA461F5DA5143969B9D" originalValue="Pacón">Pacon</normalizedToken>
,
<normalizedToken id="6C62F9C2805AE6107FE4C31B049F0FAE" originalValue="Río">Rio</normalizedToken>
Orinoco dr. in pools behind beach along
<normalizedToken id="E28AD5369660FFBAB82724FF83E73711" originalValue="Río">Rio</normalizedToken>
Ventuari about 0.5 hr above mouth,
<collectionCode id="8AFC2981AE7299F6F3767B6888B73BE9" collectionName="MBUCV">MBUCV</collectionCode>
V-23092, 1, (165.2)
</materialsCitation>
.
<materialsCitation id="7E3021BFC54EBDF69E79239E09A60A3E" ID-GBIF-Occurrence="1058481097" country="Venezuela">
<normalizedToken id="17FFDCE313A42578F61D7851D35DC3F1" originalValue="Río">Rio</normalizedToken>
Cataniapo,
<normalizedToken id="5A473A3B9EDD6DFFDCE5658C588A8013" originalValue="Río">Rio</normalizedToken>
Orinoco dr. in Las Palvas,
<collectionCode id="824A28D27E35DCEFD251329D0C9F9A47" collectionName="MBUCV">MBUCV</collectionCode>
V-24501 (none measured)
</materialsCitation>
.
<materialsCitation id="C829ADFC0A33DBB6681A4A2E5F35E6D3" ID-GBIF-Occurrence="1058481199" country="Venezuela">
<normalizedToken id="6848AEFE78791582CB2837D1C8EA5CD6" originalValue="Río">Rio</normalizedToken>
Mavaca,
<normalizedToken id="BF45013CE14D5D47A58530AC332CB4AD" originalValue="Río">Rio</normalizedToken>
Orinoco dr. river above camp base,
<collectionCode id="43F0F041179317378CB832E46E12A276" collectionName="MBUCV">MBUCV</collectionCode>
V-19239, 1, (186.1)
</materialsCitation>
.
<materialsCitation id="617C8FCDCF0E03BD3948E65A0448CACF" ID-GBIF-Occurrence="1058481204" country="Venezuela">
<normalizedToken id="5EFCAD1BC6A6C94958A7961D16542721" originalValue="Río">Rio</normalizedToken>
Siapa, Rio Casiquiare -
<normalizedToken id="15EBD565D4BE44DE43F893E4C7C4B85A" originalValue="Río">Rio</normalizedToken>
Negro dr. Departmento
<normalizedToken id="A6BAABD05095F75AFDF94913A9251D20" originalValue="Río">Rio</normalizedToken>
Negro, at campsite near Laguna Cumicapi between
<normalizedToken id="57C3CA185A2FD1BF23489AE0EF385C73" originalValue="Río">Rio</normalizedToken>
Emoni and
<normalizedToken id="BDC734D178301E682138A5BFFA872F96" originalValue="Río">Rio</normalizedToken>
Manipitare,
<collectionCode id="FF179951B77B806DF660EBDEBD9ED069" collectionName="MCNG">MCNG</collectionCode>
37041, 1, (100.3)
</materialsCitation>
.
<materialsCitation id="C77A4C3429082FD923076873F4F0C448" ID-GBIF-Occurrence="1058481086" country="Venezuela">
<normalizedToken id="D2BD01048DEA9BC02D1EE5BF82FEC9A6" originalValue="Río">Rio</normalizedToken>
Siapa,
<normalizedToken id="A9CDD5899CBEEAFAD93A6F2A1764EFB4" originalValue="Río">Rio</normalizedToken>
Casiquiare dr. near and downstream of confluence with
<normalizedToken id="516EC130F1EE5BC9F842D069B1BC8A63" originalValue="Río">Rio</normalizedToken>
Manipitare,
<geoCoordinate id="440C1CB038F066BF3880834537853C17" direction="north" orientation="latitude" value="1.9016666">01°5406”N</geoCoordinate>
,
<geoCoordinate id="4066B2CF46BA81E8C3C8FF748670DDFB" direction="west" orientation="longitude" value="-65.951385">65°5705”W</geoCoordinate>
,
<collectionCode id="581BE75393FAB42C0DE72C4DEEF5C505" collectionName="MCNG">MCNG</collectionCode>
37032, 1, (199.5)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4A01EA2A1CA5E22C099762E61871A08" type="diagnosis">
<paragraph id="8C567F12714B2302601FAAE14CECB651" lastPageNumber="31" pageNumber="30">
Diagnosis:
<taxonomicName id="B1FAD4A28F03352E71AF8A4B89C2826D" LSID-ZBK="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus hemicochliodon Armbruster 2003:29" order="Siluriformes" pageNumber="30" phylum="Chordata" rank="species" species="hemicochliodon">Hypostomus hemicochliodon</taxonomicName>
differs from all other members of the
<taxonomicName id="641E55A0BC861307053F82948F80BEBE" LSID-ZBK="urn:lsid:zoobank.org:act:7520B3CD-59C7-4425-A30E-1336A0B2AADD" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus cochliodon Kner 1854:265 [15]" order="Siluriformes" pageNumber="30" phylum="Chordata" rank="species" species="cochliodon">H. cochliodon</taxonomicName>
group except
<taxonomicName id="4E45D174C91E12552B9D3B72430DE104" LSID-ZBK="urn:lsid:zoobank.org:act:3F8E55C1-0EAD-49CA-B887-0D953CC48DA0" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus sculpodon Armbruster 2003:49" order="Siluriformes" pageNumber="30" phylum="Chordata" rank="species" species="sculpodon">H. sculpodon</taxonomicName>
by the presence of a medium-sized buccal papilla (condition intermediate between those shown in figures 4A and 4B), intermediately developed teeth (Fig. 1B; juvenile
<taxonomicName id="64DD70617EFC171B36D7FCAFDD7E5C15" LSID-ZBK="urn:lsid:zoobank.org:act:7520B3CD-59C7-4425-A30E-1336A0B2AADD" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus cochliodon Kner 1854:265 [15]" order="Siluriformes" pageNumber="30" phylum="Chordata" rank="species" species="cochliodon">H. cochliodon</taxonomicName>
also occasionally have intermediate teeth), and a large patch of odontodes on the opercle.
<taxonomicName id="E65516FE63B8E6702047ECDF2EA9001B" LSID-ZBK="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus hemicochliodon Armbruster 2003:29" order="Siluriformes" pageNumber="30" phylum="Chordata" rank="species" species="hemicochliodon">Hypostomus hemicochliodon</taxonomicName>
differs from
<taxonomicName id="1259E67FED431E59C53FC16DFCA91C4C" LSID-ZBK="urn:lsid:zoobank.org:act:3F8E55C1-0EAD-49CA-B887-0D953CC48DA0" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus sculpodon Armbruster 2003:49" order="Siluriformes" pageNumber="30" phylum="Chordata" rank="species" species="sculpodon">H. sculpodon</taxonomicName>
by coloration (dark brown with closely spaced spots vs. brownish-red with widely spaced spots), by having a greater dorsal-anal length to anal-fin length ratio (average= 111.1 ± 8.4%, 92.5-132.6% vs. average = 79.2 ± 7.6%, 69.6-89.7%), and almost completely by having a larger interorbital width/HL ratio (average = 49.9 ± 2.7%, 44.1- 55.6% vs. average = 51.4 ± 3.1%, 38.5-45.4%).
</paragraph>
</subSubSection>
<subSubSection id="683192B40C371BA7EB5CB8E3813FCB2B" type="description">
<paragraph id="7D4886DA2C8C56E23D7BF86F461D1255" pageNumber="31">
Description: See description of the
<taxonomicName id="BCD112270DB00126B254A5BCE109BEC2" LSID-ZBK="urn:lsid:zoobank.org:act:7520B3CD-59C7-4425-A30E-1336A0B2AADD" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus cochliodon Kner 1854:265 [15]" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="cochliodon">Hypostomus cochliodon</taxonomicName>
group for more details. Morphometric data given in Table 2. Color brown with medium to large spots. Spots become larger posteriorly, often coalescing on caudal peduncle to form large, longitudinally oval dashes or stripes. Some adults with spots on body absent. Spots on abdomen variable; fairly large specimens often have few or no spots on the abdomen; abdomen often significantly lighter than sides, almost white. Four faint dorsal saddles often present: first below anterior rays of dorsal fin, second below posterior rays of and slightly posterior to dorsal fin, third slightly anterior to and ventral to adipose fin, and fourth at base of caudal fin. One captive
<taxonomicName id="A194151B1305EA180DB9560450F52696" LSID-ZBK="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus hemicochliodon Armbruster 2003:29" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="hemicochliodon">Hypostomus hemicochliodon</taxonomicName>
demonstrated well-developed ability to change color, particularly ability to lighten coloration of abdomen and intensify lateral spots or dorsal saddles (either spots or the saddles dominant, but never both dark at same time).
</paragraph>
<paragraph id="3ADFE899EDC4F7CD6D74AB67FCEB3975" pageNumber="31">Dorsal fin fairly short, usually falls short of preadipose plate when depressed. Depressed pectoral-fin spine ventral to pelvic fin reaches 2-3 plates beyond bases of pelvic-fin rays. Tip of pectoral-fin spine of nuptial males with stout, recurved, hypertrophied odontodes.</paragraph>
<paragraph id="0D9A77C1C13CB9C612D4398695EB8855" pageNumber="31">Keels strong, sharp. Orbit forming ridge distinctly raised above medial surface of head; ridges of dorsal and lateral aspect of head well-developed. Longitudinal ridge formed of raised bone and slightly larger odontodes present on pterotic-supracleithrum beginning at postdorsal corner of orbit. Opercle broadly exposed, always supporting much more than ten odontodes. Nuptial body odontodes absent (Fig. 2A). Plates in skin anterior to dorsal-fin spine absent or few (Fig. 6A; Table 7).</paragraph>
<paragraph id="A27EC58F2284DFE2D9D00A6C5C9E2DD8" pageNumber="31">Each jaw with 7-23 teeth (mode = 15), teeth of intermediate size (Fig. 1B). Average angle between dentaries 63° (SD = 7°; range: 45°-75°; N=23). Lateral line plates 28-30; dorsal plates 8-9; interdorsal plates 6-8; adipose caudal plates 8-10.</paragraph>
</subSubSection>
<subSubSection id="5271573A947F9E5B0ED95CA68BFCC02D" type="distribution">
<paragraph id="22073C61615447C6944A73B6E9CBB56B" pageNumber="31">
Range: Has a wide range across central South America in the
<normalizedToken id="BD25ADF0F23FA616888393BCBE07E37E" originalValue="Ríos">Rios</normalizedToken>
<normalizedToken id="E3CC7CD006CCDBA3DD9A8165B22F048C" originalValue="Marañon">Maranon</normalizedToken>
, Itaya, Nanay, Napo, and Orosa of the upper
<normalizedToken id="D901C95C6C73D53893C31F20597268E3" originalValue="Río">Rio</normalizedToken>
Amazonas of Peru, the
<normalizedToken id="A4DEBC94179A961F8749009A62C13F6B" originalValue="Río">Rio</normalizedToken>
Napo drainage of the Upper
<normalizedToken id="3ACA3C19BEF9E739033D0D76B5C91D9E" originalValue="Río">Rio</normalizedToken>
Amazonas drainage of Ecuador, the Rios Tapajos and Xingu of the lower Rio Amazonas drainage of Brazil, and the upper
<normalizedToken id="D4CAD1EE5A64321289B98C51FBCCAFFD" originalValue="Río">Rio</normalizedToken>
Negro drainage (middle
<normalizedToken id="150440F725F890FB82D02F035563BC10" originalValue="Río">Rio</normalizedToken>
Amazonas drainage) and upper
<normalizedToken id="BE5B7F3F0AAB5D31C322F92B3E1BDCE9" originalValue="Río">Rio</normalizedToken>
Orinoco drainage of Venezuela. (Fig. 8).
<taxonomicName id="15350F088D42C3F71B1019FBDF02A326" LSID-ZBK="urn:lsid:zoobank.org:act:4125CE0B-B466-47CA-B90F-CF9073FEE6F8" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus hemicochliodon Armbruster 2003:29" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="hemicochliodon">Hypostomus hemicochliodon</taxonomicName>
is sympatric with
<taxonomicName id="3962EF94A4AB2BC65F9812485110017F" LSID-ZBK="urn:lsid:zoobank.org:act:C7C2629A-1686-4E4E-9E15-59B1D1162CBE" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus ericius Armbruster 2003:25" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="ericius">H. ericius</taxonomicName>
,
<taxonomicName id="0A269EB918B672729DD0BFDB90762ED0" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="oculeus">H. oculeus</taxonomicName>
,
<taxonomicName id="6570E43B4A3E4D606A0B271121D066AA" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="pyrineusi">H. pyrineusi</taxonomicName>
, and
<taxonomicName id="D3255FE085A454B17B9E13B2AA3F316D" LSID-ZBK="urn:lsid:zoobank.org:act:3F8E55C1-0EAD-49CA-B887-0D953CC48DA0" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus sculpodon Armbruster 2003:49" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="sculpodon">H. sculpodon</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="ED9F135540BF8283064AEA12FB601CDD" type="etymology">
<paragraph id="184560CCE5292F2582494179542E0E43" pageNumber="31">
Etymology: From the Latin, hemi, for half, the Latin, coclea (coch-), for spoon, and the Greek, odon, for tooth in reference to the fact that the teeth are about half as spoonshaped as those of wood-specializing members of the
<taxonomicName id="554E376607ECD35DD40320C6D7C4970B" LSID-ZBK="urn:lsid:zoobank.org:act:7520B3CD-59C7-4425-A30E-1336A0B2AADD" family="Loricariidae" genus="Hypostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hypostomus cochliodon Kner 1854:265 [15]" order="Siluriformes" pageNumber="31" phylum="Chordata" rank="species" species="cochliodon">Hypostomus cochliodon</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>