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<document id="ED76C6890A5655530A63E13BB42C40AC" ID-CLB-Dataset="25414" ID-DOI="10.11646/zootaxa.4674.2.1" ID-GBIF-Dataset="a3c9f2fa-30d7-48b9-9a2e-3f81cc0a3e3e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3458836" ID-ZooBank="B6626FE9-6490-4BC9-BACB-A4437AACDF25" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569309377345" checkinUser="plazi" docAuthor="Bollino, Maurizio, Uliana, Marco &amp; Sabatinelli, Guido" docDate="2019" docId="03F087D8C15C8424FF0CF93DFCF18E94" docLanguage="en" docName="zootaxa.4674.2.1.pdf.imf" docOrigin="Zootaxa 4674 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pygopleurus pseudopsilotrichius" docType="treatment" docVersion="9" lastPageNumber="166" masterDocId="FFC9FFA0C150842BFF9BFFD0FF8D8D17" masterDocTitle="The Pygopleurus Motschulsky (Coleoptera: Scarabaeoidea: Glaphyridae) of mainland Greece: taxonomy, faunistics and ecological notes" masterLastPageNumber="202" masterPageNumber="151" pageNumber="163" updateTime="1698756305895" updateUser="ExternalLinkService">
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<mods:title id="9CA02F3998D70BF8E312E4DC04DE43B2">The Pygopleurus Motschulsky (Coleoptera: Scarabaeoidea: Glaphyridae) of mainland Greece: taxonomy, faunistics and ecological notes</mods:title>
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<mods:namePart id="3BC8C6EFD3DC120A8ABEEA3C6C059374">Bollino, Maurizio</mods:namePart>
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<mods:namePart id="98DC685E50C0C47CC1028DCDC2FF8431">Uliana, Marco</mods:namePart>
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<mods:date id="A11DDC1C36D7DA5BA61BE94955686C6E">2019</mods:date>
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<subSubSection id="C3436545C15C8427FF0CF93DFBCD8A10" box="[151,1088,1773,1800]" pageId="12" pageNumber="163" type="nomenclature">
<paragraph id="8BE636CEC15C8427FF0CF93DFBCD8A10" blockId="12.[151,1436,1773,1944]" box="[151,1088,1773,1800]" pageId="12" pageNumber="163">
<heading id="D0AE81A2C15C8427FF0CF93DFBCD8A10" bold="true" box="[151,1088,1773,1800]" fontSize="11" level="1" pageId="12" pageNumber="163" reason="1">
<emphasis id="B92DEADCC15C8427FF0CF93DFBCD8A10" bold="true" box="[151,1088,1773,1800]" pageId="12" pageNumber="163">
<taxonomicName id="4C594D4DC15C8427FF0CF93DFD7B8A10" ID-CoL="4QPFL" authority="(Petrovitz, 1958)" baseAuthorityName="Petrovitz" baseAuthorityYear="1958" box="[151,758,1773,1800]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudopsilotrichius">
<emphasis id="B92DEADCC15C8427FF0CF93DFDAC8A1F" bold="true" box="[151,545,1773,1800]" italics="true" pageId="12" pageNumber="163">Pygopleurus pseudopsilotrichius</emphasis>
(
<bibRefCitation id="EFC84B3FC15C8427FDABF93DFD638A10" author="Petrovitz, R." box="[560,750,1773,1799]" pageId="12" pageNumber="163" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz, 1958</bibRefCitation>
)
</taxonomicName>
valid species (
<figureCitation id="13622A4BC15C8427FC32F93DFBB58A10" box="[937,1080,1773,1799]" captionStart-0="FIGURES 25–31" captionStart-1="FIGURE 32" captionStartId-0="13.[151,264,1869,1894]" captionStartId-1="14.[151,250,1266,1291]" captionTargetBox-0="[163,1408,194,1804]" captionTargetBox-1="[170,1408,181,1235]" captionTargetId-0="figure@13.[151,1436,181,1844]" captionTargetId-1="figure@14.[170,1416,181,1241]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 25–31. Pygopleurus pseudopsilotrichius (Petrovitz, 1958), habitus and genitalia. 25, male (Lakonia, Gorani); 26, male (Arcadia, Karitena); 27, female (Lakonia, Chanià); 28, female (Lakonia, Gorani); 29, left paramere in lateral and dorsal view (Laconia, Chanià); 30, everted endophallus in left lateral view (Laconia, Chanià); 31, left genital sclerites of female (Laconia, Dafni)." captionText-1="FIGURE 32. Pygopleurus anemoninus (Brullé, 1832) and P. pseudopsilotrichius (Petrovitz, 1958): geographical, altitudinal, and temporal distribution of records. Red curves showing the expected distribution if the species was uniformly present at all heights and all times (see Material and methods)." figureDoi-0="http://doi.org/10.5281/zenodo.3458852" figureDoi-1="http://doi.org/10.5281/zenodo.3458854" httpUri-0="https://zenodo.org/record/3458852/files/figure.png" httpUri-1="https://zenodo.org/record/3458854/files/figure.png" pageId="12" pageNumber="163">Figs. 25–32</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3436545C15C8427FF5CF8C1FF5A8A8F" pageId="12" pageNumber="163" type="reference_group">
<paragraph id="8BE636CEC15C8427FF5CF8C1FC3D8A3B" blockId="12.[151,1436,1773,1944]" box="[199,944,1809,1836]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB4911E0C15C8427FF5CF8C1FC3D8A3B" box="[199,944,1809,1836]" pageId="12" pageNumber="163">
<taxonomicName id="4C594D4DC15C8427FF5CF8C1FC268A3B" ID-CoL="5DGT7" authority="Petrovitz, 1958: 50" authorityName="Petrovitz" authorityPageNumber="50" authorityYear="1958" box="[199,939,1809,1836]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudopsilotrichia" subGenus="Pygopleurus">
<emphasis id="B92DEADCC15C8427FF5CF8C1FEC08A3C" box="[199,333,1809,1835]" italics="true" pageId="12" pageNumber="163">Amphicoma</emphasis>
(
<emphasis id="B92DEADCC15C8427FEC6F8C1FE668A3C" box="[349,491,1809,1835]" italics="true" pageId="12" pageNumber="163">Pygopleurus</emphasis>
)
<emphasis id="B92DEADCC15C8427FE60F8C1FD468A3C" box="[507,715,1809,1835]" italics="true" pageId="12" pageNumber="163">pseudopsilotrichia</emphasis>
<treatmentCitation id="0AF810DFC15C8427FD48F8C1FC268A3B" author="Petrovitz, R." box="[723,939,1809,1836]" page="50" pageId="12" pageNumber="163" year="1958">
<bibRefCitation id="EFC84B3FC15C8427FD48F8C1FC268A3B" author="Petrovitz, R." box="[723,939,1809,1836]" pageId="12" pageNumber="163" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz, 1958: 50</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE636CEC15C8427FF5CF8E6FA058A47" blockId="12.[151,1436,1773,1944]" box="[199,1416,1845,1872]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB4911E0C15C8427FF5CF8E6FA058A47" box="[199,1416,1845,1872]" pageId="12" pageNumber="163">
=
<taxonomicName id="4C594D4DC15C8427FF45F8E5FD6F8A58" ID-CoL="5DGT7" authorityName="Petrovitz" authorityYear="1958" box="[222,738,1845,1872]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudopsilotrichia" subGenus="Pygopleurus">
<emphasis id="B92DEADCC15C8427FF45F8E5FEE98A58" box="[222,356,1845,1871]" italics="true" pageId="12" pageNumber="163">Amphicoma</emphasis>
(
<emphasis id="B92DEADCC15C8427FEEFF8E5FD8F8A58" box="[372,514,1845,1871]" italics="true" pageId="12" pageNumber="163">Pygopleurus</emphasis>
)
<emphasis id="B92DEADCC15C8427FD89F8E5FD6F8A58" box="[530,738,1845,1871]" italics="true" pageId="12" pageNumber="163">pseudopsilotrichia</emphasis>
</taxonomicName>
f.
<emphasis id="B92DEADCC15C8427FC9AF8E5FCC48A58" box="[769,841,1845,1871]" italics="true" pageId="12" pageNumber="163">viridis</emphasis>
<treatmentCitation id="0AF810DFC15C8427FCD4F8E5FBA88A47" author="Petrovitz, R." box="[847,1061,1845,1872]" page="50" pageId="12" pageNumber="163" year="1958">
<bibRefCitation id="EFC84B3FC15C8427FCD4F8E5FBA88A47" author="Petrovitz, R." box="[847,1061,1845,1872]" pageId="12" pageNumber="163" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz, 1958: 50</bibRefCitation>
</treatmentCitation>
[infrasubspecific, unavailable].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE636CEC15C8427FF5CF88AFF5A8A8F" blockId="12.[151,1436,1773,1944]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB4911E0C15C8427FF5CF88AFF5A8A8F" pageId="12" pageNumber="163">
=
<taxonomicName id="4C594D4DC15C8427FF7AF889FD668A64" ID-CoL="5DGT7" authorityName="Petrovitz" authorityYear="1958" box="[225,747,1881,1908]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudopsilotrichia" subGenus="Pygopleurus">
<emphasis id="B92DEADCC15C8427FF7AF889FEEA8A64" box="[225,359,1881,1907]" italics="true" pageId="12" pageNumber="163">Amphicoma</emphasis>
(
<emphasis id="B92DEADCC15C8427FEE1F889FD858A64" box="[378,520,1881,1907]" italics="true" pageId="12" pageNumber="163">Pygopleurus</emphasis>
)
<emphasis id="B92DEADCC15C8427FD80F889FD668A64" box="[539,747,1881,1907]" italics="true" pageId="12" pageNumber="163">pseudopsilotrichia</emphasis>
</taxonomicName>
f.
<emphasis id="B92DEADCC15C8427FC96F889FC0E8A64" box="[781,899,1881,1907]" italics="true" pageId="12" pageNumber="163">albopilosa</emphasis>
<treatmentCitation id="0AF810DFC15C8427FC16F889FBE48A63" author="Petrovitz, R." box="[909,1129,1881,1908]" page="51" pageId="12" pageNumber="163" year="1958">
<bibRefCitation id="EFC84B3FC15C8427FC16F889FBE48A63" author="Petrovitz, R." box="[909,1129,1881,1908]" pageId="12" pageNumber="163" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz, 1958: 51</bibRefCitation>
</treatmentCitation>
[infrasubspecific, unavailable].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3436545C15C8425FF0CF816FC6B8AFD" lastPageId="14" lastPageNumber="165" pageId="12" pageNumber="163" type="materials_examined">
<paragraph id="8BE636CEC15C8427FF0CF816FD9C8AC8" blockId="12.[151,529,1989,2016]" box="[151,529,1989,2016]" pageId="12" pageNumber="163">
<emphasis id="B92DEADCC15C8427FF0CF816FEAE8AF7" bold="true" box="[151,291,1989,2016]" pageId="12" pageNumber="163">Type series.</emphasis>
<materialsCitation id="3B313C93C15C8427FEB1F815FD808AC8" ID-GBIF-Occurrence="2417904779" box="[298,525,1989,2015]" collectionCode="MHNG" pageId="12" pageNumber="163" specimenCount="1" typeStatus="syntype">
<typeStatus id="54E2886CC15C8427FEB1F815FE1C8AC8" box="[298,401,1989,2015]" pageId="12" pageNumber="163" type="syntype">Syntypes</typeStatus>
in
<collectionCode id="ED48AE0BC15C8427FE2FF815FD808AC8" box="[436,525,1989,2015]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="163" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF266646C15D8426FF0CF89DFEA98AC5" ID-DOI="http://doi.org/10.5281/zenodo.3458852" ID-Zenodo-Dep="3458852" httpUri="https://zenodo.org/record/3458852/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,264,1869,1894]" targetBox="[163,1408,194,1804]" targetPageId="13">
<paragraph id="8BE636CEC15D8426FF0CF89DFEA98AC5" blockId="13.[151,1436,1869,2002]" pageId="13" pageNumber="164">
<emphasis id="B92DEADCC15D8426FF0CF89DFEDB8A71" bold="true" box="[151,342,1869,1894]" pageId="13" pageNumber="164">FIGURES 25–31.</emphasis>
<taxonomicName id="4C594D4DC15D8426FEC5F89EFCE98A71" authority="(Petrovitz, 1958)" authorityName="Petrovitz" authorityYear="1958" baseAuthorityName="Petrovitz" baseAuthorityYear="1958" box="[350,868,1869,1894]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pseudopsilotrichius">
<emphasis id="B92DEADCC15D8426FEC5F89EFD218A71" box="[350,684,1870,1894]" italics="true" pageId="13" pageNumber="164">Pygopleurus pseudopsilotrichius</emphasis>
(
<bibRefCitation id="EFC84B3FC15D8426FD20F89DFCD08A71" author="Petrovitz, R." box="[699,861,1869,1894]" pageId="13" pageNumber="164" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz, 1958</bibRefCitation>
)
</taxonomicName>
, habitus and genitalia. 25, male (Lakonia, Gorani); 26, male (Arcadia, Karitena); 27, female (Lakonia, Chanià); 28, female (Lakonia, Gorani); 29, left paramere in lateral and dorsal view (Laconia, Chanià); 30, everted endophallus in left lateral view (Laconia, Chanià); 31, left genital sclerites of female (Laconia, Dafni).
</paragraph>
</caption>
<caption id="DF266646C15E8425FF0CFB22FD078844" ID-DOI="http://doi.org/10.5281/zenodo.3458854" ID-Zenodo-Dep="3458854" httpUri="https://zenodo.org/record/3458854/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,250,1266,1291]" targetBox="[170,1408,181,1235]" targetPageId="14">
<paragraph id="8BE636CEC15E8425FF0CFB22FD078844" blockId="14.[151,1437,1266,1363]" pageId="14" pageNumber="165">
<emphasis id="B92DEADCC15E8425FF0CFB22FEAF881C" bold="true" box="[151,290,1266,1291]" pageId="14" pageNumber="165">FIGURE 32.</emphasis>
<taxonomicName id="4C594D4DC15E8425FEB2FB23FD48881C" authority="(Brulle, 1832)" authorityName="Brulle" authorityYear="1832" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[297,709,1266,1291]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="anemoninus">
<emphasis id="B92DEADCC15E8425FEB2FB23FDA6881C" box="[297,555,1267,1291]" italics="true" pageId="14" pageNumber="165">Pygopleurus anemoninus</emphasis>
(
<bibRefCitation id="EFC84B3FC15E8425FDA0FB22FD33881C" author="Brulle, M." box="[571,702,1266,1291]" pageId="14" pageNumber="165" pagination="165 - 187" refId="ref29938" refString="Brulle, M. (1832) Expedition scientifique de Moree. Section des Sciences physiques. Ie Partie: Zoologie 2 e section: Des Animaux articules. F. G. Levrault, Paris, 400 pp. [165 - 187]" type="book chapter" year="1832">Brullé, 1832</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C594D4DC15E8425FD62FB24FB1E881C" authority="(Petrovitz, 1958)" authorityName="Petrovitz" authorityYear="1958" baseAuthorityName="Petrovitz" baseAuthorityYear="1958" box="[761,1171,1266,1291]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="pseudopsilotrichius">
<emphasis id="B92DEADCC15E8425FD62FB24FC57881C" box="[761,986,1267,1291]" italics="true" pageId="14" pageNumber="165">P. pseudopsilotrichius</emphasis>
(
<bibRefCitation id="EFC84B3FC15E8425FC72FB22FB06881C" author="Petrovitz, R." box="[1001,1163,1266,1291]" pageId="14" pageNumber="165" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz, 1958</bibRefCitation>
)
</taxonomicName>
: geographical, altitudinal, and temporal distribution of records. Red curves showing the expected distribution if the species was uniformly present at all heights and all times (see Material and methods).
</paragraph>
</caption>
<paragraph id="8BE636CEC15E8425FF5CFAACFD978B31" blockId="14.[151,1437,1403,1575]" pageId="14" pageNumber="165">
Although
<bibRefCitation id="EFC84B3FC15E8425FEA3FAACFD948881" author="Petrovitz, R." box="[312,537,1403,1430]" pageId="14" pageNumber="165" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz (1958: 50)</bibRefCitation>
clearly states “
<emphasis id="B92DEADCC15E8425FD5FFAADFBF18881" box="[708,1148,1404,1430]" italics="true" pageId="14" pageNumber="165">Туре und Kotypen in meiner Sammlung</emphasis>
” [type and cotypes in my collection], his collection (in
<collectionCode id="ED48AE0BC15E8425FE45FA70FDB888AD" box="[478,565,1440,1466]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="165" type="Museum">MHNG</collectionCode>
) includes only
<specimenCount id="9D5FFD47C15E8425FD44FA70FCFB88AD" box="[735,886,1440,1466]" pageId="14" pageNumber="165" type="generic">49 specimens</specimenCount>
all labelled as “cotype” (currently,
<typeStatus id="54E2886CC15E8425FB64FA71FAEB88AC" box="[1279,1382,1441,1467]" pageId="14" pageNumber="165" type="syntype">syntypes</typeStatus>
). We do not know if the type was actually selected by Petrovitz and is currently elsewhere, or if it has never been selected at all. In any case, as the species is easily identifiable using the characters indicated here below, we do not deem it necessary to designate a
<typeStatus id="54E2886CC15E8425FE37F9DCFD9B8B31" box="[428,534,1548,1574]" pageId="14" pageNumber="165" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BE636CEC15E8425FF0CF99FFD2D8B7E" blockId="14.[151,672,1614,1641]" box="[151,672,1614,1641]" pageId="14" pageNumber="165">
<materialsCitation id="3B313C93C15E8425FF0CF99FFD168B7E" ID-GBIF-Occurrence="2417904693" box="[151,667,1614,1641]" country="Greece" location="Gythion" pageId="14" pageNumber="165" specimenCount="1" stateProvince="Peloponnisos" typeStatus="holotype">
<emphasis id="B92DEADCC15E8425FF0CF99FFEBB8B7E" bold="true" box="[151,310,1614,1641]" pageId="14" pageNumber="165">
<typeStatus id="54E2886CC15E8425FF0CF99FFF5F8B7E" box="[151,210,1615,1641]" pageId="14" pageNumber="165">Type</typeStatus>
locality.
</emphasis>
[
<collectingCountry id="F34E765EC15E8425FED8F99EFE178B7E" box="[323,410,1614,1641]" name="Greece" pageId="14" pageNumber="165">Greece</collectingCountry>
],
<collectingRegion id="499DF82CC15E8425FE37F99FFDBF8B7E" box="[428,562,1615,1641]" country="Greece" name="Peloponnisos" pageId="14" pageNumber="165">Peloponnes</collectingRegion>
:
<location id="8E866015C15E8425FDA7F99FFD168B7E" LSID="urn:lsid:plazi:treatment:03F087D8C15C8424FF0CF93DFCF18E94:8E866015C15E8425FDA7F99FFD168B7E" box="[572,667,1615,1641]" country="Greece" name="Gythion" pageId="14" pageNumber="165" stateProvince="Peloponnisos">Gythion</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE636CEC15E8425FF0CF941FC688BBC" blockId="14.[151,1162,1681,1744]" box="[151,997,1681,1708]" pageId="14" pageNumber="165">
<emphasis id="B92DEADCC15E8425FF0CF941FE0C8BBB" bold="true" box="[151,385,1681,1708]" pageId="14" pageNumber="165">Material examined.</emphasis>
<specimenCount id="9D5FFD47C15E8425FE1CF942FE698BBB" box="[391,484,1682,1708]" pageId="14" pageNumber="165" type="male">199 ♂♂</specimenCount>
,
<specimenCount id="9D5FFD47C15E8425FE6BF942FDC18BBB" box="[496,588,1682,1708]" pageId="14" pageNumber="165" type="female">181 ♀♀</specimenCount>
, detailed in supplementary material.
</paragraph>
<paragraph id="8BE636CEC15E8425FF0CF966FB078BC7" blockId="14.[151,1162,1681,1744]" box="[151,1162,1717,1744]" pageId="14" pageNumber="165">
<materialsCitation id="3B313C93C15E8425FF0CF966FB088BC7" ID-GBIF-Occurrence="2417904717" box="[151,1157,1717,1744]" collectionCode="MHNG" pageId="14" pageNumber="165" specimenCount="98" specimenCount-female="31" specimenCount-male="18" typeStatus="holotype">
<emphasis id="B92DEADCC15E8425FF0CF966FE5B8BC7" bold="true" box="[151,470,1717,1744]" pageId="14" pageNumber="165">
<typeStatus id="54E2886CC15E8425FF0CF966FF5F8BC7" box="[151,210,1718,1744]" pageId="14" pageNumber="165">Type</typeStatus>
specimens examined:
</emphasis>
<specimenCount id="9D5FFD47C15E8425FE46F966FDF88BD8" box="[477,629,1717,1744]" pageId="14" pageNumber="165" type="generic">49 specimens</specimenCount>
(
<specimenCount id="9D5FFD47C15E8425FD1FF966FD598BC7" box="[644,724,1718,1744]" pageId="14" pageNumber="165" type="male">18 ♂♂</specimenCount>
,
<specimenCount id="9D5FFD47C15E8425FD7BF966FCA68BC7" box="[736,811,1718,1744]" pageId="14" pageNumber="165" type="female">31 ♀♀</specimenCount>
) labeled as cotypes in
<collectionCode id="ED48AE0BC15E8425FBB7F966FB088BC7" box="[1068,1157,1718,1744]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="165" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE636CEC15E8425FF0CF927FE1F8A05" blockId="14.[151,1437,1783,2027]" box="[151,402,1783,1810]" pageId="14" pageNumber="165">
<heading id="D0AE81A2C15E8425FF0CF927FE1F8A05" bold="true" box="[151,402,1783,1810]" fontSize="11" level="1" pageId="14" pageNumber="165" reason="1">
<emphasis id="B92DEADCC15E8425FF0CF927FE1F8A05" bold="true" box="[151,402,1783,1810]" pageId="14" pageNumber="165">Verified distribution.</emphasis>
</heading>
</paragraph>
<paragraph id="8BE636CEC15E8425FF0CF8CBFF7F8A21" blockId="14.[151,1437,1783,2027]" box="[151,242,1819,1846]" pageId="14" pageNumber="165">
<heading id="D0AE81A2C15E8425FF0CF8CBFF7F8A21" bold="true" box="[151,242,1819,1846]" fontSize="11" level="1" pageId="14" pageNumber="165" reason="1">
<emphasis id="B92DEADCC15E8425FF0CF8CBFF7F8A21" bold="true" box="[151,242,1819,1846]" pageId="14" pageNumber="165">
<collectingCountry id="F34E765EC15E8425FF0CF8CBFF618A21" box="[151,236,1819,1846]" name="Greece" pageId="14" pageNumber="165">Greece</collectingCountry>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE636CEC15E8425FF0CF890FC6B8AFD" blockId="14.[151,1437,1783,2027]" pageId="14" pageNumber="165">
<emphasis id="B92DEADCC15E8425FF0CF890FE2B8A4D" bold="true" box="[151,422,1855,1882]" pageId="14" pageNumber="165">
<collectingRegion id="499DF82CC15E8425FF0CF890FEA08A4D" box="[151,301,1856,1882]" country="Greece" name="Peloponnisos" pageId="14" pageNumber="165">Peloponnese</collectingRegion>
. Arcadia:
</emphasis>
near Vourvoura; near
<collectingRegion id="499DF82CC15E8425FD29F890FD7B8A4D" box="[690,758,1856,1882]" country="Kosovo" name="Viti" pageId="14" pageNumber="165">Vitina</collectingRegion>
(Menalo Mts); Kandila,
<quantity id="4CA19B2BC15E8425FBBAF890FBE08A4D" box="[1057,1133,1856,1882]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="165" unit="km" value="14.0">14 km</quantity>
N of Levidhion;
<quantity id="4CA19B2BC15E8425FAD8F890FA0C8A4D" box="[1347,1409,1856,1882]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="165" unit="km" value="3.0">3 km</quantity>
S of Alepochori; Dirrahi; near Karitena; Prosilion; Stenò.
<emphasis id="B92DEADCC15E8425FCD1F8B3FC268A69" bold="true" box="[842,939,1891,1918]" pageId="14" pageNumber="165">Argolis:</emphasis>
Pass between Tripoli and Argo.
<emphasis id="B92DEADCC15E8425FAB4F8B3FA118A69" bold="true" box="[1327,1436,1891,1918]" pageId="14" pageNumber="165">Lakonia:</emphasis>
Polovitsa;
<quantity id="4CA19B2BC15E8425FEAEF858FEFD8AB5" box="[309,368,1928,1954]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="165" unit="km" value="2.0">2 km</quantity>
SW of Polovitsa;
<quantity id="4CA19B2BC15E8425FDDAF858FDF18AB5" box="[577,636,1928,1954]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="165" unit="km" value="4.0">4 km</quantity>
S of Trapezonti; Agios Nikolaos (Taygetos); Arna (Taygetos); Gorani, Moni Golas; Gythion; Gythion, beach East; Mavrovouni (Gythion, Stomio Bay); Mystras; Chania; near Dafni; Paleopanagia (Taygetos); Loggastra; Karvelas; Kosiarion; Monemvasia;
</paragraph>
</subSubSection>
<subSubSection id="C3436545C15F8424FF0CFF48FE118C54" pageId="15" pageNumber="166" type="diagnosis">
<paragraph id="8BE636CEC15F8424FF0CFF48FE118C54" blockId="15.[151,1436,152,323]" pageId="15" pageNumber="166">
<emphasis id="B92DEADCC15F8424FF0CFF48FE9F8DA4" bold="true" box="[151,274,152,179]" pageId="15" pageNumber="166">Diagnosis.</emphasis>
Characters of the
<taxonomicName id="4C594D4DC15F8424FE7BFF4AFDEA8DA4" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[480,615,154,179]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="anemoninus">
<emphasis id="B92DEADCC15F8424FE7BFF4AFDEA8DA4" box="[480,615,154,179]" italics="true" pageId="15" pageNumber="166">anemoninus</emphasis>
</taxonomicName>
species-group described above, plus the following characters. Male with lateral setation of abdomen and setation of elytra black. Head and pronotum usually with dark and unsaturated hues of violet or green. Paramera similar to those of
<taxonomicName id="4C594D4DC15F8424FD36FF32FCB98DEC" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[685,820,226,251]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="anemoninus">
<emphasis id="B92DEADCC15F8424FD36FF32FCB98DEC" box="[685,820,226,251]" italics="true" pageId="15" pageNumber="166">anemoninus</emphasis>
</taxonomicName>
and not diagnostic, everted endophallus similar to that of
<taxonomicName id="4C594D4DC15F8424FF2DFED6FEB08C08" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[182,317,262,287]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="anemoninus">
<emphasis id="B92DEADCC15F8424FF2DFED6FEB08C08" box="[182,317,262,287]" italics="true" pageId="15" pageNumber="166">anemoninus</emphasis>
</taxonomicName>
but diagnostic (
<figureCitation id="13622A4BC15F8424FE6FFED5FDF28C08" box="[500,639,261,287]" captionStart="FIGURES 25–31" captionStartId="13.[151,264,1869,1894]" captionTargetBox="[163,1408,194,1804]" captionTargetId="figure@13.[151,1436,181,1844]" captionTargetPageId="13" captionText="FIGURES 25–31. Pygopleurus pseudopsilotrichius (Petrovitz, 1958), habitus and genitalia. 25, male (Lakonia, Gorani); 26, male (Arcadia, Karitena); 27, female (Lakonia, Chanià); 28, female (Lakonia, Gorani); 29, left paramere in lateral and dorsal view (Laconia, Chanià); 30, everted endophallus in left lateral view (Laconia, Chanià); 31, left genital sclerites of female (Laconia, Dafni)." figureDoi="http://doi.org/10.5281/zenodo.3458852" httpUri="https://zenodo.org/record/3458852/files/figure.png" pageId="15" pageNumber="166">Figs. 29–30</figureCitation>
). Female similar to that of
<taxonomicName id="4C594D4DC15F8424FC2BFED6FBDE8C08" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[944,1107,262,287]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="anemoninus">
<emphasis id="B92DEADCC15F8424FC2BFED6FBDE8C08" box="[944,1107,262,287]" italics="true" pageId="15" pageNumber="166">P. anemoninus</emphasis>
</taxonomicName>
, reliably identifiable only by association with males.
</paragraph>
</subSubSection>
<subSubSection id="C3436545C15F8424FF0CFEA0FAA98C9B" box="[151,1316,368,396]" pageId="15" pageNumber="166" type="distribution">
<paragraph id="8BE636CEC15F8424FF0CFEA0FAA98C9B" blockId="15.[151,1316,368,396]" box="[151,1316,368,396]" pageId="15" pageNumber="166">
<emphasis id="B92DEADCC15F8424FF0CFEA0FE168C9C" bold="true" box="[151,411,368,395]" pageId="15" pageNumber="166">
Distribution (
<figureCitation id="13622A4BC15F8424FEA3FEA0FE038C9C" box="[312,398,368,395]" captionStart="FIGURE 32" captionStartId="14.[151,250,1266,1291]" captionTargetBox="[170,1408,181,1235]" captionTargetId="figure@14.[170,1416,181,1241]" captionTargetPageId="14" captionText="FIGURE 32. Pygopleurus anemoninus (Brullé, 1832) and P. pseudopsilotrichius (Petrovitz, 1958): geographical, altitudinal, and temporal distribution of records. Red curves showing the expected distribution if the species was uniformly present at all heights and all times (see Material and methods)." figureDoi="http://doi.org/10.5281/zenodo.3458854" httpUri="https://zenodo.org/record/3458854/files/figure.png" pageId="15" pageNumber="166">Fig. 32</figureCitation>
).
</emphasis>
Restricted to Central and Southern
<collectingRegion id="499DF82CC15F8424FCB7FEA1FC378C9C" box="[812,954,369,395]" country="Greece" name="Peloponnisos" pageId="15" pageNumber="166">Peloponnese</collectingRegion>
(Lakonia, Arcadia and Argolis).
</paragraph>
</subSubSection>
<subSubSection id="C3436545C15F8424FF0CFE68FE4C8F90" pageId="15" pageNumber="166" type="biology_ecology">
<paragraph id="8BE636CEC15F8424FF0CFE68FE4C8F90" blockId="15.[151,1436,440,647]" pageId="15" pageNumber="166">
<emphasis id="B92DEADCC15F8424FF0CFE68FD898CC4" bold="true" box="[151,516,440,467]" pageId="15" pageNumber="166">
Eco-ethological notes (
<figureCitation id="13622A4BC15F8424FE39FE68FE7A8CC4" box="[418,503,440,467]" captionStart="FIGURE 32" captionStartId="14.[151,250,1266,1291]" captionTargetBox="[170,1408,181,1235]" captionTargetId="figure@14.[170,1416,181,1241]" captionTargetPageId="14" captionText="FIGURE 32. Pygopleurus anemoninus (Brullé, 1832) and P. pseudopsilotrichius (Petrovitz, 1958): geographical, altitudinal, and temporal distribution of records. Red curves showing the expected distribution if the species was uniformly present at all heights and all times (see Material and methods)." figureDoi="http://doi.org/10.5281/zenodo.3458854" httpUri="https://zenodo.org/record/3458854/files/figure.png" pageId="15" pageNumber="166">Fig. 32</figureCitation>
).
</emphasis>
The adults of this species are active between the middle of April (
<date id="FFE7100EC15F8424FB7FFE69FADD8CC4" box="[1252,1360,440,467]" pageId="15" pageNumber="166">
12
<superScript id="7C2C9B86C15F8424FA9AFE68FA838CD1" attach="left" box="[1281,1294,440,454]" fontSize="6" pageId="15" pageNumber="166">th</superScript>
April
</date>
, Laconia, Monemvasia, sea level) and the end of May (
<date id="FFE7100EC15F8424FD25FE0DFCAF8CE0" box="[702,802,476,503]" pageId="15" pageNumber="166">
27
<superScript id="7C2C9B86C15F8424FD40FE0CFD658CFD" attach="left" box="[731,744,476,490]" fontSize="6" pageId="15" pageNumber="166">th</superScript>
May
</date>
, Arcadia, Vytina, altitude not recorded, but close to
<quantity id="4CA19B2BC15F8424FAFFFE0DFF3C8F0C" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="166" unit="m" value="1000.0">1000 m</quantity>
). The species, which is usually abundant, is found from sea level up to
<quantity id="4CA19B2BC15F8424FC24FDD1FB988F0C" box="[959,1045,513,539]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="166" unit="m" value="1000.0">1000 m</quantity>
, both elevations being documented in the mountain ranges of Parnon and Taygetos. Like its vicariant species
<taxonomicName id="4C594D4DC15F8424FC6EFDF6FB158F28" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[1013,1176,550,575]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="anemoninus">
<emphasis id="B92DEADCC15F8424FC6EFDF6FB158F28" box="[1013,1176,550,575]" italics="true" pageId="15" pageNumber="166">P. anemoninus</emphasis>
</taxonomicName>
, the adults are mainly found feeding or mating on
<taxonomicName id="4C594D4DC15F8424FE4AFD9AFDBD8F74" box="[465,560,586,611]" class="Magnoliopsida" family="Papaveraceae" genus="Papaver" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="B92DEADCC15F8424FE4AFD9AFDBD8F74" box="[465,560,586,611]" italics="true" pageId="15" pageNumber="166">Papaver</emphasis>
</taxonomicName>
,
<taxonomicName id="4C594D4DC15F8424FDA4FD99FD6E8F74" box="[575,739,585,611]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="incanus">
<emphasis id="B92DEADCC15F8424FDA4FD99FD6E8F74" box="[575,739,585,611]" italics="true" pageId="15" pageNumber="166">Cistus incanus</emphasis>
</taxonomicName>
, and Asteracee (white or yellow Tubuliflorae and Liguliflorae like
<taxonomicName id="4C594D4DC15F8424FF53FDBDFE9F8F90" box="[200,274,621,647]" class="Gymnolaemata" family="Cymuloporidae" genus="Crepis" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="166" phylum="Bryozoa" rank="genus">
<emphasis id="B92DEADCC15F8424FF53FDBDFE9F8F90" box="[200,274,621,647]" italics="true" pageId="15" pageNumber="166">Crepis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C594D4DC15F8424FED3FDBDFE3D8F90" box="[328,432,621,647]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="B92DEADCC15F8424FED3FDBDFE3D8F90" box="[328,432,621,647]" italics="true" pageId="15" pageNumber="166">Anthemis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3436545C15F8424FF0CFD64FCF18E94" pageId="15" pageNumber="166" type="discussion">
<paragraph id="8BE636CEC15F8424FF0CFD64FCF18E94" blockId="15.[151,1437,692,899]" pageId="15" pageNumber="166">
<emphasis id="B92DEADCC15F8424FF0CFD64FE1D8FD8" bold="true" box="[151,400,692,719]" pageId="15" pageNumber="166">Taxonomic remarks.</emphasis>
<taxonomicName id="4C594D4DC15F8424FE01FD65FC818FD8" baseAuthorityName="Petrovitz" baseAuthorityYear="1958" box="[410,780,693,719]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="pseudopsilotrichius">
<emphasis id="B92DEADCC15F8424FE01FD65FC818FD8" box="[410,780,693,719]" italics="true" pageId="15" pageNumber="166">Pygopleurus pseudopsilotrichius</emphasis>
</taxonomicName>
was described as a distinct species, but placed into synonymy with
<taxonomicName id="4C594D4DC15F8424FEBFFD0AFE4A8FE4" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[292,455,730,755]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="anemoninus">
<emphasis id="B92DEADCC15F8424FEBFFD0AFE4A8FE4" box="[292,455,730,755]" italics="true" pageId="15" pageNumber="166">P. anemoninus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC84B3FC15F8424FE6AFD09FD1B8FE4" author="Baraud, J." box="[497,662,729,755]" pageId="15" pageNumber="166" pagination="331 - 375" refId="ref29740" refString="Baraud, J. (1989) Revision du genre Pygopleurus Motschulsky (Coleoptera, Scarabaoidea: Glaphyridae). Annales de la Societe Entomologique de France, 25 (3), 331 - 375, 89 figs." type="book chapter" year="1989">Baraud (1989)</bibRefCitation>
. Comparing more than
<specimenCount id="9D5FFD47C15F8424FC05FD09FBCE8FE4" box="[926,1091,729,755]" pageId="15" pageNumber="166" type="generic">720 specimens</specimenCount>
of both nominal taxa, we have ascertained that examined populations can be consistently distinguished by the characters mentioned in the diagnosis. In addition, populations distinguished on morphological basis are distributed on two parapatric ranges (fig. 32). In spite of the rather limited morphological differences encountered, the mentioned observations and the absence of intermediate populations support treating these as distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>
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