452 lines
52 KiB
XML
452 lines
52 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/vz.72.e77702" ID-GBIF-Dataset="756d7aec-8ed2-473e-83cf-8779c96fa6c1" ID-GBIF-Taxon="193278647" ID-Pensoft-Pub="2625-8498-72-47" ID-Pensoft-UUID="66A14C58EB6659F29033A6E475A606B4" ID-ZooBank="3A5B7AD25A8142E9B0136A95ED03E7B6" ModsDocID="2625-8498-72-47" checkinTime="1644016157858" checkinUser="pensoft" docAuthor="Grismer, L. Lee, del Pinto, Lelani, Quah, Evan S. H., Anuar, Shahrul, Cota, Michael, McGuire, Jimmy A., Iskandar, Djoko T., Wood Jr, Perry L. & Grismer, Jesse L." docDate="2022" docId="FBEECBA893B75AC494CC2E029A6C3342" docLanguage="en" docName="VertZool 72: 47-80" docOrigin="Vertebrate Zoology 72" docPubDate="2022-02-03" docSource="http://dx.doi.org/10.3897/vz.72.e77702" docTitle="Gekko albofasciolatus" docType="treatment" docVersion="3" id="66A14C58EB6659F29033A6E475A606B4" lastPageNumber="47" masterDocId="66A14C58EB6659F29033A6E475A606B4" masterDocTitle="Phylogenetic and multivariate analyses of Gekko smithii Gray, 1842 recover a new species from Peninsular Malaysia and support the resurrection of G. albomaculatus (Giebel, 1861) from Sumatra" masterLastPageNumber="80" masterPageNumber="47" pageId="0" pageNumber="47" updateTime="1644020671795" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Phylogenetic and multivariate analyses of Gekko smithii Gray, 1842 recover a new species from Peninsular Malaysia and support the resurrection of G. albomaculatus (Giebel, 1861) from Sumatra</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Grismer, L. Lee</mods:namePart>
|
||
<mods:affiliation>Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA</mods:affiliation>
|
||
<mods:nameIdentifier type="email">lgrismer@lasierra.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>del Pinto, Lelani</mods:namePart>
|
||
<mods:affiliation>Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Quah, Evan S. H.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5357-1953</mods:nameIdentifier>
|
||
<mods:affiliation>Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Sabah, Malaysia & School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Anuar, Shahrul</mods:namePart>
|
||
<mods:affiliation>School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia & Center for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Malaysia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Cota, Michael</mods:namePart>
|
||
<mods:affiliation>Natural History Museum, National Science Museum, Thailand</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>McGuire, Jimmy A.</mods:namePart>
|
||
<mods:affiliation>Museum of Vertebrate Zoology and Department of Integrative Biology, University of California, Berkeley, California 94720, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Iskandar, Djoko T.</mods:namePart>
|
||
<mods:affiliation>The Indonesian Academy of Sciences, Jakarta 10110 Indonesia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Wood Jr, Perry L.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3767-5274</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan 48109 - 1085, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Grismer, Jesse L.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2542-5430</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Vertebrate Zoology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2022</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2022-02-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>72</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>47</mods:start>
|
||
<mods:end>80</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/vz.72.e77702</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/vz.72.e77702</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">2625-8498-72-47</mods:identifier>
|
||
<mods:identifier type="ZooBank">3A5B7AD25A8142E9B0136A95ED03E7B6</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">66A14C58EB6659F29033A6E475A606B4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="193278647" LSID="urn:lsid:plazi:treatment:FBEECBA893B75AC494CC2E029A6C3342" httpUri="http://treatment.plazi.org/id/FBEECBA893B75AC494CC2E029A6C3342" lastPageNumber="47" pageId="0" pageNumber="47">
|
||
<subSubSection pageId="0" pageNumber="47" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<taxonomicName LSID="FBEECBA8-93B7-5AC4-94CC-2E029A6C3342" authority="(Guenther 1867)" baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">
|
||
Gekko albofasciolatus (
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
1867)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="47" type="">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<bibRefCitation author="Roesler, H" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="1 - 50" refId="B117" refString="Roesler, H, Bauer, AM, Heinicke, MP, Greenbaum, E, Jackman, T, Nguyen, TQ, Ziegler, T, 2011. Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae). Zootaxa 2989: 1 - 50" title="Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae)." volume="2989" year="2011">
|
||
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
||
et al. (2011)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Wood, PL" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="47" refId="B135" refString="Wood, PL, Guo, X, Travers, SL, Su, YC, Olson, KV, Bauer, AM, Grismer, LL, Siler, CD, Moyle, RG, Andersen, MJ, Brown, RF, 2020a. . Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731." title=". Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731." year="2020 a">Wood et al. (2020a)</bibRefCitation>
|
||
considered
|
||
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Gekko albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
a valid species within
|
||
<taxonomicName genus="G." lsidName="G." pageId="0" pageNumber="47" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G.</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Grismer & del Pinto & Quah & Anuar & Cota & McGuire & Iskandar & Wood Jr & Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Gekko</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation author="Roesler, H" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="1 - 50" refId="B117" refString="Roesler, H, Bauer, AM, Heinicke, MP, Greenbaum, E, Jackman, T, Nguyen, TQ, Ziegler, T, 2011. Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae). Zootaxa 2989: 1 - 50" title="Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae)." volume="2989" year="2011">
|
||
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
||
et al. (2011)
|
||
</bibRefCitation>
|
||
provide a detailed history of the taxonomy
|
||
<taxonomicName genus="G." lsidName="G. albofaciolatus" pageId="0" pageNumber="47" rank="species" species="albofaciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofaciolatus</emphasis>
|
||
</taxonomicName>
|
||
which is best repeated here in its original form: "In the original description of
|
||
<taxonomicName authority="Brongniart, 1800" authorityName="Gunther" authorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko albo-fasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albo-fasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Gecko albo-fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Gekko albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Sauria" pageId="0" pageNumber="47" refId="B58" refString="Guenther, A, 1867. Additions to the knowledge of Australian reptiles and fishes. Annals and Magazine of Natural History Series 3, 320: 45-68." title="Additions to the knowledge of Australian reptiles and fishes. Annals and Magazine of Natural History Series 3, 320: 45 - 68." year="1867">
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
(1867)
|
||
</bibRefCitation>
|
||
only provided a vague type locality Polynesia with a question mark.
|
||
<bibRefCitation author="Smith, MA" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B123" refString="Smith, MA, 1935. The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor & Francis Ltd." title="The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor & Francis Ltd." year="1935">Smith (1935)</bibRefCitation>
|
||
subsequently stated 'type loc. unknown, probably Malay Archipelago.' Later,
|
||
<bibRefCitation author="Wermuth, H" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="47" refId="B133" refString="Wermuth, H, 1965. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Berlin, Das Tierreich, 80, Walterv de Gruyter, XXII + 246 pp." title="Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Berlin, Das Tierreich, 80, Walterv de Gruyter, XXII + 246 pp." year="1965">Wermuth (1965)</bibRefCitation>
|
||
wrote
|
||
<normalizedToken originalValue="‘Polynesia’">'Polynesia'</normalizedToken>
|
||
as the type locality with the addition (fide
|
||
<bibRefCitation author="Smith, MA" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B123" refString="Smith, MA, 1935. The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor & Francis Ltd." title="The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor & Francis Ltd." year="1935">Smith 1935</bibRefCitation>
|
||
), that the species probably occurs in the Indo-Australian Archipelago. According to
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="586 - 600" refId="B59" refString="Guenther, A, 1872. On the reptiles and amphibians of Borneo. Proceedings of the Zoological Society of London 1872: 586 - 600" title="On the reptiles and amphibians of Borneo." volume="1872" year="1872">
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
(1872)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
|
||
</taxonomicName>
|
||
occurs in the north of Borneo (Labuan, now Sabah) and
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
in the South (Banjermassin, Martapoura). We herein restrict the type locality of
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
to Banjermassin (= Banjar-masin), Kalimantan, Indonesia. While commenting on
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="586 - 600" refId="B59" refString="Guenther, A, 1872. On the reptiles and amphibians of Borneo. Proceedings of the Zoological Society of London 1872: 586 - 600" title="On the reptiles and amphibians of Borneo." volume="1872" year="1872">
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
(1872
|
||
</bibRefCitation>
|
||
:589) stated that he received three specimens from Dr. Bleeker from Borneo under different names (
|
||
<taxonomicName class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus pentonopus" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="pentonopus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Platydactylus pentonopus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus borneensis" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Platydactylus borneensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus zosterophorus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="zosterophorus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Hemidactylus zosterophorus</emphasis>
|
||
</taxonomicName>
|
||
). From
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="586 - 600" refId="B59" refString="Guenther, A, 1872. On the reptiles and amphibians of Borneo. Proceedings of the Zoological Society of London 1872: 586 - 600" title="On the reptiles and amphibians of Borneo." volume="1872" year="1872">
|
||
<normalizedToken originalValue="Günther’s">Guenther's</normalizedToken>
|
||
(1872)
|
||
</bibRefCitation>
|
||
footnotes it is obvious that he allocated all of
|
||
<normalizedToken originalValue="Bleeker’s">Bleeker's</normalizedToken>
|
||
specimens to
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
. Of
|
||
<normalizedToken originalValue="Bleeker’s">Bleeker's</normalizedToken>
|
||
species, only
|
||
<taxonomicName class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus borneensis" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Platydactylus borneensis</emphasis>
|
||
</taxonomicName>
|
||
had been recorded from Borneo. The names
|
||
<emphasis italics="true" pageId="0" pageNumber="47">
|
||
P.
|
||
<normalizedToken originalValue="borneënsis">borneensis</normalizedToken>
|
||
</emphasis>
|
||
(=
|
||
<taxonomicName genus="P." lsidName="P. borneensis" pageId="0" pageNumber="47" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. borneensis</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName genus="H." lsidName="H. zosterophorus" pageId="0" pageNumber="47" rank="species" species="zosterophorus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">H. zosterophorus</emphasis>
|
||
</taxonomicName>
|
||
were introduced by
|
||
<bibRefCitation author="Bleeker, P" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch Indie" pageId="0" pageNumber="47" pagination="470 - 475" refId="B8" refString="Bleeker, P, 1857. Berigt omtrent eenige Reptilien van Sumatra, Borneo, Batjan en Boero. Natuurkundig Tijdschrift voor Nederlandsch Indie 13: 470 - 475" title="Berigt omtrent eenige Reptilien van Sumatra, Borneo, Batjan en Boero." volume="13" year="1857">Bleeker (1857)</bibRefCitation>
|
||
without formal species descriptions (see also
|
||
<bibRefCitation author="Bleeker, P" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch Indie" pageId="0" pageNumber="47" pagination="284 - 298" refId="B9" refString="Bleeker, P, 1860. Over de Reptilien-Fauna van Sumatra. Natuurkundig Tijdschrift voor Nederlandsch Indie 21: 284 - 298" title="Over de Reptilien-Fauna van Sumatra." volume="21" year="1860">Bleeker 1860</bibRefCitation>
|
||
). The type locality of
|
||
<taxonomicName class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus borneensis" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Platydactylus borneensis</emphasis>
|
||
</taxonomicName>
|
||
(non
|
||
<taxonomicName authorityName="Gunther" authorityYear="1864" class="Reptilia" family="Gekkonidae" genus="Pentadactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pentadactylus borneensis" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Pentadactylus borneensis</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
, 1864 =
|
||
<taxonomicName baseAuthorityName="Gunther" baseAuthorityYear="1864" class="Reptilia" family="Eublepharidae" genus="Aeluroscalabotes" kingdom="Animalia" lsidName="Aeluroscalabotes felinus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="felinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Aeluroscalabotes felinus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
, 1864; non
|
||
<taxonomicName authorityName="Gray" authorityYear="1845" class="Reptilia" family="Phyllodactylidae" genus="Tarentola" kingdom="Animalia" lsidName="Tarentola borneensis" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Tarentola borneensis</emphasis>
|
||
</taxonomicName>
|
||
Gray, 1845 =
|
||
<taxonomicName class="Reptilia" family="Phyllodactylidae" genus="Tarentola" kingdom="Animalia" lsidName="Tarentola delalandii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="delalandii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Tarentola delalandii</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Duméril">Dumeril</normalizedToken>
|
||
& Bibron, 1836) is
|
||
<normalizedToken originalValue="‘Bandjermasin’">'Bandjermasin'</normalizedToken>
|
||
and the type locality of
|
||
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus zosterophorus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="zosterophorus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Hemidactylus zosterophorus</emphasis>
|
||
</taxonomicName>
|
||
is 'Padang (ook op Nias)'. In his critical review of
|
||
<normalizedToken originalValue="Bleeker’s">Bleeker's</normalizedToken>
|
||
type specimens,
|
||
<bibRefCitation author="Boulenger, GA" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch Indie" pageId="0" pageNumber="47" refId="B12" refString="Boulenger, GA, 1887. Note on some reptiles from Sumatra described by Bleeker in 1860. Annals and Magazine of Natural History 20: 152." title="Note on some reptiles from Sumatra described by Bleeker in 1860. Annals and Magazine of Natural History 20: 152." year="1887">Boulenger (1887)</bibRefCitation>
|
||
did not consider the taxa
|
||
<taxonomicName class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus borneensis" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Platydactylus borneensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus zosterophorus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="zosterophorus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Hemidactylus zosterophorus</emphasis>
|
||
</taxonomicName>
|
||
. According to
|
||
<bibRefCitation author="Bauer, AM" journalOrPublisher="Evolution" pageId="0" pageNumber="47" refId="B4" refString="Bauer, AM, 1994. Gekkonidae (Reptilia, Sauria), Part I Australia and Oceania. Das Tierreich 109 (Part). Walter de Gruyter, Berlin, XII + 306 pp." title="Gekkonidae (Reptilia, Sauria), Part I Australia and Oceania. Das Tierreich 109 (Part). Walter de Gruyter, Berlin, XII + 306 pp." year="1994">Bauer (1994)</bibRefCitation>
|
||
, both names are species inquirenda and
|
||
<bibRefCitation author="Kluge, AG" journalOrPublisher="Hamadryad" pageId="0" pageNumber="47" pagination="1 - 209" refId="B72" refString="Kluge, AG, 2001. Gekkotan lizard taxonomy. Hamadryad 26: 1 - 209" title="Gekkotan lizard taxonomy." volume="26" year="2001">Kluge (2001)</bibRefCitation>
|
||
listed them as nomina nuda (see also
|
||
<bibRefCitation author="Roesler, H" journalOrPublisher="Gekkota" pageId="0" pageNumber="47" pagination="28 - 153" refId="B115" refString="Roesler, H, 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Gecko -Taxa (Reptilia: Gekkonomorpha). Gekkota 2: 28 - 153" title="Kommentierte Liste der rezent, subrezent und fossil bekannten Gecko - Taxa (Reptilia: Gekkonomorpha)." volume="2" year="2000">
|
||
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
||
2000
|
||
</bibRefCitation>
|
||
). Therefore, the names
|
||
<taxonomicName class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus borneensis" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Platydactylus borneensis</emphasis>
|
||
</taxonomicName>
|
||
Bleeker, 1857 and
|
||
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus zosterophorus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="zosterophorus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Hemidactylus zosterophorus</emphasis>
|
||
</taxonomicName>
|
||
Bleeker, 1857, which are according to
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="586 - 600" refId="B59" refString="Guenther, A, 1872. On the reptiles and amphibians of Borneo. Proceedings of the Zoological Society of London 1872: 586 - 600" title="On the reptiles and amphibians of Borneo." volume="1872" year="1872">
|
||
<normalizedToken originalValue="Günther’s">Guenther's</normalizedToken>
|
||
(1872)
|
||
</bibRefCitation>
|
||
statements younger, subjective synonyms of
|
||
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Gekko albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
, 1867, are not available. The same concerns
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="586 - 600" refId="B59" refString="Guenther, A, 1872. On the reptiles and amphibians of Borneo. Proceedings of the Zoological Society of London 1872: 586 - 600" title="On the reptiles and amphibians of Borneo." volume="1872" year="1872">
|
||
<normalizedToken originalValue="Günther’s">Guenther's</normalizedToken>
|
||
(1872)
|
||
</bibRefCitation>
|
||
name
|
||
<taxonomicName class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus pentonopus" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="pentonopus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Platydactylus pentonopus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Boulenger, GA" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch Indie" pageId="0" pageNumber="47" refId="B11" refString="Boulenger, GA, 1885. Catalogue of the Lizards in the British Museum, vol. 2 Trustees of the British Museum, London." title="Catalogue of the Lizards in the British Museum, vol. 2 Trustees of the British Museum, London." year="1885">Boulenger (1885)</bibRefCitation>
|
||
synonymized
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation author="Wermuth, H" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="47" refId="B133" refString="Wermuth, H, 1965. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Berlin, Das Tierreich, 80, Walterv de Gruyter, XXII + 246 pp." title="Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Berlin, Das Tierreich, 80, Walterv de Gruyter, XXII + 246 pp." year="1965">Wermuth 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Kluge, AG" journalOrPublisher="Smithsonian Herpetological Information Service" pageId="0" pageNumber="47" pagination="1 - 35" refId="B70" refString="Kluge, AG, 1991. Checklist of gekkonoid lizards. Smithsonian Herpetological Information Service 85: 1 - 35" title="Checklist of gekkonoid lizards." volume="85" year="1991">Kluge 1991</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Kluge, AG" journalOrPublisher="International Gecko Society, San Diego" pageId="0" pageNumber="47" refId="B71" refString="Kluge, AG, 1993. Gekkonoid Lizard Taxonomy. International Gecko Society, San Diego" title="Gekkonoid Lizard Taxonomy." year="1993">1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Bauer, AM" journalOrPublisher="Evolution" pageId="0" pageNumber="47" refId="B4" refString="Bauer, AM, 1994. Gekkonidae (Reptilia, Sauria), Part I Australia and Oceania. Das Tierreich 109 (Part). Walter de Gruyter, Berlin, XII + 306 pp." title="Gekkonidae (Reptilia, Sauria), Part I Australia and Oceania. Das Tierreich 109 (Part). Walter de Gruyter, Berlin, XII + 306 pp." year="1994">Bauer 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Roesler, H" journalOrPublisher="Gekkota" pageId="0" pageNumber="47" pagination="28 - 153" refId="B115" refString="Roesler, H, 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Gecko -Taxa (Reptilia: Gekkonomorpha). Gekkota 2: 28 - 153" title="Kommentierte Liste der rezent, subrezent und fossil bekannten Gecko - Taxa (Reptilia: Gekkonomorpha)." volume="2" year="2000">
|
||
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
||
2000
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation author="de Rooij, N" journalOrPublisher="Systematic Biology" pageId="0" pageNumber="47" refId="B29" refString="de Rooij, N, 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden, E. J. Brill Ltd., xiv + 384 pp." title="The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden, E. J. Brill Ltd., xiv + 384 pp." year="1915">De Rooij (1915)</bibRefCitation>
|
||
obviously also followed
|
||
<bibRefCitation author="Boulenger, GA" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch Indie" pageId="0" pageNumber="47" refId="B11" refString="Boulenger, GA, 1885. Catalogue of the Lizards in the British Museum, vol. 2 Trustees of the British Museum, London." title="Catalogue of the Lizards in the British Museum, vol. 2 Trustees of the British Museum, London." year="1885">Boulenger (1885)</bibRefCitation>
|
||
, because she listed the distribution of
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
based on
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="586 - 600" refId="B59" refString="Guenther, A, 1872. On the reptiles and amphibians of Borneo. Proceedings of the Zoological Society of London 1872: 586 - 600" title="On the reptiles and amphibians of Borneo." volume="1872" year="1872">
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
(1872)
|
||
</bibRefCitation>
|
||
under the species
|
||
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
|
||
</taxonomicName>
|
||
. Recently,
|
||
<bibRefCitation author="Kluge, AG" journalOrPublisher="Hamadryad" pageId="0" pageNumber="47" pagination="1 - 209" refId="B72" refString="Kluge, AG, 2001. Gekkotan lizard taxonomy. Hamadryad 26: 1 - 209" title="Gekkotan lizard taxonomy." volume="26" year="2001">Kluge (2001)</bibRefCitation>
|
||
revalidated
|
||
<taxonomicName genus="G." lsidName="G. albo-fasciolatus" pageId="0" pageNumber="47" rank="species" species="albo-fasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albo-fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
at the specific rank, but
|
||
<bibRefCitation author="Malkmus, R" journalOrPublisher="Hamadryad" pageId="0" pageNumber="47" refId="B86" refString="Malkmus, R, Manthey, U, Vogel, G, Hoffman, P, Kosuch, J, 2002. Amphibians and Reptiles of Mount Kinabalu (North Borneo). Ganter Verlag, Ruggell, Liechtenstein." title="Amphibians and Reptiles of Mount Kinabalu (North Borneo). Ganter Verlag, Ruggell, Liechtenstein." year="2002">Malkmus et al. (2002)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Das, I" journalOrPublisher="Philosophical Transactions B of The Royal Society" pageId="0" pageNumber="47" refId="B24" refString="Das, I, 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, vi + 83 pp." title="Lizards of Borneo. Natural History Publications, Kota Kinabalu, vi + 83 pp." year="2004">Das (2004)</bibRefCitation>
|
||
did not consider
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
as part of the Bornean herpetofauna."
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Sauria" pageId="0" pageNumber="47" refId="B58" refString="Guenther, A, 1867. Additions to the knowledge of Australian reptiles and fishes. Annals and Magazine of Natural History Series 3, 320: 45-68." title="Additions to the knowledge of Australian reptiles and fishes. Annals and Magazine of Natural History Series 3, 320: 45 - 68." year="1867">
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
(1867)
|
||
</bibRefCitation>
|
||
noted that
|
||
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Gekko albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
had a reddish-olive dorsum marbled with grayish and a uniformly whitish venter. Specimens with a reddish-olive dorsum with gray marbling also occur in Sabah (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution and color pattern variation in Gekko albofasciolatus and G. cf. albofasciolatus in Borneo A. LSUDPC 12916 from Gunung Mulu, Sarawak, East Malaysia. B. LSUDPC 12918 from Gunung Mulu, Sarawak, East Malaysia. C. LSUDPC 12919) from Kelay Subdistrict, Kalimantan, Indonesia. D. LSUDPC 1293 from Kalimantan, Indonesia. E. LSUDPC 12949 from Kinabatangan River, Sabah, East Malaysia. F. LSUDPC 12924 from Lambir, Sarawak, East Malaysia. G. LSUDPC 12925 from Lambir, Sarawak, East Malaysia. H. LSUDPC 12929 from Gunung Gading, Sarawak, East Malaysia. Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature. Photographs by Alan Watson www. alanwatsonfeatherstone. com (A), Wolfgang Grossmann (B, H), C. J. Franklin (C), Mistar Kamsi (D), Chien C. Lee (E), and Nick Baker (F, G)." figureDoi="10.3897/vz.72.e77702.figure9" httpUri="https://binary.pensoft.net/fig/641485" pageId="0" pageNumber="47">9E</figureCitation>
|
||
) and preserved material examined from western and eastern Borneo all have uniform whitish venters, although in life, some have faint-yellow marbling which presumably fades after preservation.
|
||
<bibRefCitation author="Guenther, A" journalOrPublisher="Sauria" pageId="0" pageNumber="47" refId="B58" refString="Guenther, A, 1867. Additions to the knowledge of Australian reptiles and fishes. Annals and Magazine of Natural History Series 3, 320: 45-68." title="Additions to the knowledge of Australian reptiles and fishes. Annals and Magazine of Natural History Series 3, 320: 45 - 68." year="1867">
|
||
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
|
||
(1867)
|
||
</bibRefCitation>
|
||
also noted his specimen had "a narrow horseshoe-shaped band across the neck, the convexity being directed backwards". This is clearly in reference to the nuchal band which was not present in specimens we examined or in photographs we acquired (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution and color pattern variation in Gekko albofasciolatus and G. cf. albofasciolatus in Borneo A. LSUDPC 12916 from Gunung Mulu, Sarawak, East Malaysia. B. LSUDPC 12918 from Gunung Mulu, Sarawak, East Malaysia. C. LSUDPC 12919) from Kelay Subdistrict, Kalimantan, Indonesia. D. LSUDPC 1293 from Kalimantan, Indonesia. E. LSUDPC 12949 from Kinabatangan River, Sabah, East Malaysia. F. LSUDPC 12924 from Lambir, Sarawak, East Malaysia. G. LSUDPC 12925 from Lambir, Sarawak, East Malaysia. H. LSUDPC 12929 from Gunung Gading, Sarawak, East Malaysia. Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature. Photographs by Alan Watson www. alanwatsonfeatherstone. com (A), Wolfgang Grossmann (B, H), C. J. Franklin (C), Mistar Kamsi (D), Chien C. Lee (E), and Nick Baker (F, G)." figureDoi="10.3897/vz.72.e77702.figure9" httpUri="https://binary.pensoft.net/fig/641485" pageId="0" pageNumber="47">9</figureCitation>
|
||
). Contra to
|
||
<bibRefCitation author="Das, I" journalOrPublisher="Philosophical Transactions B of The Royal Society" pageId="0" pageNumber="47" refId="B24" refString="Das, I, 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, vi + 83 pp." title="Lizards of Borneo. Natural History Publications, Kota Kinabalu, vi + 83 pp." year="2004">Das (2004</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Das, I" journalOrPublisher="Philosophical Transactions B of The Royal Society" pageId="0" pageNumber="47" refId="B25" refString="Das, I, 2007. Amphibians and Reptiles of Brunei. Natural History Publications, Kota Kinabalu, viii + 200 pp" title="Amphibians and Reptiles of Brunei. Natural History Publications, Kota Kinabalu, viii + 200 pp" year="2007">2007</bibRefCitation>
|
||
), all Bornean specimens we examined had the characteristic dark Y-shaped marking on the head common to
|
||
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
|
||
</taxonomicName>
|
||
and geckos of the eastern peninsular lineage (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution and color pattern variation in Gekko smithii and G. hulk sp. nov. on the Thai-Malay Peninsula. A. Gekko smithii (LSUHC 15052) from Perlis State Park, Perlis, Peninsular Malaysia. B. Gekko smithii (LSUDPC 12884) from Puncak Janing, Kedah, Peninsualr Malaysia. C. Gekko smithii (LSUHC 9626) from Sedim, Kedah, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12869) from Penang Hill (type locality), Penang, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12886) from Lata Kedondong, Selangor, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12887) from Kota Damansara, Selangor, Peninsular Malaysia. G. Gekko smithii (LSUDPC 11174) from Khaopu Khaoya, Nakhon Si Thamarat Province, Thailand. H. Gekko smithii (LSUDPC 11173) from Bacho Falls, Narathiwat Province, Thailand. I. Gekko smithii (LSUDPC 12878) from the Weang District, Narathiwat Province, Thailand. J. Gekko hulk sp. nov. (LSUDPC 12880) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 8696) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUDPC 7991) from Gunung Tebu, Terengganu, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12897) from Merapoh, Pahang, Peninsular Malaysia. N. Gekko hulk sp. nov. (LSUDPC 12897) from Pulau Tioman, Pahang, Peninsular Malaysia. Photographs by L. Lee Grismer (A, K, L, N), Evan S. H. Quah (B, C, D), Kurt H. P. Guek (E), Steven Wong (F), Henrick Bringsoe (G, H); Parinya Pawangkhanant (I), Ian Dugdale (J), and Nick Baker (M). Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature." figureDoi="10.3897/vz.72.e77702.figure7" httpUri="https://binary.pensoft.net/fig/641483" pageId="0" pageNumber="47">7</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution and color pattern variation in Gekko albofasciolatus and G. cf. albofasciolatus in Borneo A. LSUDPC 12916 from Gunung Mulu, Sarawak, East Malaysia. B. LSUDPC 12918 from Gunung Mulu, Sarawak, East Malaysia. C. LSUDPC 12919) from Kelay Subdistrict, Kalimantan, Indonesia. D. LSUDPC 1293 from Kalimantan, Indonesia. E. LSUDPC 12949 from Kinabatangan River, Sabah, East Malaysia. F. LSUDPC 12924 from Lambir, Sarawak, East Malaysia. G. LSUDPC 12925 from Lambir, Sarawak, East Malaysia. H. LSUDPC 12929 from Gunung Gading, Sarawak, East Malaysia. Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature. Photographs by Alan Watson www. alanwatsonfeatherstone. com (A), Wolfgang Grossmann (B, H), C. J. Franklin (C), Mistar Kamsi (D), Chien C. Lee (E), and Nick Baker (F, G)." figureDoi="10.3897/vz.72.e77702.figure9" httpUri="https://binary.pensoft.net/fig/641485" pageId="0" pageNumber="47">9</figureCitation>
|
||
).
|
||
<bibRefCitation author="Grossmann, W" journalOrPublisher="Natur und Tier Verlag GmbH, Muenster" pageId="0" pageNumber="47" refId="B53" refString="Grossmann, W, 2006. Gruenaugengeckos Gekko smithii & Gekko siamensis. Natur und Tier Verlag GmbH, Muenster" title="Gruenaugengeckos Gekko smithii & Gekko siamensis." year="2006">Grossmann (2006)</bibRefCitation>
|
||
illustrated a specimen from eastern Kalimantan (i.e. eastern Borneo) with a brown vertebral stripe and small white dorsal flecks. All adult Bornean specimens examined herein, have small white flecks and variable degrees of dorsal striping and ground color (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution and color pattern variation in Gekko albofasciolatus and G. cf. albofasciolatus in Borneo A. LSUDPC 12916 from Gunung Mulu, Sarawak, East Malaysia. B. LSUDPC 12918 from Gunung Mulu, Sarawak, East Malaysia. C. LSUDPC 12919) from Kelay Subdistrict, Kalimantan, Indonesia. D. LSUDPC 1293 from Kalimantan, Indonesia. E. LSUDPC 12949 from Kinabatangan River, Sabah, East Malaysia. F. LSUDPC 12924 from Lambir, Sarawak, East Malaysia. G. LSUDPC 12925 from Lambir, Sarawak, East Malaysia. H. LSUDPC 12929 from Gunung Gading, Sarawak, East Malaysia. Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature. Photographs by Alan Watson www. alanwatsonfeatherstone. com (A), Wolfgang Grossmann (B, H), C. J. Franklin (C), Mistar Kamsi (D), Chien C. Lee (E), and Nick Baker (F, G)." figureDoi="10.3897/vz.72.e77702.figure9" httpUri="https://binary.pensoft.net/fig/641485" pageId="0" pageNumber="47">9</figureCitation>
|
||
). Therefore, dorsal color pattern is not a reliable character separating populations from Borneo west of the Iran Mountains and populations east of the Iran Mountains.
|
||
<bibRefCitation author="Roesler, H" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="1 - 50" refId="B117" refString="Roesler, H, Bauer, AM, Heinicke, MP, Greenbaum, E, Jackman, T, Nguyen, TQ, Ziegler, T, 2011. Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae). Zootaxa 2989: 1 - 50" title="Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae)." volume="2989" year="2011">
|
||
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
||
et al. (2011)
|
||
</bibRefCitation>
|
||
noted that northern Bornean populations might be different on the basis of
|
||
<normalizedToken originalValue="Günther’s">Guenther's</normalizedToken>
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
(eastern Borneo) having 26 rows of ventral scales versus 29-39 in
|
||
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
|
||
</taxonomicName>
|
||
s.l. However, specimens examined from northern Borneo (N = 7) have 23-33 scale rows and a specimen examined from eastern Kalimantan at Kelay (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution and color pattern variation in Gekko albofasciolatus and G. cf. albofasciolatus in Borneo A. LSUDPC 12916 from Gunung Mulu, Sarawak, East Malaysia. B. LSUDPC 12918 from Gunung Mulu, Sarawak, East Malaysia. C. LSUDPC 12919) from Kelay Subdistrict, Kalimantan, Indonesia. D. LSUDPC 1293 from Kalimantan, Indonesia. E. LSUDPC 12949 from Kinabatangan River, Sabah, East Malaysia. F. LSUDPC 12924 from Lambir, Sarawak, East Malaysia. G. LSUDPC 12925 from Lambir, Sarawak, East Malaysia. H. LSUDPC 12929 from Gunung Gading, Sarawak, East Malaysia. Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature. Photographs by Alan Watson www. alanwatsonfeatherstone. com (A), Wolfgang Grossmann (B, H), C. J. Franklin (C), Mistar Kamsi (D), Chien C. Lee (E), and Nick Baker (F, G)." figureDoi="10.3897/vz.72.e77702.figure9" httpUri="https://binary.pensoft.net/fig/641485" pageId="0" pageNumber="47">9C</figureCitation>
|
||
) has 27 scale rows. Therefore, at this point, other than the spotted nuchal band, there are no morphological or color pattern differences between
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
populations east of the Iran Mountains and populations west of the Iran Mountains (i.e. from the Malaysian states of Sarawak and Sabah and the Sultanate of Brunei). The molecular data clearly indicate that the Bornean samples from Sarawak are not conspecific with
|
||
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
|
||
</taxonomicName>
|
||
s.s. or any other lineage (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Maximum likelihood topology of the Gekko smithii complex based on 1267 base pairs of ND 2 and its flanking tRNAs with BPP and UFB values, respectively, at the nodes." figureDoi="10.3897/vz.72.e77702.figure2" httpUri="https://binary.pensoft.net/fig/641478" pageId="0" pageNumber="47">2</figureCitation>
|
||
). Therefore, the name
|
||
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Gekko albofasciolatus</emphasis>
|
||
</taxonomicName>
|
||
, is available for Bornean populations. However, owing to an absence of molecular data from eastern populations, all populations west of the Iran Mountains are referred to as
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">G. cf. albofasciolatus</taxonomicName>
|
||
pending the outcome of a molecular analysis. We believe this approach is justified based on the fact that the Banjaran Titiwangsa of Peninsular Malaysia separate
|
||
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
|
||
</taxonomicName>
|
||
and the east peninsular lineage and the Iran Mountains may be doing the same with populations from Borneo. Investigations on Bornean populations are currently underway (Grismer et al. unpubl.)
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.72.e77702.figure9" httpUri="https://binary.pensoft.net/fig/641485" pageId="0" pageNumber="47" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 9.</emphasis>
|
||
Distribution and color pattern variation in
|
||
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">Gekko albofasciolatus</taxonomicName>
|
||
and
|
||
<taxonomicName genus="G." lsidName="G. albofasciolatus" pageId="0" pageNumber="47" rank="species" species="albofasciolatus">G. cf. albofasciolatus</taxonomicName>
|
||
in Borneo A. LSUDPC 12916 from Gunung Mulu, Sarawak, East Malaysia. B. LSUDPC 12918 from Gunung Mulu, Sarawak, East Malaysia. C. LSUDPC 12919) from Kelay Subdistrict, Kalimantan, Indonesia. D. LSUDPC 1293 from Kalimantan, Indonesia. E. LSUDPC 12949 from Kinabatangan River, Sabah, East Malaysia. F. LSUDPC 12924 from Lambir, Sarawak, East Malaysia. G. LSUDPC 12925 from Lambir, Sarawak, East Malaysia. H. LSUDPC 12929 from Gunung Gading, Sarawak, East Malaysia. Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature. Photographs by Alan Watson www.alanwatsonfeatherstone.com (A), Wolfgang Grossmann (B, H), C. J. Franklin (C), Mistar Kamsi (D), Chien C. Lee (E), and Nick Baker (F,G).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |