treatments-xml/data/3F/3E/87/3F3E87D4FFF9661435FFA2AFA9DB7532.xml

<document id="80BD5DEE11F97AE667860E6A0004F20D" ID-CLB-Dataset="26830" ID-DOI="10.11646/zootaxa.4604.2.6" ID-GBIF-Dataset="6ffbbdb0-5acc-4333-a3f1-50129a629849" ID-ISSN="1175-5326" ID-Zenodo-Dep="2807287" ID-ZooBank="150B04BB-EE8B-4FC5-A908-462FEFCD0530" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557813017847" checkinUser="plazi" docAuthor="Oliver, Paul M., Günther, Rainer &amp; Richards, Stephen J." docDate="2019" docId="3F3E87D4FFF9661435FFA2AFA9DB7532" docLanguage="en" docName="zootaxa.4604.2.6.pdf" docOrigin="Zootaxa 4604 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Litoria pronimia Menzies 1993" docType="treatment" docVersion="9" lastPageNumber="341" masterDocId="C307FFACFFFA66123568A273AA267472" masterDocTitle="Systematics of New Guinea treefrogs (Litoria: Pelodryadidae) with erectile rostral spikes: an extended description of Litoria pronimia and a new species from the Foja Mountains" masterLastPageNumber="348" masterPageNumber="335" pageNumber="338" updateTime="1698728582715" updateUser="ExternalLinkService">
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<mods:title id="227EEA5E21A5AEA0403B0694414F88B2">Systematics of New Guinea treefrogs (Litoria: Pelodryadidae) with erectile rostral spikes: an extended description of Litoria pronimia and a new species from the Foja Mountains</mods:title>
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<taxonomicName id="70974D41FFF9661135FFA2AFAB467484" authorityName="Menzies" authorityYear="1993" box="[151,352,220,246]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="338" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFF9661135FFA2AFAB467484" bold="true" box="[151,352,220,246]" italics="true" pageId="3" pageNumber="338">Litoria pronimia</emphasis>
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<paragraph id="B72836C2FFF9661135FFA28CABF8756A" blockId="3.[151,478,220,352]" box="[151,478,255,280]" pageId="3" pageNumber="338">Peppered Pinocchio Treefrog</paragraph>
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<paragraph id="B72836C2FFF9661135FFA357AB1A754E" blockId="3.[151,478,220,352]" box="[151,316,292,317]" pageId="3" pageNumber="338">
<bibRefCitation id="D3064B33FFF9661135FFA357AB1A754E" author="Menzies, J." box="[151,316,292,317]" pageId="3" pageNumber="338" pagination="225 - 255" refId="ref7574" refString="Menzies, J. (1993) Systematics of Litoria iris (Anura, Hylidae) and its allies in New-Guinea and a note on sexual dimorphism in the group. Australian Journal of Zoology, 41, 225 - 255. https: // doi. org / 10.1071 / ZO 9930225" type="journal article" year="1993">Menzies, 1993</bibRefCitation>
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<figureCitation id="2FAC2A47FFF9661135FFA334AAF17512" box="[151,215,327,352]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1894]" captionTargetBox="[151,1435,195,1848]" captionTargetId="figure@2.[151,1435,195,1848]" captionTargetPageId="2" captionText="FIGURE 1. Papuan Litoria with a rostral spike: A) Litoria pronimia, male, Tabubil, PNG, with spike erect; B) Litoria pronimia, male, Tabubil, PNG, with spike not erect; C) Litoria pronimia, male Kikori River Basin, PNG; D) Litoria pronimia, male, Timika, Indonesia; E) Litoria havina, male, Kikori River Basin, PNG; F) Litoria mucro, male, Foja Mountains, Indonesia; G) Litoria prora, male and female, Kikori River Basin; H) Litoria chrisdahli, Wamangu, PNG. Photographs: S. Richards." httpUri="https://zenodo.org/record/2807292/files/figure.png" pageId="3" pageNumber="338">Figs 1</figureCitation>
A–D, 2
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<emphasis id="85E3EAD0FFF9661135FFA3FCA8BF75DA" bold="true" box="[151,665,399,424]" pageId="3" pageNumber="338">Material examined for extended description.</emphasis>
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amph. 13775–13380 
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<emphasis id="85E3EAD0FFF9661135AFA035AB67762D" bold="true" box="[199,321,582,607]" pageId="3" pageNumber="338">Diagnosis.</emphasis>
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<emphasis id="85E3EAD0FFF966113427A03BA834762D" box="[335,530,584,607]" italics="true" pageId="3" pageNumber="338">Litoria pronimia</emphasis>
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can be diagnosed from all other 
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<emphasis id="85E3EAD0FFF9661136DFA03BAE23762D" box="[951,1029,584,607]" italics="true" pageId="3" pageNumber="338">Litoria</emphasis>
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by the following combination of characters: a long (up to at least 
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) erectile rostral spike in males that is circular in cross section; relatively small size (SVL up to 
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); slender build (HW/SVL 0.22–0.26); tibia approximately half length of body (TL/ SVL 0.049–0.053); moderately large eyes (EYE/SVL 0.10–0.12); moderately small tympanum (TYM/SVL 0.42– 0.53); small, rounded and often green tubercles extending across the mid-dorsum in life; brown to yellowish-brown dorsal colouration with, or without, distinct green transverse bands; and orange colouration in concealed areas of thighs and axilla.
</paragraph>
</subSubSection>
<caption id="E3E8664AFFF9661135FFA115ABDE77CF" ID-Table-UUID="E3E8664AFFF9661135FFA115ABDE77CF" httpUri="http://table.plazi.org/id/E3E8664AFFF9661135FFA115ABDE77CF" pageId="3" pageNumber="338" targetBox="[159,1413,985,1998]" targetIsTable="true" targetPageId="3">
<paragraph id="B72836C2FFF9661135FFA115ABDE77CF" blockId="3.[151,1436,869,958]" pageId="3" pageNumber="338">
<emphasis id="85E3EAD0FFF9661135FFA115AB2D770F" bold="true" box="[151,267,870,893]" pageId="3" pageNumber="338">TABLE 1.</emphasis>
Summary of meristic data from the holotype of 
<taxonomicName id="70974D41FFF966113676A114A9FC770F" authority="Oliver &amp; Günther &amp; Richards, 2019" authorityName="Oliver &amp; Günther &amp; Richards" authorityYear="2019" box="[798,986,869,894]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="338" phylum="Chordata" rank="species" species="pinocchio" status="sp. nov.">
<emphasis id="85E3EAD0FFF966113676A114A9FC770F" box="[798,986,869,894]" italics="true" pageId="3" pageNumber="338">Litoria pinocchio</emphasis>
</taxonomicName>
<emphasis id="85E3EAD0FFF96611368AA115AE1F770F" bold="true" box="[994,1081,870,893]" pageId="3" pageNumber="338">
<taxonomicNameLabel id="9ED057ABFFF96611368AA115AE1F770F" box="[994,1081,870,893]" pageId="3" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and male 
<taxonomicName id="70974D41FFF9661131C4A114AF46770C" authorityName="Menzies" authorityYear="1993" box="[1196,1376,871,894]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="338" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFF9661131C4A114AF46770C" box="[1196,1376,871,894]" italics="true" pageId="3" pageNumber="338">Litoria pronimia</emphasis>
</taxonomicName>
from locations across southern New Guinea. Ratios for which 
<taxonomicName id="70974D41FFF966113677A1F4A9F877EF" authority="Oliver &amp; Günther &amp; Richards, 2019" authorityName="Oliver &amp; Günther &amp; Richards" authorityYear="2019" box="[799,990,901,926]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="338" phylum="Chordata" rank="species" species="pinocchio" status="sp. nov.">
<emphasis id="85E3EAD0FFF966113677A1F4A9F877EF" box="[799,990,901,926]" italics="true" pageId="3" pageNumber="338">Litoria pinocchio</emphasis>
</taxonomicName>
<emphasis id="85E3EAD0FFF966113682A1F5AE6377EF" bold="true" box="[1002,1093,902,925]" pageId="3" pageNumber="338">
<taxonomicNameLabel id="9ED057ABFFF966113682A1F5AE6377EF" box="[1002,1093,902,925]" pageId="3" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
does not overlap with 
<taxonomicName id="70974D41FFF96611303AA1F4AADE77CC" authorityName="Menzies" authorityYear="1993" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="338" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFF96611303AA1F4AADE77CC" italics="true" pageId="3" pageNumber="338">Litoria pronimia</emphasis>
</taxonomicName>
are highlighted in bold.
</paragraph>
</caption>
<paragraph id="B72836C2FFF966113444A1AAAFA373BC" pageId="3" pageNumber="338">
<table id="C597C462FFF999ED35F7A1AAAFA373BC" box="[159,1413,985,1998]" gridcols="6" gridrows="25" pageId="3" pageNumber="338">
<tr id="09A73480FFF999ED35F7A1AAAFA3707E" box="[159,1413,985,1036]" gridrow="0" pageId="3" pageNumber="338" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
<th id="4A765DFCFFF999ED3444A1AAABF2707E" box="[300,468,985,1036]" gridcol="1" gridrow="0" pageId="3" pageNumber="338">
<taxonomicName id="70974D41FFF966113444A1AAABF2779C" authority="Oliver &amp; Günther &amp; Richards, 2019" authorityName="Oliver &amp; Günther &amp; Richards" authorityYear="2019" box="[300,468,985,1006]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="338" phylum="Chordata" rank="species" species="pinocchio" status="sp. nov.">
<emphasis id="85E3EAD0FFF966113444A1AAABF2779C" box="[300,468,985,1006]" italics="true" pageId="3" pageNumber="338">Litoria pinocchio</emphasis>
</taxonomicName>
<emphasis id="85E3EAD0FFF966113432A185AB80707E" bold="true" box="[346,422,1014,1036]" pageId="3" pageNumber="338">
<taxonomicNameLabel id="9ED057ABFFF966113432A185AB80707E" box="[346,422,1014,1036]" pageId="3" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="4A765DFCFFF999ED36B3A1AAAEBF707E" box="[987,1177,985,1036]" gridcol="4" gridrow="0" pageId="3" pageNumber="338">
<taxonomicName id="70974D41FFF96611368EA1AAAEA3779C" authorityName="Menzies" authorityYear="1993" box="[998,1157,985,1006]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="338" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFF96611368EA1AAAEA3779C" box="[998,1157,985,1006]" italics="true" pageId="3" pageNumber="338">Litoria pronimia</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="09A73480FFF999ED35F7A66DAFA37041" box="[159,1413,1054,1075]" gridrow="1" pageId="3" pageNumber="338" rowspan-0="1">
<td id="4A765DFCFFF999ED3444A66DABF27041" box="[300,468,1054,1075]" gridcol="1" gridrow="1" pageId="3" pageNumber="338">Foja Mountains</td>
<td id="4A765DFCFFF999ED349FA66DA8907041" box="[503,694,1054,1075]" gridcol="2" gridrow="1" pageId="3" pageNumber="338">Fakfak Mountains</td>
<td id="4A765DFCFFF999ED378FA66DA98C7041" box="[743,938,1054,1075]" gridcol="3" gridrow="1" pageId="3" pageNumber="338">Timika area</td>
<td id="4A765DFCFFF999ED36B3A66DAEBF7041" box="[987,1177,1054,1075]" gridcol="4" gridrow="1" pageId="3" pageNumber="338">Tabubil</td>
<td id="4A765DFCFFF999ED31A0A66DAFA37041" box="[1224,1413,1054,1075]" gridcol="5" gridrow="1" pageId="3" pageNumber="338">Gulf Province</td>
</tr>
<tr id="09A73480FFF999ED35F7A63AAFA3702C" box="[159,1413,1097,1118]" gridrow="2" pageId="3" pageNumber="338" rowspan-0="1">
<td id="4A765DFCFFF999ED3444A63AABF2702C" box="[300,468,1097,1118]" gridcol="1" gridrow="2" pageId="3" pageNumber="338">n=1</td>
<td id="4A765DFCFFF999ED349FA63AA890702C" box="[503,694,1097,1118]" gridcol="2" gridrow="2" pageId="3" pageNumber="338">n=2</td>
<td id="4A765DFCFFF999ED378FA63AA98C702C" box="[743,938,1097,1118]" gridcol="3" gridrow="2" pageId="3" pageNumber="338">n=6</td>
<td id="4A765DFCFFF999ED36B3A63AAEBF702C" box="[987,1177,1097,1118]" gridcol="4" gridrow="2" pageId="3" pageNumber="338">n=8</td>
<td id="4A765DFCFFF999ED31A0A63AAFA3702C" box="[1224,1413,1097,1118]" gridcol="5" gridrow="2" pageId="3" pageNumber="338">n=7</td>
</tr>
<tr id="09A73480FFF999ED35F7A602AFA370F4" box="[159,1413,1137,1158]" gridrow="3" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A602AB2170F4" box="[159,263,1137,1158]" gridcol="0" gridrow="3" pageId="3" pageNumber="338">SVL</th>
<td id="4A765DFCFFF999ED3444A602ABF270F4" box="[300,468,1137,1158]" gridcol="1" gridrow="3" pageId="3" pageNumber="338">29.2</td>
<td id="4A765DFCFFF999ED349FA602A89070F4" box="[503,694,1137,1158]" gridcol="2" gridrow="3" pageId="3" pageNumber="338">28.9(28.6–29.1)</td>
<td id="4A765DFCFFF999ED378FA602A98C70F4" box="[743,938,1137,1158]" gridcol="3" gridrow="3" pageId="3" pageNumber="338">28.1(26.9–29.2)</td>
<td id="4A765DFCFFF999ED36B3A602AEBF70F4" box="[987,1177,1137,1158]" gridcol="4" gridrow="3" pageId="3" pageNumber="338">28.9(28.1–30.4)</td>
<td id="4A765DFCFFF999ED31A0A602AFA370F4" box="[1224,1413,1137,1158]" gridcol="5" gridrow="3" pageId="3" pageNumber="338">28.7(26.5–30.1)</td>
</tr>
<tr id="09A73480FFF999ED35F7A6EAAFA370DC" box="[159,1413,1177,1198]" gridrow="4" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A6EAAB2170DC" box="[159,263,1177,1198]" gridcol="0" gridrow="4" pageId="3" pageNumber="338">TL</th>
<td id="4A765DFCFFF999ED3444A6EAABF270DC" box="[300,468,1177,1198]" gridcol="1" gridrow="4" pageId="3" pageNumber="338">14.1</td>
<td id="4A765DFCFFF999ED349FA6EAA89070DC" box="[503,694,1177,1198]" gridcol="2" gridrow="4" pageId="3" pageNumber="338">15.0(14.9–15.0)</td>
<td id="4A765DFCFFF999ED378FA6EAA98C70DC" box="[743,938,1177,1198]" gridcol="3" gridrow="4" pageId="3" pageNumber="338">14.2(13.4–14.8)</td>
<td id="4A765DFCFFF999ED36B3A6EAAEBF70DC" box="[987,1177,1177,1198]" gridcol="4" gridrow="4" pageId="3" pageNumber="338">14.7(14.2–15.9)</td>
<td id="4A765DFCFFF999ED31A0A6EAAFA370DC" box="[1224,1413,1177,1198]" gridcol="5" gridrow="4" pageId="3" pageNumber="338">14.4(13.2–15.2)</td>
</tr>
<tr id="09A73480FFF999ED35F7A6B2AFA370A4" box="[159,1413,1217,1238]" gridrow="5" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A6B2AB2170A4" box="[159,263,1217,1238]" gridcol="0" gridrow="5" pageId="3" pageNumber="338">HW</th>
<td id="4A765DFCFFF999ED3444A6B2ABF270A4" box="[300,468,1217,1238]" gridcol="1" gridrow="5" pageId="3" pageNumber="338">7.6</td>
<td id="4A765DFCFFF999ED349FA6B2A89070A4" box="[503,694,1217,1238]" gridcol="2" gridrow="5" pageId="3" pageNumber="338">7.2(7.2–7.2)</td>
<td id="4A765DFCFFF999ED378FA6B2A98C70A4" box="[743,938,1217,1238]" gridcol="3" gridrow="5" pageId="3" pageNumber="338">7.0(6.7–7.3)</td>
<td id="4A765DFCFFF999ED36B3A6B2AEBF70A4" box="[987,1177,1217,1238]" gridcol="4" gridrow="5" pageId="3" pageNumber="338">7.0(6.7–7.3)</td>
<td id="4A765DFCFFF999ED31A0A6B2AFA370A4" box="[1224,1413,1217,1238]" gridcol="5" gridrow="5" pageId="3" pageNumber="338">7.0(6.6–7.3)</td>
</tr>
<tr id="09A73480FFF999ED35F7A69AAFA3708C" box="[159,1413,1257,1278]" gridrow="6" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A69AAB21708C" box="[159,263,1257,1278]" gridcol="0" gridrow="6" pageId="3" pageNumber="338">HL</th>
<td id="4A765DFCFFF999ED3444A69AABF2708C" box="[300,468,1257,1278]" gridcol="1" gridrow="6" pageId="3" pageNumber="338">10.1</td>
<td id="4A765DFCFFF999ED349FA69AA890708C" box="[503,694,1257,1278]" gridcol="2" gridrow="6" pageId="3" pageNumber="338">10.2(9.8–10.6)</td>
<td id="4A765DFCFFF999ED378FA69AA98C708C" box="[743,938,1257,1278]" gridcol="3" gridrow="6" pageId="3" pageNumber="338">8.8(8.3–9.1)</td>
<td id="4A765DFCFFF999ED36B3A69AAEBF708C" box="[987,1177,1257,1278]" gridcol="4" gridrow="6" pageId="3" pageNumber="338">10.0(9.4–10.7)</td>
<td id="4A765DFCFFF999ED31A0A69AAFA3708C" box="[1224,1413,1257,1278]" gridcol="5" gridrow="6" pageId="3" pageNumber="338">9.5(8.7–10.3)</td>
</tr>
<tr id="09A73480FFF999ED35F7A762AFA37154" box="[159,1413,1297,1318]" gridrow="7" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A762AB217154" box="[159,263,1297,1318]" gridcol="0" gridrow="7" pageId="3" pageNumber="338">EYE</th>
<td id="4A765DFCFFF999ED3444A762ABF27154" box="[300,468,1297,1318]" gridcol="1" gridrow="7" pageId="3" pageNumber="338">3.6</td>
<td id="4A765DFCFFF999ED349FA762A8907154" box="[503,694,1297,1318]" gridcol="2" gridrow="7" pageId="3" pageNumber="338">3.2(2.9–3.5)</td>
<td id="4A765DFCFFF999ED378FA762A98C7154" box="[743,938,1297,1318]" gridcol="3" gridrow="7" pageId="3" pageNumber="338">3.3(3.2–3.3)</td>
<td id="4A765DFCFFF999ED36B3A762AEBF7154" box="[987,1177,1297,1318]" gridcol="4" gridrow="7" pageId="3" pageNumber="338">3.4(3.4–3.6)</td>
<td id="4A765DFCFFF999ED31A0A762AFA37154" box="[1224,1413,1297,1318]" gridcol="5" gridrow="7" pageId="3" pageNumber="338">3.1(3.0–3.2)</td>
</tr>
<tr id="09A73480FFF999ED35F7A74AAFA3713C" box="[159,1413,1337,1358]" gridrow="8" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A74AAB21713C" box="[159,263,1337,1358]" gridcol="0" gridrow="8" pageId="3" pageNumber="338">TYM</th>
<td id="4A765DFCFFF999ED3444A74AABF2713C" box="[300,468,1337,1358]" gridcol="1" gridrow="8" pageId="3" pageNumber="338">1.6</td>
<td id="4A765DFCFFF999ED349FA74AA890713C" box="[503,694,1337,1358]" gridcol="2" gridrow="8" pageId="3" pageNumber="338">1.3(1.2–1.3)</td>
<td id="4A765DFCFFF999ED378FA74AA98C713C" box="[743,938,1337,1358]" gridcol="3" gridrow="8" pageId="3" pageNumber="338">1.3(1.2–1.4)</td>
<td id="4A765DFCFFF999ED36B3A74AAEBF713C" box="[987,1177,1337,1358]" gridcol="4" gridrow="8" pageId="3" pageNumber="338">1.4(1.2–1.5)</td>
<td id="4A765DFCFFF999ED31A0A74AAFA3713C" box="[1224,1413,1337,1358]" gridcol="5" gridrow="8" pageId="3" pageNumber="338">1.3(1.2–1.4)</td>
</tr>
<tr id="09A73480FFF999ED35F7A712AFA37104" box="[159,1413,1377,1398]" gridrow="9" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A712AB217104" box="[159,263,1377,1398]" gridcol="0" gridrow="9" pageId="3" pageNumber="338">IN</th>
<td id="4A765DFCFFF999ED3444A712ABF27104" box="[300,468,1377,1398]" gridcol="1" gridrow="9" pageId="3" pageNumber="338">2.6</td>
<td id="4A765DFCFFF999ED349FA712A8907104" box="[503,694,1377,1398]" gridcol="2" gridrow="9" pageId="3" pageNumber="338">2.3(2.2–2.4)</td>
<td id="4A765DFCFFF999ED378FA712A98C7104" box="[743,938,1377,1398]" gridcol="3" gridrow="9" pageId="3" pageNumber="338">2.2(2.1–2.3)</td>
<td id="4A765DFCFFF999ED36B3A712AEBF7104" box="[987,1177,1377,1398]" gridcol="4" gridrow="9" pageId="3" pageNumber="338">2.4(2.2–2.6)</td>
<td id="4A765DFCFFF999ED31A0A712AFA37104" box="[1224,1413,1377,1398]" gridcol="5" gridrow="9" pageId="3" pageNumber="338">2.4(2.1–2.6)</td>
</tr>
<tr id="09A73480FFF999ED35F7A7FAAFA371EC" box="[159,1413,1417,1438]" gridrow="10" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A7FAAB2171EC" box="[159,263,1417,1438]" gridcol="0" gridrow="10" pageId="3" pageNumber="338">EN</th>
<td id="4A765DFCFFF999ED3444A7FAABF271EC" box="[300,468,1417,1438]" gridcol="1" gridrow="10" pageId="3" pageNumber="338">2.6</td>
<td id="4A765DFCFFF999ED349FA7FAA89071EC" box="[503,694,1417,1438]" gridcol="2" gridrow="10" pageId="3" pageNumber="338">2.5(2.3–2.7)</td>
<td id="4A765DFCFFF999ED378FA7FAA98C71EC" box="[743,938,1417,1438]" gridcol="3" gridrow="10" pageId="3" pageNumber="338">2.4(2.3–2.5)</td>
<td id="4A765DFCFFF999ED36B3A7FAAEBF71EC" box="[987,1177,1417,1438]" gridcol="4" gridrow="10" pageId="3" pageNumber="338">2.5(2.3–2.6)</td>
<td id="4A765DFCFFF999ED31A0A7FAAFA371EC" box="[1224,1413,1417,1438]" gridcol="5" gridrow="10" pageId="3" pageNumber="338">2.3(2.0–2.5)</td>
</tr>
<tr id="09A73480FFF999ED35F7A7C2AFA371B4" box="[159,1413,1457,1478]" gridrow="11" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A7C2AB2171B4" box="[159,263,1457,1478]" gridcol="0" gridrow="11" pageId="3" pageNumber="338">3FD</th>
<td id="4A765DFCFFF999ED3444A7C2ABF271B4" box="[300,468,1457,1478]" gridcol="1" gridrow="11" pageId="3" pageNumber="338">1.5</td>
<td id="4A765DFCFFF999ED349FA7C2A89071B4" box="[503,694,1457,1478]" gridcol="2" gridrow="11" pageId="3" pageNumber="338">1.4(1.4–1.4)</td>
<td id="4A765DFCFFF999ED378FA7C2A98C71B4" box="[743,938,1457,1478]" gridcol="3" gridrow="11" pageId="3" pageNumber="338">1.2(1.0–1.3)</td>
<td id="4A765DFCFFF999ED36B3A7C2AEBF71B4" box="[987,1177,1457,1478]" gridcol="4" gridrow="11" pageId="3" pageNumber="338">1.4(1.1–1.5)</td>
<td id="4A765DFCFFF999ED31A0A7C2AFA371B4" box="[1224,1413,1457,1478]" gridcol="5" gridrow="11" pageId="3" pageNumber="338">1.4(1.2–1.5)</td>
</tr>
<tr id="09A73480FFF999ED35F7A7AAAFA3719C" box="[159,1413,1497,1518]" gridrow="12" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A7AAAB21719C" box="[159,263,1497,1518]" gridcol="0" gridrow="12" pageId="3" pageNumber="338">3FP</th>
<td id="4A765DFCFFF999ED3444A7AAABF2719C" box="[300,468,1497,1518]" gridcol="1" gridrow="12" pageId="3" pageNumber="338">1.2</td>
<td id="4A765DFCFFF999ED349FA7AAA890719C" box="[503,694,1497,1518]" gridcol="2" gridrow="12" pageId="3" pageNumber="338">1.2(1.2–1.2)</td>
<td id="4A765DFCFFF999ED378FA7AAA98C719C" box="[743,938,1497,1518]" gridcol="3" gridrow="12" pageId="3" pageNumber="338">0.9(0.8–1.0)</td>
<td id="4A765DFCFFF999ED36B3A7AAAEBF719C" box="[987,1177,1497,1518]" gridcol="4" gridrow="12" pageId="3" pageNumber="338">1.1(0.9–1.6)</td>
<td id="4A765DFCFFF999ED31A0A7AAAFA3719C" box="[1224,1413,1497,1518]" gridcol="5" gridrow="12" pageId="3" pageNumber="338">1.0(0.8–1.1)</td>
</tr>
<tr id="09A73480FFF999ED35F7A472AFA37264" box="[159,1413,1537,1558]" gridrow="13" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A472AB217264" box="[159,263,1537,1558]" gridcol="0" gridrow="13" pageId="3" pageNumber="338">4TD</th>
<td id="4A765DFCFFF999ED3444A472ABF27264" box="[300,468,1537,1558]" gridcol="1" gridrow="13" pageId="3" pageNumber="338">1.3</td>
<td id="4A765DFCFFF999ED349FA472A8907264" box="[503,694,1537,1558]" gridcol="2" gridrow="13" pageId="3" pageNumber="338">1.3(1.3–1.3)</td>
<td id="4A765DFCFFF999ED378FA472A98C7264" box="[743,938,1537,1558]" gridcol="3" gridrow="13" pageId="3" pageNumber="338">1.0(0.8–1.1)</td>
<td id="4A765DFCFFF999ED36B3A472AEBF7264" box="[987,1177,1537,1558]" gridcol="4" gridrow="13" pageId="3" pageNumber="338">1.3(1.2–1.4)</td>
<td id="4A765DFCFFF999ED31A0A472AFA37264" box="[1224,1413,1537,1558]" gridcol="5" gridrow="13" pageId="3" pageNumber="338">1.2(1.0–1.3)</td>
</tr>
<tr id="09A73480FFF999ED35F7A45AAFA3724C" box="[159,1413,1577,1598]" gridrow="14" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A45AAB21724C" box="[159,263,1577,1598]" gridcol="0" gridrow="14" pageId="3" pageNumber="338">4TP</th>
<td id="4A765DFCFFF999ED3444A45AABF2724C" box="[300,468,1577,1598]" gridcol="1" gridrow="14" pageId="3" pageNumber="338">1.2</td>
<td id="4A765DFCFFF999ED349FA45AA890724C" box="[503,694,1577,1598]" gridcol="2" gridrow="14" pageId="3" pageNumber="338">1.1(1.0–1.1)</td>
<td id="4A765DFCFFF999ED378FA45AA98C724C" box="[743,938,1577,1598]" gridcol="3" gridrow="14" pageId="3" pageNumber="338">0.8(0.7–0.8)</td>
<td id="4A765DFCFFF999ED36B3A45AAEBF724C" box="[987,1177,1577,1598]" gridcol="4" gridrow="14" pageId="3" pageNumber="338">0.9(0.8–1.0)</td>
<td id="4A765DFCFFF999ED31A0A45AAFA3724C" box="[1224,1413,1577,1598]" gridcol="5" gridrow="14" pageId="3" pageNumber="338">0.9(0.8–1.0)</td>
</tr>
<tr id="09A73480FFF999ED35F7A422AFA37214" box="[159,1413,1617,1638]" gridrow="15" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A422AB217214" box="[159,263,1617,1638]" gridcol="0" gridrow="15" pageId="3" pageNumber="338">SVL1</th>
<td id="4A765DFCFFF999ED3444A422ABF27214" box="[300,468,1617,1638]" gridcol="1" gridrow="15" pageId="3" pageNumber="338">31.7</td>
<td id="4A765DFCFFF999ED349FA422A8907214" box="[503,694,1617,1638]" gridcol="2" gridrow="15" pageId="3" pageNumber="338">30.4(30.1–30.6)</td>
<td id="4A765DFCFFF999ED378FA422A98C7214" box="[743,938,1617,1638]" gridcol="3" gridrow="15" pageId="3" pageNumber="338">29.1(27.4–30.5)</td>
<td id="4A765DFCFFF999ED36B3A422AEBF7214" box="[987,1177,1617,1638]" gridcol="4" gridrow="15" pageId="3" pageNumber="338">30.1(29.3–32.6)</td>
<td id="4A765DFCFFF999ED31A0A422AFA37214" box="[1224,1413,1617,1638]" gridcol="5" gridrow="15" pageId="3" pageNumber="338">30.4(27.7–32.4)</td>
</tr>
<tr id="09A73480FFF999ED35F7A40AAFA372FC" box="[159,1413,1657,1678]" gridrow="16" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A40AAB2172FC" box="[159,263,1657,1678]" gridcol="0" gridrow="16" pageId="3" pageNumber="338">RL</th>
<td id="4A765DFCFFF999ED3444A40AABF272FC" box="[300,468,1657,1678]" gridcol="1" gridrow="16" pageId="3" pageNumber="338">2.5</td>
<td id="4A765DFCFFF999ED349FA40AA89072FC" box="[503,694,1657,1678]" gridcol="2" gridrow="16" pageId="3" pageNumber="338">1.5(1.5–1.5)</td>
<td id="4A765DFCFFF999ED378FA40AA98C72FC" box="[743,938,1657,1678]" gridcol="3" gridrow="16" pageId="3" pageNumber="338">1.0(0.5–1.3)</td>
<td id="4A765DFCFFF999ED36B3A40AAEBF72FC" box="[987,1177,1657,1678]" gridcol="4" gridrow="16" pageId="3" pageNumber="338">1.9(0.7–2.8)</td>
<td id="4A765DFCFFF999ED31A0A40AAFA372FC" box="[1224,1413,1657,1678]" gridcol="5" gridrow="16" pageId="3" pageNumber="338">1.6(0.5–2.4)</td>
</tr>
<tr id="09A73480FFF999ED35F7A4D2AFA372C4" box="[159,1413,1697,1718]" gridrow="17" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A4D2AFA372C4" box="[159,1413,1697,1718]" colspan="6" colspanRight="5" gridcol="0" gridrow="17" pageId="3" pageNumber="338">Ratios</th>
</tr>
<tr id="09A73480FFF999ED35F7A4BAAFA372AC" box="[159,1413,1737,1758]" gridrow="18" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A4BAAB2172AC" box="[159,263,1737,1758]" gridcol="0" gridrow="18" pageId="3" pageNumber="338">RL/SVL</th>
<td id="4A765DFCFFF999ED3444A4BAABF272AC" box="[300,468,1737,1758]" gridcol="1" gridrow="18" pageId="3" pageNumber="338">0.086</td>
<td id="4A765DFCFFF999ED349FA4BAA89072AC" box="[503,694,1737,1758]" gridcol="2" gridrow="18" pageId="3" pageNumber="338">0.052(0.052–0.052)</td>
<td id="4A765DFCFFF999ED378FA4BAA98C72AC" box="[743,938,1737,1758]" gridcol="3" gridrow="18" pageId="3" pageNumber="338">0.034 (0.019–0.045)</td>
<td id="4A765DFCFFF999ED36B3A4BAAEBF72AC" box="[987,1177,1737,1758]" gridcol="4" gridrow="18" pageId="3" pageNumber="338">0.067(0.024–0.097)</td>
<td id="4A765DFCFFF999ED31A0A4BAAFA372AC" box="[1224,1413,1737,1758]" gridcol="5" gridrow="18" pageId="3" pageNumber="338">0.057(0.017–0.080)</td>
</tr>
<tr id="09A73480FFF999ED35F7A482AFA37374" box="[159,1413,1777,1798]" gridrow="19" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A482AB217374" box="[159,263,1777,1798]" gridcol="0" gridrow="19" pageId="3" pageNumber="338">HW/SVL</th>
<td id="4A765DFCFFF999ED3444A482ABF27374" box="[300,468,1777,1798]" gridcol="1" gridrow="19" pageId="3" pageNumber="338">0.26</td>
<td id="4A765DFCFFF999ED349FA482A8907374" box="[503,694,1777,1798]" gridcol="2" gridrow="19" pageId="3" pageNumber="338">0.25(0.25–0.25)</td>
<td id="4A765DFCFFF999ED378FA482A98C7374" box="[743,938,1777,1798]" gridcol="3" gridrow="19" pageId="3" pageNumber="338">0.25 (0.24–0.26)</td>
<td id="4A765DFCFFF999ED36B3A482AEBF7374" box="[987,1177,1777,1798]" gridcol="4" gridrow="19" pageId="3" pageNumber="338">0.24(0.22–0.25)</td>
<td id="4A765DFCFFF999ED31A0A482AFA37374" box="[1224,1413,1777,1798]" gridcol="5" gridrow="19" pageId="3" pageNumber="338">0.24(0.23–0.25)</td>
</tr>
<tr id="09A73480FFF999ED35F7A56AAFA3735C" box="[159,1413,1817,1838]" gridrow="20" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A56AAB21735C" box="[159,263,1817,1838]" gridcol="0" gridrow="20" pageId="3" pageNumber="338">EYE/SVL</th>
<td id="4A765DFCFFF999ED3444A56AABF2735C" box="[300,468,1817,1838]" gridcol="1" gridrow="20" pageId="3" pageNumber="338">0.12</td>
<td id="4A765DFCFFF999ED349FA56AA890735C" box="[503,694,1817,1838]" gridcol="2" gridrow="20" pageId="3" pageNumber="338">0.11(0.10–0.12)</td>
<td id="4A765DFCFFF999ED378FA56AA98C735C" box="[743,938,1817,1838]" gridcol="3" gridrow="20" pageId="3" pageNumber="338">0.12(0.11–0.12)</td>
<td id="4A765DFCFFF999ED36B3A56AAEBF735C" box="[987,1177,1817,1838]" gridcol="4" gridrow="20" pageId="3" pageNumber="338">0.12(0.11–0.13)</td>
<td id="4A765DFCFFF999ED31A0A56AAFA3735C" box="[1224,1413,1817,1838]" gridcol="5" gridrow="20" pageId="3" pageNumber="338">0.11(0.10–0.12)</td>
</tr>
<tr id="09A73480FFF999ED35F7A532AFA37325" box="[159,1413,1857,1879]" gridrow="21" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A532AB217325" box="[159,263,1857,1879]" gridcol="0" gridrow="21" pageId="3" pageNumber="338">TL/SVL</th>
<td id="4A765DFCFFF999ED3444A532ABF27325" box="[300,468,1857,1879]" gridcol="1" gridrow="21" pageId="3" pageNumber="338">
<emphasis id="85E3EAD0FFF966113403A532ABB37325" bold="true" box="[363,405,1857,1879]" pageId="3" pageNumber="338">0.48</emphasis>
</td>
<td id="4A765DFCFFF999ED349FA532A8907325" box="[503,694,1857,1879]" gridcol="2" gridrow="21" pageId="3" pageNumber="338">0.52(0.51–0.52)</td>
<td id="4A765DFCFFF999ED378FA532A98C7325" box="[743,938,1857,1879]" gridcol="3" gridrow="21" pageId="3" pageNumber="338">0.51(0.50–0.52)</td>
<td id="4A765DFCFFF999ED36B3A532AEBF7325" box="[987,1177,1857,1879]" gridcol="4" gridrow="21" pageId="3" pageNumber="338">0.51(0.49–0.52)</td>
<td id="4A765DFCFFF999ED31A0A532AFA37325" box="[1224,1413,1857,1879]" gridcol="5" gridrow="21" pageId="3" pageNumber="338">0.50(0.49–0.53)</td>
</tr>
<tr id="09A73480FFF999ED35F7A51AAFA3730D" box="[159,1413,1897,1919]" gridrow="22" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A51AAB21730D" box="[159,263,1897,1919]" gridcol="0" gridrow="22" pageId="3" pageNumber="338">TYM/SVL</th>
<td id="4A765DFCFFF999ED3444A51AABF2730D" box="[300,468,1897,1919]" gridcol="1" gridrow="22" pageId="3" pageNumber="338">
<emphasis id="85E3EAD0FFF96611340DA51AABBD730D" bold="true" box="[357,411,1897,1919]" pageId="3" pageNumber="338">0.055</emphasis>
</td>
<td id="4A765DFCFFF999ED349FA51AA890730D" box="[503,694,1897,1919]" gridcol="2" gridrow="22" pageId="3" pageNumber="338">0.043(0.041–0.045)</td>
<td id="4A765DFCFFF999ED378FA51AA98C730D" box="[743,938,1897,1919]" gridcol="3" gridrow="22" pageId="3" pageNumber="338">0.047(0.042–0.052)</td>
<td id="4A765DFCFFF999ED36B3A51AAEBF730D" box="[987,1177,1897,1919]" gridcol="4" gridrow="22" pageId="3" pageNumber="338">0.047(0.042–0.053)</td>
<td id="4A765DFCFFF999ED31A0A51AAFA3730D" box="[1224,1413,1897,1919]" gridcol="5" gridrow="22" pageId="3" pageNumber="338">0.045(0.042–0.049)</td>
</tr>
<tr id="09A73480FFF999ED35F7A5E2AFA373D4" box="[159,1413,1937,1958]" gridrow="23" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A5E2AB2173D4" box="[159,263,1937,1958]" gridcol="0" gridrow="23" pageId="3" pageNumber="338">4TD/SVL</th>
<td id="4A765DFCFFF999ED3444A5E2ABF273D4" box="[300,468,1937,1958]" gridcol="1" gridrow="23" pageId="3" pageNumber="338">0.045</td>
<td id="4A765DFCFFF999ED349FA5E2A89073D4" box="[503,694,1937,1958]" gridcol="2" gridrow="23" pageId="3" pageNumber="338">0.045(0.045–0.045)</td>
<td id="4A765DFCFFF999ED378FA5E2A98C73D4" box="[743,938,1937,1958]" gridcol="3" gridrow="23" pageId="3" pageNumber="338">0.034(0.029–0.038)</td>
<td id="4A765DFCFFF999ED36B3A5E2AEBF73D4" box="[987,1177,1937,1958]" gridcol="4" gridrow="23" pageId="3" pageNumber="338">0.045(0.041–0.049)</td>
<td id="4A765DFCFFF999ED31A0A5E2AFA373D4" box="[1224,1413,1937,1958]" gridcol="5" gridrow="23" pageId="3" pageNumber="338">0.042(0.038–0.045)</td>
</tr>
<tr id="09A73480FFF999ED35F7A5CAAFA373BC" box="[159,1413,1977,1998]" gridrow="24" pageId="3" pageNumber="338">
<th id="4A765DFCFFF999ED35F7A5CAAB2173BC" box="[159,263,1977,1998]" gridcol="0" gridrow="24" pageId="3" pageNumber="338">3FD/SVL</th>
<td id="4A765DFCFFF999ED3444A5CAABF273BC" box="[300,468,1977,1998]" gridcol="1" gridrow="24" pageId="3" pageNumber="338">0.051</td>
<td id="4A765DFCFFF999ED349FA5CAA89073BC" box="[503,694,1977,1998]" gridcol="2" gridrow="24" pageId="3" pageNumber="338">0.049(0.048–0.049)</td>
<td id="4A765DFCFFF999ED378FA5CAA98C73BC" box="[743,938,1977,1998]" gridcol="3" gridrow="24" pageId="3" pageNumber="338">0.042(0.037–0.045)</td>
<td id="4A765DFCFFF999ED36B3A5CAAEBF73BC" box="[987,1177,1977,1998]" gridcol="4" gridrow="24" pageId="3" pageNumber="338">0.048(0.039–0.051)</td>
<td id="4A765DFCFFF999ED31A0A5CAAFA373BC" box="[1224,1413,1977,1998]" gridcol="5" gridrow="24" pageId="3" pageNumber="338">0.048(0.045–0.052)</td>
</tr>
</table>
</paragraph>
<subSubSection id="FF8D6549FFFE661435AFA2E4A9DB7532" lastPageId="6" lastPageNumber="341" pageId="4" pageNumber="339" type="description">
<paragraph id="B72836C2FFFE661635AFA2E4AF77764F" blockId="4.[151,1437,151,1833]" pageId="4" pageNumber="339">
<emphasis id="85E3EAD0FFFE661635AFA2E4AB7274C2" bold="true" box="[199,340,151,176]" pageId="4" pageNumber="339">Description.</emphasis>
Summary measurements and ratios for samples from four disjunct groups of sites are given in 
<tableCitation id="FA150379FFFE66163008A2E4AA8374A6" captionStart="TABLE 1" captionStartId="3.[151,239,870,893]" captionTargetPageId="3" captionText="TABLE 1. Summary of meristic data from the holotype of Litoria pinocchio sp. nov. and male Litoria pronimia from locations across southern New Guinea. Ratios for which Litoria pinocchio sp. nov. does not overlap with Litoria pronimia are highlighted in bold." httpUri="http://table.plazi.org/id/E3E8664AFFF9661135FFA115ABDE77CF" pageId="4" pageNumber="339" tableUuid="E3E8664AFFF9661135FFA115ABDE77CF">Table 1</tableCitation>
. Body very slender and elongate, tibia approximately half length of body; head wider than body in dorsal profile, clearly distinct from neck. Snout rounded in dorsal view and truncate in lateral view. Rostral spike in males extending from tip of upper jaw; mean spikelength varies among populations sampled from different localities (see 
<tableCitation id="FA150379FFFE66163051A377AFAA756E" box="[1337,1420,260,285]" captionStart="TABLE 1" captionStartId="3.[151,239,870,893]" captionTargetPageId="3" captionText="TABLE 1. Summary of meristic data from the holotype of Litoria pinocchio sp. nov. and male Litoria pronimia from locations across southern New Guinea. Ratios for which Litoria pinocchio sp. nov. does not overlap with Litoria pronimia are highlighted in bold." httpUri="http://table.plazi.org/id/E3E8664AFFF9661135FFA115ABDE77CF" pageId="4" pageNumber="339" tableUuid="E3E8664AFFF9661135FFA115ABDE77CF">Table 1</tableCitation>
). Canthus rostralis moderately well defined, slightly curved; loreal region concave. Nares closer to tip of snout (excluding spike) than to eye, oriented laterally. Eyes moderately large, protruding in both dorsal and lateral views; pupil horizontal. Upper jaw protruding marginally beyond lower jaw. Tympanum small with distinct to indistinct annulus, bordered dorsally by a fleshy supratympanic fold extending to the superior edge of the insertion of the upper arm. Choanae small and circular, situated close to anterior and lateral edge of palate; no vomerine teeth visible; tongue fleshy and ovoid and usually with a slightly indented posterior edge. Vocal slits present. Dorsal skin tuberculate; ventral skin finely to coarsely granular on throat, abdomen and tibia; remaining ventral surfaces of limbs smooth; additional tubercles also present in clusters around the vent, and to a variable extent on sides of ankles and upper forearms.
</paragraph>
<paragraph id="B72836C2FFFE661635AFA034A84D772E" blockId="4.[151,1437,151,1833]" pageId="4" pageNumber="339">
Fingers with relative lengths III&gt;IV&gt;II&gt;I; fleshy, whitish to translucent webbing between all digits, forming a narrow basal strip between I and II, extending to disc on distal edge of II and proximal edge of IV, and to penultimate phalanx on both sides on III. Terminal finger discs expanded (narrower in the six samples from Timika: see 
<tableCitation id="FA150379FFFE6616344DA0C7AB5F76BE" box="[293,377,692,717]" captionStart="TABLE 1" captionStartId="3.[151,239,870,893]" captionTargetPageId="3" captionText="TABLE 1. Summary of meristic data from the holotype of Litoria pinocchio sp. nov. and male Litoria pronimia from locations across southern New Guinea. Ratios for which Litoria pinocchio sp. nov. does not overlap with Litoria pronimia are highlighted in bold." httpUri="http://table.plazi.org/id/E3E8664AFFF9661135FFA115ABDE77CF" pageId="4" pageNumber="339" tableUuid="E3E8664AFFF9661135FFA115ABDE77CF">Table 1</tableCitation>
), wider than toe discs and with distinct circum-marginal grooves. Nuptial pads dark brown, roughly tear-shaped with point of tear oriented in posteroventral direction. Indistinct unpigmented bifid subarticular tubercles usually visible at base of penultimate phalanx on all fingers, further indistinct unpigmented subarticular tubercles in series on finger IV, indistinct proximal metacarpal tubercles at base of I, and small distal metacarpal tubercles at base of III and IV.
</paragraph>
<paragraph id="B72836C2FFFE661635AFA11BA8DB7025" blockId="4.[151,1437,151,1833]" pageId="4" pageNumber="339">
Toes moderately long, relative lengths IV&gt;III&gt;V&gt;II&gt;I. All digits with extensive fleshy, opaque webbing, basal between I and II, extending to anterior end of penultimate phalanx on distal edge of II and III and proximal edge of V, to halfway along penultimate phalanx on proximal edge of III and IV, and base of penultimate tubercle on distal edge of IV. Terminal discs slightly expanded (narrower in the six samples from Timika: see 
<tableCitation id="FA150379FFFE661631FFA1A7AECE779E" box="[1175,1256,980,1005]" captionStart="TABLE 1" captionStartId="3.[151,239,870,893]" captionTargetPageId="3" captionText="TABLE 1. Summary of meristic data from the holotype of Litoria pinocchio sp. nov. and male Litoria pronimia from locations across southern New Guinea. Ratios for which Litoria pinocchio sp. nov. does not overlap with Litoria pronimia are highlighted in bold." httpUri="http://table.plazi.org/id/E3E8664AFFF9661135FFA115ABDE77CF" pageId="4" pageNumber="339" tableUuid="E3E8664AFFF9661135FFA115ABDE77CF">Table 1</tableCitation>
), narrower than finger discs and with distinct circum-marginal grooves. Indistinct unpigmented subarticular tubercles on penultimate phalanx of all toes, single on I-III, bifid on IV and V. No other tubercles clearly apparent. Small, indistinct unpigmented metatarsal tubercle at base of I.
</paragraph>
<paragraph id="B72836C2FFFE661635AFA616A81471EF" blockId="4.[151,1437,151,1833]" pageId="4" pageNumber="339">
In preservative, ground colour of all dorsal surfaces medium brown, with extensive blueish green and darker brown flecking, maculations, vermiculations and/or blotches across all dorsal surfaces, blueish pattern elements sometimes coalesce into three indistinct transverse blotches, overall darker pigmentation elements tend to be densest on body, and less concentrated and more finely reticulated towards distal extremities of limbs (
<figureCitation id="2FAC2A47FFFE6616302EA6BCAFA9709A" box="[1350,1423,1231,1256]" captionStart="FIGURE 2" captionStartId="5.[151,250,912,934]" captionTargetBox="[151,1416,239,898]" captionTargetId="figure@5.[151,1436,188,898]" captionTargetPageId="5" captionText="FIGURE 2. Litoria pronimia in preservative from near the type locality at Tabubil (SAMA R71135–6, 71140) and the Kikori River Basin (SAMA R77131). Photograph by Peter Waddington, Queensland Museum." httpUri="https://zenodo.org/record/2807296/files/figure.png" pageId="4" pageNumber="339">Fig. 2</figureCitation>
). Rostral spike light brown, with extensive darker brown flecking at base, tending towards unpatterned at tip. White patches often, but not always, present on either side of head, extending from posterolateral edge of eye, below tympanum and around axilla. Venter predominantly buff, internal organs sometimes visible, throat sometimes offwhite, and/or with two clusters of dark brown maculations laterally, tubercles around vent and sometimes those along outer edge of limbs, off-white.
</paragraph>
<paragraph id="B72836C2FFFE661635AFA7D4A90A735B" blockId="4.[151,1437,151,1833]" pageId="4" pageNumber="339">
<emphasis id="85E3EAD0FFFE661635AFA7D4AB8E71B2" bold="true" box="[199,424,1447,1472]" pageId="4" pageNumber="339">Appearance in life.</emphasis>
The following description is based on photographs of specimens from 
<collectingCountry id="CF807652FFFE661631D6A7DAAFBE71B2" box="[1214,1432,1447,1472]" name="Papua New Guinea" pageId="4" pageNumber="339">Papua New Guinea</collectingCountry>
; Tabubil (Western Province) (
<figureCitation id="2FAC2A47FFFE661634B5A7BFA83B7196" box="[477,541,1484,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1894]" captionTargetBox="[151,1435,195,1848]" captionTargetId="figure@2.[151,1435,195,1848]" captionTargetPageId="2" captionText="FIGURE 1. Papuan Litoria with a rostral spike: A) Litoria pronimia, male, Tabubil, PNG, with spike erect; B) Litoria pronimia, male, Tabubil, PNG, with spike not erect; C) Litoria pronimia, male Kikori River Basin, PNG; D) Litoria pronimia, male, Timika, Indonesia; E) Litoria havina, male, Kikori River Basin, PNG; F) Litoria mucro, male, Foja Mountains, Indonesia; G) Litoria prora, male and female, Kikori River Basin; H) Litoria chrisdahli, Wamangu, PNG. Photographs: S. Richards." httpUri="https://zenodo.org/record/2807292/files/figure.png" pageId="4" pageNumber="339">Fig. 1</figureCitation>
A–B), Gobe (
<collectingRegion id="7553F820FFFE661637D2A7BFA9797197" box="[698,863,1484,1509]" country="Papua New Guinea" name="Gulf" pageId="4" pageNumber="339">Gulf Province</collectingRegion>
), Agogo (
<collectingRegion id="7553F820FFFE661636A7A7BFAF397197" box="[975,1311,1484,1509]" country="Papua New Guinea" name="Southern Highlands" pageId="4" pageNumber="339">Southern Highlands Province</collectingRegion>
) (
<figureCitation id="2FAC2A47FFFE6616305EA7BFAFAB7196" box="[1334,1421,1484,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1894]" captionTargetBox="[151,1435,195,1848]" captionTargetId="figure@2.[151,1435,195,1848]" captionTargetPageId="2" captionText="FIGURE 1. Papuan Litoria with a rostral spike: A) Litoria pronimia, male, Tabubil, PNG, with spike erect; B) Litoria pronimia, male, Tabubil, PNG, with spike not erect; C) Litoria pronimia, male Kikori River Basin, PNG; D) Litoria pronimia, male, Timika, Indonesia; E) Litoria havina, male, Kikori River Basin, PNG; F) Litoria mucro, male, Foja Mountains, Indonesia; G) Litoria prora, male and female, Kikori River Basin; H) Litoria chrisdahli, Wamangu, PNG. Photographs: S. Richards." httpUri="https://zenodo.org/record/2807292/files/figure.png" pageId="4" pageNumber="339">Fig. 1C</figureCitation>
), Crater Mountain (
<collectingRegion id="7553F820FFFE6616340CA79CA817727A" box="[356,561,1519,1544]" country="Papua New Guinea" name="Chimbu" pageId="4" pageNumber="339">Chimbu Province</collectingRegion>
), and 
<collectingCountry id="CF807652FFFE66163710A79CA8CF727A" box="[632,745,1519,1544]" name="Indonesia" pageId="4" pageNumber="339">Indonesia</collectingCountry>
; Timika (
<figureCitation id="2FAC2A47FFFE66163630A79CA997727A" box="[856,945,1519,1544]" captionStart="FIGURE 1" captionStartId="2.[151,250,1872,1894]" captionTargetBox="[151,1435,195,1848]" captionTargetId="figure@2.[151,1435,195,1848]" captionTargetPageId="2" captionText="FIGURE 1. Papuan Litoria with a rostral spike: A) Litoria pronimia, male, Tabubil, PNG, with spike erect; B) Litoria pronimia, male, Tabubil, PNG, with spike not erect; C) Litoria pronimia, male Kikori River Basin, PNG; D) Litoria pronimia, male, Timika, Indonesia; E) Litoria havina, male, Kikori River Basin, PNG; F) Litoria mucro, male, Foja Mountains, Indonesia; G) Litoria prora, male and female, Kikori River Basin; H) Litoria chrisdahli, Wamangu, PNG. Photographs: S. Richards." httpUri="https://zenodo.org/record/2807292/files/figure.png" pageId="4" pageNumber="339">Fig. 1D</figureCitation>
) and the Bomberai Peninsula. Dorsal base colouration light to dark brown, with variable amounts of green and darker brown spotting, mottling or banding on body, limbs and head. Dorsal and lateral tubercles on torso and limbs often, but not always, green and contrasting against brown base colouration. Head brown, often with extensive green pigmentation, usually including a ring around the eyes, and more variably a green transverse band between the eyes. Dense clusters of dark-brown to almost black maculations extend to a variable degree across the limbs, lateral portions of torso, and in patches across the back. Off-white patches sometimes present below tympanum and on exposed surfaces of lower hindlimbs. Iris pattern complex; base colouration light brownish, with extensive darker brown vermiculations; rim of pupil orange. Hidden regions of axilla and groin orange.
</paragraph>
<caption id="E3E8664AFFFF661735FFA1E3AE2B77B6" httpUri="https://zenodo.org/record/2807296/files/figure.png" pageId="5" pageNumber="340" startId="5.[151,250,912,934]" targetBox="[151,1416,239,898]" targetPageId="5">
<paragraph id="B72836C2FFFF661735FFA1E3AE2B77B6" blockId="5.[151,1436,912,964]" pageId="5" pageNumber="340">
<emphasis id="85E3EAD0FFFF661735FFA1E3AB3577D7" bold="true" box="[151,275,912,934]" pageId="5" pageNumber="340">FIGURE 2.</emphasis>
<taxonomicName id="70974D41FFFF66173472A1E2ABE277D4" authorityName="Menzies" authorityYear="1993" box="[282,452,913,934]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF66173472A1E2ABE277D4" box="[282,452,913,934]" italics="true" pageId="5" pageNumber="340">Litoria pronimia</emphasis>
</taxonomicName>
in preservative from near the type locality at Tabubil (SAMA R71135–6, 71140) and the Kikori River Basin (SAMA R77131). Photograph by Peter Waddington, Queensland Museum.
</paragraph>
</caption>
<paragraph id="B72836C2FFFF661735AFA19FAB8771C7" blockId="5.[151,1436,1004,2037]" pageId="5" pageNumber="340">
<emphasis id="85E3EAD0FFFF661735AFA19FA8767077" bold="true" box="[199,592,1004,1029]" pageId="5" pageNumber="340">Comparisons with other species.</emphasis>
The combination of small adult size (&lt;
<quantity id="706F9B27FFFF66173171A19FAE497076" box="[1049,1135,1004,1029]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="340" unit="mm" value="35.0">35 mm</quantity>
), slender build, and males with a rostral spike that is rounded in cross-section will distinguish 
<taxonomicName id="70974D41FFFF661736B0A662AE47705A" authorityName="Menzies" authorityYear="1993" box="[984,1121,1041,1064]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF661736B0A662AE47705A" box="[984,1121,1041,1064]" italics="true" pageId="5" pageNumber="340">L. pronimia</emphasis>
</taxonomicName>
from all other previously recognised Melanesian 
<taxonomicName id="70974D41FFFF661734C9A646ABC9703E" box="[417,495,1077,1100]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="genus">
<emphasis id="85E3EAD0FFFF661734C9A646ABC9703E" box="[417,495,1077,1100]" italics="true" pageId="5" pageNumber="340">Litoria</emphasis>
</taxonomicName>
except 
<taxonomicName id="70974D41FFFF66173720A646A894703E" authorityName="Menzies" authorityYear="1993" box="[584,690,1076,1100]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="havina">
<emphasis id="85E3EAD0FFFF66173720A646A894703E" box="[584,690,1076,1100]" italics="true" pageId="5" pageNumber="340">L. havina</emphasis>
</taxonomicName>
, 
<taxonomicName id="70974D41FFFF661737A9A646A915703E" authorityName="Günther" authorityYear="2008" box="[705,819,1076,1100]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="mareku">
<emphasis id="85E3EAD0FFFF661737A9A646A915703E" box="[705,819,1076,1100]" italics="true" pageId="5" pageNumber="340">L. mareku</emphasis>
</taxonomicName>
and 
<taxonomicName id="70974D41FFFF66173603A646A9F6703E" authorityName="Menzies" authorityYear="1993" box="[875,976,1077,1100]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="mucro">
<emphasis id="85E3EAD0FFFF66173603A646A9F6703E" box="[875,976,1077,1100]" italics="true" pageId="5" pageNumber="340">L. mucro</emphasis>
</taxonomicName>
. 
<taxonomicName id="70974D41FFFF661736B7A646AEBC703E" authorityName="Menzies" authorityYear="1993" box="[991,1178,1077,1100]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF661736B7A646AEBC703E" box="[991,1178,1077,1100]" italics="true" pageId="5" pageNumber="340">Litoria pronimia</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="70974D41FFFF66173059A646AFBD703E" authorityName="Menzies" authorityYear="1993" box="[1329,1435,1076,1100]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="havina">
<emphasis id="85E3EAD0FFFF66173059A646AFBD703E" box="[1329,1435,1076,1100]" italics="true" pageId="5" pageNumber="340">L. havina</emphasis>
</taxonomicName>
in having a complex brown and green dorsal pattern (vs uniform green or brown) and orange groin colouration (vs red); from 
<taxonomicName id="70974D41FFFF6617347BA60EABA370E6" authorityName="Günther" authorityYear="2008" box="[275,389,1148,1172]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="mareku">
<emphasis id="85E3EAD0FFFF6617347BA60EABA370E6" box="[275,389,1148,1172]" italics="true" pageId="5" pageNumber="340">L. mareku</emphasis>
</taxonomicName>
in its larger size (male SVL 
<quantity id="706F9B27FFFF661737BAA60FA95170E6" box="[722,887,1148,1173]" metricMagnitude="-2" metricUnit="m" metricValue="2.8649999999999998" metricValueMax="3.04" metricValueMin="2.69" pageId="5" pageNumber="340" unit="mm" value="28.65" valueMax="30.4" valueMin="26.9">26.9–30.4 mm</quantity>
vs 
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), long erectile rostral spike in males (vs effectively a short stub that is wider than long), and light to medium-brown base colouration on dorsum (vs at least some regions of very dark chocolate brown); and from 
<taxonomicName id="70974D41FFFF661736F7A6B6AE2370AE" authorityName="Menzies" authorityYear="1993" box="[927,1029,1221,1244]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="mucro">
<emphasis id="85E3EAD0FFFF661736F7A6B6AE2370AE" box="[927,1029,1221,1244]" italics="true" pageId="5" pageNumber="340">L. mucro</emphasis>
</taxonomicName>
by its more complex dorsal colour pattern including extensive green flecking and banding (vs predominantly olive-green to brown dorsum with at most a small amount of light-green flecking). Males of 
<taxonomicName id="70974D41FFFF66173665A77EA95E7156" authorityName="Menzies" authorityYear="1993" box="[781,888,1292,1316]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="havina">
<emphasis id="85E3EAD0FFFF66173665A77EA95E7156" box="[781,888,1292,1316]" italics="true" pageId="5" pageNumber="340">L. havina</emphasis>
</taxonomicName>
and 
<taxonomicName id="70974D41FFFF661736D8A77EAE337156" authorityName="Menzies" authorityYear="1993" box="[944,1045,1293,1316]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="mucro">
<emphasis id="85E3EAD0FFFF661736D8A77EAE337156" box="[944,1045,1293,1316]" italics="true" pageId="5" pageNumber="340">L. mucro</emphasis>
</taxonomicName>
tend to have rostral spikes shorter than 
<taxonomicName id="70974D41FFFF661735A6A742AB74713A" authorityName="Menzies" authorityYear="1993" box="[206,338,1329,1352]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF661735A6A742AB74713A" box="[206,338,1329,1352]" italics="true" pageId="5" pageNumber="340">L. pronimia</emphasis>
</taxonomicName>
(&lt;
<quantity id="706F9B27FFFF66173419A743ABEE713A" box="[369,456,1328,1353]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="340" unit="mm" value="1.5">1.5 mm</quantity>
vs&gt; 
<quantity id="706F9B27FFFF66173496A743A87E713A" box="[510,600,1328,1353]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="340" unit="mm" value="1.5">1.5 mm</quantity>
), however as noted above the length of the rostral spike may vary between populations and samples of 
<taxonomicName id="70974D41FFFF661734B9A726A873711E" authorityName="Menzies" authorityYear="1993" box="[465,597,1365,1388]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF661734B9A726A873711E" box="[465,597,1365,1388]" italics="true" pageId="5" pageNumber="340">L. pronimia</emphasis>
</taxonomicName>
, reducing the utility of this character. Even though there are relatively few diagnostic characters seperating 
<taxonomicName id="70974D41FFFF6617376BA70AA84E71E2" authorityName="Menzies" authorityYear="1993" box="[515,616,1401,1424]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="mucro">
<emphasis id="85E3EAD0FFFF6617376BA70AA84E71E2" box="[515,616,1401,1424]" italics="true" pageId="5" pageNumber="340">L. mucro</emphasis>
</taxonomicName>
and 
<taxonomicName id="70974D41FFFF661737F5A70AA90771E2" authorityName="Menzies" authorityYear="1993" box="[669,801,1401,1424]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF661737F5A70AA90771E2" box="[669,801,1401,1424]" italics="true" pageId="5" pageNumber="340">L. pronimia</emphasis>
</taxonomicName>
, genetic data also indicate that they are deeply divergent (Oliver 
<emphasis id="85E3EAD0FFFF66173587A7EEAB0171C6" box="[239,295,1436,1460]" italics="true" pageId="5" pageNumber="340">et al.</emphasis>
accepted).
</paragraph>
<paragraph id="B72836C2FFFF661735AFA7B2A91372FF" blockId="5.[151,1436,1004,2037]" pageId="5" pageNumber="340">
Four other species of 
<taxonomicName id="70974D41FFFF661734D4A7B2A82C71AA" box="[444,522,1473,1496]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="genus">
<emphasis id="85E3EAD0FFFF661734D4A7B2A82C71AA" box="[444,522,1473,1496]" italics="true" pageId="5" pageNumber="340">Litoria</emphasis>
</taxonomicName>
from New 
<collectingCountry id="CF807652FFFF661737E4A7B3A8FB71AB" box="[652,733,1472,1497]" name="Guinea-Bissau" pageId="5" pageNumber="340">Guinea</collectingCountry>
have a rostral spike: 
<taxonomicName id="70974D41FFFF661736A6A7B2AEED71AB" authority="Richards" authorityName="Richards" authorityYear="2007" box="[974,1227,1472,1497]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="chrisdahli">
<emphasis id="85E3EAD0FFFF661736A6A7B2AE7B71AA" box="[974,1117,1472,1496]" italics="true" pageId="5" pageNumber="340">L. chrisdahli</emphasis>
Richards
</taxonomicName>
, 
<taxonomicName id="70974D41FFFF661731B0A7B2AADB718F" authority="Hiaso and Richards" authorityName="Hiaso and Richards" authorityYear="2006" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="hilli">
<emphasis id="85E3EAD0FFFF661731B0A7B2AF0571AA" box="[1240,1315,1472,1496]" italics="true" pageId="5" pageNumber="340">L. hilli</emphasis>
Hiaso and Richards
</taxonomicName>
, 
<taxonomicName id="70974D41FFFF66173463A796A81F718F" authority="Gunther" authorityName="Gunther" box="[267,569,1508,1533]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="humboldtorum">
<emphasis id="85E3EAD0FFFF66173463A796ABF7718E" box="[267,465,1508,1532]" italics="true" pageId="5" pageNumber="340">L. humboldtorum</emphasis>
Günther
</taxonomicName>
, and 
<taxonomicName id="70974D41FFFF66173711A796A94F718E" authority="Menzies. In" authorityName="Menzies. In" box="[633,873,1508,1533]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="prora">
<emphasis id="85E3EAD0FFFF66173711A796A8FE718E" box="[633,728,1509,1532]" italics="true" pageId="5" pageNumber="340">L. prora</emphasis>
Menzies. In
</taxonomicName>
addition to their larger size (male SVL between 37.0– 
<quantity id="706F9B27FFFF661735BEA47BAB1A7252" box="[214,316,1544,1568]" metricMagnitude="-2" metricUnit="m" metricValue="4.9399999999999995" pageId="5" pageNumber="340" unit="mm" value="49.4">49.4 mm</quantity>
) these species can be diagnosed from 
<taxonomicName id="70974D41FFFF66173787A47AA9557252" authorityName="Menzies" authorityYear="1993" box="[751,883,1545,1568]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF66173787A47AA9557252" box="[751,883,1545,1568]" italics="true" pageId="5" pageNumber="340">L. pronimia</emphasis>
</taxonomicName>
by an array of characters largely summarised by 
<bibRefCitation id="D3064B33FFFF661735FFA45EAB097236" author="Kraus, F." box="[151,303,1580,1604]" pageId="5" pageNumber="340" pagination="365 - 374" refId="ref7468" refString="Kraus, F. (2007) A new species of treefrog of the genus Litoria (Hylidae) from the Louisiade Islands, Papua New Guinea. Herpetologica, 63, 365 - 374. https: // doi. org / 10.1655 / 0018 - 0831 (2007) 63 [365: ANSOTO] 2.0. CO; 2" type="journal article" year="2007">Kraus (2007)</bibRefCitation>
: fleshy rostral spike present in both sexes, moderate to extensive dermal crenulations along the distal edge of the limbs and fourth finger, fully webbed hands, papillae along the chin margin, and posterior abdomen and ventral surfaces of thighs dark purplish with white pustules.
</paragraph>
<paragraph id="B72836C2FFFF661735AFA4EBABE3736E" blockId="5.[151,1436,1004,2037]" pageId="5" pageNumber="340">
<emphasis id="85E3EAD0FFFF661735AFA4EBAB3C72C3" bold="true" box="[199,282,1688,1713]" pageId="5" pageNumber="340">Range.</emphasis>
<taxonomicName id="70974D41FFFF6617344AA4EAABFB72C2" authorityName="Menzies" authorityYear="1993" box="[290,477,1689,1712]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF6617344AA4EAABFB72C2" box="[290,477,1689,1712]" italics="true" pageId="5" pageNumber="340">Litoria pronimia</emphasis>
</taxonomicName>
is now known from a broad band of lowland and foothill forest spanning over 
<quantity id="706F9B27FFFF6617300CA4EBAA9D72A7" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="340" unit="km" value="1000.0">1000 km</quantity>
along the the southern edge of the Central Cordillera from the Crater Mountain Wildlife Management Area in the east, to at least as far west as Timika in 
<collectingRegion id="7553F820FFFF661737EBA492A8E1728A" box="[643,711,1761,1784]" country="Indonesia" name="Papua Barat" pageId="5" pageNumber="340">Papua</collectingRegion>
<collectingCountry id="CF807652FFFF661737A7A493A919728B" box="[719,831,1760,1785]" name="Indonesia" pageId="5" pageNumber="340">Indonesia</collectingCountry>
, with a further (potentially disjunct) population in the Fakfak Mountains (
<figureCitation id="2FAC2A47FFFF6617341BA577AB91736E" box="[371,439,1796,1821]" captionStart="FIGURE 3" captionStartId="6.[151,250,1306,1328]" captionTargetBox="[178,1410,451,1259]" captionTargetId="figure@6.[151,1436,375,1284]" captionTargetPageId="6" captionText="FIGURE 3. Distribution of Litoria pronimia (squares) across southern New Guinea and Litoria pinocchio sp. nov. in the Foja Mountains (asterisk)." httpUri="https://zenodo.org/record/2807298/files/figure.png" pageId="5" pageNumber="340">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="B72836C2FFFF661435AFA55BAF1E74A7" blockId="5.[151,1436,1004,2037]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="341" pageId="5" pageNumber="340">
<emphasis id="85E3EAD0FFFF661735AFA55BABB57333" bold="true" box="[199,403,1832,1857]" pageId="5" pageNumber="340">Ecological notes.</emphasis>
<taxonomicName id="70974D41FFFF661734F7A55AA8797332" authorityName="Menzies" authorityYear="1993" box="[415,607,1833,1856]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF661734F7A55AA8797332" box="[415,607,1833,1856]" italics="true" pageId="5" pageNumber="340">Litoria pronimia</emphasis>
</taxonomicName>
is known from lowland, hill and lower montane forests at elevations between ~ 
<quantity id="706F9B27FFFF66173462A53FABBF7316" box="[266,409,1868,1893]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="11.5" metricValueMin="2.0" pageId="5" pageNumber="340" unit="m" value="675.0" valueMax="1150.0" valueMin="200.0">200–1150 m</quantity>
a.s.l. where it forms breeding choruses around ponds or other waterbodies in disturbed or primary rainforest. One of the authors (R. Günther) found a small gelatinous egg cluster containing less than 10 white eggs on a leaf approximately 
<quantity id="706F9B27FFFF66173743A5E7A85973DE" box="[555,639,1940,1965]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="340" unit="cm" value="150.0">150 cm</quantity>
above a roadside ditch, at 
<quantity id="706F9B27FFFF661736C7A5E7A9D173DE" box="[943,1015,1940,1964]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="340" unit="m" value="800.0">800 m</quantity>
a.s.l. in the Fakfak Mountains on the 
<date id="C3291002FFFF661735FFA5CBABA473A2" box="[151,386,1976,2001]" pageId="5" pageNumber="340" value="2008-09-07">7th September 2008</date>
. According to investigations of mitochondrial 12S genes these eggs belong to 
<taxonomicName id="70974D41FFFF66173067A5CAAFB273A2" authorityName="Menzies" authorityYear="1993" box="[1295,1428,1977,2000]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFF66173067A5CAAFB273A2" box="[1295,1428,1977,2000]" italics="true" pageId="5" pageNumber="340">L. pronimia</emphasis>
</taxonomicName>
, with sequences the same as ZMB 85620–21 (R. Günther, unpublished). At the same site calling males of 
<taxonomicName id="70974D41FFFF66143026A5AEAB5574C2" authorityName="Günther" authorityYear="2008" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" lastPageId="6" lastPageNumber="341" order="Anura" pageId="5" pageNumber="340" phylum="Chordata" rank="species" species="christianbergmanni">
<emphasis id="85E3EAD0FFFF66143026A5AEAB5574C2" italics="true" lastPageId="6" lastPageNumber="341" pageId="5" pageNumber="340">Litoria christianbergmanni</emphasis>
</taxonomicName>
and 
<taxonomicName id="70974D41FFFC661434C6A2EAA83C74C2" authorityName="Menzies" authorityYear="1993" box="[430,538,152,176]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="6" pageNumber="341" phylum="Chordata" rank="species" species="havina">
<emphasis id="85E3EAD0FFFC661434C6A2EAA83C74C2" box="[430,538,152,176]" italics="true" pageId="6" pageNumber="341">L. havina</emphasis>
</taxonomicName>
were found, in addition to many eggs and larvae of the latter species. 
<taxonomicName id="70974D41FFFC66143026A2EBAADB74A6" authorityName="Menzies" authorityYear="1993" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="6" pageNumber="341" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFC66143026A2EBAADB74A6" italics="true" pageId="6" pageNumber="341">Litoria pronimia</emphasis>
</taxonomicName>
occurs in sympatry with, and uses the same breeding sites as 
<taxonomicName id="70974D41FFFC661436D8A2CEAE3C74A6" authorityName="Menzies" authorityYear="1993" box="[944,1050,188,212]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="6" pageNumber="341" phylum="Chordata" rank="species" species="havina">
<emphasis id="85E3EAD0FFFC661436D8A2CEAE3C74A6" box="[944,1050,188,212]" italics="true" pageId="6" pageNumber="341">L. havina</emphasis>
</taxonomicName>
across much of its range.
</paragraph>
<paragraph id="B72836C2FFFC661435AFA2ACA9DB7532" blockId="6.[151,1436,151,320]" pageId="6" pageNumber="341">
<emphasis id="85E3EAD0FFFC661435AFA2ACABE4748A" bold="true" box="[199,450,223,248]" pageId="6" pageNumber="341">Conservation status.</emphasis>
<taxonomicName id="70974D41FFFC661434B8A293A8B47485" authorityName="Menzies" authorityYear="1993" box="[464,658,224,247]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="6" pageNumber="341" phylum="Chordata" rank="species" species="pronimia">
<emphasis id="85E3EAD0FFFC661434B8A293A8B47485" box="[464,658,224,247]" italics="true" pageId="6" pageNumber="341">Litoria pronimia</emphasis>
</taxonomicName>
is currently listed as Least Concern by the IUCN based on an assessment in 2004. Subsequent records extend its known distribution 1000 kilometres to the west, including large areas of relatively little-disturbed and continuous forest at suitable elevations.
</paragraph>
</subSubSection>
</treatment>
</document>