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<document id="16469162464158D29BBF2C231112226D" ID-CLB-Dataset="264211" ID-DOI="10.5852/ejt.2023.889.2245" ID-GBIF-Dataset="db71145d-18b1-4ad1-a2a4-72d7b8928a7d" ID-ISSN="2118-9773" ID-Zenodo-Dep="8269451" ID-ZooBank="52E11625-6E26-4E4F-8B15-05195087BAB4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692626252704" checkinUser="felipe" docAuthor="Domahovski, Alexandre C. &amp; Cavichioli, Rodney R." docDate="2023" docId="66347308576DFFBDFE779152FC9AFEC3" docLanguage="en" docName="ejt-889_1-53.pdf" docOrigin="European Journal of Taxonomy 889 (1)" docSource="http://zoobank.org/52e11625-6e26-4e4f-8b15-05195087bab4" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Costanana helvacosta DeLong &amp; Freytag 1972" docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="9A0D0B70574FFF9EFF8A935BFF99FFB7" masterDocTitle="Eight new species of Costanana DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini): taxonomic changes, key to males, and description of Metacostana gen. nov." masterLastPageNumber="53" masterPageNumber="1" pageNumber="35" updateTime="1699595446126" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="57E73A4D3A4C83AE598E99201822F189">Eight new species of Costanana DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini): taxonomic changes, key to males, and description of Metacostana gen. nov.</mods:title>
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<mods:namePart id="F03D6E17B0B6C369C138E3877EF92D23">Domahovski, Alexandre C.</mods:namePart>
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<mods:affiliation id="789E2D24465809B6D052432F13A407A3">Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ, Brazil.</mods:affiliation>
<mods:nameIdentifier id="67B7487DD31D87F0CAC32217DD23A1AB" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="3E3BCFF9B1FFD9FCE1B08EDE90F32850">Cavichioli, Rodney R.</mods:namePart>
<mods:nameIdentifier id="C2CB90CAEFA25FC689B522556FB3DF5A" type="LSID">DFC42ABD-5602-4D6E-B691-49F716ECDFDC</mods:nameIdentifier>
<mods:affiliation id="0161D09E4A51F2E979655DA3A8E023F6">Departamento de Zoologia, Universidade Federal do Paraná. Caixa Postal 19020, Curitiba, 81531 - 980, PR, Brazil.</mods:affiliation>
<mods:nameIdentifier id="14196ED08DBEF7FA11708CA0ACF43622" type="email">cavich@ufpr.br</mods:nameIdentifier>
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<mods:title id="696D1AC8C3BFB925AA064FBF33D586CE">European Journal of Taxonomy</mods:title>
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<mods:part id="30BA35E5230332B1438F12BE8731A18E">
<mods:date id="322D1B6031E09CB6099043BF85BA7734">2023</mods:date>
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<mods:number id="8D89962FEC6D468782FFF366996CB2ED">889</mods:number>
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<mods:number id="03774BE89719CE04661FADF828F6202C">1</mods:number>
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<treatment id="66347308576DFFBDFE779152FC9AFEC3" ID-DOI="http://doi.org/10.5281/zenodo.8269489" ID-GBIF-Taxon="212449058" ID-Zenodo-Dep="8269489" LSID="urn:lsid:plazi:treatment:66347308576DFFBDFE779152FC9AFEC3" httpUri="http://treatment.plazi.org/id/66347308576DFFBDFE779152FC9AFEC3" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="A6879195576DFFBCFE779152FBAFFD94" box="[509,1078,520,547]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="EE22C21E576DFFBCFE779152FBAFFD94" blockId="34.[509,1078,520,582]" box="[509,1078,520,547]" pageId="34" pageNumber="35">
<heading id="B56A7572576DFFBCFE779152FBAFFD94" box="[509,1078,520,547]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="35" reason="2">
<taxonomicName id="299DB99D576DFFBCFE779152FBAFFD94" ID-CoL="YW9N" authority="DeLong &amp; Freytag, 1972" authorityName="DeLong &amp; Freytag" authorityYear="1972" box="[509,1078,520,547]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="helvacosta">
<emphasis id="DCE91E0C576DFFBCFE779152FC9FFD94" bold="true" box="[509,774,520,547]" italics="true" pageId="34" pageNumber="35">Costanana helvacosta</emphasis>
DeLong &amp; Freytag, 1972
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A6879195576DFFBCFD7A9177FCDAFDF1" box="[752,835,556,582]" pageId="34" pageNumber="35" type="description">
<paragraph id="EE22C21E576DFFBCFD7A9177FCDAFDF1" blockId="34.[509,1078,520,582]" box="[752,835,556,582]" pageId="34" pageNumber="35">
<figureCitation id="76A6DE9B576DFFBCFD7A9177FCDAFDF1" box="[752,835,556,582]" captionStart="Fig" captionStartId="34.[189,232,1963,1989]" captionTargetBox="[211,1377,995,1924]" captionTargetId="figure-90@34.[211,1377,995,1924]" captionTargetPageId="34" captionText="Fig. 18. Costanana helvacosta DeLong &amp; Freytag, 1972, holotype, ♂ (CMNH 724,310). A. Habitus, dorsal view. B. Habitus, lateral view. C. Habitus, ventral view. D. Labels." figureDoi="http://doi.org/10.5281/zenodo.8269505" httpUri="https://zenodo.org/record/8269505/files/figure.png" pageId="34" pageNumber="35">Fig. 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A6879195576DFFBCFF379136FCDAFD31" box="[189,835,620,647]" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="EE22C21E576DFFBCFF379136FCDAFD31" blockId="34.[189,835,620,647]" box="[189,835,620,647]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="CE8DE530576DFFBCFF379136FCDAFD31" box="[189,835,620,647]" pageId="34" pageNumber="35">
<treatmentCitation id="6F3CE40F576DFFBCFF379136FCA6FD31" author="DeLong D. M. &amp; Freytag P. H." box="[189,831,620,647]" page="496" pageId="34" pageNumber="35" year="1972">
<taxonomicName id="299DB99D576DFFBCFF379136FCA6FD31" ID-CoL="YW9N" authority="DeLong &amp; Freytag, 1972 b: 496" authorityName="DeLong &amp; Freytag" authorityPageNumber="496" authorityYear="1972" box="[189,831,620,647]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="helvacosta">
<emphasis id="DCE91E0C576DFFBCFF379136FE59FD31" box="[189,448,620,647]" italics="true" pageId="34" pageNumber="35">Costanana helvacosta</emphasis>
<bibRefCitation id="8A0CBFEF576DFFBCFE429136FCA6FD31" author="DeLong D. M. &amp; Freytag P. H." box="[456,831,620,647]" pageId="34" pageNumber="35" pagination="491 - 500" refId="ref23276" refString="DeLong D. M. &amp; Freytag P. H. 1972 b. Studies of the Gyponinae: the genus Costanana and five new species (Homoptera: Cicadellidae). Journal of the Kansas Entomological Society 45 (4): 491 - 500." type="journal article" year="1972">DeLong &amp; Freytag, 1972b: 496</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A6879195576DFFBCFF3791F6FE84FC8C" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="EE22C21E576DFFBCFF3791F6FE33FD70" blockId="34.[189,426,685,711]" box="[189,426,685,711]" pageId="34" pageNumber="35">
<heading id="B56A7572576DFFBCFF3791F6FE33FD70" bold="true" box="[189,426,685,711]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="DCE91E0C576DFFBCFF3791F6FE33FD70" bold="true" box="[189,426,685,711]" pageId="34" pageNumber="35">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="EE22C21E576DFFBCFF609187FE84FC8C" blockId="34.[189,1398,732,827]" pageId="34" pageNumber="35">
<materialsCitation id="5EF5C843576DFFBCFF609187FE84FC8C" ID-GBIF-Occurrence="4399003316" collectingDate="1898-05" collectionCode="CMNH" country="Colombia" location="Colombia" municipality="Minca" pageId="34" pageNumber="35" specimenCode="CMNH 724310" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="DCE91E0C576DFFBCFF609187FEC5FD41" bold="true" box="[234,348,732,758]" pageId="34" pageNumber="35">
<typeStatus id="31267CBC576DFFBCFF609187FEC5FD41" box="[234,348,732,758]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="968A828E576DFFBCFF3791A5FEC1FCAF" box="[189,344,766,792]" name="Colombia" pageId="34" pageNumber="35">COLOMBIA</collectingCountry>
•
<specimenCount id="F89B0997576DFFBCFEFA91A4FE10FCAE" box="[368,393,767,793]" pageId="34" pageNumber="35" type="male">♂</specimenCount>
; “
<collectingMunicipality id="0E465864576DFFBCFE2A91A5FE77FCAF" box="[416,494,766,792]" pageId="34" pageNumber="35">Minca</collectingMunicipality>
\
<location id="EB4294C5576DFFBCFD8E91A5FDE1FCAF" LSID="urn:lsid:plazi:treatment:66347308576DFFBDFE779152FC9AFEC3:EB4294C5576DFFBCFD8E91A5FDE1FCAF" box="[516,632,766,792]" country="Colombia" municipality="Minca" name="Colombia" pageId="34" pageNumber="35">Colombia</location>
\ Acc. No. 1999”, “
<collectingDate id="8A671D36576DFFBCFCD791A5FC72FCAF" box="[861,1003,766,792]" pageId="34" pageNumber="35">May \ 1898</collectingDate>
”, “CMNH-IZ \ 724,310”;
<specimenCode id="BE3B6A65576DFFBCFA9491A5FE80FC8C" collectionCode="CMNH" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="34" pageNumber="35" type="Museum">CMNH 724310</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="BAE29296576DFFBCFF3794F0FB83F85F" ID-DOI="http://doi.org/10.5281/zenodo.8269505" ID-Zenodo-Dep="8269505" httpUri="https://zenodo.org/record/8269505/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1963,1989]" targetBox="[211,1377,995,1924]" targetPageId="34">
<paragraph id="EE22C21E576DFFBCFF3794F0FB83F85F" blockId="34.[189,1398,1963,2025]" pageId="34" pageNumber="35">
<emphasis id="DCE91E0C576DFFBCFF3794F0FE85F872" bold="true" box="[189,284,1963,1989]" pageId="34" pageNumber="35">Fig. 18.</emphasis>
<taxonomicName id="299DB99D576DFFBCFEAF94F7FCFCF872" authority="DeLong &amp; Freytag, 1972" authorityName="DeLong &amp; Freytag" authorityYear="1972" box="[293,869,1963,1990]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="helvacosta">
<emphasis id="DCE91E0C576DFFBCFEAF94F7FDB2F872" box="[293,555,1963,1990]" italics="true" pageId="34" pageNumber="35">Costanana helvacosta</emphasis>
DeLong &amp; Freytag, 1972
</taxonomicName>
, holotype, ♂ (CMNH 724,310).
<emphasis id="DCE91E0C576DFFBCFB6494F0FA9AF872" bold="true" box="[1262,1283,1963,1989]" pageId="34" pageNumber="35">A</emphasis>
. Habitus, dorsal view.
<emphasis id="DCE91E0C576DFFBCFED89495FEFFF85F" bold="true" box="[338,358,1998,2024]" pageId="34" pageNumber="35">B</emphasis>
. Habitus, lateral view.
<emphasis id="DCE91E0C576DFFBCFDFE9495FD10F85F" bold="true" box="[628,649,1998,2024]" pageId="34" pageNumber="35">C</emphasis>
. Habitus, ventral view.
<emphasis id="DCE91E0C576DFFBCFC2B9495FC2FF85F" bold="true" box="[929,950,1998,2024]" pageId="34" pageNumber="35">D</emphasis>
. Labels.
</paragraph>
</caption>
<subSubSection id="A6879195576CFFBDFF379252FC9AFEC3" pageId="35" pageNumber="36" type="discussion">
<paragraph id="EE22C21E576CFFBDFF379252FEB6FE94" blockId="35.[189,303,265,291]" box="[189,303,265,291]" pageId="35" pageNumber="36">
<heading id="B56A7572576CFFBDFF379252FEB6FE94" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="DCE91E0C576CFFBDFF379252FEB6FE94" bold="true" box="[189,303,265,291]" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="EE22C21E576CFFBDFF37926CFC9AFEC3" blockId="35.[189,1399,311,373]" pageId="35" pageNumber="36">
<taxonomicName id="299DB99D576CFFBDFF37926CFE58FEE6" authorityName="DeLong &amp; Freytag" authorityYear="1972" box="[189,449,311,337]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="helvacosta">
<emphasis id="DCE91E0C576CFFBDFF37926CFE58FEE6" box="[189,449,311,337]" italics="true" pageId="35" pageNumber="36">Costanana helvacosta</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="299DB99D576CFFBDFD2E926CFC9BFEE6" baseAuthorityName="Fowler" baseAuthorityYear="1903" box="[676,770,311,337]" class="Insecta" family="Cicadellidae" genus="Gypona" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="DCE91E0C576CFFBDFD2E926CFC9BFEE6" box="[676,770,311,337]" italics="true" pageId="35" pageNumber="36">C. nana</emphasis>
</taxonomicName>
,
<taxonomicName id="299DB99D576CFFBDFC9B926CFC08FEE6" authorityName="DeLong &amp; Wolda" authorityYear="1983" box="[785,913,311,337]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="santana">
<emphasis id="DCE91E0C576CFFBDFC9B926CFC08FEE6" box="[785,913,311,337]" italics="true" pageId="35" pageNumber="36">C. santana</emphasis>
</taxonomicName>
and
<taxonomicName id="299DB99D576CFFBDFC47926CFB32FEE6" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" box="[973,1195,311,337]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rubromarginata" status="sp. nov.">
<emphasis id="DCE91E0C576CFFBDFC47926CFB32FEE6" box="[973,1195,311,337]" italics="true" pageId="35" pageNumber="36">C. rubromarginata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7DAA377576CFFBDFB399263FA97FEE5" box="[1203,1294,312,338]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
and was described based on a single male from
<collectingCountry id="968A828E576CFFBDFD0D9201FD66FEC3" box="[647,767,346,372]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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