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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="50964DE4152A5003D190E847CC3C46EA" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus minutus Caterino &amp; Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="139" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="136" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040966" LSID="urn:lsid:zoobank.org:act:B7D20E38-5376-4CD7-933B-127318B88B1B" httpUri="http://treatment.plazi.org/id/50964DE4152A5003D190E847CC3C46EA" lastPageId="138" lastPageNumber="139" pageId="135" pageNumber="136">
<subSubSection pageId="135" pageNumber="136" type="nomenclature">
<paragraph pageId="135" pageNumber="136">
<taxonomicName LSID="urn:lsid:zoobank.org:act:B7D20E38-5376-4CD7-933B-127318B88B1B" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus minutus" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="minutus">Operclipygus minutus</taxonomicName>
<taxonomicNameLabel pageId="135" pageNumber="136">sp. n.</taxonomicNameLabel>
Figs 35
<normalizedToken originalValue="GH38ADMap">G-H38A-DMap</normalizedToken>
13
</paragraph>
</subSubSection>
<subSubSection pageId="135" pageNumber="136" type="type locality">
<paragraph pageId="135" pageNumber="136">Type locality.</paragraph>
<paragraph pageId="135" pageNumber="136">
BRASIL:
<normalizedToken originalValue="Pará:Belém">Para:Belem</normalizedToken>
, Utinga [
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.45">1°27'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.433334">48°26'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="135" pageNumber="136" type="type material">
<paragraph pageId="135" pageNumber="136">Type material.</paragraph>
<paragraph pageId="135" pageNumber="136">
Holotype male: &quot;BRASIL:
<normalizedToken originalValue="Pará">Para</normalizedToken>
:
<normalizedToken originalValue="Belém">Belem</normalizedToken>
, Utinga (IPEAN),
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.45">1°27'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.433334">48°26'W</geoCoordinate>
.
<normalizedToken originalValue="Piège">Piege</normalizedToken>
<normalizedToken originalValue="dinterception">d'interception</normalizedToken>
. v.1985&quot; / &quot;Caterino/Tishechkin
<taxonomicName lsidName="" pageId="135" pageNumber="136" rank="tribe" tribe="Exosternini">Exosternini</taxonomicName>
Voucher EXO-01776&quot; (MNHN). Paratypes (5): BRAZIL:
<normalizedToken originalValue="Pará">Para</normalizedToken>
: 1:IPEAN, Utinga,
<normalizedToken originalValue="Belém">Belem</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.45">1°27'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.433334">48°26'W</geoCoordinate>
, viii.1985, FIT (FMNH), 1: xi.1984, FIT (CHND); 2: Altamira -
<normalizedToken originalValue="Marabá">Maraba</normalizedToken>
: km 18,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.15">3°09'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.05">52°03'W</geoCoordinate>
, v.1984, FIT (CHND, MSCC); 1: Barcarena,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.5">1°30'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.616665">48°37'W</geoCoordinate>
, 13-25.vi.1991, FIT (AKTC).
</paragraph>
</subSubSection>
<subSubSection lastPageId="138" lastPageNumber="139" pageId="135" pageNumber="136" type="description">
<paragraph pageId="135" pageNumber="136">Diagnostic description.</paragraph>
<paragraph lastPageId="138" lastPageNumber="139" pageId="135" pageNumber="136">
Length: 1.15-1.25 mm, width: 0.81-0.87 mm; body very small, rather elongate, parallel-sided; frons very shallowly depressed, frontal stria complete, transverse across front; supraorbital stria fine, complete, meeting sides of frontal stria; epistoma shallowly emarginate anteriorly, flat, coplanar with labrum; labrum with sides narrowed, apex weakly emarginate; left mandible with fairly prominent bifid tooth at base; right mandible with single acute tooth; pronotal disk with prescutellar impression linear, almost 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as scutellum; lateral submarginal pronotal stria absent; anterior marginal pronotal stria interrupted for width of head, but meeting corner of anterior submarginal stria, which is close to margin, recurved posterad for approximately one-third pronotal length; median pronotal gland open
<pageBreakToken pageId="136" pageNumber="137" start="start">ings</pageBreakToken>
situated alongside anterior submarginal stria, about 4 puncture widths from anterior margin; elytra with outer subhumeral stria present in apical half, inner subhumeral stria absent, striae 1-4 complete, 5th stria present in apical half, sutural stria present in apical three-fourths; ground punctation of elytral disks sparse but conspicuous; prosternal keel narrowly produced at base, carinal striae close, subparallel anteriorly, slightly diverging but connected basally, united in anterior arch about one-fourth from presternal suture; mesoventral margin emarginate, marginal stria abbreviated; mesometaventral stria narrowly arched forward to near mesoventral margin, displacing marginal stria; lateral metaventral stria extending toward middle of metacoxa, not abbreviated apically; 1st abdominal ventrite with two complete lateral striae close and parallel; propygidium with ground punctation sparse, well impressed, conspicuous, with small, shallow, round punctures separated by about their diameters; pygidium with fine ground punctation very dense, with coarser punctures fairly sparsely interspersed; marginal pygidial sulcus complete, fine but well impressed. Male genitalia (Figs 38
<normalizedToken originalValue="AD">A-D</normalizedToken>
): accessory sclerites present; T8 with sides subparallel, roundly converging to apex, apical emargination shallow, but with larger, distinctly desclerotized region of roughly the same shape surrounding it, basal emargination broad, shallow, basal membrane
<pageBreakToken pageId="137" pageNumber="138" start="start">attachment</pageBreakToken>
line distad emargination by about one-third its depth; S8 widest at base, sides rounded, convergent to narrow subacute apices, apical guides narrow, evenly developed throughout length, halves separate, diverging from base; T9 wider, subpar
<pageBreakToken pageId="138" pageNumber="139" start="start">allel</pageBreakToken>
in middle half, narrowed to basal apodemes, with prominent angles where narrowed to apex, apices curved inward, weakly subacute; T10 with halves separate; S9 desclerotized along midline, broad, narrowest near apex, widened to broad, truncate base, lateral flanges wide, widened laterally to prominent basal and apical corners, apex with narrow median emargination, apical flanges low and separate; tegmen with sides rounded, widest at middle, narrowed to subacute apices, curved downward in apical fourth, with ventral subapical cleft, medioventral process long, with truncate apex, projecting strongly beneath about one-third from base; basal piece about one-third tegmen length; median lobe short, only about one-fourth tegmen length.
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="139" type="remarks">
<paragraph pageId="138" pageNumber="139">Remarks.</paragraph>
<paragraph pageId="138" pageNumber="139">
This species is unusual in its small and narrowly elongate body form (Fig. 35G), as well as its lack of lateral submarginal pronotal stria. It is very similar to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impositus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="impositus">Operclipygus impositus</taxonomicName>
, but is generally slightly more convex, with slightly rounder sides. Male genitalia should be examined for unambiguous identification.
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="139" type="etymology">
<paragraph pageId="138" pageNumber="139">Etymology.</paragraph>
<paragraph pageId="138" pageNumber="139">
Even by
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
group standards this is a small species, and is named to reflect this fact.
</paragraph>
</subSubSection>
</treatment>
</document>