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<document id="828CA9547D8F410254976229207F3085" ID-CLB-Dataset="56250" ID-DOI="10.11646/phytotaxa.603.2.5" ID-GBIF-Dataset="c60d3785-c9d5-4545-9c23-f9cbbb70b72c" ID-ISSN="1179-3163" ID-Zenodo-Dep="8157964" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1689662273530" checkinUser="plazi" docAuthor="Murillo-A, José &amp; Triana-Moreno, Luz A." docDate="2023" docId="B90D5721BB75372BFF018D34FEFDE8AD" docLanguage="en" docName="phytotaxa.603.2.5.pdf" docOrigin="Phytotaxa 603 (2)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Parablechnum prostratum J. Murillo &amp; L. A. Triana 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="188" masterDocId="45342F59BB75372FFF898A75FFA7EA1D" masterDocTitle="A new species of Parablechnum (Blechnaceae) with prostrate leaves" masterLastPageNumber="190" masterPageNumber="184" pageNumber="184" updateTime="1699541105910" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D4323D5AFD5917C535D561A205518D8C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A0319725C4FA84FBABD21E9FE4E04690">
<mods:title id="6E06B8A93D083FF3763F6FC6B2B0AA93">A new species of Parablechnum (Blechnaceae) with prostrate leaves</mods:title>
</mods:titleInfo>
<mods:name id="30A6D7DD09DAE92E9FED2A6B9CF5AF03" type="personal">
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<mods:namePart id="E423AA058903E6FFA2F9FA4AB1534724">Murillo-A, José</mods:namePart>
<mods:affiliation id="2D737FF2BF7CB6102B224AEFEADDA993">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, Colombia.</mods:affiliation>
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<mods:role id="D0076E5FFC288D4BF1101E434356FA07">
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<mods:namePart id="3A918D931F99FEDD0118681C6D017C65">Triana-Moreno, Luz A.</mods:namePart>
<mods:affiliation id="007F5A6E8C805BBA0D649E2E9E33A983">Departamento de Ciencias Biológicas, Universidad de Caldas, Manizales, Colombia.</mods:affiliation>
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<mods:title id="24EFB2C5497C4EDC9424D1FD487F46D3">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="661ADECE86A876E5F931A9F48E1F9AD7">
<mods:date id="2DC5B0067049E590813101B9822C4621">2023</mods:date>
<mods:detail id="7046833BB78971F935A7CDD008C75140" type="pubDate">
<mods:number id="A25B9841637125E6C470E7325B8AE6EA">2023-07-18</mods:number>
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<mods:number id="5F8C2C4A1BB820D929DA113B189C373C">603</mods:number>
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<mods:number id="E050ED0CC5D418A44F8DE9C35F47460F">2</mods:number>
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<mods:url id="E35E10C6C4E6F6641DAA7A8A78648ED1">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
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<mods:classification id="A76488D5325232F2C9CA03271D598D6D">journal article</mods:classification>
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<mods:identifier id="8F1682E877CBADBB5D30D7F0D3D92846" type="DOI">10.11646/phytotaxa.603.2.5</mods:identifier>
<mods:identifier id="2C671ADCBF587D720F34F8DBD9C2997B" type="GBIF-Dataset">c60d3785-c9d5-4545-9c23-f9cbbb70b72c</mods:identifier>
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<treatment id="B90D5721BB75372BFF018D34FEFDE8AD" ID-DOI="http://doi.org/10.5281/zenodo.8164795" ID-GBIF-Taxon="212415708" ID-Zenodo-Dep="8164795" LSID="urn:lsid:plazi:treatment:B90D5721BB75372BFF018D34FEFDE8AD" httpUri="http://treatment.plazi.org/id/B90D5721BB75372BFF018D34FEFDE8AD" lastPageId="4" lastPageNumber="188" pageId="0" pageNumber="184">
<subSubSection id="79BEB5BCBB75372FFF018D34FCDEED41" box="[136,889,1857,1884]" pageId="0" pageNumber="184" type="nomenclature">
<paragraph id="311BE637BB75372FFF018D34FCDEED41" blockId="0.[136,1451,1857,1953]" box="[136,889,1857,1884]" pageId="0" pageNumber="184">
<emphasis id="03D03A25BB75372FFF018D34FCDEED41" bold="true" box="[136,889,1857,1884]" pageId="0" pageNumber="184">
<taxonomicName id="F6A49DB4BB75372FFF018D34FCABED46" ID-CoL="BTDN6" authority="J. Murillo &amp; L.A. Triana" authorityName="J. Murillo &amp; L. A. Triana" authorityYear="2023" box="[136,780,1857,1884]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="0" pageNumber="184" phylum="Tracheophyta" rank="species" species="prostratum" status="sp. nov.">
<emphasis id="03D03A25BB75372FFF018D34FE67ED41" bold="true" box="[136,448,1857,1884]" italics="true" pageId="0" pageNumber="184">Parablechnum prostratum</emphasis>
J. Murillo &amp; L.A. Triana
</taxonomicName>
,
<emphasis id="03D03A25BB75372FFC978D37FCDEED41" bold="true" box="[798,889,1858,1884]" italics="true" pageId="0" pageNumber="184">
<taxonomicNameLabel id="18E3875EBB75372FFC978D37FCDEED41" box="[798,889,1858,1884]" pageId="0" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="79BEB5BCBB75372FFC0F8D37FC5BED41" box="[902,1020,1858,1884]" pageId="0" pageNumber="184" type="description">
<paragraph id="311BE637BB75372FFC0F8D37FC5BED41" blockId="0.[136,1451,1857,1953]" box="[902,1020,1858,1884]" pageId="0" pageNumber="184">(Fig. 1,2).</paragraph>
</subSubSection>
<subSubSection id="79BEB5BCBB75372FFB818D37FB51EDBC" pageId="0" pageNumber="184" type="materials_examined">
<paragraph id="311BE637BB75372FFB818D37FB51EDBC" blockId="0.[136,1451,1857,1953]" pageId="0" pageNumber="184">
<materialsCitation id="81CCEC6ABB75372FFB818D37FC79EDBC" ID-GBIF-Occurrence="4161799301" collectingDate="2023-02-22" collectionCode="COL!" collectorName=" J. Murillo" country="Colombia" county="Type" elevation="2218" latitude="1.0851945" location="Mocoa" longLatPrecision="1" longitude="-76.832886" municipality="San Francisco" pageId="0" pageNumber="184" specimenCount="1" stateProvince="Putumayo" typeStatus="holotype">
<collectingCounty id="D87A9EBBBB75372FFB818D37FBE3ED41" box="[1032,1092,1858,1884]" pageId="0" pageNumber="184">Type</collectingCounty>
:
<collectingCountry id="49B3A6A7BB75372FFBDD8D37FB4AED41" box="[1108,1261,1858,1884]" name="Colombia" pageId="0" pageNumber="184">COLOMBIA</collectingCountry>
.
<collectingRegion id="F36028D5BB75372FFB758D37FAD7ED41" box="[1276,1392,1858,1884]" country="Colombia" name="Putumayo" pageId="0" pageNumber="184">Putumayo</collectingRegion>
,
<collectingMunicipality id="D17F7C4DBB75372FFA088D37FF5FED63" pageId="0" pageNumber="184">San Francisco</collectingMunicipality>
, carretera
<collectorName id="9C5183E1BB75372FFEF88D11FE75ED62" box="[369,466,1892,1919]" pageId="0" pageNumber="184">de Pasto</collectorName>
a
<location id="347BB0ECBB75372FFE788D10FDE5ED62" LSID="urn:lsid:plazi:treatment:B90D5721BB75372BFF018D34FEFDE8AD:347BB0ECBB75372FFE788D10FDE5ED62" box="[497,578,1893,1919]" country="Colombia" county="Type" latitude="1.0851945" longLatPrecision="1" longitude="-76.832886" municipality="San Francisco" name="Mocoa" pageId="0" pageNumber="184" stateProvince="Putumayo">Mocoa</location>
,
<location id="347BB0ECBB75372FFDC68D11FC8CED62" LSID="urn:lsid:plazi:treatment:B90D5721BB75372BFF018D34FEFDE8AD:347BB0ECBB75372FFDC68D11FC8CED62" box="[591,811,1892,1919]" country="Colombia" county="Type" latitude="1.0851945" longLatPrecision="1" longitude="-76.832886" municipality="San Francisco" name="Vereda Los Monos" pageId="0" pageNumber="184" stateProvince="Putumayo">Vereda Los Monos</location>
, km 93–94, cerca al restaurante
<collectorName id="9C5183E1BB75372FFB2A8D11FA85ED63" box="[1187,1314,1892,1918]" pageId="0" pageNumber="184">El Refugio</collectorName>
,
<geoCoordinate id="549080F0BB75372FFAB98D11FA01ED62" box="[1328,1446,1892,1919]" degrees="1" direction="north" minutes="5" orientation="latitude" pageId="0" pageNumber="184" precision="1" seconds="6.7" value="1.0851945">1°5’6.7”N</geoCoordinate>
,
<geoCoordinate id="549080F0BB75372FFF018DF3FE8AEDBC" box="[136,301,1926,1953]" degrees="76" direction="west" minutes="49" orientation="longitude" pageId="0" pageNumber="184" precision="1" seconds="58.4" value="-76.832886">76°49’58.4”W</geoCoordinate>
,
<quantity id="F65C4BD2BB75372FFEB18DF2FE28EDBC" box="[312,399,1927,1953]" metricMagnitude="3" metricUnit="m" metricValue="2.218" pageId="0" pageNumber="184" unit="m" value="2218.0">
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<collectorName id="9C5183E1BB75372FFDA78DF2FD38EDBD" box="[558,671,1926,1952]" pageId="0" pageNumber="184">J. Murillo</collectorName>
et al. 4901
</emphasis>
(
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,
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<materialsCitation id="81CCEC6ABB75372FFC6C8DF2FB54EDBC" ID-GBIF-Occurrence="4161799307" box="[997,1267,1927,1953]" collectingDate="2023-02-22" collectionCode="FAUC, PSO" collectorName="J. Murillo" country="Colombia" elevation="2218" latitude="1.0851945" location="Mocoa" longLatPrecision="1" longitude="-76.832886" municipality="San Francisco" pageId="0" pageNumber="184" specimenCount="1" stateProvince="Putumayo" typeStatus="holotype">Isotypes FAUC!, PSO!)</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="79BEB5BCBB75372DFF018DBEFA94EE99" lastPageId="2" lastPageNumber="186" pageId="0" pageNumber="184" type="diagnosis">
<paragraph id="311BE637BB75372EFF018DBEFD0FEAC6" blockId="0.[136,1451,1995,2091]" lastBlockId="1.[136,1451,158,219]" lastPageId="1" lastPageNumber="185" pageId="0" pageNumber="184">
Diagnosis:—
<taxonomicName id="F6A49DB4BB75372FFE958DBEFDE1EDF8" authorityName="J. Murillo &amp; L. A. Triana" authorityYear="2023" box="[284,582,1995,2021]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="0" pageNumber="184" phylum="Tracheophyta" rank="species" species="prostratum">
<emphasis id="03D03A25BB75372FFE958DBEFDE1EDF8" box="[284,582,1995,2021]" italics="true" pageId="0" pageNumber="184">Parablechnum prostratum</emphasis>
</taxonomicName>
is recognized by having prostrate, rosulate sterile leaves, appressed to the substrate, petioles and rachises densely covered with scales of variable size and shape and entire to dentate margin; generally cylindrical aerophores (1–)
<quantity id="F65C4BD2BB75372FFDBD8265FD35E236" box="[564,658,2064,2091]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="0" pageNumber="184" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long; pinnae revolute, close to each other, touching on their margins, and abaxially covered with abundant scales on the middle vein. The fertile leaf is erect, the pinnae linear with cordate, bulliform bases, wider than the rest of the pinna.
</paragraph>
<caption id="65DBB6BFBB74372EFF018D55FCB9ED47" ID-DOI="http://doi.org/10.5281/zenodo.8157966" ID-Zenodo-Dep="8157966" httpUri="https://zenodo.org/record/8157966/files/figure.png" pageId="1" pageNumber="185" startId="1.[136,234,1824,1846]" targetBox="[185,1408,275,1793]" targetPageId="1" targetType="figure">
<paragraph id="311BE637BB74372EFF018D55FCB9ED47" blockId="1.[136,1452,1824,1882]" pageId="1" pageNumber="185">
<emphasis id="03D03A25BB74372EFF018D55FEA3ED28" bold="true" box="[136,260,1824,1846]" pageId="1" pageNumber="185">FIGURE 1.</emphasis>
<taxonomicName id="F6A49DB4BB74372EFE838D55FDAFED2B" authorityName="J. Murillo &amp; L. A. Triana" authorityYear="2023" box="[266,520,1824,1846]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="185" phylum="Tracheophyta" rank="species" species="prostratum">
<emphasis id="03D03A25BB74372EFE838D55FDAFED2B" box="[266,520,1824,1846]" italics="true" pageId="1" pageNumber="185">Parablechnum prostratum</emphasis>
</taxonomicName>
. A. Habit, B. Detail of the fertile pinnae, C. Rhizome scale, D. Petiole scale, E-F. Pinnae scales, G. Aerophore (
<emphasis id="03D03A25BB74372EFE908D30FE42ED44" box="[281,485,1860,1882]" italics="true" pageId="1" pageNumber="185">J. Murillo et al. 4901</emphasis>
COL). Drawing by M. Morales.
</paragraph>
</caption>
<paragraph id="311BE637BB77372DFF348AEBFA94EE99" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="186">
Terrestrial or rupicolous, rhizome erect,
<emphasis id="03D03A25BB77372DFD018AEAFD05EAA5" box="[648,674,159,184]" italics="true" pageId="2" pageNumber="186">ca</emphasis>
.
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long; densely covered with ovate 6–11 (–25) × 2–4 (–5) mm, brown, with long-acuminate apex, basifixed, entire, opaque scales. Dimorphic leaves, the sterile ones prostrate, rosulate; petioles 3–11 (–27) cm in length, 1.5–2 (–10) mm in diameter, adaxially grooved, light brown to dark brown, transversely papillose, sometimes smooth; densely covered with ovate to ovate-lanceolate scales of two sizes, some (6–) 9–10 (–25) × (1.5) 2–2.5 (–5) mm, others 1–2 × 0.2–0.6 (–1) mm, brown, with acuminate to long-acuminate apex, margins entire to denticulate, with sparse teeth
<emphasis id="03D03A25BB77372DFD6A8B26FD5AEB71" box="[739,765,339,364]" italics="true" pageId="2" pageNumber="186">ca</emphasis>
.
<quantity id="F65C4BD2BB77372DFC858B27FCC4EB70" box="[780,867,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="186" unit="mm" value="0.2">0.2 mm</quantity>
long; laminae 23– 32 (–40) × 6.5–10.5 (–13.5) cm, pinnate, elliptical; rachis stramineous, sometimes atropurpureous, transversely papillose; densely covered with brown, ovate scales of
<specimenCount id="27A22DBEBB77372DFEB18BEFFE10EBA8" box="[312,439,410,437]" pageId="2" pageNumber="186" type="generic" typeStatus="types">three types</specimenCount>
: (i) linear-ovate, 7–9 ×
<quantity id="F65C4BD2BB77372DFD4F8BEFFC9CEBA8" box="[710,827,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="186" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
, with filiform apex, margin entire or with sparse teeth, (ii) ovate, 3–3.5 ×
<quantity id="F65C4BD2BB77372DFED28BCBFE43EBC4" box="[347,484,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="2" pageNumber="186" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
, with dentate margin; (iii) linear, 1–1.5 ×
<quantity id="F65C4BD2BB77372DFC348BCBFBE1EBC4" box="[957,1094,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="186" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
, with sparse, long teeth
<quantity id="F65C4BD2BB77372DFADE8BCBFF13EBE0" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="186" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
long; buds absent, aerophores present, cylindrical or sometimes bacillate or reniform,
<emphasis id="03D03A25BB77372DFB2D8B96FB61EBE1" box="[1188,1222,483,508]" italics="true" pageId="2" pageNumber="186">ca.</emphasis>
(1–)
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long, shorter distally; pinnae (11–) 23–32 (–41) pairs, close to each other, usually touching each other, perpendicular to the rachis, sometimes the basal pair slightly deflected; oblong, falcate, 3–5 ×
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, chartaceous, with rounded apex, cordate base, equilateral, margin entire, sometimes serrated, revolute, sometimes strongly revolute when dry; adaxially covered with abundant to sparse, linear to rounded scales
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, with long marginal teeth
<quantity id="F65C4BD2BB77372DFB568807FACEE890" box="[1247,1385,626,653]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="2" pageNumber="186" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
long; middle vein abaxially densely covered with ovate scales, 2–3 × (0.5–)
<quantity id="F65C4BD2BB77372DFC2B88E3FBB1E8AC" box="[930,1046,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="186" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
, apex acuminate to long-acuminate, with sparse teeth, distally covered with abundant scales similar to those adaxially present; petiolules stramineous, sometimes adaxially atropurpureous in basal pinnae,
<quantity id="F65C4BD2BB77372DFD7888ABFCF6E8E4" box="[753,849,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="186" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long; veins free, simple, sometimes furcate at their bases, sometimes adaxially elevated when dry, flat to immersed abaxially, ending in conspicuous hydathodes. Fertile leaves erect, with petioles proximally dark brown, yellowish with dark brown blotches in the middle portions, and stramineous distally, sometimes dark brown through their entire length, 7–10 (–26) cm long, 3–4.5 (–12) mm diam., adaxially grooved, densely covered with scales similar to those on the petioles of sterile leaves; laminae 17–23 (–41) × 4–5.5 (–9) cm, pinnate, elliptical, with truncate base, covered with abundant scales similar to those of sterile laminae; rachis stramineous, adaxially grooved, with cylindrical aerophores
<quantity id="F65C4BD2BB77372DFCF989C3FC62E9CC" box="[880,965,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="186" unit="mm" value="2.2">2.2 mm</quantity>
long, distal ones
<quantity id="F65C4BD2BB77372DFB0C89C3FB7EE9CC" box="[1157,1241,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="186" unit="mm" value="1.2">1.2 mm</quantity>
long; pinnae (16–) 19–29 (–35) pairs; linear, 2–3 ×
<quantity id="F65C4BD2BB77372DFE7389AFFDDCE9E8" box="[506,635,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="186" unit="cm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 cm</quantity>
, chartaceous, with obtuse to rounded apex, base cordate, wider than the rest of the pinna, bulliform,
<quantity id="F65C4BD2BB77372DFE4E898BFD82EE04" box="[455,549,1022,1049]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="186" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
wide, equilateral; margin entire, densely covered with scales similar to those of the sterile pinnae; petiolules dark brown
<quantity id="F65C4BD2BB77372DFDC38E57FD01EE20" box="[586,678,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="186" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long; veins free, inconspicuous, ending in inconspicuous hydathodes. Indusium elongated, brown, erose, glabrous to abundantly covered with linear scales
<quantity id="F65C4BD2BB77372DFBE58E33FB50EE7C" box="[1132,1271,1094,1121]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="2" pageNumber="186" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
long, with few medial and basal prolongations
<emphasis id="03D03A25BB77372DFE658E1EFDA9EE99" box="[492,526,1131,1156]" italics="true" pageId="2" pageNumber="186">ca.</emphasis>
<quantity id="F65C4BD2BB77372DFD9C8E1FFDCCEE98" box="[533,619,1130,1157]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="186" unit="mm" value="0.5">0.5 mm</quantity>
long. Spores light brown, perine reticulated, with large areoles.
</paragraph>
</subSubSection>
<subSubSection id="79BEB5BCBB77372DFF348EFBFBEDEF24" pageId="2" pageNumber="186" type="biology_ecology">
<paragraph id="311BE637BB77372DFF348EFBFBEDEF24" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="186">
<emphasis id="03D03A25BB77372DFF348EFBFE4CEEB5" bold="true" box="[189,491,1166,1192]" pageId="2" pageNumber="186">Habitat and distribution:</emphasis>
—
<location id="347BB0ECBB77372DFD818EFBFD9EEEB5" LSID="urn:lsid:plazi:treatment:B90D5721BB75372BFF018D34FEFDE8AD:347BB0ECBB77372DFD818EFBFD9EEEB5" box="[520,569,1166,1192]" country="Colombia" county="Andean forest" municipality="Southern" name="This" pageId="2" pageNumber="186" stateProvince="Cauca">This</location>
species has been recorded in the
<collectingRegion id="F36028D5BB77372DFC248EFBFC51EEB5" box="[941,1014,1166,1192]" country="Colombia" name="Cauca" pageId="2" pageNumber="186">Cauca</collectingRegion>
,
<collectingRegion id="F36028D5BB77372DFB888EFBFBF5EEB5" box="[1025,1106,1166,1192]" country="Colombia" name="Narino" pageId="2" pageNumber="186">Nariño</collectingRegion>
, and
<collectingRegion id="F36028D5BB77372DFB058EFAFB59EEB4" box="[1164,1278,1167,1193]" country="Colombia" name="Putumayo" pageId="2" pageNumber="186">Putumayo</collectingRegion>
departments of
<collectingMunicipality id="D17F7C4DBB77372DFF018EC7FF4AEED1" box="[136,237,1202,1228]" pageId="2" pageNumber="186">Southern</collectingMunicipality>
<collectingCountry id="49B3A6A7BB77372DFF7D8EC7FEC5EED1" box="[244,354,1202,1228]" name="Colombia" pageId="2" pageNumber="186">Colombia</collectingCountry>
(
<figureCitation id="A99FFAB2BB77372DFEFB8EC7FE12EED1" box="[370,437,1202,1228]" captionStart="FIGURE 3" captionStartId="4.[136,229,2058,2080]" captionTargetBox="[158,1430,713,2034]" captionTargetId="figure-283@4.[158,1430,713,2034]" captionTargetPageId="4" captionText="FIGURE 3. Distribution map of Parablechnum prostratum. Modified from QGIS v 3.18.3-Zürich (http://qgis.org)." figureDoi="http://doi.org/10.5281/zenodo.8157970" httpUri="https://zenodo.org/record/8157970/files/figure.png" pageId="2" pageNumber="186">Fig. 3</figureCitation>
), between the
<collectingCounty id="D87A9EBBBB77372DFDD18EC7FD51EED1" box="[600,758,1202,1228]" pageId="2" pageNumber="186">Andean forest</collectingCounty>
and paramo, at elevations from
<quantity id="F65C4BD2BB77372DFBD68EC7FA86EED0" box="[1119,1313,1202,1229]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="3.4" metricValueMin="1.5" pageId="2" pageNumber="186" unit="m" value="2450.0" valueMax="3400.0" valueMin="1500.0">
<elevation id="BA890104BB77372DFBD68EC7FA86EED0" box="[1119,1313,1202,1229]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="3.4" metricValueMin="1.5" pageId="2" pageNumber="186" unit="m" value="2450.0" valueMax="3400.0" valueMin="1500.0">1,500 to 3,400 m</elevation>
</quantity>
. It grows on cliffs in zones with high humidity and associated with a dense cover of bryophytes. The rhizome is erect,
<emphasis id="03D03A25BB77372DFA998EA2FA8DEEED" box="[1296,1322,1239,1264]" italics="true" pageId="2" pageNumber="186">ca</emphasis>
.
<quantity id="F65C4BD2BB77372DFABE8EA3FACAEEEC" box="[1335,1389,1238,1265]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="186" unit="cm" value="5.0">5 cm</quantity>
long, making its way between associated bryophytes, so that leaves grow prostrate on the bryophyte layer. It is possible that leaves look pendulous, but this is due to the strong slope of the places where it grows.
</paragraph>
</subSubSection>
<subSubSection id="79BEB5BCBB77372DFF348F37FF76EF9C" pageId="2" pageNumber="186" type="etymology">
<paragraph id="311BE637BB77372DFF348F37FF76EF9C" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="186">
<emphasis id="03D03A25BB77372DFF348F37FEEFEF41" bold="true" box="[189,328,1346,1372]" pageId="2" pageNumber="186">Etymology:</emphasis>
—The species epithet refers to the prostrate growth of sterile leaves, a unique character within the genus.
</paragraph>
</subSubSection>
<subSubSection id="79BEB5BCBB77372DFF348FFFFF6BECBD" pageId="2" pageNumber="186" type="materials_examined">
<paragraph id="311BE637BB77372DFF348FFFFF6BECBD" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="186">
<emphasis id="03D03A25BB77372DFF348FFFFE63EFB9" bold="true" box="[189,452,1418,1444]" pageId="2" pageNumber="186">Examined specimens:</emphasis>
—
<materialsCitation id="81CCEC6ABB77372DFE688FFFFF7DEFD5" ID-GBIF-Occurrence="4161799305" collectingDate="1961-10-04" collectingDateMax="1967-09-22" collectingDateMin="1961-10-04" collectionCode="COL" collectorName="W. Hagemann &amp; R. Tryon" country="Colombia" county="Paramo de Purace" elevation="3400" location="Popayan" municipality="Purace" pageId="2" pageNumber="186" specimenCount="1" stateProvince="Cauca">
<collectingCountry id="49B3A6A7BB77372DFE688FFFFDDEEFB9" box="[481,633,1418,1444]" name="Colombia" pageId="2" pageNumber="186">COLOMBIA</collectingCountry>
.
<emphasis id="03D03A25BB77372DFD0F8FFFFD7BEFB9" bold="true" box="[646,732,1418,1444]" pageId="2" pageNumber="186">
<collectingRegion id="F36028D5BB77372DFD0F8FFFFD71EFB9" box="[646,726,1418,1444]" country="Colombia" name="Cauca" pageId="2" pageNumber="186">Cauca</collectingRegion>
:
</emphasis>
<collectingCounty id="D87A9EBBBB77372DFD6F8FFFFC1DEFB9" box="[742,954,1418,1444]" pageId="2" pageNumber="186">Páramo de Puracé</collectingCounty>
,
<quantity id="F65C4BD2BB77372DFC418FFFFB84EFB8" box="[968,1059,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="186" unit="m" value="3400.0">
<elevation id="BA890104BB77372DFC418FFFFB84EFB8" box="[968,1059,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="186" unit="m" value="3400.0">3400 m</elevation>
</quantity>
,
<date id="451AC0F7BB77372DFBBB8FFFFB6FEFB9" box="[1074,1224,1418,1444]" pageId="2" pageNumber="186" value="1967-09-22">
<collectingDate id="555E391FBB77372DFBBB8FFFFB6FEFB9" box="[1074,1224,1418,1444]" pageId="2" pageNumber="186" value="1967-09-22">22 Sep 1967</collectingDate>
</date>
,
<emphasis id="03D03A25BB77372DFB5F8FFEFA0CEFB9" box="[1238,1451,1418,1444]" italics="true" pageId="2" pageNumber="186">
<collectorName id="9C5183E1BB77372DFB5F8FFEFAD0EFB9" box="[1238,1399,1419,1444]" pageId="2" pageNumber="186">W. Hagemann</collectorName>
561
</emphasis>
(
<collectionCode id="57B57EF2BB77372DFF198FDBFF69EFD5" box="[144,206,1454,1480]" pageId="2" pageNumber="186">COL</collectionCode>
);
</materialsCitation>
<materialsCitation id="81CCEC6ABB77372DFF6B8FDBFB7DEFD5" ID-GBIF-Occurrence="4161799308" box="[226,1242,1454,1481]" collectingDate="1961-10-04" collectingDateMax="1967-09-22" collectingDateMin="1961-10-04" collectionCode="COL" collectorName="W. Hagemann &amp; R. Tryon" country="Colombia" county="Paramo de Purace" elevation="3400" location="Popayan" municipality="Purace" pageId="2" pageNumber="186" specimenCount="1" stateProvince="Cauca">
<quantity id="F65C4BD2BB77372DFF6B8FDBFE8EEFD5" box="[226,297,1454,1480]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="186" unit="km" value="10.0">
<locationDeviation id="20A9FF15BB77372DFF6B8FDBFE8EEFD5" box="[226,297,1454,1480]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="186" unit="km" value="10.0">10 km</locationDeviation>
</quantity>
east of
<collectingMunicipality id="D17F7C4DBB77372DFE088FDBFE6AEFD5" box="[385,461,1454,1480]" pageId="2" pageNumber="186">Puracé</collectingMunicipality>
(
<quantity id="F65C4BD2BB77372DFE558FDBFD83EFD5" box="[476,548,1454,1480]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="186" unit="km" value="27.0">
<locationDeviation id="20A9FF15BB77372DFE558FDBFD83EFD5" box="[476,548,1454,1480]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="186" unit="km" value="27.0">27 km</locationDeviation>
</quantity>
east of
<location id="347BB0ECBB77372DFDF58FDBFD46EFD5" LSID="urn:lsid:plazi:treatment:B90D5721BB75372BFF018D34FEFDE8AD:347BB0ECBB77372DFDF58FDBFD46EFD5" box="[636,737,1454,1480]" country="Colombia" county="Paramo de Purace" municipality="Purace" name="Popayan" pageId="2" pageNumber="186" stateProvince="Cauca">Popayán</location>
),
<quantity id="F65C4BD2BB77372DFD7D8FDBFCECEFD4" box="[756,843,1454,1481]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="186" unit="m" value="3200.0">
<elevation id="BA890104BB77372DFD7D8FDBFCECEFD4" box="[756,843,1454,1481]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="186" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="451AC0F7BB77372DFCDE8FDBFC71EFD5" box="[855,982,1454,1480]" pageId="2" pageNumber="186" value="1961-10-04">
<collectingDate id="555E391FBB77372DFCDE8FDBFC71EFD5" box="[855,982,1454,1480]" pageId="2" pageNumber="186" value="1961-10-04">4 Oct 1961</collectingDate>
</date>
,
<emphasis id="03D03A25BB77372DFC6B8FDAFB26EFD5" box="[994,1153,1454,1480]" italics="true" pageId="2" pageNumber="186">
<collectorName id="9C5183E1BB77372DFC6B8FDAFBE5EFD5" box="[994,1090,1455,1480]" pageId="2" pageNumber="186">R. Tryon</collectorName>
5961
</emphasis>
(
<collectionCode id="57B57EF2BB77372DFB198FDBFB69EFD5" box="[1168,1230,1454,1480]" pageId="2" pageNumber="186">COL</collectionCode>
).
</materialsCitation>
<materialsCitation id="81CCEC6ABB77372DFB688FDBFF7DEC0D" ID-GBIF-Occurrence="4161799309" collectingDate="1943-10-04" collectingDateMax="1943-10-06" collectingDateMin="1943-10-04" collectionCode="COL" collectorName="F. Fosberg" elevation="3200" location="Cerro Gualcala" municipality="Cerro Gualcala" pageId="2" pageNumber="186" specimenCount="1" stateProvince="Narino">
<emphasis id="03D03A25BB77372DFB688FDBFA99EFD5" bold="true" box="[1249,1342,1454,1480]" pageId="2" pageNumber="186">
<collectingRegion id="F36028D5BB77372DFB688FDBFA9EEFD5" box="[1249,1337,1454,1480]" country="Colombia" name="Narino" pageId="2" pageNumber="186">Nariño</collectingRegion>
:
</emphasis>
NE ridge of
<collectingMunicipality id="D17F7C4DBB77372DFF218FA7FEF2EFF1" box="[168,341,1490,1516]" country="Colombia" name="Cerro Gualcala" pageId="2" pageNumber="186" stateProvince="Narino">Cerro Gualcalá</collectingMunicipality>
,
<locationDeviation id="20A9FF15BB77372DFEEB8FA7FE41EFF1" bearing="0.0" box="[354,486,1490,1517]" distance="18000.0" pageId="2" pageNumber="186">
<quantity id="F65C4BD2BB77372DFEEB8FA7FE0DEFF1" box="[354,426,1490,1516]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="186" unit="km" value="18.0">18 km</quantity>
N of
</locationDeviation>
<locationDeviation id="20A9FF15BB77372DFE678FA6FD01EFF1" box="[494,678,1490,1517]" location="Piedrancha" pageId="2" pageNumber="186">E of Piedrancha</locationDeviation>
, 1º 7–11’N–77º 43–45’W,
<quantity id="F65C4BD2BB77372DFC5E8FA7FB88EFF0" box="[983,1071,1490,1517]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="186" unit="m" value="3200.0">
<elevation id="BA890104BB77372DFC5E8FA7FB88EFF0" box="[983,1071,1490,1517]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="186" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="451AC0F7BB77372DFBB58FA7FB7DEFF1" box="[1084,1242,1490,1516]" pageId="2" pageNumber="186" value="1943-10-04" valueMax="1943-10-06" valueMin="1943-10-04">
<collectingDate id="555E391FBB77372DFBB58FA7FB7DEFF1" box="[1084,1242,1490,1516]" pageId="2" pageNumber="186" value="1943-10-04" valueMax="1943-10-06" valueMin="1943-10-04">4–6 Oct 1943</collectingDate>
</date>
,
<emphasis id="03D03A25BB77372DFB6F8FA6FA0CEFF1" box="[1254,1451,1490,1516]" italics="true" pageId="2" pageNumber="186">
<collectorName id="9C5183E1BB77372DFB6F8FA6FAF8EFF1" box="[1254,1375,1490,1516]" pageId="2" pageNumber="186">F. Fosberg</collectorName>
21163
</emphasis>
(
<collectionCode id="57B57EF2BB77372DFF198F83FF69EC0D" box="[144,206,1526,1552]" pageId="2" pageNumber="186">COL</collectionCode>
).
</materialsCitation>
<materialsCitation id="81CCEC6ABB77372DFF688F83FAB7EC0D" ID-GBIF-Occurrence="4161799302" box="[225,1296,1526,1553]" collectingDate="1972-11-20" collectionCode="COL, PSO" collectorName="W. Hagemann" elevation="2050" location="El Pepino" municipality="El Pepino" pageId="2" pageNumber="186" specimenCount="1" stateProvince="Putumayo">
<emphasis id="03D03A25BB77372DFF688F83FEC2EC0D" bold="true" box="[225,357,1526,1552]" pageId="2" pageNumber="186">
<collectingRegion id="F36028D5BB77372DFF688F83FEC7EC0D" box="[225,352,1526,1552]" country="Colombia" name="Putumayo" pageId="2" pageNumber="186">Putumayo</collectingRegion>
:
</emphasis>
<quantity id="F65C4BD2BB77372DFEE48F83FE13EC0D" box="[365,436,1526,1553]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="186" unit="km" value="50.0">
<locationDeviation id="20A9FF15BB77372DFEE48F83FE13EC0D" box="[365,436,1526,1553]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="186" unit="km" value="50.0">50 km</locationDeviation>
</quantity>
west of
<collectingMunicipality id="D17F7C4DBB77372DFD9C8F83FD21EC0D" box="[533,646,1526,1552]" pageId="2" pageNumber="186">El Pepino</collectingMunicipality>
,
<quantity id="F65C4BD2BB77372DFD1B8F83FD4EEC0C" box="[658,745,1526,1553]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="2" pageNumber="186" unit="m" value="2050.0">
<elevation id="BA890104BB77372DFD1B8F83FD4EEC0C" box="[658,745,1526,1553]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="2" pageNumber="186" unit="m" value="2050.0">2050 m</elevation>
</quantity>
,
<date id="451AC0F7BB77372DFD7C8F83FC2CEC0D" box="[757,907,1526,1553]" pageId="2" pageNumber="186" value="1972-11-20">
<collectingDate id="555E391FBB77372DFD7C8F83FC2CEC0D" box="[757,907,1526,1553]" pageId="2" pageNumber="186" value="1972-11-20">20 Nov 1972</collectingDate>
</date>
,
<emphasis id="03D03A25BB77372DFC1E8F82FBD3EC0D" box="[919,1140,1526,1552]" italics="true" pageId="2" pageNumber="186">
<collectorName id="9C5183E1BB77372DFC1E8F82FB92EC0D" box="[919,1077,1527,1552]" pageId="2" pageNumber="186">W. Hagemann</collectorName>
1548
</emphasis>
(
<collectionCode id="57B57EF2BB77372DFB0D8F83FB67EC0D" box="[1156,1216,1526,1552]" pageId="2" pageNumber="186">COL</collectionCode>
,
<collectionCode id="57B57EF2BB77372DFB458F83FAA3EC0D" box="[1228,1284,1526,1552]" pageId="2" pageNumber="186">PSO</collectionCode>
);
</materialsCitation>
<materialsCitation id="81CCEC6ABB77372DFA918F82FABBEC29" ID-GBIF-Occurrence="4161799306" collectingDate="1967-11-27" collectionCode="COL" collectorName="Pasto-Mocoa &amp; L. E. Mora" elevation="1850" pageId="2" pageNumber="186" specimenCount="1">
<collectorName id="9C5183E1BB77372DFA918F82FA0BEC0C" box="[1304,1452,1527,1553]" pageId="2" pageNumber="186">Pasto-Mocoa</collectorName>
road,
<locationDeviation id="20A9FF15BB77372DFF4E8C6FFDDEEC29" box="[199,633,1562,1588]" firstLocation="El Mirador" pageId="2" pageNumber="186" secondLocation="San Francisco">between El Mirador and San Francisco</locationDeviation>
,
<quantity id="F65C4BD2BB77372DFD0A8C6FFCB8EC28" box="[643,799,1562,1589]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="2" pageNumber="186" unit="m" value="1850.0" valueMax="2200.0" valueMin="1500.0">
<elevation id="BA890104BB77372DFD0A8C6FFCB8EC28" box="[643,799,1562,1589]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="2" pageNumber="186" unit="m" value="1850.0" valueMax="2200.0" valueMin="1500.0">1500–2200 m</elevation>
</quantity>
,
<date id="451AC0F7BB77372DFCA38C6FFC19EC29" box="[810,958,1562,1589]" pageId="2" pageNumber="186" value="1967-11-27">
<collectingDate id="555E391FBB77372DFCA38C6FFC19EC29" box="[810,958,1562,1589]" pageId="2" pageNumber="186" value="1967-11-27">27 Nov 1967</collectingDate>
</date>
,
<emphasis id="03D03A25BB77372DFC418C6EFB63EC29" box="[968,1220,1562,1588]" italics="true" pageId="2" pageNumber="186">
<collectorName id="9C5183E1BB77372DFC418C6EFBE7EC29" box="[968,1088,1563,1588]" pageId="2" pageNumber="186">L. E. Mora</collectorName>
4455, 4467
</emphasis>
(
<collectionCode id="57B57EF2BB77372DFB5B8C6FFAB7EC29" box="[1234,1296,1562,1588]" pageId="2" pageNumber="186">COL</collectionCode>
);
</materialsCitation>
<materialsCitation id="81CCEC6ABB77372DFAAB8C6FFD43EC45" ID-GBIF-Occurrence="4161799303" collectingDate="1969-11-26" collectionCode="COL" collectorName="Mirador &amp; W. Schwabe" elevation="2000" pageId="2" pageNumber="186" specimenCount="1">
<collectorName id="9C5183E1BB77372DFAAB8C6FFA27EC29" box="[1314,1408,1562,1588]" pageId="2" pageNumber="186">Mirador</collectorName>
,
<emphasis id="03D03A25BB77372DFA038C6EFA03EC29" box="[1418,1444,1563,1588]" italics="true" pageId="2" pageNumber="186">ca</emphasis>
.
<quantity id="F65C4BD2BB77372DFF018C4BFF78EC44" box="[136,223,1598,1625]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="186" unit="m" value="2000.0">
<elevation id="BA890104BB77372DFF018C4BFF78EC44" box="[136,223,1598,1625]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="186" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="451AC0F7BB77372DFF638C4BFED8EC45" box="[234,383,1598,1625]" pageId="2" pageNumber="186" value="1969-11-26">
<collectingDate id="555E391FBB77372DFF638C4BFED8EC45" box="[234,383,1598,1625]" pageId="2" pageNumber="186" value="1969-11-26">26 Nov 1969</collectingDate>
</date>
,
<emphasis id="03D03A25BB77372DFE038C4AFD2CEC45" box="[394,651,1598,1624]" italics="true" pageId="2" pageNumber="186">
<collectorName id="9C5183E1BB77372DFE038C4AFDB7EC45" box="[394,528,1598,1624]" pageId="2" pageNumber="186">W. Schwabe</collectorName>
C1 69/077
</emphasis>
(
<collectionCode id="57B57EF2BB77372DFD138C4BFD7FEC45" box="[666,728,1598,1624]" pageId="2" pageNumber="186">COL</collectionCode>
);
</materialsCitation>
<materialsCitation id="81CCEC6ABB77372DFD628C4BFF6BECBD" ID-GBIF-Occurrence="4161799304" collectionCode="COL, FAUC, PSO" collectorName=" J. Murillo" elevation="2196" latitude="1.0844445" location="San Francisco" longLatPrecision="21" longitude="-76.83083" municipality="San Francisco" pageId="2" pageNumber="186" specimenCount="1">
<collectingMunicipality id="D17F7C4DBB77372DFD628C4BFC2BEC45" box="[747,908,1598,1624]" country="Colombia" latitude="1.0844445" longLatPrecision="21" longitude="-76.83083" municipality="El Pepino" name="San Francisco" pageId="2" pageNumber="186" stateProvince="Putumayo">San Francisco</collectingMunicipality>
, carretera
<collectorName id="9C5183E1BB77372DFC768C4BFBFAEC44" box="[1023,1117,1598,1625]" pageId="2" pageNumber="186">de Pasto</collectorName>
a
<collectorName id="9C5183E1BB77372DFBFE8C4AFB6FEC44" box="[1143,1224,1599,1625]" pageId="2" pageNumber="186">Mocoa</collectorName>
,
<collectorName id="9C5183E1BB77372DFB588C4BFA0FEC44" box="[1233,1448,1598,1625]" pageId="2" pageNumber="186">Vereda Los Monos</collectorName>
, km 93–94, cerca al restaurante
<collectorName id="9C5183E1BB77372DFE718C17FDDEEC61" box="[504,633,1634,1660]" pageId="2" pageNumber="186">El Refugio</collectorName>
,
<geoCoordinate id="549080F0BB77372DFD018C17FD4EEC60" box="[648,745,1634,1661]" degrees="1" direction="north" minutes="5" orientation="latitude" pageId="2" pageNumber="186" precision="15" seconds="4" value="1.0844445">1°5’4”N</geoCoordinate>
,
<geoCoordinate id="549080F0BB77372DFD708C17FC2FEC60" box="[761,904,1634,1661]" degrees="76" direction="west" minutes="49" orientation="longitude" pageId="2" pageNumber="186" precision="15" seconds="51" value="-76.83083">76°49’51”W</geoCoordinate>
,
<quantity id="F65C4BD2BB77372DFC1E8C17FC55EC60" box="[919,1010,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="2.196" pageId="2" pageNumber="186" unit="m" value="2196.0">
<elevation id="BA890104BB77372DFC1E8C17FC55EC60" box="[919,1010,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="2.196" pageId="2" pageNumber="186" unit="m" value="2196.0">2196 m</elevation>
</quantity>
,
<emphasis id="03D03A25BB77372DFB888C16FB59EC61" box="[1025,1278,1634,1660]" italics="true" pageId="2" pageNumber="186">
<collectorName id="9C5183E1BB77372DFB888C16FBD2EC61" box="[1025,1141,1634,1660]" pageId="2" pageNumber="186">J. Murillo</collectorName>
et al. 4900
</emphasis>
(
<collectionCode id="57B57EF2BB77372DFA988C17FAEAEC61" box="[1297,1357,1634,1660]" pageId="2" pageNumber="186">COL</collectionCode>
,
<collectionCode id="57B57EF2BB77372DFAD58C17FA0FEC61" box="[1372,1448,1634,1660]" pageId="2" pageNumber="186">FAUC</collectionCode>
,
<collectionCode id="57B57EF2BB77372DFF018CF3FF67ECBD" box="[136,192,1670,1696]" pageId="2" pageNumber="186">PSO</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="79BEB5BCBB77372BFF348CDFFEFDE8AD" lastPageId="4" lastPageNumber="188" pageId="2" pageNumber="186" type="discussion">
<paragraph id="311BE637BB77372CFF348CDFFB60EBC4" blockId="2.[136,1452,158,2093]" lastBlockId="3.[136,1452,158,473]" lastPageId="3" lastPageNumber="187" pageId="2" pageNumber="186">
<emphasis id="03D03A25BB77372DFF348CDFFEA7ECD9" bold="true" box="[189,256,1706,1732]" pageId="2" pageNumber="186">Notes</emphasis>
:—The new species described herein undoubtedly belongs to
<taxonomicName id="F6A49DB4BB77372DFC248CDFFBF5ECD9" box="[941,1106,1706,1732]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="186" phylum="Tracheophyta" rank="genus">
<emphasis id="03D03A25BB77372DFC248CDFFBF5ECD9" box="[941,1106,1706,1732]" italics="true" pageId="2" pageNumber="186">Parablechnum</emphasis>
</taxonomicName>
. It features defining characters for the genus: it has pinnate laminae, with petiolulate pinnae and conform apical pinnae, and lacks reduced pinnae at the base (
<figureCitation id="A99FFAB2BB77372DFF7A8C87FEE7ED11" box="[243,320,1778,1804]" captionStart="FIGURE 2" captionStartId="3.[136,229,1996,2018]" captionTargetBox="[151,1436,497,1971]" captionTargetId="figure-174@3.[151,1436,497,1971]" captionTargetPageId="3" captionText="FIGURE 2. Parablechnum prostratum. A. Habit, B. Juvenile sterile leaf, C. Fertile leaf, D. Petiole and lamina scales, E. Detail of the rachis showing the aerophores (white circles), F. Cross section of petiole, showing six meristeles (white arrows), G. Detail of the abaxial surface of fertile pinnae, H. Rhizome scale, I. Petiole scale, J-K. Pinna scales. Photos by J. Murillo and L.A. Triana." figureDoi="http://doi.org/10.5281/zenodo.8157968" httpUri="https://zenodo.org/record/8157968/files/figure.png" pageId="2" pageNumber="186">Figs. 2</figureCitation>
A-D) (
<bibRefCitation id="55359BC6BB77372DFE068C87FDC5ED11" author="Gasper, A. L. &amp; Dittrich, V. A. O. &amp; Salino, A." box="[399,610,1778,1804]" pageId="2" pageNumber="186" pagination="191 - 227" refId="ref3111" refString="Gasper, A. L., Dittrich, V. A. O. &amp; Salino, A. (2016) A classification for Blechnaceae (Polypodiales: Polypodiopsida): New genera, resurrected names, and combinations. Phytotaxa 275: 191 - 227. https: // doi. org / 10.11646 / phytotaxa. 275.3.1" type="journal article" year="2016">
Gasper
<emphasis id="03D03A25BB77372DFE618C86FD87ED11" box="[488,544,1778,1804]" italics="true" pageId="2" pageNumber="186">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="55359BC6BB77372DFDE48C87FCA3ED11" author="Molino, S." box="[621,772,1778,1804]" pageId="2" pageNumber="186" refId="ref3384" refString="Molino, S. (2022) Sinopsis global y aproximacion a la filogenia molecular del genero Parablechnum C. Presl (Blechnaceae, Polypodiopsida). Ph. D. Thesis, Universidad Complutense de Madrid. Madrid. 445 pp. [https: // eprints. ucm. es / id / eprint / 75407 / 1 / T 43509. pdf]" type="book" year="2022">Molino 2022</bibRefCitation>
). Another character that has also been found in some species of
<taxonomicName id="F6A49DB4BB77372DFF2F8D63FEECED2D" box="[166,331,1814,1840]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="186" phylum="Tracheophyta" rank="genus">
<emphasis id="03D03A25BB77372DFF2F8D63FEECED2D" box="[166,331,1814,1840]" italics="true" pageId="2" pageNumber="186">Parablechnum</emphasis>
</taxonomicName>
, and that is also in the new species proposed here, is the presence of six meristeles at the base of the petiole, two enlarged on the adaxial side and the remaining on the abaxial side smaller and arranged in an arc (
<figureCitation id="A99FFAB2BB77372DFAC28D4FFA04ED49" box="[1355,1443,1850,1876]" captionStart="FIGURE 2" captionStartId="3.[136,229,1996,2018]" captionTargetBox="[151,1436,497,1971]" captionTargetId="figure-174@3.[151,1436,497,1971]" captionTargetPageId="3" captionText="FIGURE 2. Parablechnum prostratum. A. Habit, B. Juvenile sterile leaf, C. Fertile leaf, D. Petiole and lamina scales, E. Detail of the rachis showing the aerophores (white circles), F. Cross section of petiole, showing six meristeles (white arrows), G. Detail of the abaxial surface of fertile pinnae, H. Rhizome scale, I. Petiole scale, J-K. Pinna scales. Photos by J. Murillo and L.A. Triana." figureDoi="http://doi.org/10.5281/zenodo.8157968" httpUri="https://zenodo.org/record/8157968/files/figure.png" pageId="2" pageNumber="186">Fig. 2 F</figureCitation>
) (
<bibRefCitation id="55359BC6BB77372DFF198D2BFEF8ED65" author="Rolleri, C. &amp; Prada, C. &amp; Passarelli, L. &amp; Galan, J. M." box="[144,351,1886,1912]" pageId="2" pageNumber="186" pagination="149 - 161" refId="ref3703" refString="Rolleri, C., Prada, C., Passarelli, L. &amp; Gabriel y Galan, J. M. (2012) Revision de dos especies de Blechnum de las regiones montanosas tropicales e intertropicales de Centroamerica y Sudamerica. Actes du Colloque Les Fougeres d'Alsace, d'Europe et du Monde 1: 149 - 161." type="journal article" year="2012">
Rolleri
<emphasis id="03D03A25BB77372DFF6C8D2AFEBBED65" box="[229,284,1886,1912]" italics="true" pageId="2" pageNumber="186">et al.</emphasis>
2012
</bibRefCitation>
).
<taxonomicName id="F6A49DB4BB77372DFEF98D2BFD31ED65" authorityName="J. Murillo &amp; L. A. Triana" authorityYear="2023" box="[368,662,1886,1912]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="186" phylum="Tracheophyta" rank="species" species="prostratum">
<emphasis id="03D03A25BB77372DFEF98D2BFD31ED65" box="[368,662,1886,1912]" italics="true" pageId="2" pageNumber="186">Parablechnum prostratum</emphasis>
</taxonomicName>
is unique within the genus because of the prostrate and rosulate growth of its sterile leaves, which are appressed to the substrate. The other species in
<taxonomicName id="F6A49DB4BB77372DFC728DF7FB07ED81" box="[1019,1184,1922,1948]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="186" phylum="Tracheophyta" rank="genus">
<emphasis id="03D03A25BB77372DFC728DF7FB07ED81" box="[1019,1184,1922,1948]" italics="true" pageId="2" pageNumber="186">Parablechnum</emphasis>
</taxonomicName>
with prostrate leaves is
<taxonomicName id="F6A49DB4BB77372DFF018DD2FE7BEDDD" authority="(G. Forst.) C. Presl" authorityName="C. Presl" baseAuthorityName="G. Forst." box="[136,476,1958,1984]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="186" phylum="Tracheophyta" rank="species" species="procerum">
<emphasis id="03D03A25BB77372DFF018DD2FEA8EDDD" box="[136,271,1959,1984]" italics="true" pageId="2" pageNumber="186">P. procerum</emphasis>
(G.Forst.) C.Presl
</taxonomicName>
from Oceania, with leaves with 1 to 12 pairs of pinnae (
<bibRefCitation id="55359BC6BB77372DFBC58DD3FAC0EDDD" author="Brownsey, P. J. &amp; Perrie, L. R." box="[1100,1383,1958,1984]" pageId="2" pageNumber="186" pagination="1 - 91" refId="ref2987" refString="Brownsey, P. J. &amp; Perrie, L. R. (2021) Blechnaceae In: Breitwieser, I. (Ed.) Flora of New Zealand: ferns and lycophytes, Fascicle 29. Manaaki Whenua Press, Lincoln, pp. 1 - 91." type="book chapter" year="2021">Brownsey &amp; Perrie 2021</bibRefCitation>
). This species could sometimes be prostrate, but in general it has suberect and arching leaves, even on the same plant both prostrate and suberect leaves can occur (
<bibRefCitation id="55359BC6BB77372DFDC38D9BFCC0E215" author="Brownsey, P. J. &amp; Perrie, L. R." box="[586,871,2030,2056]" pageId="2" pageNumber="186" pagination="1 - 91" refId="ref2987" refString="Brownsey, P. J. &amp; Perrie, L. R. (2021) Blechnaceae In: Breitwieser, I. (Ed.) Flora of New Zealand: ferns and lycophytes, Fascicle 29. Manaaki Whenua Press, Lincoln, pp. 1 - 91." type="book chapter" year="2021">Brownsey &amp; Perrie 2021</bibRefCitation>
). All other species of
<taxonomicName id="F6A49DB4BB77372DFBD18D9BFB5AE215" box="[1112,1277,2030,2056]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="186" phylum="Tracheophyta" rank="genus">
<emphasis id="03D03A25BB77372DFBD18D9BFB5AE215" box="[1112,1277,2030,2056]" italics="true" pageId="2" pageNumber="186">Parablechnum</emphasis>
</taxonomicName>
feature erect to arched leaves (
<bibRefCitation id="55359BC6BB77372DFEB98267FE6DE231" author="Molino, S." box="[304,458,2066,2092]" pageId="2" pageNumber="186" refId="ref3384" refString="Molino, S. (2022) Sinopsis global y aproximacion a la filogenia molecular del genero Parablechnum C. Presl (Blechnaceae, Polypodiopsida). Ph. D. Thesis, Universidad Complutense de Madrid. Madrid. 445 pp. [https: // eprints. ucm. es / id / eprint / 75407 / 1 / T 43509. pdf]" type="book" year="2022">Molino 2022</bibRefCitation>
). Within
<taxonomicName id="F6A49DB4BB77372DFDB88267FD66E231" box="[561,705,2066,2092]" class="Polypodiopsida" family="Blechnaceae" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="186" phylum="Tracheophyta" rank="family">Blechnaceae</taxonomicName>
, the prostrate distribution of the leaves has also been described in
<taxonomicName id="F6A49DB4BB76372CFF018AEBFEDCEAA5" box="[136,379,158,184]" class="Polypodiopsida" family="Blechnaceae" genus="Struthiopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="spicant">
<emphasis id="03D03A25BB76372CFF018AEBFEDCEAA5" box="[136,379,158,184]" italics="true" pageId="3" pageNumber="187">Struthiopteris spicant</emphasis>
</taxonomicName>
(L.) Weiss (
<bibRefCitation id="55359BC6BB76372CFD8F8AEBFD58EAA5" author="Rolleri, C. &amp; Prada, C." box="[518,767,158,184]" pageId="3" pageNumber="187" pagination="7 - 50" refId="ref3648" refString="Rolleri, C. &amp; Prada, C. (2006) Revision de los grupos de especies del genero Blechnum (Blechnaceae-Pteridophyta): El grupo B. pennamarina. Acta Botanica Malacitanica 31: 7 - 50. https: // doi. org / 10.24310 / abm. v 31 i 31.7119" type="journal article" year="2006">Rolleri &amp; Prada 2006</bibRefCitation>
), apparently a character that is not constant, because
<bibRefCitation id="55359BC6BB76372CFAD08AEBFEB3EAC1" author="Molino, S. &amp; Galan, J. M. &amp; Sessa, E. &amp; Wasowicz, P." pageId="3" pageNumber="187" pagination="185 - 198" refId="ref3323" refString="Molino, S. Gabriel y Galan, J. M., Sessa, E. &amp; Wasowicz, P. (2019) A multi-character analysis of Struthiopteris leads to the rescue of Spicantopsis (Blechnaceae, Polypodiopsida). Taxon 68: 185 - 198. https: // doi. org / 10.1002 / tax. 12036" type="journal article" year="2019">
Molino
<emphasis id="03D03A25BB76372CFF018AB6FF66EAC1" box="[136,193,194,220]" italics="true" pageId="3" pageNumber="187">et al.</emphasis>
(2019)
</bibRefCitation>
do not mention it for this species, but they do for
<taxonomicName id="F6A49DB4BB76372CFCDB8AB6FC47EAC0" box="[850,992,195,221]" class="Polypodiopsida" family="Blechnaceae" genus="Spicantopsis" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="genus">
<emphasis id="03D03A25BB76372CFCDB8AB6FC47EAC0" box="[850,992,195,221]" italics="true" pageId="3" pageNumber="187">Spicantopsis</emphasis>
</taxonomicName>
. Additionally, several species from
<collectingCountry id="49B3A6A7BB76372CFAFF8AB6FF45EB1D" name="New Zealand" pageId="3" pageNumber="187">New Zealand</collectingCountry>
have been described by
<bibRefCitation id="55359BC6BB76372CFD828A92FCE5EB1D" author="Brownsey, P. J. &amp; Perrie, L. R." box="[523,834,230,257]" pageId="3" pageNumber="187" pagination="1 - 91" refId="ref2987" refString="Brownsey, P. J. &amp; Perrie, L. R. (2021) Blechnaceae In: Breitwieser, I. (Ed.) Flora of New Zealand: ferns and lycophytes, Fascicle 29. Manaaki Whenua Press, Lincoln, pp. 1 - 91." type="book chapter" year="2021">Brownsey &amp; Perrie (2021)</bibRefCitation>
with sterile leaves in a prostrate to suberect rosette, including
<taxonomicName id="F6A49DB4BB76372CFF718B7FFC34EB39" authority="(Hook. f.) Gasper &amp; V. A. O. Dittrich" authorityName="Gasper &amp; V. A. O. Dittrich" baseAuthorityName="Hook. f." box="[248,915,266,292]" class="Polypodiopsida" family="Blechnaceae" genus="Austroblechnum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="banksii">
<emphasis id="03D03A25BB76372CFF718B7FFDA1EB39" box="[248,518,266,292]" italics="true" pageId="3" pageNumber="187">Austroblechnum banksii</emphasis>
(Hook.f.) Gasper &amp; V.A.O.Dittrich
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFC178B7FFE97EB55" authority="(R. Br.) Gasper &amp; V. A. O. Dittrich" authorityName="Gasper &amp; V. A. O. Dittrich" baseAuthorityName="R. Br." class="Polypodiopsida" family="Blechnaceae" genus="Austroblechnum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis id="03D03A25BB76372CFC178B7FFB42EB39" box="[926,1253,266,292]" italics="true" pageId="3" pageNumber="187">Austroblechnum lanceolatum</emphasis>
(R.Br.) Gasper &amp; V.A.O.Dittrich
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFEB48B5BFB1AEB55" authority="(Colenso ex Hook.) Gasper &amp; V. A. O. Dittrich" authorityName="Gasper &amp; V. A. O. Dittrich" baseAuthorityName="Hook." box="[317,1213,302,329]" class="Polypodiopsida" family="Blechnaceae" genus="Austroblechnum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="membranaceum">
<emphasis id="03D03A25BB76372CFEB48B5BFD08EB55" box="[317,687,302,328]" italics="true" pageId="3" pageNumber="187">Austroblechnum membranaceum</emphasis>
(Colenso ex Hook.) Gasper &amp; V.A.O.Dittrich
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFB438B5BFE5EEB71" authority="(R. Br.) Gasper &amp; V. A. O. Dittrich" authorityName="Gasper &amp; V. A. O. Dittrich" baseAuthorityName="R. Br." class="Polypodiopsida" family="Blechnaceae" genus="Cranfillia" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="fluviatilis">
<emphasis id="03D03A25BB76372CFB438B5BFA0BEB55" box="[1226,1452,302,328]" italics="true" pageId="3" pageNumber="187">Cranfillia fluviatilis</emphasis>
(R.Br.) Gasper &amp; V.A.O.Dittrich
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFD8F8B27FBF4EB71" authority="(Colenso) Gasper &amp; V. A. O. Dittrich" authorityName="Gasper &amp; V. A. O. Dittrich" baseAuthorityName="Colenso" box="[518,1107,338,364]" class="Polypodiopsida" family="Blechnaceae" genus="Cranfillia" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="03D03A25BB76372CFD8F8B27FD1EEB71" box="[518,697,338,364]" italics="true" pageId="3" pageNumber="187">Cranfillia nigra</emphasis>
(Colenso) Gasper &amp; V.A.O.Dittrich
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFBD68B27FAECEB71" authority="Parris" authorityName="Parris" box="[1119,1355,338,364]" class="Polypodiopsida" family="Blechnaceae" genus="Doodia" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="03D03A25BB76372CFBD68B27FB58EB71" box="[1119,1279,338,364]" italics="true" pageId="3" pageNumber="187">Doodia mollis</emphasis>
Parris
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFADE8B27FEC7EB8D" authority="Colenso" authorityName="Colenso" class="Polypodiopsida" family="Blechnaceae" genus="Doodia" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="squarrosa">
<emphasis id="03D03A25BB76372CFADE8B27FF5EEB8D" italics="true" pageId="3" pageNumber="187">Doodia squarrosa</emphasis>
Colenso
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFEE48B02FCC0EB8C" authority="(G. Forst.) Willd. However" authorityName="Willd. However" baseAuthorityName="G. Forst." box="[365,871,374,401]" class="Polypodiopsida" family="Blechnaceae" genus="Lomaria" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="03D03A25BB76372CFEE48B02FD95EB8D" box="[365,562,374,400]" italics="true" pageId="3" pageNumber="187">Lomaria discolor</emphasis>
(G.Forst.) Willd. However
</taxonomicName>
, in the analysis of the images of live plants shown by
<bibRefCitation id="55359BC6BB76372CFF228BEEFE71EBA9" author="Brownsey, P. J. &amp; Perrie, L. R." box="[171,470,410,437]" pageId="3" pageNumber="187" pagination="1 - 91" refId="ref2987" refString="Brownsey, P. J. &amp; Perrie, L. R. (2021) Blechnaceae In: Breitwieser, I. (Ed.) Flora of New Zealand: ferns and lycophytes, Fascicle 29. Manaaki Whenua Press, Lincoln, pp. 1 - 91." type="book chapter" year="2021">Brownsey &amp; Perrie (2021)</bibRefCitation>
only the prostrate growth and in contact with the substrate is observed in
<taxonomicName id="F6A49DB4BB76372CFA9C8BEEFA22EBA9" box="[1301,1413,410,437]" class="Polypodiopsida" family="Blechnaceae" genus="Austroblechnum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="banksii">
<emphasis id="03D03A25BB76372CFA9C8BEEFA22EBA9" box="[1301,1413,410,437]" italics="true" pageId="3" pageNumber="187">A. banksii</emphasis>
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB76372CFA1A8BEEFE9CEBC5" class="Polypodiopsida" family="Blechnaceae" genus="Austroblechnum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="membranaceum">
<emphasis id="03D03A25BB76372CFA1A8BEEFE9CEBC5" italics="true" pageId="3" pageNumber="187">A. membranaceum</emphasis>
</taxonomicName>
and
<taxonomicName id="F6A49DB4BB76372CFEF88BCAFE68EBC5" box="[369,463,447,473]" class="Polypodiopsida" family="Blechnaceae" genus="Cranfillia" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="03D03A25BB76372CFEF88BCAFE68EBC5" box="[369,463,447,473]" italics="true" pageId="3" pageNumber="187">C. nigra</emphasis>
</taxonomicName>
. Undoubtedly,
<taxonomicName id="F6A49DB4BB76372CFDF28BCAFCACEBC5" box="[635,779,447,472]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="prostraum">
<emphasis id="03D03A25BB76372CFDF28BCAFCACEBC5" box="[635,779,447,472]" italics="true" pageId="3" pageNumber="187">P. prostraum</emphasis>
</taxonomicName>
does not belong to any of these genera.
</paragraph>
<caption id="65DBB6BFBB76372CFF018DB9FB46E237" ID-DOI="http://doi.org/10.5281/zenodo.8157968" ID-Zenodo-Dep="8157968" httpUri="https://zenodo.org/record/8157968/files/figure.png" pageId="3" pageNumber="187" startId="3.[136,229,1996,2018]" targetBox="[151,1436,497,1971]" targetPageId="3" targetType="figure">
<paragraph id="311BE637BB76372CFF018DB9FB46E237" blockId="3.[136,1451,1996,2090]" pageId="3" pageNumber="187">
<emphasis id="03D03A25BB76372CFF018DB9FF58EDFC" bold="true" box="[136,255,1996,2018]" pageId="3" pageNumber="187">FIGURE 2.</emphasis>
<emphasis id="03D03A25BB76372CFE8F8DB9FDABEDFC" box="[262,524,1996,2017]" italics="true" pageId="3" pageNumber="187">
<taxonomicName id="F6A49DB4BB76372CFE8F8DB9FDAFEDFC" authorityName="J. Murillo &amp; L. A. Triana" authorityYear="2023" box="[262,520,1996,2017]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="187" phylum="Tracheophyta" rank="species" species="prostratum">Parablechnum prostratum</taxonomicName>
.
</emphasis>
A. Habit, B. Juvenile sterile leaf, C. Fertile leaf, D. Petiole and lamina scales, E. Detail of the rachis showing the aerophores (white circles), F. Cross section of petiole, showing six meristeles (white arrows), G. Detail of the abaxial surface of fertile pinnae, H. Rhizome scale, I. Petiole scale, J-K. Pinna scales. Photos by J. Murillo and L.A. Triana.
</paragraph>
</caption>
<paragraph id="311BE637BB71372BFF348AEBFEFDE8AD" blockId="4.[136,1452,158,688]" pageId="4" pageNumber="188">
Other character present in
<taxonomicName id="F6A49DB4BB71372BFE6E8AEAFDD9EAA5" authorityName="J. Murillo &amp; L. A. Triana" authorityYear="2023" box="[487,638,159,184]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="prostratum">
<emphasis id="03D03A25BB71372BFE6E8AEAFDD9EAA5" box="[487,638,159,184]" italics="true" pageId="4" pageNumber="188">P. prostratum</emphasis>
</taxonomicName>
is that the bases of fertile pinnae are bulliform, making them wider than the rest of the pinna. Several species from Oceania, such as
<taxonomicName id="F6A49DB4BB71372BFCBC8AB7FA2DEAC1" authority="(Tindale) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="Tindale" box="[821,1418,194,220]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="camfieldii">
<emphasis id="03D03A25BB71372BFCBC8AB7FBF4EAC1" box="[821,1107,194,220]" italics="true" pageId="4" pageNumber="188">Parablechnum camfieldii</emphasis>
(Tindale) Gasper &amp; Salino
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB71372BFA1E8AB6FDE7EB1D" authority="(E. Fourn.) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="E. Fourn." class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="confusum">
<emphasis id="03D03A25BB71372BFA1E8AB6FF54EB1D" italics="true" pageId="4" pageNumber="188">P. confusum</emphasis>
(E.Fourn.) Gasper &amp; Salino
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB71372BFDC68A92FBB6EB1D" authority="(Brause) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="Brause" box="[591,1041,230,256]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="dilatatum">
<emphasis id="03D03A25BB71372BFDC68A92FD7EEB1D" box="[591,729,230,256]" italics="true" pageId="4" pageNumber="188">P. dilatatum</emphasis>
(Brause) Gasper &amp; Salino
</taxonomicName>
,
<taxonomicName id="F6A49DB4BB71372BFBA98A92FE5EEB39" authority="(T. C. Chambers &amp; P. A. Farrant) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="T. C. Chambers &amp; P. A. Farrant" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="novae-zelandiae">
<emphasis id="03D03A25BB71372BFBA98A92FB50EB1D" box="[1056,1271,230,256]" italics="true" pageId="4" pageNumber="188">P. novae-zelandiae</emphasis>
(T.C.Chambers &amp; P.A.Farrant) Gasper &amp; Salino
</taxonomicName>
, and
<taxonomicName id="F6A49DB4BB71372BFDBE8B7EFB8BEB39" authority="(E. Fourn.) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="E. Fourn." box="[567,1068,266,292]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="subcordatum">
<emphasis id="03D03A25BB71372BFDBE8B7EFD42EB39" box="[567,741,266,292]" italics="true" pageId="4" pageNumber="188">P. subcordatum</emphasis>
(E.Fourn.) Gasper &amp; Salino
</taxonomicName>
, as well as
<taxonomicName id="F6A49DB4BB71372BFB3B8B7EFE7AEB55" authority="(Lorence &amp; A. R. Sm.) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="Lorence &amp; A. R. Sm." class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="pacificum">
<emphasis id="03D03A25BB71372BFB3B8B7EFA9AEB39" box="[1202,1341,266,292]" italics="true" pageId="4" pageNumber="188">P. pacificum</emphasis>
(Lorence &amp; A.R.Sm.) Gasper &amp; Salino
</taxonomicName>
, from the Southern Pacific, and sometimes
<taxonomicName id="F6A49DB4BB71372BFC518B5AFA00EB55" authority="K. Kessler &amp; A. R. Sm." authorityName="K. Kessler &amp; A. R. Sm." box="[984,1447,302,329]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="wohlgemuthii">
<emphasis id="03D03A25BB71372BFC518B5AFB28EB55" box="[984,1167,302,328]" italics="true" pageId="4" pageNumber="188">P. wohlgemuthii</emphasis>
K. Kessler &amp; A.R. Sm.
</taxonomicName>
, from
<collectingCountry id="49B3A6A7BB71372BFF4E8B27FEBBEB71" box="[199,284,338,364]" name="Bolivia" pageId="4" pageNumber="188">Bolivia</collectingCountry>
, also have fertile pinnae with broadened bases (
<bibRefCitation id="55359BC6BB71372BFCC98B27FC46EB71" author="Molino, S." box="[832,993,338,364]" pageId="4" pageNumber="188" refId="ref3384" refString="Molino, S. (2022) Sinopsis global y aproximacion a la filogenia molecular del genero Parablechnum C. Presl (Blechnaceae, Polypodiopsida). Ph. D. Thesis, Universidad Complutense de Madrid. Madrid. 445 pp. [https: // eprints. ucm. es / id / eprint / 75407 / 1 / T 43509. pdf]" type="book" year="2022">Molino, 2022</bibRefCitation>
). However, in these species the bases of pinnae are flattened (
<emphasis id="03D03A25BB71372BFEF38B02FE3EEB8D" box="[378,409,375,400]" italics="true" pageId="4" pageNumber="188">vs.</emphasis>
bulliform) and leaves are erect to arched (
<emphasis id="03D03A25BB71372BFC0B8B02FC06EB8D" box="[898,929,375,400]" italics="true" pageId="4" pageNumber="188">vs.</emphasis>
prostrate). The new species is very similar to
<taxonomicName id="F6A49DB4BB71372BFF018BEEFDD3EBA9" authority="(A. Rojas) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="A. Rojas" box="[136,628,410,436]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="moranianum">
<emphasis id="03D03A25BB71372BFF018BEEFE94EBA9" box="[136,307,411,436]" italics="true" pageId="4" pageNumber="188">P. moranianum</emphasis>
(A. Rojas) Gasper &amp; Salino
</taxonomicName>
from
<collectingCountry id="49B3A6A7BB71372BFD308BEFFC94EBA9" box="[697,819,410,436]" name="Costa Rica" pageId="4" pageNumber="188">Costa Rica</collectingCountry>
(
<bibRefCitation id="55359BC6BB71372BFCCB8BEFFC62EBA9" author="Rojas, A." box="[834,965,410,436]" pageId="4" pageNumber="188" pagination="388 - 394" refId="ref3595" refString="Rojas, A. (2006) Two New Species of Blechnum (Blechnaceae) from the Neotropics. Brittonia 58: 388 - 394. https: // doi. org / 10.1663 / 0007 - 196 X (2006) 58 [388: TNSOBB] 2.0. CO; 2" type="journal article" year="2006">Rojas 2006</bibRefCitation>
) in leaf size and shape, as well as the scale shape and distribution on the lamina. However, the latter species has erect and arched sterile leaves (
<emphasis id="03D03A25BB71372BFB658BCAFAADEBC5" box="[1260,1290,447,472]" italics="true" pageId="4" pageNumber="188">vs.</emphasis>
prostrate) and sterile pinnae are separated (
<emphasis id="03D03A25BB71372BFE438B96FE4FEBE1" box="[458,488,483,508]" italics="true" pageId="4" pageNumber="188">vs.</emphasis>
touching by their margins). Cylindrical aerophores in the new species resemble those of
<taxonomicName id="F6A49DB4BB71372BFF218873FC85E83D" authority="(Broadh.) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="Broadh." box="[168,802,518,544]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="chiriquanum">
<emphasis id="03D03A25BB71372BFF218873FE43E83D" box="[168,484,518,544]" italics="true" pageId="4" pageNumber="188">Parablechnum chiriquanum</emphasis>
(Broadh.) Gasper &amp; Salino
</taxonomicName>
, but in this species sterile leaves are erect (
<emphasis id="03D03A25BB71372BFA9B8872FA96E83D" box="[1298,1329,519,544]" italics="true" pageId="4" pageNumber="188">vs.</emphasis>
prostrate), pinnae are flattened (
<emphasis id="03D03A25BB71372BFEF1885EFE30E859" box="[376,407,555,580]" italics="true" pageId="4" pageNumber="188">vs.</emphasis>
revolute) and leaves are generally larger (&gt;
<quantity id="F65C4BD2BB71372BFC0D885FFC6BE858" box="[900,972,554,581]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="188" unit="cm" value="70.0">70 cm</quantity>
<emphasis id="03D03A25BB71372BFC5D885EFC4CE859" box="[980,1003,555,580]" italics="true" pageId="4" pageNumber="188">vs</emphasis>
. &lt;
<quantity id="F65C4BD2BB71372BFB83885FFBF2E859" box="[1034,1109,554,580]" metricMagnitude="-1" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="188" unit="cm" value="67.0">67 cm</quantity>
). Laminae are similar to those of
<taxonomicName id="F6A49DB4BB71372BFF2E883BFD60E875" authority="(Rosenst.) Gasper &amp; Salino" authorityName="Gasper &amp; Salino" baseAuthorityName="Rosenst." box="[167,711,590,616]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="lima">
<emphasis id="03D03A25BB71372BFF2E883BFE21E875" box="[167,390,590,616]" italics="true" pageId="4" pageNumber="188">Parablechnum lima</emphasis>
(Rosenst.) Gasper &amp; Salino
</taxonomicName>
, but the latter has tuberculiform aerophores and erect leaves. The leaves also resemble those of
<taxonomicName id="F6A49DB4BB71372BFE558806FDFFE891" box="[476,600,626,652]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="188" phylum="Tracheophyta" rank="species" species="stuebelii">
<emphasis id="03D03A25BB71372BFE558806FDFFE891" box="[476,600,626,652]" italics="true" pageId="4" pageNumber="188">P. stuebelii</emphasis>
</taxonomicName>
, but this species has largely reptant rhizomes, with erect leaves distributed along the rhizome.
</paragraph>
</subSubSection>
</treatment>
</document>
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