166 lines
22 KiB
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166 lines
22 KiB
XML
<document id="76CB6CA30476FE61E84408DA53E8F3AD" ID-DOI="http://dx.doi.org/10.3897/zookeys.689.12021" ID-GBIF-Dataset="907713bd-c7f2-4f8e-a75e-b7f9c958865f" ID-PMC="PMC5672588" ID-Pensoft-Pub="1313-2970-689-1" ID-PubMed="29200920" ID-ZooBank="2F40BF4AD35F4CC697D5976EC201E652" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-689-1" ModsDocOrigin="ZooKeys 689" ModsDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" checkinTime="1502744367222" checkinUser="pensoft" docAuthor="Lackner, Tomas & Leschen, Richard A. B." docDate="2017" docId="5051FB48CF9DC350B7C61B7A832371DF" docLanguage="en" docName="ZooKeys 689: 1-263" docOrigin="ZooKeys 689" docSource="http://dx.doi.org/10.3897/zookeys.689.12021" docTitle="Tomogenius kuscheli Dahlgren 1976" docType="treatment" docVersion="6" lastPageNumber="191" masterDocId="D302FFB6FFA66345FFBAFF89311EFFDE" masterDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" masterLastPageNumber="263" masterPageNumber="1" pageNumber="187" updateTime="1701316739810" updateUser="plazi">
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<mods:title id="C9F25E9705DD8D8C748FF47B020C1BA7">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:namePart id="5962E5710768116EB428A49EC2A66DE2">Lackner, Tomas</mods:namePart>
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<mods:namePart id="9CD14F2B0BBBA69C4BBEB268EBE3E86E">Leschen, Richard A. B.</mods:namePart>
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<mods:typeOfResource id="7228023E2698B22A1A4BD6659740F6A9">text</mods:typeOfResource>
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<mods:date id="4F92792CEDB08769B06C0B9E962EB61D">2017</mods:date>
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<mods:number id="AC627F9A27C189C77321B7D8E09B9CF6">689</mods:number>
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<mods:classification id="D6DDDC2627812839FD2DD29CE05BF668">journal article</mods:classification>
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<mods:identifier id="B0BB218A1A31779F1600A19099EB019A" type="DOI">http://dx.doi.org/10.3897/zookeys.689.12021</mods:identifier>
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<treatment id="5051FB48CF9DC350B7C61B7A832371DF" ID-GBIF-Taxon="132648697" LSID="urn:lsid:plazi:treatment:5051FB48CF9DC350B7C61B7A832371DF" httpUri="http://treatment.plazi.org/id/5051FB48CF9DC350B7C61B7A832371DF" lastPageId="190" lastPageNumber="191" pageId="186" pageNumber="187">
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<subSubSection id="FE6EA461F83EF74A78DF26C0AC155A13" pageId="186" pageNumber="187" type="nomenclature">
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<paragraph id="67071F3A4B524AFF780793C9E7A06A13" pageId="186" pageNumber="187">
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<taxonomicName id="8AB478EE99A10B09C7ADD50D48919C67" ID-CoL="7CH2L" authority="Dahlgren, 1976" authorityName="Dahlgren" authorityYear="1976" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius kuscheli" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="kuscheli">Tomogenius kuscheli Dahlgren, 1976</taxonomicName>
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Figs 660, 661-669, 670-678, 766
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</paragraph>
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</subSubSection>
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<subSubSection id="EE6F46DA36941C72310B03A9DD76156B" pageId="186" pageNumber="187" type="reference_group">
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<paragraph id="142940A0E6D469DDFF79891F1D12213D" pageId="186" pageNumber="187">
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<taxonomicName id="C34716F065ECAD6ABC1220EC4F4C5CA9" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius kuscheli" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="kuscheli">Tomogenius kuscheli</taxonomicName>
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Dahlgren, 1976: 409, fig. 22B.
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</paragraph>
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</subSubSection>
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<subSubSection id="FF446C232619F3DE2EFD9B4EACA9A4C1" pageId="186" pageNumber="187" type="type locality">
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<paragraph id="5762DFDCB8F7879A414240348360337C" pageId="186" pageNumber="187">Type locality.</paragraph>
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<paragraph id="79193424298DFD4E34A9C01AB40D6C98" pageId="186" pageNumber="187">New Zealand: Stephens Island.</paragraph>
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<caption id="E6627307E1F015D4689008A7206EC501" pageId="186" pageNumber="187">
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<paragraph id="446D361812D4ABAC3B6532F59536D59E" pageId="186" pageNumber="187">
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Figure 660.
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<taxonomicName id="E34FE56B32BA7FDBD86233ED9E640DB5" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius kuscheli" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="kuscheli">Tomogenius kuscheli</taxonomicName>
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Dahlgren, 1976 habitus, dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="6CF745EC74C6982E4DEDBA9EBD4A768F" pageId="186" pageNumber="187" type="type material">
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<paragraph id="E7C849BAD83537C312598063B68886F1" pageId="186" pageNumber="187">Type material examined.</paragraph>
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<paragraph id="FE9FBE958DA57FCBE80774AD1A1F79BA" pageId="186" pageNumber="187">
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<taxonomicName id="DFA1DCBEB6AD43ADFF39F56A514DFF57" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius kuscheli" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="kuscheli">Tomogenius kuscheli</taxonomicName>
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Dahlgren, 1976: holotype, ♀, side-mounted on a triangular card, both protarsi broken off, with the following labels: "Stephens I / Nelson 16 Feb. 76 / G.W. Ramsay" (printed); followed by:
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<normalizedToken id="5D9A42A0964FE47687D554FFAF338319" originalValue="“Litter”">"Litter"</normalizedToken>
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(printed); followed by: "
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<taxonomicName id="3DFF38A8CBC3B137F25D6C3A07358DC8" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="genus">Tomogenius</taxonomicName>
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/ sp. n. 1 / Kuschel det. 1974" (written-printed); followed by: "HOLOTYPE / TOMOGENIUS / KUSCHELI / G. DAHLGREN / 30.12.1975" (written); followed by:
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<normalizedToken id="25283D3A055A7D4F88682AE88011D332" originalValue="“HOLOTYPE”">"HOLOTYPE"</normalizedToken>
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(red label, printed); followed by: "Entomology / Division / D.S.I.R / New Zealand" (olive label, printed) (NZAC).
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</paragraph>
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</subSubSection>
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<subSubSection id="BCB884B3D6FC02494DBAA84D949A7DE0" pageId="186" pageNumber="187" type="materials_examined">
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<paragraph id="D1CEF4AF244EDF1E751710BCA99C540B" pageId="186" pageNumber="187">Additional material examined.</paragraph>
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<paragraph id="7BF783B21E058AFF61F8870293186631" pageId="186" pageNumber="187">
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NEW ZEALAND. South Island. SD: 1 ♀, Motuara Island, 23.i.1971, J. R. Jackson (ex
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<taxonomicName id="BECC2D1496BC6B03A34AA6B667CC5FE4" class="Aves" family="Procellariidae" genus="Puffinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Puffinus gavia" order="Procellariiformes" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="gavia">Puffinus gavia</taxonomicName>
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nest) (AMNZ); 3 ♂♂ & 2 ♀♀, ditto, but LUNZ (1 ♂ in coll. TLAN); 1 spec., Stephens Island, Keepers Bush, 20.ii.1994, J.W.M. Marris (ex petrel burrow) (LUNZ).
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</paragraph>
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</subSubSection>
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<subSubSection id="1D23065C08122013A87BC745031935F3" pageId="186" pageNumber="187" type="biology_ecology">
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<paragraph id="E7F8FF8F6F91840C6B9FF626DE99491F" pageId="186" pageNumber="187">Biology.</paragraph>
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<paragraph id="418A1A978CFBB663415DB0FB93F0D92E" pageId="186" pageNumber="187">
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Found in the burrow of a petrel and a shearwater (
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<taxonomicName id="18ECBFA2D0BF22B216FBCB362C42F3A5" class="Aves" family="Procellariidae" genus="Puffinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Puffinus gavia" order="Procellariiformes" pageId="186" pageNumber="187" phylum="Chordata" rank="species" species="gavia">Puffinus gavia</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="8876D4C9920A948D0BDC732A676D7A72" pageId="186" pageNumber="187" type="distribution">
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<paragraph id="49CB1BC508C58F264687E6F90C79CEF3" pageId="186" pageNumber="187">Distribution.</paragraph>
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<paragraph id="D11BDA5D5E30F299967F4ED28B74512D" pageId="186" pageNumber="187">New Zealand, South Island (SD) Stephens and Motuara Islands (Fig. 766).</paragraph>
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</subSubSection>
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<subSubSection id="35076936DFC5666A17FF75657427D63F" pageId="186" pageNumber="187" type="remarks">
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<paragraph id="3D94777A14634EB66B75138B2288F976" pageId="186" pageNumber="187">Remarks.</paragraph>
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<paragraph id="C0BE003F6F6446880626950C5501BA1C" pageId="186" pageNumber="187">
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Several specimens from the Motuara Island have a short stria next to humeral elytral stria, which may be homologous with the first dorsal elytral stria. However, in the
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<taxonomicName id="9777A8F13E5D0F6B43C6E2F12DE7730D" lsidName="" pageId="186" pageNumber="187" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
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, often the humeral elytral stria is doubled and it is therefore difficult to determine exact homology of this stria. Its basal end is not hooked as in the other dorsal elytral striae; it is less deeply impressed and is not on par with the humeral elytral stria regarding its length. This species is most similar to
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<taxonomicName id="E2D7E11EA6A3B5648E3C66A40E644D10" lsidName="T. australis" pageId="186" pageNumber="187" rank="species" species="australis">T. australis</taxonomicName>
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, especially by the structure of prosternal keel, elytral punctation, or the doubled first striae (compare Figs 622 and 665 or 619 and 660).
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</paragraph>
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</subSubSection>
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<subSubSection id="0768FD0764574FBCF77D2FA1F00CCBE3" lastPageId="190" lastPageNumber="191" pageId="186" pageNumber="187" type="description">
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<paragraph id="6A235104303BC380C643C922B582EA3C" pageId="186" pageNumber="187">Re-description.</paragraph>
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<paragraph id="55054AF5F88175B0396126BF89473F53" pageId="186" pageNumber="187">Body length: PEL: 4.00-4.40 mm; EL: 2.25-2.75 mm; APW: 1.25-1.50 mm; PPW: 2.75-3.00 mm; EW: 3.25-3.50 mm.</paragraph>
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<paragraph id="2524FDFEF22E3E9C8CD5FE38838B61B4" pageId="186" pageNumber="187">Body (Fig. 660) ovoid, moderately convex, pronotum narrower than elytra; cuticle without metallic luster, dark brown to black; legs, antenna and mouthparts dark brown.</paragraph>
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<paragraph id="26097C15AC0883458E25C47A77A9BC3D" pageId="186" pageNumber="187">Antennal scape (Fig. 661) not particularly thickened, lower margin carinate, with few short setae; club (Fig. 662) rounded, rather small, entirely covered in dense short sensilla, intermingled with sparser longer erect sensilla; ventral side of antennal club with two pairs of slit-like orifices, sensory structures situated within antennal club not examined.</paragraph>
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<caption id="11903D964DB49892ADD27E140C03C65E" pageId="186" pageNumber="187">
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<paragraph id="9332D4DD2C88AED99CB6D82FB7F9091B" pageId="186" pageNumber="187">
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Figures 661-669. 661
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<taxonomicName id="6FB3F6DE66F3DE9C3640B30E790BA9D5" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius kuscheli" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="kuscheli">Tomogenius kuscheli</taxonomicName>
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Dahlgren, 1976 head, dorsal view 662 antennal club, dorsal view 663 mentum, ventral view 664 propygidium + pygidium 665 prosternum 666 lateral disc of metaventrite + metepisternum 667 protibia, dorsal view 668 ditto, ventral view 669 metatibia, ventral view.
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</paragraph>
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</caption>
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<paragraph id="71A43AF6D2BB0A3A1814D4108564A6B1" lastPageId="187" lastPageNumber="188" pageId="186" pageNumber="187">
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Mandibles finely and sparsely punctate, rounded, acutely pointed, sub-apical tooth on inner margin of left mandible rather small, obtuse; labrum slightly convex, almost
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<pageBreakToken id="99AF078D29FBF2776EDBAA9763B25C34" pageId="187" pageNumber="188" start="start">even</pageBreakToken>
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, not depressed medially, with several punctures (especially on basal half); each labral pit with two moderately long labral setae; terminal labial palpomere elongated, its width about one-fourth its length; mentum (Fig. 663) sub-trapezoid, anterior angles slightly produced, anterior margin with a deep median emargination, surface around it with several longer setae, lateral margins with a single one row of much shorter sparse ramose setae, disc with few scattered setae, median part of mentum almost smooth; cardo of maxilla with few short setae on lateral margin; stipes triangular, with several setae; terminal maxillary palpomere elongated, its width about one-fourth its length, about three times as long as penultimate palpomere; rest of mouthparts not examined.
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</paragraph>
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<paragraph id="8DD248FFEC86AEF8C34508E75BBA6EA1" pageId="187" pageNumber="188">Clypeus (Fig. 661) large, rectangular, even, rounded laterally, with very dense shallow elongate punctures, separated by less than half their diameter; frontal and supraorbital striae absent; frontal disc (Fig. 661) with punctures similar to those of clypeus, becoming more rounded and sparser posteriorly; eyes flattened, visible from above.</paragraph>
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<paragraph id="1FA23F7E41596A5CF7556105594060A2" lastPageId="188" lastPageNumber="189" pageId="187" pageNumber="188">
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Pronotal sides (Fig. 660) feebly convergent anteriorly, apical angles prominent, marginal pronotal stria complete, thin, slightly carinate; disc entirely with punctation, becoming coarser and denser laterally where punctures very coarse and dense
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<pageBreakToken id="9EAE4A85A67EE83DBC517FEEDDF7F21C" pageId="188" pageNumber="189" start="start">separated</pageBreakToken>
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by less than their diameter, interspaces with isodiametric structures, medially punctures becoming finer and sparser separated by several times their diameter; along pronotal base an irregular row of larger punctures present; distinct pre-scutellar depression present; pronotal hypomeron glabrous; scutellum small.
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</paragraph>
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<paragraph id="C746DD95A59E201E081B6834D3C2DD11" pageId="188" pageNumber="189">Elytral epipleura with small punctures and irregular strioles; marginal epipleural stria double, both striae well impressed, complete; marginal elytral stria well impressed, shortly present also along elytral base, apically continuous along elytral apex as fine, complete apical elytral stria; humeral elytral stria deeply impressed on basal third (occasionally doubled), surface around it deeply striolate; inner subhumeral stria absent; elytral disc with three dorsal elytral striae 2-4, first dorsal elytral stria absent, second to fourth striae well impressed, impunctate, slightly surpassing elytral half; between fourth dorsal elytral and sutural striae a characteristic hooked appendix present; sutural elytral stria present on basal eighth as a short fragment. Elytral disc on basal half (roughly) with punctures separated several times their diameter (fourth elytral interval almost impunctate), on apical half (roughly) punctures becoming larger and denser, laterally extremely aciculate, confluent, interspaces imbricate.</paragraph>
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<paragraph id="3EAD5AF5082D9A89E07B4F21AF1AD94E" pageId="188" pageNumber="189">Propygidium (Fig. 664) transverse, about four times as broad as long, partially covered by elytra, with dense and coarse punctures separated by less than their diameter; pygidium (Fig. 664) with similar round punctures, becoming sparser and finer towards apex.</paragraph>
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<paragraph id="BDB25D77BC45A984BD40655932941FD8" pageId="188" pageNumber="189">Anterior margin of median portion of prosternum (Fig. 665) rounded; marginal prosternal stria present only laterally; prosternal process between carinal prosternal striae flattened, broad, anterior third elevated, slightly projecting, laterally with sparse shallow oval punctures, intermingled with alutaceous microsculpture, apically and between carinal prosternal striae punctures sparse and microscopic; carinal prosternal striae (Fig. 665) slightly bisinuate, terminating in large and deep apical foveae separated by apex of prosternal keel (Fig. 665); lateral prosternal striae carinate, bisinuate, convergent anteriorly, apically terminating near apical foveae.</paragraph>
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<paragraph id="4AA53A3ADB1DD53E024E5408648DE720" pageId="188" pageNumber="189">Discal marginal mesoventral stria well impressed, somewhat carinate, anteriorly medially projected; disc flattened, laterally with deep large punctures separated several times their diameter becoming finer and even sparser, intermingled with sparse microscopic punctures; meso-metaventral suture distinct, meso-metaventral sutural stria well impressed, undulate, slightly distanced from meso-metaventral suture.</paragraph>
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<paragraph id="23B5C37D822A1602528D978E87FD90F7" pageId="188" pageNumber="189">Intercoxal disc of metaventrite convex, finely and sparsely punctate, punctures in apical and basal corners becoming larger and coarser. Lateral metaventral stria well impressed, carinate, almost straight, not reaching metacoxa; lateral disc of metaventrite (Fig. 666) flattened, with sparse shallow punctures; metepisternum + fused metepimeron (Fig. 666) evenly with much coarser and denser punctation; lateral metepisternal stria absent.</paragraph>
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<paragraph id="3AB395686FDFB473BCBA20AE2968834D" pageId="188" pageNumber="189">Intercoxal disc of first abdominal ventrite completely striate laterally; disc laterally with dense and coarse punctation becoming sparser and finer medio-apically.</paragraph>
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<paragraph id="633475E59511E9070322E54990F6E4AA" lastPageId="189" lastPageNumber="190" pageId="188" pageNumber="189">
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Protibia (Fig. 667) almost identical to that of
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<taxonomicName id="941E90740C0893C77F3849912AB1B575" lsidName="T. australis" pageId="188" pageNumber="189" rank="species" species="australis">T. australis</taxonomicName>
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, differing from it only by areolate outer part of posterior surface of protibia (Fig. 668) (as opposed to smooth in
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<taxonomicName id="72FD60825576694EAC27E99A945E0847" lsidName="T. australis" pageId="189" pageNumber="190" rank="species" species="australis">
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<pageBreakToken id="62F6FF7276871C489240F8E191F232EE" pageId="189" pageNumber="190" start="start">T</pageBreakToken>
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. australis
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</taxonomicName>
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) and by the presence of two-three irregular rows of short setae on median part of posterior surface of protibia (as opposed to two dense rows of minuscule setae in
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<taxonomicName id="058F71A1BDC1ACE2FD37D6F4F7E2B1C5" lsidName="T. australis" pageId="189" pageNumber="190" rank="species" species="australis">T. australis</taxonomicName>
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). Meso- and metatibiae (Fig. 669) basically identical to those of
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<taxonomicName id="E051589AD22C4D2DDA556567952F81BB" lsidName="T. australis" pageId="189" pageNumber="190" rank="species" species="australis">T. australis</taxonomicName>
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.
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</paragraph>
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<paragraph id="F05720D46A3F485801F04E9712FAC18A" lastPageId="190" lastPageNumber="191" pageId="189" pageNumber="190">
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Male genitalia. Eighth sternite (Figs 670-671) widely separated medially, their apices with a tuft of long regular setae arranged in two rows; eighth tergite (Fig. 671) only slightly larger than sternite, apically faintly inwardly arcuate; basal margin strongly inwardly arcuate. Basal margin of eithth tergite medially with a curious oval structure;
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<pageBreakToken id="D3B8542C78A0F2313FB4010306C41B41" pageId="190" pageNumber="191" start="start">8</pageBreakToken>
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th sternite and tergite not fused laterally (Fig. 672). Ninth tergite (Figs 673-674) medially narrowly separated; apical margin of tenth tergite straight. Spiculum gastrale (Figs 675-676) with almost parallel body and dilated ends: apical end strongly sclerotized; basal margin cordate. Aedeagus (Figs 677-678) almost parallel-sided, apical fifth roundly dilated; parameres fused almost along their entire length. Basal piece of aedeagus rather short, its ratio to ratio of parameres approximately 1:4.
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</paragraph>
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<caption id="8B218D4A99B1703A2D3AC06084A705C7" pageId="190" pageNumber="191">
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<paragraph id="85F1AC629AB896C28BA0A57B8D663F74" pageId="190" pageNumber="191">
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Figures 670-678. 670
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<taxonomicName id="B0B37FF5921AB71A84C0CC2D6626C00E" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius kuscheli" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="kuscheli">Tomogenius kuscheli</taxonomicName>
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Dahlgren, 1976 male terminalia: 8th sternite + 8th tergite, ventral view 671 ditto, dorsal view 672 ditto, lateral view 673 male terminalia: 9th + 10th tergites, dorsal view 674 ditto, lateral view 675 male terminalia: spiculum gastrale, ventral view 676 ditto, lateral view 677 male terminalia: aedeagus, dorsal view 678 ditto, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |