146 lines
16 KiB
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146 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.220.2178" ID-GBIF-Dataset="07cc435f-a623-4a17-b510-a38bf0569844" ID-PMC="PMC3459032" ID-Pensoft-Pub="1313-2970-220-1" ID-PubMed="23077429" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-220-1" ModsDocOrigin="ZooKeys 220" ModsDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" checkinTime="1451248758882" checkinUser="pensoft" docAuthor="Henry, Thomas J." docDate="2012" docId="6535A145B562CAB5A369758F7AE40255" docLanguage="en" docName="ZooKeys 220: 1-114" docOrigin="ZooKeys 220" docSource="http://dx.doi.org/10.3897/zookeys.220.2178" docTitle="Tytthus femoralis Henry, sp. n." docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="FFA2D657D204FFEEFF8DF82F5143FF91" masterDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="26" updateTime="1668154436991" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Henry, Thomas J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>220</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>114</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.220.2178</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.220.2178</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-220-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152036991" LSID="urn:lsid:zoobank.org:act:B264F11F-69AD-4E03-8C44-70DE7060E461" httpUri="http://treatment.plazi.org/id/6535A145B562CAB5A369758F7AE40255" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
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<subSubSection pageId="25" pageNumber="26" type="nomenclature">
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<paragraph pageId="25" pageNumber="26">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:B264F11F-69AD-4E03-8C44-70DE7060E461" authority="Henry" class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus femoralis" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="femoralis">Tytthus femoralis Henry</taxonomicName>
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<taxonomicNameLabel pageId="25" pageNumber="26">sp. n.</taxonomicNameLabel>
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Figs 13, 14128-131
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph pageId="25" pageNumber="26">Diagnosis.</paragraph>
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<paragraph lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
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This species is distinguished by the combination of a black head, pronotum, and scutellum; translucent hemelytra usually marked with dark brown through middle of the corium; the fuscous antennal segment I, with only the apex pale; the pale antennal segment II having a fuscous basal band; the fuscous hind femur contrasting
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<pageBreakToken pageId="26" pageNumber="27" start="start">with</pageBreakToken>
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the pale yellow pro- and mesofemora; and the pale tibiae with the bases or
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<normalizedToken originalValue="“knees”">"knees"</normalizedToken>
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fuscous. All known specimens of this species are macropterous.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus femoralis" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="femoralis">Tytthus femoralis</taxonomicName>
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keys to
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<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus entrerianus" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="entrerianus">Tytthus entrerianus</taxonomicName>
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based on the pale tibiae with knee spots and the apically infuscated hind femora. It can be distinguished from
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<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus entrerianus" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="entrerianus">Tytthus entrerianus</taxonomicName>
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by the pale antennal segment II and the more extensively infuscated hind femur.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
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<paragraph pageId="26" pageNumber="27">Description.</paragraph>
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<paragraph pageId="26" pageNumber="27">Male (n =10, plus holotype in parentheses) (Fig. 13): Length to apex of hemelytron 2.56-3.42 mm (2.83 mm), length to base of cuneus 1.90-2.25 mm (2.08 mm), width across hemelytra 0.80-0.99 mm (0.88 mm). Head: Length 0.30-0.34 mm (0.35 mm), width across eyes 0.61-0.67 mm (0.63 mm), interocular width 0.27-0.29 mm (0.29 mm). Labium: Length 1.20-1.38 mm (1.28 mm). Antenna: Segment I length 0.38-0.50 mm (0.38 mm), II 0.94-1.21 mm (1.10 mm), III 0.58-0.72 mm (0.66 mm), IV 0.40-0.59 mm (0.58 mm). Pronotum: Length 0.34-0.38 mm (0.35 mm), basal width 0.75-0.90 mm (0.78 mm).</paragraph>
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<paragraph lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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Coloration: Head: Uniformly fuscous to black, with a large, yellow, interocular spot near inner margin of each eye, spots nearly converging on some specimens; eyes fuscous to dark
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<pageBreakToken pageId="27" pageNumber="28" start="start">reddish</pageBreakToken>
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brown. Labium: Pale yellow, with apical half of segment IV brown. Antenna: Segment I fuscous to black, with apical one quarter pale or yellowish; segment II pale yellowish brown, with a distinct fuscous band at base and sometimes with apex infuscated; segments III and IV black. Pronotum: Uniformly black. Mesoscutum: Uniformly black. Scutellum: Uniformly black. Hemelytron: Translucent, with a dark brown cloud or patch on apical half of cor- ium and inner apical half of clavus ranging from somewhat indistinct to a definite dark pattern (Figs 13, 14); membrane pale translucent brown. Ostiolar evaporative area: Dark brown to fuscous, with a yellow patch on posterior edge. Ventral surface: Thorax and abdomen uniformly fuscous to black. Legs: Coxae pale yellow, with basal thirds to halves dark brown; pro- and meso femora pale yellow, metafemur dark brown to fuscous, with basal third and apex pale yellow; tibiae pale, each with a fuscous base or
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<normalizedToken originalValue="“knee”">"knee"</normalizedToken>
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; tarsi, and claws pale yellow.
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</paragraph>
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<paragraph pageId="27" pageNumber="28">Structure, texture, and vestiture: Head: Shiny, impunctate; buccula relatively broad, tapering posteriorly, ending near level with middle of eye; set with scattered, semierect setae. Labium: Extending to metacoxe or base of abdomen; segment I extending beyond base of head onto xyphus to bases of procoxae. Antenna: Segment I sparsely set with recumbent setae and two erect, subapical bristlelike setae; segment II set with only very short, recumbent setae. Pronotum: Shiny, impunctate; anterior angles rounded; lateral margins weakly concave, widening at posterior angles; posterior margin weakly sinuate; calli weakly swollen; set with relatively long, recumbent setae. Mesoscutum: Weakly shining, impunctate, broadly exposed; set with a few scattered semierect setae. Scutellum: Shiny, impunctate; equilateral; set with scattered erect and semierect setae. Hemelytron: Macropterous, with cuneus and membrane fully developed, extending well beyond apex of abdomen; evenly set with relatively long, recumbent setae.</paragraph>
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<paragraph pageId="27" pageNumber="28">Male genitalia: Left paramere (Fig. 128): Mitt-shaped; right arm long, broad, acute apically; left arm short, apically acute. Right paramere (Fig. 129): Endosoma (Fig. 130): S-shaped. Phallotheca (Fig. 131): Slender, apically acute.</paragraph>
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<paragraph pageId="27" pageNumber="28">Female (n =5) (Fig. 14): Length to apex of hemelytron 2.82-3.52 mm mm, length to base of cuneus 2.05-2.50 mm, width across hemelytra 1.00-1.14 mm. Head: Length 0.30-0.35 mm, width across eyes 0.62-0.69 mm, interocular width 0.30-0.32 mm. Labium: Length 1.12-1.50 mm. Antenna: Segment I length 0.32-0.50 mm, II 0.85-1.17 mm, III 0.53-0.75 mm, IV 0.54-0.59 mm.</paragraph>
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<paragraph pageId="27" pageNumber="28">Pronotum: Length 0.40-0.41 mm, basal width 0.91-0.99 mm.</paragraph>
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<paragraph pageId="27" pageNumber="28">Similar to males in general appearance and coloration, differing primarily in the overall broader form. The one color exception is that the ventral area of the abdomen in many females is pale yellow, whereas in males the abdomen is always uniformly fuscous to black.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="etymology">
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<paragraph pageId="27" pageNumber="28">Etymology.</paragraph>
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<paragraph pageId="27" pageNumber="28">The specific epithet A "femoralis" denotes the dark brown to fuscous hind femur in contrast to the uniformly pale pro- and mesofemora.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="hosts">
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<paragraph pageId="27" pageNumber="28">Hosts.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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Four specimens from San Carlos, Ecuador, with the label a "Host-Eggs of
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<taxonomicName class="Insecta" family="Delphacidae" genus="Perkinsiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perkinsiella" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Perkinsiella</taxonomicName>
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spp. " Three specimens intercepted on
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<taxonomicName class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">Musa</taxonomicName>
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sp. certainly represent incidental or sitting records.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="distribution">
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<paragraph pageId="27" pageNumber="28">Distribution.</paragraph>
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<paragraph pageId="27" pageNumber="28">So far recorded from Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, Guatemala, Honduras, Jamaica, Mexico, Panama, and Peru.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="type material">
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<paragraph pageId="27" pageNumber="28">Type material.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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Holotype ♂ (00161890) (USNM): ECUADOR: Manabi: Bahia de Caraquez (35 kms SE), 10 May 1975, Ashley B. Gurney. Paratypes: BOLIVIA:El Beni: Capivara on Rio Itenez, approx. 20 km E. of Versalles, 22 Jul 1964, J. K. Bouseman, J. Lussenhop, 1 ♂ (00165937) (AMNH). BRAZIL:Amazonas: Reserva Ducke, 25 km NNE Manaus,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-2.9136">2.9136°S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-59.9464">59.9464°W</geoCoordinate>
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, 120 m, 26 Jul 1973, R. T. Schuh, 1 ♀ (00165831) (AMNH). COLOMBIA:Valle del Cauca: Jamundi,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="3.25833">3.25833°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="555" value="-76.54">76.54°W</geoCoordinate>
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, 13 Jul 1985, F. Garcia, on Jameo arroz (
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<taxonomicName genus="Poaceae" lsidName="Poaceae" pageId="27" pageNumber="28" rank="genus">Poaceae</taxonomicName>
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), 6 ♂♂ (00161854 - 00161859), 2 ♀♀ (00161860, 00161861) (USNM). COSTA RICA:San Jose: San Jose,
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="9.9333">9.9333°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-84.0833">84.0833°W</geoCoordinate>
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, 1147 m, 1932, H. Schmidt, 1 ♂ (00165935), 1 ♀ (00165936) (AMNH). CUBA: Origin unknown, intercepted at Baltimore, Maryland by APHIS/PPQ, 26 Nov 1925, in
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<normalizedToken originalValue="“ship’s">"ship's</normalizedToken>
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light socket"), 1 ♀ (00161862) (USNM). ECUADOR:Chimborazo: Huigra, 15 Jun 1914, H. S. Parish, 1 ♂ (00161899) (USNM). El Oro: Victoria-Arenillas, 150 m, 18 Aug 1977 - 19 Aug 1977, L. Pena G., 1 ♀ (00161931) (USNM). Guayas: Duran, 91 m, 23 Jun 1914, H. S. Parish, 2 ♂♂ (00161900, 00161901) (USNM), 2 ♂♂ (CU). Ingenio San Carlos, 07 May 1982, Robert Morey,
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<normalizedToken originalValue="host–eggs">host-eggs</normalizedToken>
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of
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<taxonomicName class="Insecta" family="Delphacidae" genus="Perkinsiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perkinsiella" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Perkinsiella</taxonomicName>
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spp., 1 ♂ (00161898), 3 ♀♀ (00161930, 00161936 - 00161937) (USNM). Los Rios: Babahoyo, 21 Jun 1975, Cohen, Langley & Monnig, at blacklight, 1 ♂ (00161934) (USNM). Chone,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.6833">0.6833°S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5555" value="-80.1">80.1°W</geoCoordinate>
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, 110 m, 09 May 1975, Ashley B. Gurney, 2 ♂ (00161929, 00161935) (USNM). Manabi: Chone,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.6833">0.6833°S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5555" value="-80.1">80.1°W</geoCoordinate>
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, 110 m, 09 May 1975, Ashley B. Gurney, 2 ♂♂ (00161929, 00161935) (USNM). Napo: Baeza (72 Km E), 1280 m, 14 Apr 1977, Elaine R. Hodges, near cut trees and bamboo, 1 ♂ (00161933) (USNM). Origin unknown, intercepted at Long Beach, California, by APHIS/PPQ, 12 May 1997, on
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<taxonomicName class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">Musa</taxonomicName>
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sp., 1 ♂ (00161891), 1 ♀ (00161932) (USNM). Origin unknown, intercepted at San Diego, California by APHIS/PPQ,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="32.71528">32.71528°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-117.15639">117.15639°W</geoCoordinate>
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, 20 May 2008, on
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<taxonomicName class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">Musa</taxonomicName>
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sp. (fruit), 1 ♂ (00161902) (USNM). GUATEMALA: Locality unknown, 13 Sep 2007, P. Perez,
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<taxonomicName class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">Musa</taxonomicName>
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sp. (
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<taxonomicName genus="Musaceae" lsidName="Musaceae" pageId="27" pageNumber="28" rank="genus">Musaceae</taxonomicName>
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), 1 ♀ (00161864) (USNM). HONDURAS:Atlantida: Lancetilla, 1900 - 1900, M. Bates, Paratype, 2 ♂♂ (00165933, 00165934) (AMNH). JAMAICA:St. Ann Parish: 5 mi S of St Anns Bay, 05 Jul 1971, J. A. Slater & R. M. Baranowski & J. E. Harrington, 1 ♂ (00165830) (AMNH). MEXICO: Origin unknown, 03 Nov 1988, intercepted at Elpaso, Texas, by APHIS/PPQ, on
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<taxonomicName class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">Musa</taxonomicName>
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sp. (
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<taxonomicName genus="Musaceae" lsidName="Musaceae" pageId="27" pageNumber="28" rank="genus">Musaceae</taxonomicName>
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), 1 ♀ (00161863) (USNM). PANAMA:Pinogana: El Real, 08 Aug 1952, F. S. Blanton, 4 ♀♀ (00161850 - 00161853), 1 ♀ (00161849) (USNM). PERU:Huanuco:Leoncio Prado Co.: Tingo Maria, 671 m, 19 Apr 1969-24 Apr 1969, P. & P. Spangler, 1 ♂ (00161938), 1 ♀ (00161939) (USNM).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |