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<document id="9D4C9BD4BB42167F6B3B425FADDC0BCA" ID-CLB-Dataset="45591" ID-DOI="10.5281/zenodo.212706" ID-GBIF-Dataset="545ed44c-044a-4be2-94c9-4b8d96f5bdff" ID-ISSN="1175-5326" ID-Zenodo-Dep="212706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460431779058" checkinUser="plazi" docAuthor="Beasley, Clark W. &amp; Miller, William R." docDate="2012" docId="36479C4DFFE1FFB261ADF8C3B681FDC9" docLanguage="en" docName="zt03170p063.pdf" docOrigin="Zootaxa 3170" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pseudechiniscus papillosus Li, Wang, Liu &amp; Su 2005" docType="treatment" docVersion="11" lastPageNumber="58" masterDocId="CA7EE435FFE0FFB1613AFFECB44EFFDE" masterDocTitle="Additional Tardigrada from Hubei Province, China, with the description of Doryphoribius barbarae sp. nov. (Eutardigrada: Parachela: Hypsibiidae)" masterLastPageNumber="63" masterPageNumber="55" pageNumber="56" updateTime="1698331490213" updateUser="ExternalLinkService">
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<mods:title id="7CAFA02734A8EEA81472E58C279B1D25">Additional Tardigrada from Hubei Province, China, with the description of Doryphoribius barbarae sp. nov. (Eutardigrada: Parachela: Hypsibiidae)</mods:title>
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<mods:name id="66AE935F71F8E47088F4D794169C39A7" type="personal">
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<mods:namePart id="643B49A496BA83D1240ED7BB2F489333">Beasley, Clark W.</mods:namePart>
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<mods:namePart id="A3F41E251EB73406E20C980964A3AE0A">Miller, William R.</mods:namePart>
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<mods:typeOfResource id="BAFAB9148949B839B1B9B87CBE5A998A">text</mods:typeOfResource>
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<mods:title id="9DB839B7DF2D57B01119587408630489">Zootaxa</mods:title>
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<mods:part id="3633B46929F5A7B401054CB7094D4F6A">
<mods:date id="571B642EBDB2F1CFDC0083E2A33164A4">2012</mods:date>
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<mods:number id="09D634DD3A379967093786926B4FFCD9">3170</mods:number>
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<mods:identifier id="C51637B4069439D65F0252CCE54B53CC" type="DOI">10.5281/zenodo.212706</mods:identifier>
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<treatment id="36479C4DFFE1FFB261ADF8C3B681FDC9" ID-DOI="http://doi.org/10.5281/zenodo.5672288" ID-GBIF-Taxon="119430497" ID-Zenodo-Dep="5672288" LSID="urn:lsid:plazi:treatment:36479C4DFFE1FFB261ADF8C3B681FDC9" httpUri="http://treatment.plazi.org/id/36479C4DFFE1FFB261ADF8C3B681FDC9" lastPageId="3" lastPageNumber="58" pageId="1" pageNumber="56">
<subSubSection id="F6F47ED0FFE1FFB061ADF8C3B51CF8B4" pageId="1" pageNumber="56" type="nomenclature">
<paragraph id="BE512D5BFFE1FFB061ADF8C3B703F897" blockId="1.[151,845,1839,1898]" box="[151,845,1839,1865]" pageId="1" pageNumber="56">
<heading id="E5199A37FFE1FFB061ADF8C3B703F897" bold="true" box="[151,845,1839,1865]" fontSize="11" level="1" pageId="1" pageNumber="56" reason="1">
<taxonomicName id="79EE56D8FFE1FFB061ADF8C3B703F897" ID-CoL="4NM29" authority="Li, Wang, Liu &amp; Su, 2005" authorityName="Li, Wang, Liu &amp; Su" authorityYear="2005" box="[151,845,1839,1865]" class="Heterotardigrada" family="Echiniscidae" genus="Pseudechiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="1" pageNumber="56" phylum="Tardigrada" rank="species" species="papillosus">
<emphasis id="8C9AF149FFE1FFB061ADF8C3B703F897" bold="true" box="[151,845,1839,1865]" pageId="1" pageNumber="56">
<emphasis id="8C9AF149FFE1FFB061ADF8C3B511F897" bold="true" box="[151,351,1839,1865]" italics="true" pageId="1" pageNumber="56">Pseudechiniscus</emphasis>
cf.
<emphasis id="8C9AF149FFE1FFB060B6F8C3B648F897" bold="true" box="[396,518,1839,1865]" italics="true" pageId="1" pageNumber="56">papillosus</emphasis>
<bibRefCitation id="DA7F50AAFFE1FFB06337F8C3B703F897" author="Li" box="[525,845,1839,1865]" pageId="1" pageNumber="56" refString="Li, X., Wang, L., Liu, Y. &amp; Su, L. (2005) A new species and five new records of the family Echiniscidae (Tardigrada) from China. Zootaxa, 1093, 25 - 33." type="journal article" year="2005">Li, Wang, Liu &amp; Su, 2005</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="BE512D5BFFE1FFB061ADF8BEB51CF8B4" blockId="1.[151,845,1839,1898]" box="[151,338,1874,1898]" pageId="1" pageNumber="56">
<figureCitation id="26D531DEFFE1FFB061ADF8BEB555F8B4" box="[151,283,1874,1898]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1960]" captionTargetBox="[240,1349,203,1891]" captionTargetId="figure@2.[228,1351,193,1898]" captionTargetPageId="2" captionText="FIGURE 1. Pseudechiniscus papillosus. A, habitus with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10 µm." httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="1" pageNumber="56">FIGURES 1</figureCitation>
AF
</paragraph>
</subSubSection>
<subSubSection id="F6F47ED0FFE1FFB061ADF876B064F86C" box="[151,1066,1946,1971]" pageId="1" pageNumber="56" type="materials_examined">
<paragraph id="BE512D5BFFE1FFB061ADF876B064F86C" blockId="1.[151,1066,1946,1971]" box="[151,1066,1946,1971]" pageId="1" pageNumber="56">
<emphasis id="8C9AF149FFE1FFB061ADF876B5CEF86D" bold="true" box="[151,384,1946,1971]" pageId="1" pageNumber="56">Material examined.</emphasis>
Eleven specimens were found in a moss sample from Site 3.
</paragraph>
</subSubSection>
<caption id="EA917DD3FFE2FFB361ADF87DB0F9F83B" httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="2" pageNumber="57" targetBox="[240,1349,203,1891]" targetPageId="2">
<paragraph id="BE512D5BFFE2FFB361ADF87DB0F9F83B" blockId="2.[151,1436,1937,2022]" pageId="2" pageNumber="57">
<emphasis id="8C9AF149FFE2FFB361ADF87DB55AF879" bold="true" box="[151,276,1937,1960]" pageId="2" pageNumber="57">FIGURE 1.</emphasis>
<taxonomicName id="79EE56D8FFE2FFB36026F87EB67AF876" box="[284,564,1938,1960]" class="Heterotardigrada" family="Echiniscidae" genus="Pseudechiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="57" phylum="Tardigrada" rank="species" species="papillosus">
<emphasis id="8C9AF149FFE2FFB36026F87EB67AF876" box="[284,564,1938,1960]" italics="true" pageId="2" pageNumber="57">Pseudechiniscus papillosus</emphasis>
</taxonomicName>
. A,
<emphasis id="8C9AF149FFE2FFB36359F87EB6E3F876" box="[611,685,1938,1960]" italics="true" pageId="2" pageNumber="57">habitus</emphasis>
with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10µm.
</paragraph>
</caption>
<subSubSection id="F6F47ED0FFE3FFB261FFFF74B681FDC9" pageId="3" pageNumber="58" type="discussion">
<paragraph id="BE512D5BFFE3FFB261FFFF74B681FDC9" blockId="3.[151,1436,151,536]" pageId="3" pageNumber="58">
<emphasis id="8C9AF149FFE3FFB261FFFF74B577FF6E" bold="true" box="[197,313,152,176]" pageId="3" pageNumber="58">Remarks.</emphasis>
This is the second report of this species, the original description from Shaanxi Province (
<bibRefCitation id="DA7F50AAFFE3FFB26405FF7BB49AFF0A" author="Li" pageId="3" pageNumber="58" refString="Li, X., Wang, L., Liu, Y. &amp; Su, L. (2005) A new species and five new records of the family Echiniscidae (Tardigrada) from China. Zootaxa, 1093, 25 - 33." type="journal article" year="2005">
Li
<emphasis id="8C9AF149FFE3FFB26458FF75B1DBFF6E" box="[1378,1429,152,176]" italics="true" pageId="3" pageNumber="58">et al</emphasis>
. 2005
</bibRefCitation>
). The specimens from this study possessed a pair of pointed lobes on the posterior margin of the pseudosegmental plate (
<figureCitation id="26D531DEFFE3FFB2600AFF33B521FF29" box="[304,367,223,247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1960]" captionTargetBox="[240,1349,203,1891]" captionTargetId="figure@2.[228,1351,193,1898]" captionTargetPageId="2" captionText="FIGURE 1. Pseudechiniscus papillosus. A, habitus with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10 µm." httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
A) and, as reported by
<bibRefCitation id="DA7F50AAFFE3FFB2634EFF33B76FFF29" box="[628,801,223,248]" pageId="3" pageNumber="58" refString="Li, X., Wang, L., Liu, Y. &amp; Su, L. (2005) A new species and five new records of the family Echiniscidae (Tardigrada) from China. Zootaxa, 1093, 25 - 33." type="journal article">
Li
<emphasis id="8C9AF149FFE3FFB263AEFF0DB683FF26" box="[660,717,224,248]" italics="true" pageId="3" pageNumber="58">et al.</emphasis>
(2005)
</bibRefCitation>
, the cephalic papillae were short and broad (
<figureCitation id="26D531DEFFE3FFB2642FFF33B11BFF29" box="[1301,1365,223,247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1960]" captionTargetBox="[240,1349,203,1891]" captionTargetId="figure@2.[228,1351,193,1898]" captionTargetPageId="2" captionText="FIGURE 1. Pseudechiniscus papillosus. A, habitus with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10 µm." httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
B, C). The Hubei specimens closely conform to the species description, but with the following differences: neck plate was present between the cephalic and scapular plates (
<figureCitation id="26D531DEFFE3FFB263ECFECBB756FEE1" box="[726,792,295,319]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1960]" captionTargetBox="[240,1349,203,1891]" captionTargetId="figure@2.[228,1351,193,1898]" captionTargetPageId="2" captionText="FIGURE 1. Pseudechiniscus papillosus. A, habitus with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10 µm." httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
B); the
<typeStatus id="615593F9FFE3FFB2624EFEC4B7EAFE9E" box="[884,932,296,320]" pageId="3" pageNumber="58">type</typeStatus>
specimens all had relatively short cirrus A, ranging from 28.7 to 36.5 µm, while the Hubei specimens showed more variation in cirrus A (a range of 28.770.3 µm for eight specimens, with an average of 53.4 µm); the original description stated spurs were absent on all claws, but a small, short, slender spur, usually perpendicular to the axis of the claw, was visible on the external claws of leg IV for some of the Hubei specimens (
<figureCitation id="26D531DEFFE3FFB2635FFE5BB6EAFE11" box="[613,676,439,463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1960]" captionTargetBox="[240,1349,203,1891]" captionTargetId="figure@2.[228,1351,193,1898]" captionTargetPageId="2" captionText="FIGURE 1. Pseudechiniscus papillosus. A, habitus with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10 µm." httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
D); leg IV dentate collar consists of a single broad tooth on the dorsal surface (
<figureCitation id="26D531DEFFE3FFB26026FE30B515FE2A" box="[284,347,476,500]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1960]" captionTargetBox="[240,1349,203,1891]" captionTargetId="figure@2.[228,1351,193,1898]" captionTargetPageId="2" captionText="FIGURE 1. Pseudechiniscus papillosus. A, habitus with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10 µm." httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
E); a spine on the front legs is not mentioned in the original description but a small spine (2.73.1 µm) was present in the Hubei specimens (
<figureCitation id="26D531DEFFE3FFB26357FE13B6E2FDC9" box="[621,684,511,535]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1960]" captionTargetBox="[240,1349,203,1891]" captionTargetId="figure@2.[228,1351,193,1898]" captionTargetPageId="2" captionText="FIGURE 1. Pseudechiniscus papillosus. A, habitus with pseudosegmental lobes visible; B, anterior of body with neck plate visible; C, cephalic papilla between buccal cirri; D, slender spur (arrow) on external claw of leg IV; E, dentate collar of leg IV composed of a single, wide point (arrow); F, spine on leg I. Scale line A, 20 µm; other scale lines 10 µm." httpUri="https://zenodo.org/record/212707/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
F).
</paragraph>
</subSubSection>
</treatment>
</document>