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<document ID-CLB-Dataset="22281" ID-DOI="http://dx.doi.org/10.3897/zookeys.1144.95619" ID-GBIF-Dataset="ec52f039-1a9b-443c-a427-e8d905fead36" ID-PMC="PMC10208442" ID-Pensoft-Pub="1313-2970-1144-1" ID-Pensoft-UUID="9485B6EA2F8E54B49F02E975297B6536" ID-PubMed="37234380" ID-ZooBank="565B46A4C01B454296356F3ED6472747" ModsDocID="1313-2970-1144-1" checkinTime="1675346952171" checkinUser="pensoft" docAuthor="Feijen, Hans R. &amp; Feijen, Cobi" docDate="2023" docId="50EF0FE744CA5020B028348CD8E248ED" docLanguage="en" docName="ZooKeys 1144: 1-93" docOrigin="ZooKeys 1144" docPubDate="2023-02-01" docSource="http://dx.doi.org/10.3897/zookeys.1144.95619" docTitle="Centrioncus copelandi Feijen &amp; Feijen 2023, sp. nov." docType="treatment" docUuid="BC6DFC78-D93E-4A68-B5FE-24EE081A4227" docUuidSource="ZooBank" docVersion="7" id="9485B6EA2F8E54B49F02E975297B6536" lastPageNumber="1" masterDocId="9485B6EA2F8E54B49F02E975297B6536" masterDocTitle="A revision of Centrioncus Speiser (Diptera, Diopsidae, Centrioncinae) with descriptions of new species from Angola, Burundi, and Kenya" masterLastPageNumber="93" masterPageNumber="1" pageNumber="1" updateTime="1720466234016" updateUser="ExternalLinkService">
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<mods:title id="6DEFCC3E659FA8744D7B26B5E798313D">A revision of Centrioncus Speiser (Diptera, Diopsidae, Centrioncinae) with descriptions of new species from Angola, Burundi, and Kenya</mods:title>
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<mods:namePart id="11F1F080122FBC9896F9AF9CBC10672E">Feijen, Hans R.</mods:namePart>
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<mods:affiliation id="2D1401CF33A99ABE107BE22B52CDE201">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
<mods:nameIdentifier id="D7176B210874081A84DF1E7282DA45D3" type="email">hans.feijen@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="5650310AA5BDF3D2D5B2D714071737E9">Feijen, Cobi</mods:namePart>
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<mods:affiliation id="7091C2A4BD0A44256D8D78CAB90ED174">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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<treatment id="50EF0FE744CA5020B028348CD8E248ED" ID-GBIF-Taxon="209152263" LSID="urn:lsid:zoobank.org:act:BC6DFC78-D93E-4A68-B5FE-24EE081A4227" httpUri="http://treatment.plazi.org/id/50EF0FE744CA5020B028348CD8E248ED" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="CA57E82ACBCDDCCB3D0360984CA6350F" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="A57E1D115173A2BD00861AFB2DDE8B23" pageId="0" pageNumber="1">
<taxonomicName id="1AEB0E689B2C92FE19D26BEC14C85146" ID-CoL="BDM77" LSID="https://zoobank.org/BC6DFC78-D93E-4A68-B5FE-24EE081A4227" authority="Feijen &amp; Feijen, 2023" authorityName="Feijen &amp; Feijen" authorityYear="2023" class="Insecta" family="Diopsidae" genus="Centrioncus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Centrioncus copelandi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="copelandi" status="sp. nov.">Centrioncus copelandi</taxonomicName>
<taxonomicNameLabel id="3EEC4D7F80582CE0D1E32B54A2F55F03" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F73514DF433EE5CABF4364DDA45994CB" pageId="0" pageNumber="1" type="description">
<paragraph id="74AD393FB8E8B28DA42A983AC2BF70ED" pageId="0" pageNumber="1">
<figureCitation id="CE1F8C40F4543BA35EB2CA83D375DD73" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Centrioncus, dorsal view of wings 1 ♂, C. aberrans, Timboroa Forest 2 ♀, paratype, C. bururiensis sp. nov., Bururi Nat. Forest 3 ♂ C. bytebieri, Ngangao Forest 4 ♀, paratype, C. copelandi sp. nov., Kasigau Mtn. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures1-4" httpUri="https://binary.pensoft.net/fig/803914" pageId="0" pageNumber="1">Figs 4</figureCitation>
<figureCitation id="7E1027E7415FE4E5DC4F9CE16F2062EA" captionStart="Figure 29" captionStartId="F8" captionText="Figure 29. Distribution map of the seven Eastern African Centrioncus species." figureDoi="10.3897/zookeys.1144.95619.figure29" httpUri="https://binary.pensoft.net/fig/803921" pageId="0" pageNumber="1">, 29</figureCitation>
<figureCitation id="9BCF3F6134E8DCCAE9750B04B1F4C1B8" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">, 74-79</figureCitation>
<figureCitation id="156CFDA8AD60ABAFB792BB8D2BF1E28A" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">, 80-83</figureCitation>
<figureCitation id="E2773E8DE0A7B60AE63B9015719EA365" captionStart="Figures 8487" captionStartId="F21" captionText="Figures 84 - 87. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 84 subanal plate, ventral view 85 spermathecae 86 egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (84, 85, 87); 0.2 mm (86)." figureDoi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1">, 84-87</figureCitation>
<figureCitation id="F0019EBA4F4B87CF5524A3DD27247AF5" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">, 88-91</figureCitation>
<figureCitation id="612394F6E30A25807CE2EBF80A1047CB" captionStart="Figure 92" captionStartId="F23" captionText="Figure 92. Centrioncus copelandi sp. nov., Kenya, Kasigau Mtn, collection site at 1117 m. Photograph by Robert Copeland." figureDoi="10.3897/zookeys.1144.95619.figure92" httpUri="https://binary.pensoft.net/fig/803936" pageId="0" pageNumber="1">, 92</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D31CE479146DA741AB8FAC6D80B02C9A" pageId="0" pageNumber="1" type="type material">
<paragraph id="723B196BDB95C5A544BA2B0E6238B1AD" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="531A09F1E07A8B1EEE43A5FA1F2A6335" pageId="0" pageNumber="1">
Kenya,
<emphasis id="B1F5874FD744102F660599032DB0CB28" bold="true" italics="true" pageId="0" pageNumber="1">holotype</emphasis>
, ♂, Coastal Prov., Kasigau Mtn,
<geoCoordinate id="181717B292B8896AFD6E8A4892DAD57C" degrees="3.82700" direction="south" orientation="latitude" precision="1" value="-3.827">3.82700°S</geoCoordinate>
,
<geoCoordinate id="B9035EA3D4A722B72B9D4C1A6E31190F" degrees="38.64875" direction="east" orientation="longitude" precision="1" value="38.64875">38.64875°E</geoCoordinate>
, 1065 m, 28.xii.2011-11.i.2012, R. Copeland (NMKE).
<emphasis id="E3879E54EB78FC028B25E1D11A72E1EC" bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
: 1 ♀, Coastal Prov., Kasigau Mtn,
<geoCoordinate id="96276CF3AA65A37B72525612AC64DA74" degrees="3.82667" direction="south" orientation="latitude" precision="1" value="-3.82667">3.82667°S</geoCoordinate>
,
<geoCoordinate id="AAFADA795207778DB4A4CD43CBE525D1" degrees="38.64982" direction="east" orientation="longitude" precision="1" value="38.64982">38.64982°E</geoCoordinate>
, 1117 m, 30.vi-13.vii.2011, R. Copeland (NMKE).
</paragraph>
</subSubSection>
<subSubSection id="FF8B76C7C1D4F77D3C21C147F9CF6AB9" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="C66F42C34D0964543197C904CEF75FCF" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="79A0D2CC65324C64DB9D3F0F5994D570" pageId="0" pageNumber="1">
<taxonomicName id="30FE8E84DC5AB3048B5D1848EF730C06" authorityName="Feijen &amp; Feijen" authorityYear="2023" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus copelandi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="copelandi">
<emphasis id="C87555BFDC93BA874CC7EA405A64893A" italics="true" pageId="0" pageNumber="1">Centrioncus copelandi</emphasis>
</taxonomicName>
sp. nov. can be recognised by a mesally slightly depressed, pruinose frons with glossy spots; glossy collar; pruinose blackish brown scutum; dark brown scutellum with brown edges and spines; pleura blackish brown (Figs
<figureCitation id="0FE67496C6A04E38CF0BB346296533FB" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
,
<figureCitation id="D1AD2F04F9854526E6795925A0B88ACE" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">76</figureCitation>
); scutellar spine/scutellum ratio: 0.87-1.00; apical seta/scutellar spine ratio: 0.86-0.92; brown, strongly incrassate fore femur (l/w ratio: ~ 2.65) with ~ 31.7 tubercles, dark brown stripe dorsally on apical third of inner side; very large, distinct central wing spot in distal third of cell br, basal two-fifths of cell r4+5 and distal two-thirds of cell bm+dm (Fig.
<figureCitation id="2CC9505C167FB7ED5264AB762483120F" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Centrioncus, dorsal view of wings 1 ♂, C. aberrans, Timboroa Forest 2 ♀, paratype, C. bururiensis sp. nov., Bururi Nat. Forest 3 ♂ C. bytebieri, Ngangao Forest 4 ♀, paratype, C. copelandi sp. nov., Kasigau Mtn. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures1-4" httpUri="https://binary.pensoft.net/fig/803914" pageId="0" pageNumber="1">4</figureCitation>
); tergites blackish brown; female 7th spiracle in tergite; sternite 4 rectangular; sternite 5 trapezoidal, invaginated anteriorly; sternites 4 and 5 with pair of strongly sclerotised spots; sternite 6 trapezoidal, 1.5
<normalizedToken id="29CF6C4D26E420A3DC26A78348046769" originalValue="×">x</normalizedToken>
as broad as sternites 1-5; anterior sclerite of female sternite 7 with w/l ratio: ~ 3.1 (Fig.
<figureCitation id="BDC3802E47AE688B8F100CFDB739DDAD" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">81</figureCitation>
); posterior sclerite of female sternite 7 truncated U-shaped, straight anterior and lateral edges with angular anterolateral corners; female cercus elongate, l/w ratio: ~ 4.4; subanal plate pentagonal, apically tapering into short extension; spermathecae rounded, basally flat (Fig.
<figureCitation id="0013530329D748887B7F848AF927FD2C" captionStart="Figures 8487" captionStartId="F21" captionText="Figures 84 - 87. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 84 subanal plate, ventral view 85 spermathecae 86 egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (84, 85, 87); 0.2 mm (86)." figureDoi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1">85</figureCitation>
), with small apical dimple; spermathecal ducts with constriction near spermathecae; outer and median arms of surstylus well separated, with short broad common base; outer arm constricted at base, apically broadening to 3
<normalizedToken id="A3F49F62E626F19FDCE21F8B184EA423" originalValue="×">x</normalizedToken>
width of base, sides concave, with 16 apical tubercles (Fig.
<figureCitation id="4477B6C68C47166EDA89D75A0891280C" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">88</figureCitation>
); median arm a slender rod, apically some long setulae but no spinous setae, slightly shorter than outer arm; inner arm twice as broad as median arm and slightly longer, apically with shape of cap opener; subepandrial clasper elongate, basal third constricted, apical corners angular, apically strongly convex; cercus (Fig.
<figureCitation id="7247A9283B29F4A909FCFD1C53724049" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">89</figureCitation>
) with broad lateral extension on distal third, apically convex.
</paragraph>
<caption id="F9BDAAA892916EB55F0277BA5226EC2B" doi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1" start="Figures 7479" startId="F19">
<paragraph id="62888CC235C5B0F0A8BFB1EC03A06AB0" pageId="0" pageNumber="1">
<emphasis id="03B58FA9FB198A347DEC4DCCFB301231" bold="true" pageId="0" pageNumber="1">Figures 74-79.</emphasis>
<taxonomicName id="D2DC19768E47C0B3B8D8C79C5F880158" authorityName="Feijen &amp; Feijen" authorityYear="2023" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus copelandi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="copelandi">
<emphasis id="3ADDFD791A17D510B664C25D3FD52C02" italics="true" pageId="0" pageNumber="1">Centrioncus copelandi</emphasis>
</taxonomicName>
sp. nov., Kasigau Mtn
<emphasis id="28FE7AAE51B43CBBE158325B7AC5C61B" bold="true" pageId="0" pageNumber="1">74</emphasis>
♂, holotype, lateral view (abdomen removed)
<emphasis id="8C75AEF6F91BCBA64A4F9C8E7A0CD237" bold="true" pageId="0" pageNumber="1">75-79</emphasis>
♀, paratype
<emphasis id="36A7CD99F34E61310B815DDB8B28EC59" bold="true" pageId="0" pageNumber="1">75</emphasis>
head, frontal view
<emphasis id="706D4459FF433717998098CC497E4077" bold="true" pageId="0" pageNumber="1">76</emphasis>
head posterolateral view, thorax, lateral view
<emphasis id="CDA224DA2514A7E589340999414501AD" bold="true" pageId="0" pageNumber="1">77</emphasis>
abdomen, dorsal view
<emphasis id="272A3F8F99583313D95C47FC11010F8E" bold="true" pageId="0" pageNumber="1">78</emphasis>
fore femur, ventral view
<emphasis id="B74862F0179AAE836331339F791C5A67" bold="true" pageId="0" pageNumber="1">79</emphasis>
hind femur, ventral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="3695ACF871005222B2A700EE95D4939C" pageId="0" pageNumber="1" type="description">
<paragraph id="9E0DC65DDC7E759A70DC993513F056C9" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="80F6B00C2E1A2B267D9AA6520CED3F55" pageId="0" pageNumber="1">
<emphasis id="C7AF84CA7B1C5E0BCF1AAF6A5B934D96" bold="true" pageId="0" pageNumber="1">
<emphasis id="F294320FD937BE5B31542164A62895C4" italics="true" pageId="0" pageNumber="1">Measurements</emphasis>
.
</emphasis>
Body length ♀ 5.1 mm, ♂ 5.0 mm, width of head ♀ 1.08 mm, ♂ 1.13 mm, wing length ♀ 4.3 mm, ♂ 4.3 mm, length of scutellar spine ♀ 0.34 mm, ♂ 0.31 mm.
</paragraph>
<paragraph id="F28BB47FD7038EDC0895FB220FB2724D" pageId="0" pageNumber="1">
<emphasis id="12F7D58D04505D524D2C1AA87DF75A47" bold="true" pageId="0" pageNumber="1">
<emphasis id="670AC12D6130C88B105CD5C0A62F0C21" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Frons slightly depressed mesally, brown but mesally and anteriorly paler brown, black stripe posterior to FOS; frons pruinose, anterolaterally more densely pruinose, black stripe behind FOS glossy, small glossy spots lateral to ocellar tubercle (Fig.
<figureCitation id="AACB0BAED2C1BAD4FCEA9DBB9E9BF3BE" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">75</figureCitation>
); occiput blackish brown, with lateroventral corners yellowish brown (Fig.
<figureCitation id="8E3CE90CCD0EEACF6FF0EB62877292A3" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">76</figureCitation>
), uniformly pruinose, more densely pruinose dorsomesally; face yellowish brown, pruinose, denser pruinosity along eye margins; antenna yellowish brown, funiculus with dark spot at base of arista; maxillary palpus dark; OVS 0.31 mm in ♀ and 0.29 in ♂, FOS 0.24 mm in ♀ and ♂.
</paragraph>
<paragraph id="EF706863399007C2F8DE953CF9D937EB" pageId="0" pageNumber="1">
<emphasis id="9E947F504235B1D549B0FFB14FCB851A" bold="true" pageId="0" pageNumber="1">
<emphasis id="E522926010E841E8E2E34BC1843B140B" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
.
</emphasis>
Collar glossy dark brown; scutum blackish brown, pruinose, posterior half of humeral callus glossy (Fig.
<figureCitation id="2FCAF0EBC3F76FE510A8C6425ECB1195" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">76</figureCitation>
); scutellum dark brown, pruinose, posterior and lateral edges and scutellar spines yellowish brown (Figs
<figureCitation id="E8CF6BEBCFEA19BE21435EEADAD2C79D" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
,
<figureCitation id="C6DBC8B0C7A1C88216C9DB933DACE0B1" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">76</figureCitation>
); pleura dark brown, pleura mainly pruinose, glossy sections include anterior proepisternum, ventral edge of anepisternum, ventral half of katepisternum and ventral spot on anepimeron, densely pruinose spots on posterior section of anepimeron and dorsoposterior section of katepisternum (Figs
<figureCitation id="928B51870BB61427787AD17F1C77A665" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
,
<figureCitation id="AB352FB83294B20352812B91A5B79B10" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">76</figureCitation>
); posterior notopleural, supra-alar and infra-alar setae present, infra-alar seta twice as long as posterior notopleural and supra-alar setae, supra-alar seta on inconspicuous carina (Fig.
<figureCitation id="61AB8B6512BACD18E4E129AADF99969A" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
); basiliform prosternum lanceolate and anteriorly sharply pointed and with line-like extension; scutal length/scutal width ratio: 1.1; scutellum projecting at angle of just&gt; 20° from body axis; scutellar spines almost aligned with dorsal plane of scutellum, diverging at angle of ~ 50°; scutellar spine/scutellum ratio: 1.00 in ♀ and 0.87 in ♂; scutellar spine/length of body ratio: 0.067 in ♀ and 0.063 in ♂; apical seta/scutellar spine ratio: 0.86 in ♀ and 0.92 in ♂; scutellar length/scutellar width (at base) ratio: ~ 0.70.
</paragraph>
<paragraph id="95FFA72F390F4294FCB852BC2FB2FD83" pageId="0" pageNumber="1">
<emphasis id="E3834EB83D4A803F7541C8396D89C04E" bold="true" pageId="0" pageNumber="1">
<emphasis id="0C8EF861D6CC0DB3E830C6DBF54990F0" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
.
</emphasis>
Almost transparent with very large, distinct, brownish central spot in distal third of cell br, basal two-fifths of cell r4+5, distal two-thirds of cell bm+dm and slightly extending into cells r2+3, m1 and m4 (Fig.
<figureCitation id="45C8EBD1CC773C932746E606AC4D2308" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Centrioncus, dorsal view of wings 1 ♂, C. aberrans, Timboroa Forest 2 ♀, paratype, C. bururiensis sp. nov., Bururi Nat. Forest 3 ♂ C. bytebieri, Ngangao Forest 4 ♀, paratype, C. copelandi sp. nov., Kasigau Mtn. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures1-4" httpUri="https://binary.pensoft.net/fig/803914" pageId="0" pageNumber="1">4</figureCitation>
); glabrous basal areas only include cells bc and c and small basal and subbasal spots in cell br; crossvein h distinct; cell sc closed; vein CuA+CuP from vein CuP onward extending under angle of 25° to wing margin in almost straight line (Fig.
<figureCitation id="76F5982DDA561721B34DD43FAD8A5AE1" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Centrioncus, dorsal view of wings 1 ♂, C. aberrans, Timboroa Forest 2 ♀, paratype, C. bururiensis sp. nov., Bururi Nat. Forest 3 ♂ C. bytebieri, Ngangao Forest 4 ♀, paratype, C. copelandi sp. nov., Kasigau Mtn. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures1-4" httpUri="https://binary.pensoft.net/fig/803914" pageId="0" pageNumber="1">4</figureCitation>
); cell cua triangular; vein M4 continuing distal of crossvein dm-m in almost straight line to wing margin; alula distinct; crossvein bm-m very vaguely indicated.
</paragraph>
<paragraph id="85950614702A67E406D0E4615FF30694" pageId="0" pageNumber="1">
<emphasis id="43BFFABDBC29A045353ABCA50D571BCD" bold="true" pageId="0" pageNumber="1">
<emphasis id="AC17F2C5CD8777501136DD99AC6B7C02" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
.
</emphasis>
Coxa 1 and trochanter 1 whitish (Fig.
<figureCitation id="A9C9949EBA965500C77A84A074E4F019" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">76</figureCitation>
), thinly pruinose; femur 1 glossy, yellowish brown with dark brown stripe on apical third of dorsal part of inner side (Figs
<figureCitation id="C9DA9BA09E9101BD9D5DA29500E763DF" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
,
<figureCitation id="D9478626F3689CFE4CD53CA3333368C1" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">78</figureCitation>
); tibia 1 dark brown, pruinose; metatarsus 1 yellowish brown, other tarsal segments whitish; mid and hind legs pale yellowish brown; apex of femur 2 brown; femur 3 dark brown on apical seventh (Fig.
<figureCitation id="A1BB824C4412913749C39282D5BAAE9F" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
); tibia 3 with brown spots basally and apically (Fig.
<figureCitation id="ACE02E2B744E167116B2D293E89D70C2" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
); femur 1 strongly incrassate in ♀ and ♂ (Figs
<figureCitation id="347FC1A1E2C8D84DB162F996E5BFCC3B" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">74</figureCitation>
,
<figureCitation id="2C0021167B0D7F4409CCB8F7D1579432" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">78</figureCitation>
), l/w ratio: 2.62 in ♀ (
<emphasis id="9E0929BE64D4305E6878E3E4CC428001" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 1) and 2.67 in ♂ (
<emphasis id="6F3A41445E6D91C5A34FF0B400FA7E08" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 1), inner row of spinous setae (Fig.
<figureCitation id="F541D3F7F87185DB023F9146306D8D9F" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">78</figureCitation>
) with 4-5 setae (
<emphasis id="B7E0F5BFD8F699C1BF5F4266D81BC1A0" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3), outer row with 4 setae (
<emphasis id="1F8BCB96E4EC177A2075F75DE73CF459" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3), inner row of tubercles with 15.3 tubercles (range 15-16,
<emphasis id="CBB2BC42AE63C70E41F3B4413CD444EF" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3), outer row with 16.3 tubercles (range 15-17,
<emphasis id="C8F5A873CEDEAD60CDDBFEA0B4EEAD7B" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3); femur 3 (Fig.
<figureCitation id="E5BA93A3E4EA8295032789955F66E06B" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">79</figureCitation>
) with 5.0 tubercles (range 4-6,
<emphasis id="F1FE3855C4B02D41A4906782970422F6" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3); setal formula 4.0, 4.7, 16.3, 15.3, 5.0; femora 2 and 3 without apical spurs.
</paragraph>
<paragraph id="CD45FB7B0B344AC39FE558D613CB09A5" pageId="0" pageNumber="1">
<emphasis id="C4069436AC5EB06D814984AFBFBAD33E" bold="true" pageId="0" pageNumber="1">
<emphasis id="E4048633442CDC790FF80543095F9D9E" italics="true" pageId="0" pageNumber="1">Preabdomen</emphasis>
.
</emphasis>
Tergites blackish brown (Fig.
<figureCitation id="9D5C3E51EB8180DAC47D3756AC0BD74C" captionStart="Figures 7479" captionStartId="F19" captionText="Figures 74 - 79. Centrioncus copelandi sp. nov., Kasigau Mtn 74 ♂, holotype, lateral view (abdomen removed) 75 - 79 ♀, paratype 75 head, frontal view 76 head posterolateral view, thorax, lateral view 77 abdomen, dorsal view 78 fore femur, ventral view 79 hind femur, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1144.95619.figures74-79" httpUri="https://binary.pensoft.net/fig/803932" pageId="0" pageNumber="1">77</figureCitation>
), very thinly pruinose, posterior edges of tergites 2 and 3 with more densely pruinose band; sternites dark brown; lateral edges of membranous ventral areas with dark brown spots; sternite 1 rectangular, constricted mesally (Fig.
<figureCitation id="3C4F3A5D1187AD6B227DA922697B1FC7" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">80</figureCitation>
); intersternite 1-2 very dark, laterally acuminate, with thin lateral connections to main sternite 2; sternite 2 rectangular, posterior of intersternite with small membranous area; sternite 3 rectangular with two narrow, elongate, more sclerotised areas (Fig.
<figureCitation id="85FA26076297A2186266DA8DFD383FE3" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">80</figureCitation>
); sternite 4 rectangular with convex lateral sides and two distinct, heavily sclerotised spots anteriorly; sternite 5 trapezoidal, anteriorly strongly constricted mesally, with two distinct, heavily sclerotised spots anteriorly (Fig.
<figureCitation id="08914064160FFCEF41C0BD83A62102BB" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">80</figureCitation>
); sternite 6 trapezoidal, ~ 1.5
<normalizedToken id="F1FB45292ADD52B53969D94AF941D637" originalValue="×">x</normalizedToken>
as broad as sternites 1-5; sternites 1-3 glossy, sternites 1 and 3 with a few microtrichia laterally; sternites 4-6 pruinose; 1st spiracle in membrane.
</paragraph>
<caption id="F4CF998B48BD004A1F7A0CA881A9CFAD" doi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1" start="Figures 8083" startId="F20">
<paragraph id="7DD8AE981499CB4342EC0AC7FECB7EFF" pageId="0" pageNumber="1">
<emphasis id="55F015B013CEF6E3B27B765B022F527B" bold="true" pageId="0" pageNumber="1">Figures 80-83.</emphasis>
<taxonomicName id="04DED6C68CE1EDBEE4AB2427F8B6E4EE" authorityName="Feijen &amp; Feijen" authorityYear="2023" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus copelandi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="copelandi">
<emphasis id="3B5CDD4B5187EF672A6D3BF3986B4183" italics="true" pageId="0" pageNumber="1">Centrioncus copelandi</emphasis>
</taxonomicName>
sp. nov., ♀, paratype, Kasigau Mtn
<emphasis id="EA4435B853B8556DBFCD20F97B522CC0" bold="true" pageId="0" pageNumber="1">80</emphasis>
sternites 1-6, ventral view
<emphasis id="A31EEFE041BFED3AB8C9A6A57E283939" bold="true" pageId="0" pageNumber="1">81, 82</emphasis>
sternite 7, ventral view
<emphasis id="98EAECF1D2572661531E50D229A0C082" bold="true" pageId="0" pageNumber="1">83</emphasis>
terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm
</paragraph>
</caption>
<paragraph id="8C7AC9D11A75CD85ACAA7847D9327266" pageId="0" pageNumber="1">
<emphasis id="05D0CAD01F8DA3D0226FDDFF88F5DE0F" bold="true" pageId="0" pageNumber="1">
<emphasis id="DB84022CA0EBEABB19B79DC9B95CE652" italics="true" pageId="0" pageNumber="1">Female postabdomen</emphasis>
.
</emphasis>
Tergite 7 brown, with mesal gap posteriorly, posterior gap extending to lateral sides, lateral edges curved under ventrally; suture in basal ring of segment 7 faint; 7th spiracle well into tergite (Fig.
<figureCitation id="A503F95694B4DFC2C0DC1215C5935F5F" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">82</figureCitation>
); anterior sclerite of sternite 7 rectangular, slightly tapering posteriorly (Figs
<figureCitation id="85EA1B0E2589A4C2982F6E93F8B0331C" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">81</figureCitation>
,
<figureCitation id="86F105A4CE453CCE7D8714A6422E08AB" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">82</figureCitation>
), w/l ratio: ~ 3.1 (Table
<tableCitation id="FF40BE0DC94198C6E76B49EB465DB300" captionStart="Table 8" captionStartId="T8" captionText="Table 8. Ratio width / length for the anterior sclerite of ♀ sternite 7, ratio length / width for ♀ cercus and ratio length / greatest width for ♂ cercus. Species are arranged according to three species-groups recognised." httpUri="http://table.plazi.org/id/BA654C5FE474EF511E0FDD042BFC63BE" pageId="0" pageNumber="1" tableUuid="BA654C5FE474EF511E0FDD042BFC63BE">8</tableCitation>
), dark brown, glossy on basal third, pruinose posteriorly (Figs
<figureCitation id="6715759869082A01F6F40642ABBC9970" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">81</figureCitation>
,
<figureCitation id="4B17355F188814BB09C8041CF26EE49C" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">82</figureCitation>
), with ~ 30 setulae; posterior sclerite of sternite 7 truncated U-shaped, consisting of broad anterior section and two broad lateral extensions (Figs
<figureCitation id="FDC54BCDB10AE73B7FCE1B08D7427936" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">81</figureCitation>
,
<figureCitation id="12C6C9DE8C8FADCD7A8231B13FF5B16C" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">82</figureCitation>
), anterior edge straight and parallel to posterior edge of anterior sternite, lateral edges of extensions straight and diverging posteriorly, anterolateral corners angular, posterolateral apices rounded, clothed in microtrichia and with four setulae at posterior apices; tergite 8 consisting of two oblong plates, well separated mesally (Fig.
<figureCitation id="A62AB6DE3B5956FF7608C1D1F2A95505" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">83</figureCitation>
); sternite 8 consisting of two oblong plates; tergite 10 (Fig.
<figureCitation id="EFCAE2DB0BBC9ECFBE62FDCCE6C38776" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">83</figureCitation>
) onion-shaped, extending posteriorly between cerci, clothed in microtrichia, with one pair of strong setulae; cercus elongate, somewhat tapering posteriorly (Fig.
<figureCitation id="A9566909A38022B1ED6724ECD1964286" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">83</figureCitation>
, Table
<tableCitation id="5BD204C9C26DD394D0617886ADA27B2E" captionStart="Table 8" captionStartId="T8" captionText="Table 8. Ratio width / length for the anterior sclerite of ♀ sternite 7, ratio length / width for ♀ cercus and ratio length / greatest width for ♂ cercus. Species are arranged according to three species-groups recognised." httpUri="http://table.plazi.org/id/BA654C5FE474EF511E0FDD042BFC63BE" pageId="0" pageNumber="1" tableUuid="BA654C5FE474EF511E0FDD042BFC63BE">8</tableCitation>
), l/w ratio: ~ 4.4; subanal plate (Figs
<figureCitation id="BF7E551FE5CDB6554E7291695F433DE7" captionStart="Figures 8083" captionStartId="F20" captionText="Figures 80 - 83. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 80 sternites 1 - 6, ventral view 81, 82 sternite 7, ventral view 83 terga 8, 10 and cerci, dorsal view. Scale bars: 0.2 mm" figureDoi="10.3897/zookeys.1144.95619.figures80-83" httpUri="https://binary.pensoft.net/fig/803933" pageId="0" pageNumber="1">83</figureCitation>
,
<figureCitation id="F7700BB7ECFDC6442B238A6E4611E68A" captionStart="Figures 8487" captionStartId="F21" captionText="Figures 84 - 87. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 84 subanal plate, ventral view 85 spermathecae 86 egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (84, 85, 87); 0.2 mm (86)." figureDoi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1">84</figureCitation>
) somewhat pentagonal, laterally rounded and apically tapering into short extension, clothed in microtrichia except anterior edge, with ~ 26 setulae; spermathecae heavily sclerotised, smooth, rounded, basally flattened (Fig.
<figureCitation id="985FCDCF0F56D8B3CABE92047E601768" captionStart="Figures 8487" captionStartId="F21" captionText="Figures 84 - 87. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 84 subanal plate, ventral view 85 spermathecae 86 egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (84, 85, 87); 0.2 mm (86)." figureDoi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1">85</figureCitation>
), with small apical dimple; junction of ducts of paired spermathecae V-shaped, spermathecal duct with distinct constrictions near junction with spermathecae (Fig.
<figureCitation id="18663EA46C0B939F7059FA13D2883A50" captionStart="Figures 8487" captionStartId="F21" captionText="Figures 84 - 87. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 84 subanal plate, ventral view 85 spermathecae 86 egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (84, 85, 87); 0.2 mm (86)." figureDoi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1">85</figureCitation>
).
</paragraph>
<caption id="D0A2189F3C07EB37EF0D30AAC03AF692" doi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1" start="Figures 8487" startId="F21">
<paragraph id="7BBC443B3CC3E4FDC7A9CA95DFF8259F" pageId="0" pageNumber="1">
<emphasis id="530D22ED20DCEADCA06CE1249A585172" bold="true" pageId="0" pageNumber="1">Figures 84-87.</emphasis>
<taxonomicName id="AD4382158BDD39FC609F018188F750C1" authorityName="Feijen &amp; Feijen" authorityYear="2023" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus copelandi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="copelandi">
<emphasis id="DED0E4C82904C09B89C6AABEDD62FF5F" italics="true" pageId="0" pageNumber="1">Centrioncus copelandi</emphasis>
</taxonomicName>
sp. nov., ♀, paratype, Kasigau Mtn
<emphasis id="26AA23ED7D30DCC6CB34BD1F7974E470" bold="true" pageId="0" pageNumber="1">84</emphasis>
subanal plate, ventral view
<emphasis id="1765D9AC041BD00DA927A3B291593134" bold="true" pageId="0" pageNumber="1">85</emphasis>
spermathecae
<emphasis id="0DE344B7DCBACA9A427A52BF85C265EA" bold="true" pageId="0" pageNumber="1">86</emphasis>
egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (
<emphasis id="64CE23B545B2E2753253F2DB0D6E80E9" bold="true" pageId="0" pageNumber="1">84, 85, 87</emphasis>
); 0.2 mm (
<emphasis id="7ADCFF056776FD5D727D0C7CC07E7D2D" bold="true" pageId="0" pageNumber="1">86</emphasis>
).
</paragraph>
</caption>
<paragraph id="1EAD3B399AE6AC49745477C5D1590D7F" pageId="0" pageNumber="1">
<emphasis id="FC8412BF94249D9F7DFE3703A75D91D1" bold="true" pageId="0" pageNumber="1">
<emphasis id="A147111505DA7904558DE1D6A2CD1E74" italics="true" pageId="0" pageNumber="1">Male postabdomen</emphasis>
.
</emphasis>
Tergite 6, sternite 8 and epandrium uniformly blackish brown; epandrium broad and rounded (too damaged to measure w/l ratio), clothed in microtrichia; outer and median arms of surstylus well separated, with short broad common base (Fig.
<figureCitation id="D49EDB13C99DAB0215F24C420DAC13DD" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">88</figureCitation>
); outer arm deeply constricted at base with concave lateral sides, broadening apically to ~ 3
<normalizedToken id="674FCEE1D7C89FF7B87C6646FBB4DA97" originalValue="×">x</normalizedToken>
basal width, apically with row of 16 tubercles, basally with nine setulae (2 very long) on inner side and a few small setulae along apical edge, on outer side with a few small setulae and central patch of microtrichia; median arm (Fig.
<figureCitation id="C21E2943DA2581D4E3DE82214182150F" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">88</figureCitation>
) slender, parallel-sided, rod-shaped, almost as long as outer arm, apically with four small setulae and two long setulae, no spinous setae present; inner arm twice as broad as median arm and slightly longer, apical quarter abruptly narrowed, cap opener-shaped, apically with seven setulae; subepandrial clasper (Fig.
<figureCitation id="A7D6DA38748000181AF004AA562D21EB" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">90</figureCitation>
) somewhat elongate, basal third constricted, apical corners angular, apically strongly convex, glabrous, with four long setulae on inner side; cercus (Fig.
<figureCitation id="86B3EC6A8585F54975DF7514423F39F9" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">89</figureCitation>
, Table
<tableCitation id="9F7C000821223F28A6757C82318A3371" captionStart="Table 8" captionStartId="T8" captionText="Table 8. Ratio width / length for the anterior sclerite of ♀ sternite 7, ratio length / width for ♀ cercus and ratio length / greatest width for ♂ cercus. Species are arranged according to three species-groups recognised." httpUri="http://table.plazi.org/id/BA654C5FE474EF511E0FDD042BFC63BE" pageId="0" pageNumber="1" tableUuid="BA654C5FE474EF511E0FDD042BFC63BE">8</tableCitation>
) slender, with broad lateral extension on distal third, apically convex, clothed in microtrichia and short setulae, 7 longer setulae along apical edge, length/greatest width ratio: 1.5 (Table
<tableCitation id="5D76BABD214F9282CC7FCEB8FAE36187" captionStart="Table 8" captionStartId="T8" captionText="Table 8. Ratio width / length for the anterior sclerite of ♀ sternite 7, ratio length / width for ♀ cercus and ratio length / greatest width for ♂ cercus. Species are arranged according to three species-groups recognised." httpUri="http://table.plazi.org/id/BA654C5FE474EF511E0FDD042BFC63BE" pageId="0" pageNumber="1" tableUuid="BA654C5FE474EF511E0FDD042BFC63BE">8</tableCitation>
); ejaculatory apodeme + sac (Fig.
<figureCitation id="ADE0086A32C2CCF93D1A9A530A393C50" captionStart="Figures 8891" captionStartId="F22" captionText="Figures 88 - 91. Centrioncus copelandi sp. nov., ♂, holotype, Kasigau Mtn 88 surstylus, inner view 89 cercus, posterior view 90 subepandrial clasper, posterior view 91 ejaculatory apodeme + sac. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1">91</figureCitation>
, Table
<tableCitation id="A9D5BD3785A89CC315FE9681C8787BBE" captionStart="Table 9" captionStartId="T9" captionText="Table 9. Length of ejaculatory apodeme + sac, length of body and ratio ejaculatory apodeme + sac / length of body for Centrioncus and Teloglabrus species. Measurements in mm, ratio as %. For some species, values are given for more than one specimen, partly from different locations." httpUri="http://table.plazi.org/id/A6225849E25FAE826DA6C72DEB78E285" pageId="0" pageNumber="1" tableUuid="A6225849E25FAE826DA6C72DEB78E285">9</tableCitation>
) very large, 10.1% of body length, ejaculatory apodeme slender, apically broadening.
</paragraph>
<caption id="B470B221D77C2C88F35A9FBD38426503" doi="10.3897/zookeys.1144.95619.figures88-91" httpUri="https://binary.pensoft.net/fig/803935" pageId="0" pageNumber="1" start="Figures 8891" startId="F22">
<paragraph id="E6558E9911033E7E2AA0236786F5FE89" pageId="0" pageNumber="1">
<emphasis id="19DAB231E4AADE8648EE2B56CCB07F2E" bold="true" pageId="0" pageNumber="1">Figures 88-91.</emphasis>
<taxonomicName id="295F6B1E3C993056B2E6CBB2690ED382" authorityName="Feijen &amp; Feijen" authorityYear="2023" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus copelandi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="copelandi">
<emphasis id="507031F2CA412305E1138B88DCDC8C8A" italics="true" pageId="0" pageNumber="1">Centrioncus copelandi</emphasis>
</taxonomicName>
sp. nov., ♂, holotype, Kasigau Mtn
<emphasis id="DA05869BA5ED42DB9DA66B4B642ED770" bold="true" pageId="0" pageNumber="1">88</emphasis>
surstylus, inner view
<emphasis id="5D289D6AF363731106B348358A2ABD82" bold="true" pageId="0" pageNumber="1">89</emphasis>
cercus, posterior view
<emphasis id="EEB053BA16D1B035219BC289E0C9A4AB" bold="true" pageId="0" pageNumber="1">90</emphasis>
subepandrial clasper, posterior view
<emphasis id="FAA2C6EAA36595BA6F6F1F3D9F24BBBD" bold="true" pageId="0" pageNumber="1">91</emphasis>
ejaculatory apodeme + sac. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="B198C058B678347A397639E33445F899" pageId="0" pageNumber="1">
<emphasis id="9299A5A12DDC8D8A8A923C4BABC7327A" bold="true" italics="true" pageId="0" pageNumber="1">Egg.</emphasis>
Female with two damaged full-grown eggs and some pieces of undeveloped eggs in abdomen. Eggs (Fig.
<figureCitation id="6382AA9F62A352AF38F7C23646589A17" captionStart="Figures 8487" captionStartId="F21" captionText="Figures 84 - 87. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 84 subanal plate, ventral view 85 spermathecae 86 egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (84, 85, 87); 0.2 mm (86)." figureDoi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1">86</figureCitation>
) measured 1.01 mm in length with elevated longitudinal ridges spanning from anterior pole to posterior pole with fine, roughly hexagonal microstructures between ridges (Fig.
<figureCitation id="4872BE12526B80A66E63289488BB590D" captionStart="Figures 8487" captionStartId="F21" captionText="Figures 84 - 87. Centrioncus copelandi sp. nov., ♀, paratype, Kasigau Mtn 84 subanal plate, ventral view 85 spermathecae 86 egg 87 egg, detail of external microstructure. Scale bars: 0.1 mm (84, 85, 87); 0.2 mm (86)." figureDoi="10.3897/zookeys.1144.95619.figures84-87" httpUri="https://binary.pensoft.net/fig/803934" pageId="0" pageNumber="1">87</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C7F6461A5B4D7D7DDF4B6953377B9E38" pageId="0" pageNumber="1" type="distribution">
<paragraph id="514421122731BFF7E35303EC1FE180A8" pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph id="DC7A2F243601BFC1A8261BA57C482051" pageId="0" pageNumber="1">
The localities of the type series are indicated on the map for Eastern Africa (Fig.
<figureCitation id="99DF9E0827A6B0DD804E7B09B8A4777D" captionStart="Figure 29" captionStartId="F8" captionText="Figure 29. Distribution map of the seven Eastern African Centrioncus species." figureDoi="10.3897/zookeys.1144.95619.figure29" httpUri="https://binary.pensoft.net/fig/803921" pageId="0" pageNumber="1">29</figureCitation>
). The collecting sites varied from 1056 to 1117 m in altitude. The habitat of the Kasigau site at 1117 m is shown in Fig.
<figureCitation id="62E44232809C82F4BA07412838E73A6D" captionStart="Figure 92" captionStartId="F23" captionText="Figure 92. Centrioncus copelandi sp. nov., Kenya, Kasigau Mtn, collection site at 1117 m. Photograph by Robert Copeland." figureDoi="10.3897/zookeys.1144.95619.figure92" httpUri="https://binary.pensoft.net/fig/803936" pageId="0" pageNumber="1">92</figureCitation>
. The distribution appears to be limited to the Kasigau Massif of the Taita Hills (see also under
<taxonomicName id="946E537CED14D0772232E925C4A8B722" lsidName="C. bytebieri" pageId="0" pageNumber="1" rank="species" species="bytebieri">
<emphasis id="03F8955E6CD7F0EC4343340ED04CAE38" italics="true" pageId="0" pageNumber="1">C. bytebieri</emphasis>
</taxonomicName>
).
<taxonomicName id="6D83595AEB2C2E2567C3EDE329FE6193" authorityName="De Meyer" authorityYear="2004" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus bytebieri" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bytebieri">
<emphasis id="F4DE7400BE3F47417425D7F4EA2E47E3" italics="true" pageId="0" pageNumber="1">Centrioncus bytebieri</emphasis>
</taxonomicName>
is only known from the Dabida Massif of the Taita hills. The gravid female was collected in early July, in the middle of the dry season which is unusual for
<taxonomicName id="4DA0D87B40A4ABD42B9BA19C5A1C458C" authorityName="Speiser" authorityYear="1910" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="EED29AEB45E6AA4EB8A5EDFB09BDFF47" italics="true" pageId="0" pageNumber="1">Centrioncus</emphasis>
</taxonomicName>
flies.
</paragraph>
<caption id="C2BF406AAA780F16B2E754911A75A4CC" doi="10.3897/zookeys.1144.95619.figure92" httpUri="https://binary.pensoft.net/fig/803936" pageId="0" pageNumber="1" start="Figure 92" startId="F23">
<paragraph id="1A22813E1853D300AA5C23810F4FA7F5" pageId="0" pageNumber="1">
<emphasis id="D7F486FFD535F72DA22FE496E38C415B" bold="true" pageId="0" pageNumber="1">Figure 92.</emphasis>
<taxonomicName id="FD5851F2ED4604614CF201B268E2BE22" authorityName="Feijen &amp; Feijen" authorityYear="2023" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus copelandi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="copelandi">
<emphasis id="70A8185202DC3EB052CDC0C9AC50E94C" italics="true" pageId="0" pageNumber="1">Centrioncus copelandi</emphasis>
</taxonomicName>
sp. nov., Kenya, Kasigau Mtn, collection site at 1117 m. Photograph by Robert Copeland.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="745E70B390A9FC79C7976523AD6A2D73" pageId="0" pageNumber="1" type="etymology">
<paragraph id="351D7469F3A26B7253A8000FA315CCF7" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="4CF7A789056EA34F3DE2B8DA87D2969B" pageId="0" pageNumber="1">
The specific epithet of
<taxonomicName id="A965A2EA62A435BADA10E073921000D5" lsidName="C. copelandi" pageId="0" pageNumber="1" rank="species" species="copelandi">
<emphasis id="625A87C1044EC3172ECCFEEDF7DE6FEE" italics="true" pageId="0" pageNumber="1">C. copelandi</emphasis>
</taxonomicName>
sp. nov. refers to the name of its collector Dr Robert S. Copeland (ICIPE), who contributed a significant number of
<taxonomicName id="2E6C8987F5E441A08CD02D8FA50B33DE" authorityName="Speiser" authorityYear="1910" class="Insecta" family="Diopsidae" genus="Centrioncus" kingdom="Animalia" lsidName="Centrioncus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="9EB5CF5B0DFF7253E10B3B4DFA79F12C" italics="true" pageId="0" pageNumber="1">Centrioncus</emphasis>
</taxonomicName>
specimens to this study.
</paragraph>
</subSubSection>
<subSubSection id="E6A3CB26A60E77D5CF10344D5C03AA8F" pageId="0" pageNumber="1" type="remarks">
<paragraph id="4FDAB4C500C459616CB85BE5A7C1AD44" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="0E385F43F7630D64DF80B7861E2D46F2" pageId="0" pageNumber="1">
From the shape of the outer arm of the surstylus (apically much wider than at base, with large number of 16 or 17 tubercles), absence of spinous setae on the median arm of the surstylus, the very large central wing spot and anteriorly strongly constricted sternite 5, it can be postulated that the closest known relative of
<taxonomicName id="BFF0C28798FC31BF2A4F4581C6F5A116" lsidName="C. copelandi" pageId="0" pageNumber="1" rank="species" species="copelandi">
<emphasis id="B10AA4A5FF63647A555A93F6A11EE2C9" italics="true" pageId="0" pageNumber="1">C. copelandi</emphasis>
</taxonomicName>
sp. nov. is
<taxonomicName id="FC2FE0DDD2DD4DE7D85FEE0BA99D752C" lsidName="C. prodiopsis" pageId="0" pageNumber="1" rank="species" species="prodiopsis">
<emphasis id="647615D376B165A4D972CED9B49A619E" italics="true" pageId="0" pageNumber="1">C. prodiopsis</emphasis>
</taxonomicName>
. From a geographical point of view that also makes some sense (Fig.
<figureCitation id="87D30777F38829454823AA0A39814568" captionStart="Figure 29" captionStartId="F8" captionText="Figure 29. Distribution map of the seven Eastern African Centrioncus species." figureDoi="10.3897/zookeys.1144.95619.figure29" httpUri="https://binary.pensoft.net/fig/803921" pageId="0" pageNumber="1">29</figureCitation>
), although
<taxonomicName id="D0A00A313BF349CAFEF394E488A7C066" lsidName="C. bytebieri" pageId="0" pageNumber="1" rank="species" species="bytebieri">
<emphasis id="A5DE919ACDB1C3A8E25CCD306FD702E9" italics="true" pageId="0" pageNumber="1">C. bytebieri</emphasis>
</taxonomicName>
occurs closer to
<taxonomicName id="51264B0E792CAEA8B91554228C6E29FA" lsidName="C. copelandi" pageId="0" pageNumber="1" rank="species" species="copelandi">
<emphasis id="AB9585593438D9565FED4C1A42544B81" italics="true" pageId="0" pageNumber="1">C. copelandi</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
</treatment>
</document>