302 lines
53 KiB
XML
302 lines
53 KiB
XML
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<mods:title id="09242528B5B13BA1176EDB0DB27BC663">Eight new species of Costanana DeLong & Freytag (Hemiptera: Cicadellidae: Gyponini): taxonomic changes, key to males, and description of Metacostana gen. nov.</mods:title>
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<mods:namePart id="EC17D9CCFE203705BC90EE2A937FA387">Domahovski, Alexandre C.</mods:namePart>
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<mods:affiliation id="C1F82800B124B9B0392D8E7D27D5FA3F">Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="F235654BD868478D7BA93398B07B1871" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="627CFD5DA659FB8D6A4C09CC8F66C227">Cavichioli, Rodney R.</mods:namePart>
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<mods:affiliation id="ADF95C775CCB95B2066BBF8B9889D94C">Departamento de Zoologia, Universidade Federal do Paraná. Caixa Postal 19020, Curitiba, 81531 - 980, PR, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="6185D8E00991D4512FEFAC42026CC37E" type="email">cavich@ufpr.br</mods:nameIdentifier>
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<mods:title id="B1D3B47A2BD9C014E50BFBF21914FF4E">European Journal of Taxonomy</mods:title>
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<mods:part id="A5A9BB54D3C4425FFF388A6AA97A6194">
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<mods:date id="68430D4679F225C8506BEE6B794A0D0C">2023</mods:date>
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<treatment id="66347308575AFF88FDC997C3FAEFF820" ID-DOI="http://doi.org/10.5281/zenodo.8271509" ID-GBIF-Taxon="212449056" ID-Zenodo-Dep="8271509" LSID="urn:lsid:zoobank.org:act:5120C9B2-98F9-4CE9-9FC6-C663FC76165A" httpUri="http://treatment.plazi.org/id/66347308575AFF88FDC997C3FAEFF820" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection id="A6879195575AFF8BFDC997C3FC68FB04" box="[579,1009,1176,1203]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="EE22C21E575AFF8BFDC997C3FC68FB04" blockId="21.[371,1217,1176,1273]" box="[579,1009,1176,1203]" pageId="21" pageNumber="22">
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<heading id="B56A7572575AFF8BFDC997C3FC68FB04" box="[579,1009,1176,1203]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="2">
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<taxonomicName id="299DB99D575AFF8BFDC997C3FC16FB04" ID-CoL="85f3ad6e-0f7a-4efe-87ff-0d0a91bfc97a" authority="Domahovski & Cavichioli, 2023" authorityName="Domahovski & Cavichioli" authorityYear="2023" box="[579,911,1176,1203]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="rubromarginata" status="sp. nov.">
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<emphasis id="DCE91E0C575AFF8BFDC997C3FC16FB04" bold="true" box="[579,911,1176,1203]" italics="true" pageId="21" pageNumber="22">Costanana rubromarginata</emphasis>
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<taxonomicNameLabel id="C7DAA377575AFF8BFC1C97C2FC68FB04" box="[918,1009,1177,1203]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="A6879195575AFF8BFEF997E7FCF4FB4E" pageId="21" pageNumber="22" type="description">
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<uri id="9A0CCE1C575AFF8BFEF997E7FB58FB60" box="[371,1217,1212,1239]" pageId="21" pageNumber="22">
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urn:lsid:zoobank.org:act:
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<uuid id="9A3BF8CB575AFF8BFD1997E7FB58FB60" box="[659,1217,1212,1239]" pageId="21" pageNumber="22">5120C9B2-98F9-4CE9-9FC6-C663FC76165A</uuid>
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</uri>
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</paragraph>
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<paragraph id="EE22C21E575AFF8BFD4D9784FCF4FB4E" blockId="21.[371,1217,1176,1273]" box="[711,877,1247,1273]" pageId="21" pageNumber="22">
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<figureCitation id="76A6DE9B575AFF8BFD4D9784FC8BFB4E" box="[711,786,1247,1273]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="21" pageNumber="22">
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Figs
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<geoCoordinate id="8BA9A4D9575AFF8BFC8B9784FC8BFB4E" box="[769,786,1247,1273]" degrees="9, 12" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="555" value="9.12">9</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="76A6DE9B575AFF8BFC949784FCF4FB4E" box="[798,877,1247,1273]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="21" pageNumber="22">12E–F</figureCitation>
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</paragraph>
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<subSubSection id="A6879195575AFF8BFF37967BFB12F9AE" pageId="21" pageNumber="22" type="diagnosis">
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<paragraph id="EE22C21E575AFF8BFF37967BFEAFFA8D" blockId="21.[189,310,1312,1338]" box="[189,310,1312,1338]" pageId="21" pageNumber="22">
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<heading id="B56A7572575AFF8BFF37967BFEAFFA8D" bold="true" box="[189,310,1312,1338]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
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<emphasis id="DCE91E0C575AFF8BFF37967BFEAFFA8D" bold="true" box="[189,310,1312,1338]" pageId="21" pageNumber="22">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE22C21E575AFF8BFF379614FB12F9AE" blockId="21.[186,1401,1359,1562]" pageId="21" pageNumber="22">
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Coloration (
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<figureCitation id="76A6DE9B575AFF8BFEC29614FE35FADE" box="[328,428,1359,1385]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="21" pageNumber="22">Fig. 12E</figureCitation>
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) of crown black with anterior margin red, pronotum brown with small black maculae, forewing costal margin with narrow light yellow macula on anterior third and small depigmented (white) macula on apical third. Style (
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<figureCitation id="76A6DE9B575AFF8BFDAC96CEFD1FFA18" box="[550,646,1429,1455]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="21" pageNumber="22">Fig. 9K</figureCitation>
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) slender and slightly broadened subapically, without processes near base of blade or apical portion. Aedeagus (
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<figureCitation id="76A6DE9B575AFF8BFD7F96E3FCE1FA64" box="[757,888,1464,1491]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="21" pageNumber="22">Fig. 9L–M</figureCitation>
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) shaft long and slender, cylindrical, curved anterad, without apical or subapical processes; pair of long and slender processes arising from basal third, parallel to shaft and slightly surpassing apex, abruptly enlarged at midlength.
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</paragraph>
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</subSubSection>
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<subSubSection id="A6879195575AFF8BFF37951AFC50F93E" box="[189,969,1601,1673]" pageId="21" pageNumber="22" type="etymology">
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<paragraph id="EE22C21E575AFF8BFF37951AFEDDF9EC" blockId="21.[189,324,1601,1627]" box="[189,324,1601,1627]" pageId="21" pageNumber="22">
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<heading id="B56A7572575AFF8BFF37951AFEDDF9EC" bold="true" box="[189,324,1601,1627]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
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<emphasis id="DCE91E0C575AFF8BFF37951AFEDDF9EC" bold="true" box="[189,324,1601,1627]" pageId="21" pageNumber="22">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE22C21E575AFF8BFF379534FC50F93E" blockId="21.[189,969,1647,1673]" box="[189,969,1647,1673]" pageId="21" pageNumber="22">The species epithet refers to the anterior margin of head being red.</paragraph>
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</subSubSection>
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<subSubSection id="A6879195575AFF8BFF3795EBFE65F870" pageId="21" pageNumber="22" type="materials_examined">
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<paragraph id="EE22C21E575AFF8BFF3795EBFE33F97D" blockId="21.[189,426,1712,1738]" box="[189,426,1712,1738]" pageId="21" pageNumber="22">
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<heading id="B56A7572575AFF8BFF3795EBFE33F97D" bold="true" box="[189,426,1712,1738]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
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<emphasis id="DCE91E0C575AFF8BFF3795EBFE33F97D" bold="true" box="[189,426,1712,1738]" pageId="21" pageNumber="22">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EE22C21E575AFF8BFF609584FC4AF8F7" blockId="21.[189,1398,1759,1856]" pageId="21" pageNumber="22">
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<materialsCitation id="5EF5C843575AFF8BFF609584FC4AF8F7" ID-GBIF-Occurrence="4399003303" collectingDateMax="2017-06-29" collectingDateMin="2017-06-26" collectingMethod="sweep" collectionCode="DZUP" collectorName="A. C. Domahovski" country="Brazil" county="Brasil" elevation="230" latitude="-9.8559" location="Fazenda Sao Nicolau" longLatPrecision="7" longitude="-58.2486" municipality="Cotriguacu" pageId="21" pageNumber="22" specimenCode="DZUP 215491" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso" typeStatus="holotype">
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<emphasis id="DCE91E0C575AFF8BFF609584FEC5F94E" bold="true" box="[234,348,1759,1785]" pageId="21" pageNumber="22">
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<typeStatus id="31267CBC575AFF8BFF609584FEC5F94E" box="[234,348,1759,1785]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectingCountry id="968A828E575AFF8BFF379459FEBFF8AB" box="[189,294,1794,1820]" name="Brazil" pageId="21" pageNumber="22">BRAZIL</collectingCountry>
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•
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<specimenCount id="F89B0997575AFF8BFEB49458FECEF8AA" box="[318,343,1795,1821]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
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; “Brasil, MT [
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],
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<collectingMunicipality id="0E465864575AFF8BFD3B9459FCAFF8AB" box="[689,822,1794,1820]" pageId="21" pageNumber="22">Cotriguaçu</collectingMunicipality>
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, \
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,
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,
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, \
|
||
<quantity id="29656FFB575AFF8BFF37947EFE9AF888" box="[189,259,1829,1855]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="22" unit="m" value="230.0">
|
||
<elevation id="65B0252D575AFF8BFF37947EFE9AF888" box="[189,259,1829,1855]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="22" unit="m" value="230.0">230m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="37DCBA09575AFF8BFE84947DFEC3F8F7" box="[270,346,1830,1856]" pageId="21" pageNumber="22">sweep</collectingMethod>
|
||
,
|
||
<collectingDate id="8A671D36575AFF8BFEEC947EFDBCF888" box="[358,549,1829,1855]" pageId="21" pageNumber="22">
|
||
<date id="9A23E4DE575AFF8BFEEC947EFE48F888" box="[358,465,1829,1855]" pageId="21" pageNumber="22" value="1926-06-29">26-29.vi.</date>
|
||
\ 2017
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="4368A7C8575AFF8BFDA5947EFC93F888" box="[559,778,1829,1855]" pageId="21" pageNumber="22">A.C. Domahovski</collectorName>
|
||
”;
|
||
<specimenCode id="BE3B6A65575AFF8BFCAA947DFC56F8F7" box="[800,975,1829,1856]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="21" pageNumber="22" type="University or college">DZUP 215491</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E575AFF8BFF60943CFE65F870" blockId="21.[189,1399,1895,1991]" pageId="21" pageNumber="22">
|
||
<materialsCitation id="5EF5C843575AFF8BFF60943CFE61F870" ID-GBIF-Occurrence="4399003324" collectingDate="2017-07-08" collectingMethod="Varredura" collectionCode="CZJA" collectorName="W. F. Lima" country="Brazil" location="Benfica" municipality="Benevides" pageId="21" pageNumber="22" specimenCode="00012313" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="paratype">
|
||
<emphasis id="DCE91E0C575AFF8BFF60943CFEC5F836" bold="true" box="[234,348,1895,1921]" pageId="21" pageNumber="22">
|
||
<typeStatus id="31267CBC575AFF8BFF60943CFEC5F836" box="[234,348,1895,1921]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="968A828E575AFF8BFF3794D1FEBFF813" box="[189,294,1930,1956]" name="Brazil" pageId="21" pageNumber="22">BRAZIL</collectingCountry>
|
||
–
|
||
<emphasis id="DCE91E0C575AFF8BFECB94D2FEE4F813" bold="true" box="[321,381,1929,1956]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="2C590CFC575AFF8BFECB94D2FEE4F813" box="[321,381,1929,1956]" country="Brazil" name="Para" pageId="21" pageNumber="22">Pará</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="F89B0997575AFF8BFE1F94D1FE5AF813" box="[405,451,1930,1956]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
|
||
; “Brasil
|
||
<collectingRegion id="2C590CFC575AFF8BFDA394D1FDC4F813" box="[553,605,1930,1956]" country="Brazil" name="Para" pageId="21" pageNumber="22">Pará</collectingRegion>
|
||
<collectingMunicipality id="0E465864575AFF8BFDEE94D1FD47F813" box="[612,734,1930,1956]" pageId="21" pageNumber="22">Benevides</collectingMunicipality>
|
||
\
|
||
<location id="EB4294C5575AFF8BFD7994D1FCD4F813" LSID="urn:lsid:plazi:treatment:66347308575AFF88FDC997C3FAEFF820:EB4294C5575AFF8BFD7994D1FCD4F813" box="[755,845,1930,1956]" country="Brazil" municipality="Benevides" name="Benfica" pageId="21" pageNumber="22" stateProvince="Para">Benfica</location>
|
||
\
|
||
<date id="9A23E4DE575AFF8BFCE894D1FC6BF813" box="[866,1010,1930,1956]" pageId="21" pageNumber="22" value="2017-07-08">
|
||
<collectingDate id="8A671D36575AFF8BFCE894D1FC6BF813" box="[866,1010,1930,1956]" pageId="21" pageNumber="22" value="2017-07-08">08.VII.2017</collectingDate>
|
||
</date>
|
||
<collectingMethod id="37DCBA09575AFF8BFC7D94D1FBF2F813" box="[1015,1131,1930,1956]" pageId="21" pageNumber="22">Varredura</collectingMethod>
|
||
\
|
||
<collectorName id="4368A7C8575AFF8BFB0A94D1FA9CF813" box="[1152,1285,1930,1956]" pageId="21" pageNumber="22">W. F. Lima</collectorName>
|
||
”, “CZDC UEPA \
|
||
<specimenCode id="BE3B6A65575AFF8BFE9694F6FE01F870" box="[284,408,1965,1991]" pageId="21" pageNumber="22">00012313</specimenCode>
|
||
”;
|
||
<collectionCode id="888C5ADB575AFF8BFE2494F6FE61F870" box="[430,504,1965,1991]" pageId="21" pageNumber="22">CZJA</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A68791955759FF88FF379253FEA0F92D" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="EE22C21E5759FF88FF379253FED7FE95" blockId="22.[189,334,264,290]" box="[189,334,264,290]" pageId="22" pageNumber="23">
|
||
<heading id="B56A75725759FF88FF379253FED7FE95" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
|
||
<emphasis id="DCE91E0C5759FF88FF379253FED7FE95" bold="true" box="[189,334,264,290]" pageId="22" pageNumber="23">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF60926CFEB3FEE6" blockId="22.[189,1174,311,373]" box="[234,298,311,337]" pageId="22" pageNumber="23">
|
||
<emphasis id="DCE91E0C5759FF88FF60926CFEB3FEE6" bold="true" box="[234,298,311,337]" pageId="22" pageNumber="23">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF379200FB0FFEC2" blockId="22.[189,1174,311,373]" box="[189,1174,346,373]" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="E8C454C25759FF88FF379200FEF7FEC4" baselines="366,366" box="[189,366,347,373]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="22" pageNumber="23">MEASUREMENTS</smallCapsWord>
|
||
.
|
||
<typeStatus id="31267CBC5759FF88FEF79201FE73FEC3" box="[381,490,346,372]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
|
||
male: total length 5.0 mm.
|
||
<typeStatus id="31267CBC5759FF88FCA49200FC0CFEC2" box="[814,917,347,373]" pageId="22" pageNumber="23" type="paratype">Paratype</typeStatus>
|
||
male (n = 1) 5.0 mm.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF3792C7FB0FFD1B" blockId="22.[187,1399,412,685]" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="E8C454C25759FF88FF3792C7FED1FE03" baselines="431,431" box="[189,328,412,438]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="22" pageNumber="23">COLORATION</smallCapsWord>
|
||
. Dorsal portion head and thorax reddish-brown (
|
||
<figureCitation id="76A6DE9B5759FF88FC1A92C7FB8EFE01" box="[912,1047,412,438]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Figs 9A, C</figureCitation>
|
||
,
|
||
<figureCitation id="76A6DE9B5759FF88FBAF92C7FBC1FE01" box="[1061,1112,412,438]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="22" pageNumber="23">12E</figureCitation>
|
||
); ventral portion yellow (
|
||
<figureCitation id="76A6DE9B5759FF88FF4D92E4FED5FE6E" box="[199,332,447,473]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Figs 9B–C</figureCitation>
|
||
,
|
||
<figureCitation id="76A6DE9B5759FF88FED192E4FE15FE6E" box="[347,396,447,473]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,649,1847]" captionTargetId="figure-47@27.[189,1398,649,1847]" captionTargetPageId="27" captionText="Fig. 12. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. obtusa sp. nov., holotype, ♂ (DZUP 215492). C–D. C. piraquarensis sp. nov., holotype, ♂ (DZUP 215485). E–F. C. rubromarginata sp. nov., holotype, ♂ (DZUP 215491). G–H. C. xenomorpha sp. nov., holotype, ♂ (MUSM). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269487" httpUri="https://zenodo.org/record/8269487/files/figure.png" pageId="22" pageNumber="23">12F</figureCitation>
|
||
). Crown (
|
||
<figureCitation id="76A6DE9B5759FF88FD8C92E4FDFAFE6E" box="[518,611,447,473]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
) black; anterior margin red; ocelli red. Face (
|
||
<figureCitation id="76A6DE9B5759FF88FB0792E4FB70FE6E" box="[1165,1257,447,473]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9B</figureCitation>
|
||
) with frons black; lorum black to dark brown; gena dark brown between eye and frons; clypeus dark brown except apex yellow. Pronotum (
|
||
<figureCitation id="76A6DE9B5759FF88FE57915EFDA0FDA8" box="[477,569,517,543]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
) with black irregular maculae near anterior and lateral margins; lateral margins yellow. Mesonotum (
|
||
<figureCitation id="76A6DE9B5759FF88FD9D9173FDE8FDF4" box="[535,625,552,579]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
) lateral angles black; pair of black maculae submedially. Forewing (
|
||
<figureCitation id="76A6DE9B5759FF88FF4E9117FEB8FDD1" box="[196,289,588,614]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9D</figureCitation>
|
||
) costal margin with elongate narrow light yellow macula on anterior third, small depigmented (white) macula on outer apical cell, remaining portion black; veins outlined by dark brown; outer half of corium, apex and appendix smoky. Legs (
|
||
<figureCitation id="76A6DE9B5759FF88FD2F91C9FC94FD1B" box="[677,781,658,684]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,742,1851]" captionTargetId="figure-195@26.[189,1398,742,1851]" captionTargetPageId="26" captionText="Fig. 11. Habitus of new species of Costanana DeLong & Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. alata sp. nov., holotype, ♂ (DZUP 215484). C–D. C. bifida sp. nov., holotype, ♂ (DZUP 215493). E–F. C. cifi sp. nov., holotype, ♂ (DZUP 215484). G–H. C. luzi sp. nov., holotype ♂ (DZUP 215489). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269485" httpUri="https://zenodo.org/record/8269485/files/figure.png" pageId="22" pageNumber="23">Fig. 11B</figureCitation>
|
||
) yellow, without black markings.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF37918FFBE8FC53" blockId="22.[187,1400,723,996]" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="E8C454C25759FF88FF37918FFEA3FD5B" baselines="743,743" box="[189,314,724,750]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Structure" pageId="22" pageNumber="23">STRUCTURE</smallCapsWord>
|
||
. Head in dorsal view (
|
||
<figureCitation id="76A6DE9B5759FF88FDC49188FD29FD59" box="[590,688,723,750]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
): median length of crown as long as half interocular width; transocular width of head 8.5 tenths humeral width of pronotum; in lateral view (
|
||
<figureCitation id="76A6DE9B5759FF88FBFB91ACFB54FCA6" box="[1137,1229,759,785]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9C</figureCitation>
|
||
), with anterior margin moderately thick, with 5–6 well developed carinae. Forewing (
|
||
<figureCitation id="76A6DE9B5759FF88FB819041FBF0FC83" box="[1035,1129,794,820]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9D</figureCitation>
|
||
) with inner discal cell half long as outer discal cell; apex rounded. Profemur AV row with 4–5 setae restricted to basal half and PV row with 1 apical seta. Protibia PD row with 3 long setae and apical PD
|
||
<subScript id="7219C05B5759FF88FBC09028FBCAFC36" attach="left" box="[1098,1107,883,897]" fontSize="6" pageId="22" pageNumber="23">1</subScript>
|
||
seta developed; PV row developed, with 8 setae increasing in thickness and length towards apex. Metatibia rows PD, AD, and AV with 20–23, 12, and 13 macrosetae, respectively. Metatarsomere I apex with 3 platellae. Metatarsomere II apex with 2 apical platellae. Other characteristics as in the generic description.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF379757FE35F981" blockId="22.[189,1401,1035,1590]" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="E8C454C25759FF88FF379757FF66FB93" baselines="1055,1055" box="[189,255,1036,1062]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="22" pageNumber="23">MALE</smallCapsWord>
|
||
<smallCapsWord id="E8C454C25759FF88FE8F974AFE1FFB93" baselines="1055" box="[261,390,1041,1060]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="22" pageNumber="23">TERMINALIA</smallCapsWord>
|
||
. Sternite VIII (
|
||
<figureCitation id="76A6DE9B5759FF88FDBE9750FD13FB92" box="[564,650,1035,1061]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9E</figureCitation>
|
||
) 1.1× as wide as long; lateral margins excavated at base, forming a triangular projection and then converging to rounded apex. Valve (
|
||
<figureCitation id="76A6DE9B5759FF88FB8B9775FBC5FBFE" box="[1025,1116,1070,1097]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9F</figureCitation>
|
||
) 1.6× as wide as long; posterior margin rounded. Pygofer (
|
||
<figureCitation id="76A6DE9B5759FF88FDE19709FD55FBDB" box="[619,716,1106,1132]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9G</figureCitation>
|
||
), in lateral view, 2× as long as maximum height; ventral margin broadly rounded; posterodorsal margin approximately straight; external surface near ventral margin with short filiform setae; macrosetae distributed on posterodorsal quadrant; apex slightly tapered and almost truncate. Subgenital plate (
|
||
<figureCitation id="76A6DE9B5759FF88FD0697E0FD75FB62" box="[652,748,1211,1237]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9G</figureCitation>
|
||
), in lateral view, surpassing half length of pygofer; in ventral view (
|
||
<figureCitation id="76A6DE9B5759FF88FED79785FE23FB4E" box="[349,442,1246,1273]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9H</figureCitation>
|
||
), 4× as long as wide; lateral margins slightly rounded; ventral surface and external margin with long filiform setae; apex tapered and weakly sclerotized. Connective (
|
||
<figureCitation id="76A6DE9B5759FF88FB159659FB68FAAB" box="[1183,1265,1282,1308]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9I</figureCitation>
|
||
) T-shaped; arms weakly produced laterally; stalk narrow, slightly shorter than maximum width or arms; median keel weakly developed. Style (
|
||
<figureCitation id="76A6DE9B5759FF88FDB99613FD13FAD5" box="[563,650,1352,1378]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9J</figureCitation>
|
||
), in dorsal view, with outer lobe well developed, rounded; in lateral view (
|
||
<figureCitation id="76A6DE9B5759FF88FED79630FE27FA32" box="[349,446,1387,1413]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9K</figureCitation>
|
||
), blade slender, evenly curved dorsally, slightly broadened subapically; ventral margin not serrated; apex tapered, curved dorsally and acute. Aedeagus (
|
||
<figureCitation id="76A6DE9B5759FF88FBB096D5FB59FA1E" box="[1082,1216,1422,1449]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9L–M</figureCitation>
|
||
) preatrium not developed; dorsal apodeme weakly produced laterally, dorsal margin straight; shaft long and slender, cylindrical, curved anterad; pair of long and slender processes arising from basal third, parallel to shaft and slightly surpassing apex, enlarged at midlength and thin and acute apically; shaft without apical or subapical processes.
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF609506FEDCF9C0" blockId="22.[189,325,1629,1690]" box="[234,325,1629,1655]" pageId="22" pageNumber="23">
|
||
<emphasis id="DCE91E0C5759FF88FF609506FEDCF9C0" bold="true" box="[234,325,1629,1655]" pageId="22" pageNumber="23">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF3795DBFEA0F92D" blockId="22.[189,325,1629,1690]" box="[189,313,1664,1690]" pageId="22" pageNumber="23">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A68791955759FF88FF379599FAEFF820" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="EE22C21E5759FF88FF379599FEB6F96B" blockId="22.[189,303,1730,1756]" box="[189,303,1730,1756]" pageId="22" pageNumber="23">
|
||
<heading id="B56A75725759FF88FF379599FEB6F96B" bold="true" box="[189,303,1730,1756]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
|
||
<emphasis id="DCE91E0C5759FF88FF379599FEB6F96B" bold="true" box="[189,303,1730,1756]" pageId="22" pageNumber="23">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="EE22C21E5759FF88FF3795ABFAEFF820" blockId="22.[188,1400,1776,1943]" pageId="22" pageNumber="23">
|
||
<taxonomicName id="299DB99D5759FF88FF3795ABFD99F8BD" authority="Domahovski & Cavichioli, 2023" authorityName="Domahovski & Cavichioli" authorityYear="2023" box="[189,512,1776,1802]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rubromarginata" status="sp. nov.">
|
||
<emphasis id="DCE91E0C5759FF88FF3795ABFD99F8BD" box="[189,512,1776,1802]" italics="true" pageId="22" pageNumber="23">Costanana rubromarginata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C7DAA3775759FF88FD8395ABFDFCF8BD" box="[521,613,1776,1802]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="299DB99D5759FF88FC8C95ABFC3FF8BD" authorityName="DeLong & Freytag" authorityYear="1972" box="[774,934,1776,1802]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="helvacosta">
|
||
<emphasis id="DCE91E0C5759FF88FC8C95ABFC3FF8BD" box="[774,934,1776,1802]" italics="true" pageId="22" pageNumber="23">C. helvacosta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="299DB99D5759FF88FC3C95ABFB8CF8BD" baseAuthorityName="Fowler" baseAuthorityYear="1903" box="[950,1045,1776,1802]" class="Insecta" family="Cicadellidae" genus="Gypona" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis id="DCE91E0C5759FF88FC3C95ABFB8CF8BD" box="[950,1045,1776,1802]" italics="true" pageId="22" pageNumber="23">C. nana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="299DB99D5759FF88FBD895ABFB4AF8BD" authorityName="DeLong & Wolda" authorityYear="1983" box="[1106,1235,1776,1802]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="santana">
|
||
<emphasis id="DCE91E0C5759FF88FBD895ABFB4AF8BD" box="[1106,1235,1776,1802]" italics="true" pageId="22" pageNumber="23">C. santana</emphasis>
|
||
</taxonomicName>
|
||
in having the aedeagus with the shaft long and slender, with the pair of processes arising near the base of the shaft and without processes distally (
|
||
<figureCitation id="76A6DE9B5759FF88FE7D946DFDE2F8E7" box="[503,635,1846,1872]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9L–M</figureCitation>
|
||
). The shape of the apex of the style, slightly expanded subapically (
|
||
<figureCitation id="76A6DE9B5759FF88FF4E9402FEBBF8C3" box="[196,290,1881,1908]" captionStart="Fig" captionStartId="23.[189,232,1857,1883]" captionTargetBox="[189,1398,268,1787]" captionTargetId="figure-11@23.[189,1398,267,1787]" captionTargetPageId="23" captionText="Fig. 9. Costanana rubromarginata sp. nov., holotype, ♂ (DZUP 215491). A. Head and thorax, dorsal view. B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view.H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269479" httpUri="https://zenodo.org/record/8269479/files/figure.png" pageId="22" pageNumber="23">Fig. 9K</figureCitation>
|
||
), is most similar to that of
|
||
<taxonomicName id="299DB99D5759FF88FDD29401FD2DF8C4" baseAuthorityName="Fowler" baseAuthorityYear="1903" box="[600,692,1882,1908]" class="Insecta" family="Cicadellidae" genus="Gypona" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis id="DCE91E0C5759FF88FDD29401FD2DF8C4" box="[600,692,1882,1908]" italics="true" pageId="22" pageNumber="23">C. nana</emphasis>
|
||
</taxonomicName>
|
||
, but the new species is separated in having the pair of lateral processes abruptly expanded medially and not parallel to the shaft along the entire length as in
|
||
<taxonomicName id="299DB99D5759FF88FA989426FAF6F820" baseAuthorityName="Fowler" baseAuthorityYear="1903" box="[1298,1391,1917,1943]" class="Insecta" family="Cicadellidae" genus="Gypona" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis id="DCE91E0C5759FF88FA989426FAF6F820" box="[1298,1391,1917,1943]" italics="true" pageId="22" pageNumber="23">C. nana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |