557 lines
82 KiB
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557 lines
82 KiB
XML
<document id="635467C63C1E503229F5BFF2C08F457B" ID-DOI="10.11646/zootaxa.3994.2.2" ID-GBIF-Dataset="69a42e8b-4d2a-4a8e-ad1a-da6d2fc952aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="243281" ID-ZooBank="41A8AD52-D7E1-4242-87D1-C7819433FA22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461273516540" checkinUser="plazi" docAuthor="Batista, Abel, Ponce, Marcos, Vesely, Milan, Mebert, Konrad, Hertz, Andreas, Köhler, Gunther, Carrizo, Arcadio & Lotzkat, Sebastian" docDate="2015" docId="03F087C8FFC0FF8561A2FCC76B6090D2" docLanguage="en" docName="zt03994p221.pdf" docOrigin="Zootaxa 3994 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidoblepharis emberawoundule Batista, Ponce, Vesely, Mebert, Hertz, Köhler, Carrizo & Lotzkat, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="196" masterDocId="FFC9FFB0FFC7FF8C6135FF9169259747" masterDocTitle="Revision of the genus Lepidoblepharis (Reptilia: Squamata: Sphaerodactylidae) in Central America, with the description of three new species" masterLastPageNumber="221" masterPageNumber="187" pageNumber="194" updateTime="1698612224987" updateUser="plazi">
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<mods:title id="CE98E2794FF7C748F5548A624764F1CF">Revision of the genus Lepidoblepharis (Reptilia: Squamata: Sphaerodactylidae) in Central America, with the description of three new species</mods:title>
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<mods:namePart id="6982D39BC3A3A80E5CDFF50D71AC2AB1">Batista, Abel</mods:namePart>
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<mods:namePart id="5926EB6E9CC80A08D35C3D72911C912F">Ponce, Marcos</mods:namePart>
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<mods:namePart id="1044FA9935EA2B877A93687808538426">Vesely, Milan</mods:namePart>
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<mods:namePart id="9F510E453C83B6D7AA392A90240F3A7D">Mebert, Konrad</mods:namePart>
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<mods:namePart id="79E5BA40CD807E3461FB71F6454A0C3E">Hertz, Andreas</mods:namePart>
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<mods:namePart id="39211C2156B918BB888939F02053ACA3">Köhler, Gunther</mods:namePart>
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<mods:namePart id="DB1B1764BD6CE05E779C271690E8D52A">Carrizo, Arcadio</mods:namePart>
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<mods:namePart id="FF494A9149B7D0D155086C908350E1A6">Lotzkat, Sebastian</mods:namePart>
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<mods:title id="9FE73D5ED711C782B8BC7A587276DD52">Zootaxa</mods:title>
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<mods:date id="D74B92925B53378A6DACBD4356EFF398">2015</mods:date>
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<treatment id="03F087C8FFC0FF8561A2FCC76B6090D2" ID-DOI="http://doi.org/10.5281/zenodo.5614821" ID-GBIF-Taxon="119639001" ID-Zenodo-Dep="5614821" LSID="urn:lsid:plazi:treatment:03F087C8FFC0FF8561A2FCC76B6090D2" httpUri="http://treatment.plazi.org/id/03F087C8FFC0FF8561A2FCC76B6090D2" lastPageId="9" lastPageNumber="196" pageId="7" pageNumber="194">
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<heading id="D0AE81B2FFC0FF8B61A2FCC76BB39436" bold="true" box="[151,662,854,881]" fontSize="11" level="1" pageId="7" pageNumber="194" reason="1">
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<emphasis id="B92DEACCFFC0FF8B61A2FCC76BB39436" bold="true" box="[151,662,854,881]" pageId="7" pageNumber="194">
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<taxonomicName id="4C594D5DFFC0FF8B61A2FCC76B179437" box="[151,562,854,880]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="emberawoundule" status="sp. nov.">
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<emphasis id="B92DEACCFFC0FF8B61A2FCC76B179437" bold="true" box="[151,562,854,880]" italics="true" pageId="7" pageNumber="194">Lepidoblepharis emberawoundule</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21E57B7FFC0FF8B630CFCC66BB39436" box="[569,662,855,881]" pageId="7" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE636DEFFC0FF8B61A2FCE8681594D5" blockId="7.[151,662,854,914]" box="[151,304,889,914]" pageId="7" pageNumber="194">
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<figureCitation id="13622A5BFFC0FF8B61A2FCE8682494D5" box="[151,257,889,914]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="11.[151,250,1918,1940]" captionStartId-1="12.[151,250,792,814]" captionStartId-2="13.[151,250,1562,1584]" captionStartId-3="14.[151,250,944,966]" captionStartId-4="18.[151,250,1802,1824]" captionTargetBox-0="[244,1340,193,1892]" captionTargetBox-1="[150,1436,193,770]" captionTargetBox-2="[151,1436,179,1539]" captionTargetBox-3="[216,1371,176,923]" captionTargetBox-4="[335,1251,193,1780]" captionTargetId-0="figure@11.[244,1340,193,1893]" captionTargetId-1="figure@12.[150,1436,193,771]" captionTargetId-2="figure@13.[151,1436,179,1541]" captionTargetId-3="figure@14.[216,1371,169,923]" captionTargetId-4="figure@18.[335,1251,193,1780]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="18" captionText-0="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." captionText-1="FIGURE 4. Feet and hands of Panamanian Lepidoblepharis. Lepidoblepharis emberawoundule: right foot of SMF 81954 and right hand of MHCH 2957; L. rufigularis (SMF 50659): right foot and right hand; L. sanctaemartae: left foot of SMF 97419 and right hand of MHCH 2948; L. victormartinezi: left foot and left hand of SMF 89963; L. xanthostigma: left foot of SMF 90190 and left hand of SMF 89576. Scale bars equal 1 mm, tip of fourth finger and toe marked with an asterisk." captionText-2="FIGURE 5. Escutcheon scales in males of Panamanian Lepidoblepharis. (A) L. emberawoundule: ventral escutcheon with 46 scales, 7 scales long x 11 wide, and subfemoral escutcheon with 4 scales per side, of SMF 81951 (SVL = 25 mm); (B) L. rufigularis: ventral escutcheon with 62 scales, 7 scales long x 13 wide, and subfemoral escutcheon with 3 and 4 scales per side, of SMF 50659 (SVL = 25 mm); (C) L. sanctaemartae: ventral escutcheon of SMF 97419 (SVL = 22 mm) with 92 scales, 10 scales long x 12 wide; (D) L. victormartinezi: ventral escutcheon of SMF 89963 (SVL = 27 mm) comprising 57 intact scales (an original total of 61 is assumed considering the symmetrical scale arrangement of the non-damaged portions), 7 scales long x 14 wide; (E – F) L. xanthostigma: (E) ventral escutcheon with 75 scales, 12 scales long x 11 wide, and subfemoral escutcheon with 2 scales per side (no additional scales hidden under strings), of SMF 90189 (SVL = 35 mm), and (F) ventral escutcheon with 36 scales, 6 scales long x 8 wide, and no subfemoral escutcheon of AB 1250 (SVL = 24 mm). All scale bars equal 1 mm." captionText-3="FIGURE 6. Hemipenis morphology of four Lepidoblepharis species from Panama. Sulcate-asulcate view, (A – B) L. emberawoundule SMF 50968; (C – D) L. sanctaemartae SMF 97419 (E – F) L. victormartinezi SMF 50950; (G – H) L. xanthostigma SMF 91558. All scale bars equal 1 mm." captionText-4="FIGURE 7. Comparison between Lepidoblepharis emberawoundule, L. microlepis (holotype), and L. peraccae (holotype). Mental plate and postmental scales in (A) L. microlepis and (B) L. emerawondule; (C) right foot of L. microlepis; (D) right foot of L. emberawoundule; (E) subcaudal scales in L. microlepis, each of the slightly enlarged median subcaudal scales is bordered laterodistally by only one scale, leading to a regular tail sequence of 1 ' 1 '; (F) subcaudal scales in L. emberawoundule, the larger of the enlarged subcaudals are bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1 ' 1 ''; (G) right foot and (H) right hand of the holotype of L. peraccae; scale bars equal 1 mm." httpUri-0="https://zenodo.org/record/243283/files/figure.png" httpUri-1="https://zenodo.org/record/243284/files/figure.png" httpUri-2="https://zenodo.org/record/243285/files/figure.png" httpUri-3="https://zenodo.org/record/243286/files/figure.png" httpUri-4="https://zenodo.org/record/243287/files/figure.png" pageId="7" pageNumber="194">Figs. 3–7</figureCitation>
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,
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<figureCitation id="13622A5BFFC0FF8B6038FCEB680994D5" box="[269,300,890,914]" captionStart="FIGURE 10" captionStartId="22.[151,250,1971,1993]" captionTargetBox="[188,1398,1299,1948]" captionTargetId="figure@22.[188,1398,1299,1949]" captionTargetPageId="22" captionText="FIGURE 10. Holotype of Lepidoblepharis emberawoundule (male SMF 50968) in life. (A) entire specimen; (B) dorsal view of the head; (C) lateral view of the head." httpUri="https://zenodo.org/record/243290/files/figure.png" pageId="7" pageNumber="194">10</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3436555FFC0FF8B61A2FC506D5194B1" pageId="7" pageNumber="194" type="reference_group">
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<paragraph id="8BE636DEFFC0FF8B61A2FC506D5194B1" blockId="7.[151,1437,961,1014]" pageId="7" pageNumber="194">
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<emphasis id="B92DEACCFFC0FF8B61A2FC5068FD9490" box="[151,472,961,983]" italics="true" pageId="7" pageNumber="194">
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<taxonomicName id="4C594D5DFFC0FF8B61A2FC5068F49490" box="[151,465,961,983]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="xanthostigma">Lepidoblepharis xanthostigma</taxonomicName>
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:
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</emphasis>
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all in part. (referring to certain populations in eastern
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<collectingCountry id="F34E764EFFC0FF8B652AFC536D509490" box="[1055,1141,962,983]" name="Panama" pageId="7" pageNumber="194">Panama</collectingCountry>
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):
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<bibRefCitation id="EFC84B2FFFC0FF8B65BDFC506C299490" author="Auth" box="[1160,1292,961,983]" pageId="7" pageNumber="194" refString="Auth, D. L. (1994) Checklist and bibliography of the amphibians and reptiles of Panama. Smithsonian Herpetological Information Service, 98, 1 - 59. http: // dx. doi. org / 10.5479 / si. 23317515.98.1" type="journal article" year="1994">Auth (1994)</bibRefCitation>
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;
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||
<bibRefCitation id="EFC84B2FFFC0FF8B642CFC53682B94B1" pageId="7" pageNumber="194" refString="Young, B. E., Sedaghatkish, G., Roca, E. & Fuenmayor, Q. D. (1999) El estatus de la conservacion de la herpetofauna de Panama. Resumen del primer taller internacional sobre la herpetofauna de Panama. The Nature Conservancy y Asociacion Nacional para la Conservacion de la Naturaleza (ANCON), Arlington, Virginia, 40 pp." type="book">
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Young
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||
<emphasis id="B92DEACCFFC0FF8B6453FC536CB89490" box="[1382,1437,961,983]" italics="true" pageId="7" pageNumber="194">et al.</emphasis>
|
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(1999)
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||
</bibRefCitation>
|
||
; Ibáñez
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||
<emphasis id="B92DEACCFFC0FF8B6054FC7068B094B1" box="[353,405,992,1014]" italics="true" pageId="7" pageNumber="194">et al.</emphasis>
|
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(2001);
|
||
<bibRefCitation id="EFC84B2FFFC0FF8B60DEFC716B5394B1" author="Kohler" box="[491,630,992,1014]" pageId="7" pageNumber="194" refString="Kohler, G. (2001) Anfibios y reptiles de Nicaragua. Herpeton Verlag, Offenbach, 208 pp." type="book" year="2001">Köhler (2001</bibRefCitation>
|
||
: Fig. 172; 2008: Fig. 136);
|
||
<bibRefCitation id="EFC84B2FFFC0FF8B62B9FC716D5594B1" box="[908,1136,992,1014]" pageId="7" pageNumber="194" refString="Jaramillo A., C. A., Wilson, L. D., Ibanez D., R. & Jaramillo, F. E. (2010) The herpetofauna of Panama: distribution and conservation status. In: Wilson, L. D., Townsend, J. H., Johnson, J. D. & Murphy, J. B. (Eds.) Conservation of Mesoamerican amphibians and reptiles. Eagle Mountain Press, Eagle Mountain, Utah, pp. 604 - 671." type="book chapter">
|
||
Jaramillo
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||
<emphasis id="B92DEACCFFC0FF8B62C4FC706D0194B1" box="[1009,1060,992,1014]" italics="true" pageId="7" pageNumber="194">et al.</emphasis>
|
||
(2010)
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||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C3436555FFC0FF8B61A2FBB26866921B" pageId="7" pageNumber="194" type="materials_examined">
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<paragraph id="8BE636DEFFC0FF8B61A2FBB26D0F93C3" blockId="7.[151,1437,1059,2020]" pageId="7" pageNumber="194">
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<emphasis id="B92DEACCFFC0FF8B61A2FBB2682E937B" bold="true" box="[151,267,1059,1084]" pageId="7" pageNumber="194">
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<typeStatus id="54E2887CFFC0FF8B61A2FBB26823937B" box="[151,262,1059,1084]" pageId="7" pageNumber="194" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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Adult male
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<collectionCode id="ED48AE1BFFC0FF8B60A0FBB268E8937C" LSID="urn:lsid:biocol.org:col:34838" box="[405,461,1059,1083]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="194">SMF</collectionCode>
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||
50968, original field number AB 963 (
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<figureCitation id="13622A5BFFC0FF8B62B4FBB26AF1937C" box="[897,980,1059,1084]" captionStart="FIGURE 10" captionStartId="22.[151,250,1971,1993]" captionTargetBox="[188,1398,1299,1948]" captionTargetId="figure@22.[188,1398,1299,1949]" captionTargetPageId="22" captionText="FIGURE 10. Holotype of Lepidoblepharis emberawoundule (male SMF 50968) in life. (A) entire specimen; (B) dorsal view of the head; (C) lateral view of the head." httpUri="https://zenodo.org/record/243290/files/figure.png" pageId="7" pageNumber="194">Fig. 10</figureCitation>
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), collected from leaf-litter at La Cascada trail, Burbayar private reserve (
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,
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,
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elev.), Cartí, Narganá, Comarca Guna Yala,
|
||
<collectingCountry id="F34E764EFFC0FF8B61A2FBFD69D693C4" box="[151,243,1132,1155]" name="Panama" pageId="7" pageNumber="194">Panama</collectingCountry>
|
||
, on
|
||
<date id="FFE7101EFFC0FF8B6015FBFA68DC93C4" box="[288,505,1131,1156]" pageId="7" pageNumber="194" value="2013-11-26">26 November 2013</date>
|
||
at 23:40 hrs, by Abel Batista and Konrad Mebert.
|
||
</paragraph>
|
||
<paragraph id="8BE636DEFFC0FF8B61F2FB1F6ADD9253" blockId="7.[151,1437,1059,2020]" pageId="7" pageNumber="194">
|
||
<emphasis id="B92DEACCFFC0FF8B61F2FB1F686393E0" bold="true" box="[199,326,1166,1191]" pageId="7" pageNumber="194">
|
||
<typeStatus id="54E2887CFFC0FF8B61F2FB1F686493E0" box="[199,321,1166,1191]" pageId="7" pageNumber="194" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Three adult males, two adult females, and one juvenile, all from
|
||
<collectingCountry id="F34E764EFFC0FF8B6563FB016D9793E0" box="[1110,1202,1168,1191]" name="Panama" pageId="7" pageNumber="194">Panama</collectingCountry>
|
||
. Three males (
|
||
<collectionCode id="ED48AE1BFFC0FF8B6457FB1E6CB993E0" LSID="urn:lsid:biocol.org:col:34838" box="[1378,1436,1167,1191]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="194">SMF</collectionCode>
|
||
81950–52) from Nusagandí field station and two females (
|
||
<collectionCode id="ED48AE1BFFC0FF8B6200FB226A4A938C" LSID="urn:lsid:biocol.org:col:34838" box="[821,879,1203,1227]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="194">SMF</collectionCode>
|
||
81953–54) from the nearby Sendero Nusagandí, Comarca Guna Yala, collected
|
||
<date id="FFE7101EFFC0FF8B60C0FB466BE593A8" box="[501,704,1238,1263]" pageId="7" pageNumber="194" value="2000-04-14" valueMax="2000-04-17" valueMin="2000-04-14">14–17 April 2000</date>
|
||
; one juvenile (
|
||
<collectionCode id="ED48AE1BFFC0FF8B6253FB466A9993A8" box="[870,956,1239,1263]" pageId="7" pageNumber="194">MHCH</collectionCode>
|
||
2952) from Río Terable, El Llano, Chepo,
|
||
<collectingCountry id="F34E764EFFC0FF8B61A2FB6D69D69254" box="[151,243,1276,1299]" name="Panama" pageId="7" pageNumber="194">Panama</collectingCountry>
|
||
, collected on
|
||
<date id="FFE7101EFFC0FF8B60B8FB6A6B4D9254" box="[397,616,1275,1300]" pageId="7" pageNumber="194" value="2012-11-27">27 November 2012</date>
|
||
; see Appendix I for locality details.
|
||
</paragraph>
|
||
<paragraph id="8BE636DEFFC0FF8B61F2FA8E6866921B" blockId="7.[151,1437,1059,2020]" pageId="7" pageNumber="194">
|
||
<emphasis id="B92DEACCFFC0FF8B61F2FA8E689E927F" bold="true" box="[199,443,1311,1336]" pageId="7" pageNumber="194">Referred specimens.</emphasis>
|
||
<collectionCode id="ED48AE1BFFC0FF8B60F1FA8E6B3D9270" box="[452,536,1311,1335]" pageId="7" pageNumber="194">MHCH</collectionCode>
|
||
2951, 2957;
|
||
<collectionCode id="ED48AE1BFFC0FF8B6381FA8E6BC99270" LSID="urn:lsid:biocol.org:col:34838" box="[692,748,1311,1335]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="194">SMF</collectionCode>
|
||
50969–70;
|
||
<collectionCode id="ED48AE1BFFC0FF8B6243FAB16AE29270" LSID="urn:lsid:biocol.org:col:34795" box="[886,967,1312,1335]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="194">FMNH</collectionCode>
|
||
170029, 170042–45; see Appendix I for locality details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3436555FFC0FF8B61F2FAF66CB890A3" pageId="7" pageNumber="194" type="diagnosis">
|
||
<paragraph id="8BE636DEFFC0FF8B61F2FAF66A26904C" blockId="7.[151,1437,1059,2020]" pageId="7" pageNumber="194">
|
||
<emphasis id="B92DEACCFFC0FF8B61F2FAF6686492C7" bold="true" box="[199,321,1383,1408]" pageId="7" pageNumber="194">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C594D5DFFC0FF8B6072FAF66BE39238" box="[327,710,1383,1407]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFC0FF8B6072FAF66BE39238" box="[327,710,1383,1407]" italics="true" pageId="7" pageNumber="194">Lepidoblepharis emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
(our
|
||
<emphasis id="B92DEACCFFC0FF8B6237FAF66A7E92C7" bold="true" box="[770,859,1383,1408]" pageId="7" pageNumber="194">
|
||
sp.
|
||
<date id="FFE7101EFFC0FF8B621FFAF66A7E92C7" box="[810,859,1383,1408]" pageId="7" pageNumber="194">nov.</date>
|
||
</emphasis>
|
||
1) is characterized by the following combination of characters:
|
||
<emphasis id="B92DEACCFFC0FF8B6022FA1A681D92E4" box="[279,312,1419,1443]" italics="true" pageId="7" pageNumber="194">(1)</emphasis>
|
||
dorsal body scales small, granular, and juxtaposed, ventral scales large, cycloid, flat, and imbricate;
|
||
<emphasis id="B92DEACCFFC0FF8B61A2FA3E699D9280" box="[151,184,1455,1479]" italics="true" pageId="7" pageNumber="194">(2)</emphasis>
|
||
scales on head small and granular;
|
||
<emphasis id="B92DEACCFFC0FF8B6368FA3E6B5A9280" box="[605,639,1455,1479]" italics="true" pageId="7" pageNumber="194">(3)</emphasis>
|
||
3–4 (3.1 ± 0.3) postrostral scales;
|
||
<emphasis id="B92DEACCFFC0FF8B652CFA3E6D1F9280" box="[1049,1082,1455,1479]" italics="true" pageId="7" pageNumber="194">(4)</emphasis>
|
||
a vaguely M-shaped posterior mental border with two paramedian clefts;
|
||
<emphasis id="B92DEACCFFC0FF8B63B2FA426B8D92AD" box="[647,680,1491,1514]" italics="true" pageId="7" pageNumber="194">(5)</emphasis>
|
||
3–7 (5.1 ± 1.04) postmentals, larger than the posteriorly adjacent scales on chin;
|
||
<emphasis id="B92DEACCFFC0FF8B607DFA66684C9148" box="[328,361,1527,1551]" italics="true" pageId="7" pageNumber="194">(6)</emphasis>
|
||
lamellae under fourth toe 6–9 (7.9 ± 0.85), lamellae under fourth finger 5–8 (6.7 ± 1.01);
|
||
<emphasis id="B92DEACCFFC0FF8B644EFA666CB99149" box="[1403,1436,1527,1550]" italics="true" pageId="7" pageNumber="194">(7)</emphasis>
|
||
median subcaudals conspicuously wider than long, but their width less than twice the width of the laterally adjacent scales or their own length, with straight or rounded posterior margins, arranged in a regular tail sequence of 1'1'';
|
||
<emphasis id="B92DEACCFFC0FF8B61A2F9F2699D913C" box="[151,184,1635,1659]" italics="true" pageId="7" pageNumber="194">(8)</emphasis>
|
||
ventral escutcheon consisting of 38–61 (48.4 ± 8.85) scales, 6–7 (6.8 ± 0.45) scales long and 10–13 (11.6 ± 1.14) wide;
|
||
<emphasis id="B92DEACCFFC0FF8B6029F916681B91D8" box="[284,318,1671,1695]" italics="true" pageId="7" pageNumber="194">(9)</emphasis>
|
||
subfemoral escutcheon consisting of 4–5 (4.4 ± 0.52) well-discernible scales per thigh arranged in a single row (
|
||
<figureCitation id="13622A5BFFC0FF8B6029F93A687A9183" box="[284,351,1707,1732]" captionStart="FIGURE 5" captionStartId="13.[151,250,1562,1584]" captionTargetBox="[151,1436,179,1539]" captionTargetId="figure@13.[151,1436,179,1541]" captionTargetPageId="13" captionText="FIGURE 5. Escutcheon scales in males of Panamanian Lepidoblepharis. (A) L. emberawoundule: ventral escutcheon with 46 scales, 7 scales long x 11 wide, and subfemoral escutcheon with 4 scales per side, of SMF 81951 (SVL = 25 mm); (B) L. rufigularis: ventral escutcheon with 62 scales, 7 scales long x 13 wide, and subfemoral escutcheon with 3 and 4 scales per side, of SMF 50659 (SVL = 25 mm); (C) L. sanctaemartae: ventral escutcheon of SMF 97419 (SVL = 22 mm) with 92 scales, 10 scales long x 12 wide; (D) L. victormartinezi: ventral escutcheon of SMF 89963 (SVL = 27 mm) comprising 57 intact scales (an original total of 61 is assumed considering the symmetrical scale arrangement of the non-damaged portions), 7 scales long x 14 wide; (E – F) L. xanthostigma: (E) ventral escutcheon with 75 scales, 12 scales long x 11 wide, and subfemoral escutcheon with 2 scales per side (no additional scales hidden under strings), of SMF 90189 (SVL = 35 mm), and (F) ventral escutcheon with 36 scales, 6 scales long x 8 wide, and no subfemoral escutcheon of AB 1250 (SVL = 24 mm). All scale bars equal 1 mm." httpUri="https://zenodo.org/record/243285/files/figure.png" pageId="7" pageNumber="194">Fig. 5</figureCitation>
|
||
A);
|
||
<emphasis id="B92DEACCFFC0FF8B60A6F93A68E79184" box="[403,450,1707,1731]" italics="true" pageId="7" pageNumber="194">(10)</emphasis>
|
||
16–20 (18.2 ± 1.17) longitudinal rows of ventral scales at midbody;
|
||
<emphasis id="B92DEACCFFC0FF8B65FFF93A6DD29184" box="[1226,1271,1707,1731]" italics="true" pageId="7" pageNumber="194">(11)</emphasis>
|
||
52–74 (61.8 ± 7.61) longitudinal rows of dorsal scales at midbody;
|
||
<emphasis id="B92DEACCFFC0FF8B63C3F95E6A0091A0" box="[758,805,1743,1767]" italics="true" pageId="7" pageNumber="194">(12)</emphasis>
|
||
bilobate hemipenis, with a third lobule rising from the pedicel (
|
||
<figureCitation id="13622A5BFFC0FF8B61C2F962681C904B" box="[247,313,1779,1804]" captionStart="FIGURE 6" captionStartId="14.[151,250,944,966]" captionTargetBox="[216,1371,176,923]" captionTargetId="figure@14.[216,1371,169,923]" captionTargetPageId="14" captionText="FIGURE 6. Hemipenis morphology of four Lepidoblepharis species from Panama. Sulcate-asulcate view, (A – B) L. emberawoundule SMF 50968; (C – D) L. sanctaemartae SMF 97419 (E – F) L. victormartinezi SMF 50950; (G – H) L. xanthostigma SMF 91558. All scale bars equal 1 mm." httpUri="https://zenodo.org/record/243286/files/figure.png" pageId="7" pageNumber="194">Fig. 6</figureCitation>
|
||
A–B);
|
||
<emphasis id="B92DEACCFFC0FF8B60B8F9626899904C" box="[397,444,1779,1803]" italics="true" pageId="7" pageNumber="194">(13)</emphasis>
|
||
SVL 21–30 (25 ± 2.07) mm.
|
||
</paragraph>
|
||
<paragraph id="8BE636DEFFC0FF8B61F2F8866CB890A3" blockId="7.[151,1437,1059,2020]" pageId="7" pageNumber="194">
|
||
<emphasis id="B92DEACCFFC0FF8B61F2F8866BC59077" bold="true" box="[199,736,1815,1840]" pageId="7" pageNumber="194">Comparison with other species of the genus.</emphasis>
|
||
<taxonomicName id="4C594D5DFFC0FF8B63DCF8866D4F9068" box="[745,1130,1815,1839]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFC0FF8B63DCF8866D4F9068" box="[745,1130,1815,1839]" italics="true" pageId="7" pageNumber="194">Lepidoblepharis emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from many species in the genus by its small size and its low number of lamellae under the fourth toe and finger (
|
||
<figureCitation id="13622A5BFFC0FF8B6474F8AA698D9030" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="11.[151,250,1918,1940]" captionStartId-1="12.[151,250,792,814]" captionTargetBox-0="[244,1340,193,1892]" captionTargetBox-1="[150,1436,193,770]" captionTargetId-0="figure@11.[244,1340,193,1893]" captionTargetId-1="figure@12.[150,1436,193,771]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." captionText-1="FIGURE 4. Feet and hands of Panamanian Lepidoblepharis. Lepidoblepharis emberawoundule: right foot of SMF 81954 and right hand of MHCH 2957; L. rufigularis (SMF 50659): right foot and right hand; L. sanctaemartae: left foot of SMF 97419 and right hand of MHCH 2948; L. victormartinezi: left foot and left hand of SMF 89963; L. xanthostigma: left foot of SMF 90190 and left hand of SMF 89576. Scale bars equal 1 mm, tip of fourth finger and toe marked with an asterisk." httpUri-0="https://zenodo.org/record/243283/files/figure.png" httpUri-1="https://zenodo.org/record/243284/files/figure.png" pageId="7" pageNumber="194">Figs. 3– 4</figureCitation>
|
||
). In the following, we provide comparisons to all other species within the genus, with the characteristics for
|
||
<taxonomicName id="4C594D5DFFC0FF8B64B0F8F1687D90DC" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFC0FF8B64B0F8F1687D90DC" italics="true" pageId="7" pageNumber="194">L. emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
in parentheses.
|
||
<taxonomicName id="4C594D5DFFC0FF8B632FF8126ABB90DC" box="[538,926,1923,1947]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFC0FF8B632FF8126ABB90DC" box="[538,926,1923,1947]" italics="true" pageId="7" pageNumber="194">Lepidoblepharis emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the Panamanian species
|
||
<taxonomicName id="4C594D5DFFC0FF8B61DBF839688690F8" box="[238,419,1959,1983]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="xanthostigma">
|
||
<emphasis id="B92DEACCFFC0FF8B61DBF839688690F8" box="[238,419,1959,1983]" italics="true" pageId="7" pageNumber="194">L. xanthostigma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B92DEACCFFC0FF8B6084F8396A1C90F8" box="[433,825,1959,1983]" italics="true" pageId="7" pageNumber="194">
|
||
<taxonomicName id="4C594D5DFFC0FF8B6084F8396B5390F8" box="[433,630,1960,1983]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="species" species="sanctaemartae">L. sanctaemartae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFC0FF8B63B4F8366A1C90F8" box="[641,825,1959,1983]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="genus" status="sp. nov.">Lepidoblepharis</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A21E57B7FFC0FF8B6275F8366ABC9087" box="[832,921,1959,1984]" pageId="7" pageNumber="194" rank="species">
|
||
<emphasis id="B92DEACCFFC0FF8B6275F8366ABC9087" bold="true" box="[832,921,1959,1984]" pageId="7" pageNumber="194">
|
||
sp.
|
||
<date id="FFE7101EFFC0FF8B625DF8366ABC9087" box="[872,921,1959,1984]" pageId="7" pageNumber="194">nov.</date>
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
2 (described below), and
|
||
<taxonomicName id="4C594D5DFFC0FF8B6589F8366C5190F8" box="[1212,1396,1959,1983]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="194" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B92DEACCFFC0FF8B6589F8366C5190F8" box="[1212,1396,1959,1983]" italics="true" pageId="7" pageNumber="194">Lepidoblepharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21E57B7FFC0FF8B644FF83669ED90A3" pageId="7" pageNumber="194" rank="species">
|
||
<emphasis id="B92DEACCFFC0FF8B644FF83669ED90A3" bold="true" pageId="7" pageNumber="194">
|
||
sp.
|
||
<date id="FFE7101EFFC0FF8B61A2F85A69ED90A3" box="[151,200,1995,2020]" pageId="7" pageNumber="194">nov.</date>
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
3 (described below) by uncorrected genetic p-distance (10–26% in 16S mtDNA between individuals).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3436555FFCFFF8461A2FF096DCC923F" pageId="8" pageNumber="195" type="reference_group">
|
||
<paragraph id="8BE636DEFFCFFF8461A2FF096DCC923F" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="195">
|
||
<taxonomicName id="4C594D5DFFCFFF8461A2FF0968D197F7" box="[151,500,152,176]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="xanthostigma">
|
||
<emphasis id="B92DEACCFFCFFF8461A2FF0968D197F7" box="[151,500,152,176]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis xanthostigma</emphasis>
|
||
</taxonomicName>
|
||
has greatly enlarged median subcaudal scales (slightly enlarged), and 12–16 lamellae under its fourth toe (6–9).
|
||
<taxonomicName id="4C594D5DFFCFFF846303FF2D6AB89793" box="[566,925,188,212]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="sanctaemartae">
|
||
<emphasis id="B92DEACCFFCFFF846303FF2D6AB89793" box="[566,925,188,212]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis sanctaemartae</emphasis>
|
||
</taxonomicName>
|
||
has large, flat, imbricate dorsal body scales (small granular scales).
|
||
<taxonomicName id="4C594D5DFFCFFF846094FF716B7C97BF" box="[417,601,224,248]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B92DEACCFFCFFF846094FF716B7C97BF" box="[417,601,224,248]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21E57B7FFCFFF84636AFF4E6B9D97BF" box="[607,696,223,248]" pageId="8" pageNumber="195" rank="species">
|
||
<emphasis id="B92DEACCFFCFFF84636AFF4E6B9D97BF" bold="true" box="[607,696,223,248]" pageId="8" pageNumber="195">
|
||
sp.
|
||
<date id="FFE7101EFFCFFF8463B2FF4E6B9D97BF" box="[647,696,223,248]" pageId="8" pageNumber="195">nov.</date>
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
2 (described below) has 13 lamellae under its fourth toe (6–9) and 11 under its fourth finger (5–8).
|
||
<taxonomicName id="4C594D5DFFCFFF84633BFE956BE0965B" box="[526,709,260,284]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B92DEACCFFCFFF84633BFE956BE0965B" box="[526,709,260,284]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21E57B7FFCFFF8463FAFE956A0E965A" box="[719,811,260,285]" pageId="8" pageNumber="195" rank="species">
|
||
<emphasis id="B92DEACCFFCFFF8463FAFE956A0E965A" bold="true" box="[719,811,260,285]" pageId="8" pageNumber="195">
|
||
sp.
|
||
<date id="FFE7101EFFCFFF8463CFFE956A0E965A" box="[762,811,260,285]" pageId="8" pageNumber="195">nov.</date>
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
3 (described below) has 4–5 lamellae under its fourth finger (5–8) and a unique lamellar configuration with 1–3 proximal lamellae per digit greatly enlarged,
|
||
<emphasis id="B92DEACCFFCFFF846419FEB96C739678" box="[1324,1366,296,319]" italics="true" pageId="8" pageNumber="195">i.e.,</emphasis>
|
||
about 3–4 times longer than any of the remaining lamellae, the ventral escutcheon consisting of 61–68 scales (38–61), and no discernible subfemoral escutcheon (4–5 discernible subfemoral escutcheon scales per thigh). To date, seven species of the genus
|
||
<taxonomicName id="4C594D5DFFCFFF84604AFE056B1396EB" box="[383,566,404,428]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92DEACCFFCFFF84604AFE056B1396EB" box="[383,566,404,428]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis</emphasis>
|
||
</taxonomicName>
|
||
have been reported to possess ten or fewer lamellae under the fourth toe,
|
||
<emphasis id="B92DEACCFFCFFF846446FE046CB996EB" box="[1395,1436,405,428]" italics="true" pageId="8" pageNumber="195">i.e.,</emphasis>
|
||
to be short-toed. Two of these,
|
||
<taxonomicName id="4C594D5DFFCFFF846330FE286A2A9697" authority="Lamar 1985" authorityName="Lamar" authorityYear="1985" box="[517,783,439,464]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="miyatai">
|
||
<emphasis id="B92DEACCFFCFFF846330FE286B5F9688" box="[517,634,440,464]" italics="true" pageId="8" pageNumber="195">L. miyatai</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF8463B1FE296A2A9697" author="Lamar" box="[644,783,439,464]" pageId="8" pageNumber="195" refString="Lamar, W. W. (1985) A new Lepidoblepharis (Sauria: Gekkonidae) from the north coast of Colombia. Herpetologica, 41, 128 - 132." type="journal article" year="1985">Lamar 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B92DEACCFFCFFF846279FE286D3D9697" box="[844,1048,441,464]" italics="true" pageId="8" pageNumber="195">
|
||
<taxonomicName id="4C594D5DFFCFFF846279FE286D319697" box="[844,1044,441,464]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="sanctaemartae">L. sanctaemartae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
possess large, flat, and imbricate dorsal scales (small, granular, and juxtaposed dorsals). Three others,
|
||
<taxonomicName id="4C594D5DFFCFFF846292FE4D6C5C96B3" authority="Werner 1910" authorityName="Werner" authorityYear="1910" box="[935,1401,476,500]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="buchwaldi">
|
||
<emphasis id="B92DEACCFFCFFF846292FE4D6DF996B3" box="[935,1244,476,500]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis buchwaldi</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF8465D1FE4C6C5C96B3" author="Werner" box="[1252,1401,476,500]" pageId="8" pageNumber="195" refString="Werner, F. (1910) Uber neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. ii. Eidechsen. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 27, 1 - 46." type="journal article" year="1910">Werner 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF8464B0FE4C6BB6955F" authority="Markezich & Taphorn 1994" authorityName="Markezich & Taphorn" authorityYear="1994" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="montecanoensis">
|
||
<emphasis id="B92DEACCFFCFFF8464B0FE4C686F9550" italics="true" pageId="8" pageNumber="195">L. montecanoensis</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF846066FE6E6BB6955F" author="Markezich" box="[339,659,511,536]" pageId="8" pageNumber="195" refString="Markezich, A. L. & Taphorn, D. C. (1994) A new Lepidoblepharis (Squamata: Gekkonidae) from the Paraguana Peninsula, Venezuela, with comments on its conservation status. Herpetologica, 50, 7 - 14." type="journal article" year="1994">Markezich & Taphorn 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C594D5DFFCFFF8463E6FD906D6B955F" authority="Ayala & Serna 1986" authorityName="Ayala & Serna" authorityYear="1986" box="[723,1102,511,536]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B92DEACCFFCFFF8463E6FD906A7C955F" box="[723,857,512,536]" italics="true" pageId="8" pageNumber="195">L. williamsi</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF846257FE6E6D6B955F" author="Ayala" box="[866,1102,511,536]" pageId="8" pageNumber="195" refString="Ayala, S. C. & Serna, M. A. (1986) Una nueva especie de Lepidoblepharis (Sauria, Gekkonidae) de la Cordillera Central de Colombia. Caldasia, 15, 649 - 654." type="journal article" year="1986">Ayala & Serna 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
, can be readily distinguished from
|
||
<taxonomicName id="4C594D5DFFCFFF8461E3FDB46892957B" box="[214,439,548,572]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFCFFF8461E3FDB46892957B" box="[214,439,548,572]" italics="true" pageId="8" pageNumber="195">L. emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
because the inspection of the illustrations and photographs available for the respective
|
||
<typeStatus id="54E2887CFFCFFF8461A2FDD668219527" box="[151,260,583,608]" pageId="8" pageNumber="195" type="holotype">holotypes</typeStatus>
|
||
showed clear differences between the species: In
|
||
<taxonomicName id="4C594D5DFFCFFF846202FDD86AEF9527" box="[823,970,584,608]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="buchwaldi">
|
||
<emphasis id="B92DEACCFFCFFF846202FDD86AEF9527" box="[823,970,584,608]" italics="true" pageId="8" pageNumber="195">L. buchwaldi</emphasis>
|
||
</taxonomicName>
|
||
, the enlarged subcaudals are much wider than long and at least twice as wide as the laterally adjacent subcaudals (less than twice as wide as they are long or as the neighboring subcaudals are wide), the dorsal tail scales are small,
|
||
<emphasis id="B92DEACCFFCFFF8462FAFD016ADC95E0" box="[975,1017,656,679]" italics="true" pageId="8" pageNumber="195">i.e.,</emphasis>
|
||
less than twice the size of the dorsal body scales (twice or more the size of the dorsal body scales), and the posterior border of the mental has a single median cleft (two paramedian clefts).
|
||
<taxonomicName id="4C594D5DFFCFFF846308FD496A8B95A8" box="[573,942,728,752]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="montecanoensis">
|
||
<emphasis id="B92DEACCFFCFFF846308FD496A8B95A8" box="[573,942,728,752]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis montecanoensis</emphasis>
|
||
</taxonomicName>
|
||
is a very small species with a SVL of
|
||
<quantity id="4CA19B3BFFCFFF846463FD4969E69453" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="8" pageNumber="195" unit="mm" value="19.5" valueMax="21.0" valueMin="18.0">18–21 mm</quantity>
|
||
(21–30), and lacks defined occipital marks in males (two well defined occipital marks in males) as well as distinctly enlarged median subcaudals (median subcaudals distinctly enlarged).
|
||
<taxonomicName id="4C594D5DFFCFFF84650CFCB16C41947F" box="[1081,1380,800,824]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B92DEACCFFCFFF84650CFCB16C41947F" box="[1081,1380,800,824]" italics="true" pageId="8" pageNumber="195">Lepidoblepharis williamsi</emphasis>
|
||
</taxonomicName>
|
||
also lacks enlarged median subcaudal scales (median subcaudals distinctly enlarged), and has only 25–40 ventral escutcheon scales (38–61). The
|
||
<typeStatus id="54E2887CFFCFFF8460CBFCF66B4594C7" box="[510,608,871,896]" pageId="8" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C594D5DFFCFFF8463B0FCF86AE194C7" authority="Boulenger 1908" authorityName="Boulenger" authorityYear="1908" box="[645,964,871,896]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="peraccae">
|
||
<emphasis id="B92DEACCFFCFFF8463B0FCF86A2F94C7" box="[645,778,873,896]" italics="true" pageId="8" pageNumber="195">L. peraccae</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF846224FCF66AE194C7" author="Boulenger" box="[785,964,871,896]" pageId="8" pageNumber="195" refString="Boulenger, G. A. (1908) Descriptions of new South-American reptiles. The Annals and Magazine of Natural History, 8, 111 - 115. http: // dx. doi. org / 10.1080 / 00222930808692365" type="journal article" year="1908">Boulenger 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
has eight lamellae under the fourth finger (5–8) and ten under the fourth toe (6–9), its plantar and palmar scales have ovoid and strongly imbricate posterior borders (those scales small, rounded, and juxtaposed;
|
||
<figureCitation id="13622A5BFFCFFF8463CEFC3E6A65948F" box="[763,832,943,968]" captionStart="FIGURE 7" captionStartId="18.[151,250,1802,1824]" captionTargetBox="[335,1251,193,1780]" captionTargetId="figure@18.[335,1251,193,1780]" captionTargetPageId="18" captionText="FIGURE 7. Comparison between Lepidoblepharis emberawoundule, L. microlepis (holotype), and L. peraccae (holotype). Mental plate and postmental scales in (A) L. microlepis and (B) L. emerawondule; (C) right foot of L. microlepis; (D) right foot of L. emberawoundule; (E) subcaudal scales in L. microlepis, each of the slightly enlarged median subcaudal scales is bordered laterodistally by only one scale, leading to a regular tail sequence of 1 ' 1 '; (F) subcaudal scales in L. emberawoundule, the larger of the enlarged subcaudals are bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1 ' 1 ''; (G) right foot and (H) right hand of the holotype of L. peraccae; scale bars equal 1 mm." httpUri="https://zenodo.org/record/243287/files/figure.png" pageId="8" pageNumber="195">Fig. 7</figureCitation>
|
||
). The
|
||
<typeStatus id="54E2887CFFCFFF8462BFFC3E6AC9948F" box="[906,1004,943,968]" pageId="8" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C594D5DFFCFFF846521FC206C62948F" authority="Noble 1923" authorityName="Noble" authorityYear="1923" box="[1044,1351,943,968]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="microlepis">
|
||
<emphasis id="B92DEACCFFCFFF846521FC206D82948F" box="[1044,1191,944,968]" italics="true" pageId="8" pageNumber="195">L. microlepis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC84B2FFFCFFF846582FC3E6C1A948F" author="Noble" box="[1207,1343,943,968]" pageId="8" pageNumber="195" refString="Noble, G. K. (1923) A new gekkonid lizard and a new brachycephalid frog from Colombia. American Museum Novitates, 88, 1 - 3." type="journal article" year="1923">Noble 1923</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="4C594D5DFFCFFF846039FC4468C894AB" box="[268,493,980,1004]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFCFFF846039FC4468C894AB" box="[268,493,980,1004]" italics="true" pageId="8" pageNumber="195">L. emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
, but differs in the scalation of the chin region and the ventral tail surface (
|
||
<figureCitation id="13622A5BFFCFFF84647DFC456CAB94AB" box="[1352,1422,980,1005]" captionStart="FIGURE 7" captionStartId="18.[151,250,1802,1824]" captionTargetBox="[335,1251,193,1780]" captionTargetId="figure@18.[335,1251,193,1780]" captionTargetPageId="18" captionText="FIGURE 7. Comparison between Lepidoblepharis emberawoundule, L. microlepis (holotype), and L. peraccae (holotype). Mental plate and postmental scales in (A) L. microlepis and (B) L. emerawondule; (C) right foot of L. microlepis; (D) right foot of L. emberawoundule; (E) subcaudal scales in L. microlepis, each of the slightly enlarged median subcaudal scales is bordered laterodistally by only one scale, leading to a regular tail sequence of 1 ' 1 '; (F) subcaudal scales in L. emberawoundule, the larger of the enlarged subcaudals are bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1 ' 1 ''; (G) right foot and (H) right hand of the holotype of L. peraccae; scale bars equal 1 mm." httpUri="https://zenodo.org/record/243287/files/figure.png" pageId="8" pageNumber="195">Fig. 7</figureCitation>
|
||
). The posterior margin of its mental is V-shaped and lacks conspicuous clefts (posterior margin M-shaped,
|
||
<emphasis id="B92DEACCFFCFFF846446FC696CB99348" box="[1395,1436,1016,1039]" italics="true" pageId="8" pageNumber="195">i.e.,</emphasis>
|
||
slightly convex in the middle, with two conspicuous paramedian clefts), there are six postmentals, with one medial postmental greatly enlarged and two neighboring scales slightly enlarged (3–7 postmentals, median scales slightly larger than the others), and the posteriorly adjacent chin scales are small and conical (small and flat, some slightly pointed, and juxtaposed,
|
||
<figureCitation id="13622A5BFFCFFF846085FB1668DB93E7" box="[432,510,1159,1184]" captionStart="FIGURE 3" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[244,1340,193,1892]" captionTargetId="figure@11.[244,1340,193,1893]" captionTargetPageId="11" captionText="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." httpUri="https://zenodo.org/record/243283/files/figure.png" pageId="8" pageNumber="195">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13622A5BFFCFFF84633FFB196B3F93E7" box="[522,538,1160,1184]" captionStart="FIGURE 7" captionStartId="18.[151,250,1802,1824]" captionTargetBox="[335,1251,193,1780]" captionTargetId="figure@18.[335,1251,193,1780]" captionTargetPageId="18" captionText="FIGURE 7. Comparison between Lepidoblepharis emberawoundule, L. microlepis (holotype), and L. peraccae (holotype). Mental plate and postmental scales in (A) L. microlepis and (B) L. emerawondule; (C) right foot of L. microlepis; (D) right foot of L. emberawoundule; (E) subcaudal scales in L. microlepis, each of the slightly enlarged median subcaudal scales is bordered laterodistally by only one scale, leading to a regular tail sequence of 1 ' 1 '; (F) subcaudal scales in L. emberawoundule, the larger of the enlarged subcaudals are bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1 ' 1 ''; (G) right foot and (H) right hand of the holotype of L. peraccae; scale bars equal 1 mm." httpUri="https://zenodo.org/record/243287/files/figure.png" pageId="8" pageNumber="195">7</figureCitation>
|
||
). Most decisively, each of the slightly enlarged subcaudal scales of the
|
||
<typeStatus id="54E2887CFFCFFF84640FFB166CB993E7" box="[1338,1436,1159,1184]" pageId="8" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C594D5DFFCFFF846180FB3C686C9383" box="[181,329,1196,1220]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="microlepis">
|
||
<emphasis id="B92DEACCFFCFFF846180FB3C686C9383" box="[181,329,1196,1220]" italics="true" pageId="8" pageNumber="195">L. microlepis</emphasis>
|
||
</taxonomicName>
|
||
is bordered laterodistally by only one scale, leading to a regular tail sequence of 1'1' (the larger of the enlarged subcaudals bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1'1''; see
|
||
<figureCitation id="13622A5BFFCFFF8460AEFB6568C4924B" box="[411,481,1268,1293]" captionStart="FIGURE 7" captionStartId="18.[151,250,1802,1824]" captionTargetBox="[335,1251,193,1780]" captionTargetId="figure@18.[335,1251,193,1780]" captionTargetPageId="18" captionText="FIGURE 7. Comparison between Lepidoblepharis emberawoundule, L. microlepis (holotype), and L. peraccae (holotype). Mental plate and postmental scales in (A) L. microlepis and (B) L. emerawondule; (C) right foot of L. microlepis; (D) right foot of L. emberawoundule; (E) subcaudal scales in L. microlepis, each of the slightly enlarged median subcaudal scales is bordered laterodistally by only one scale, leading to a regular tail sequence of 1 ' 1 '; (F) subcaudal scales in L. emberawoundule, the larger of the enlarged subcaudals are bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1 ' 1 ''; (G) right foot and (H) right hand of the holotype of L. peraccae; scale bars equal 1 mm." httpUri="https://zenodo.org/record/243287/files/figure.png" pageId="8" pageNumber="195">Fig. 7</figureCitation>
|
||
E–F). The remaining species of the genus,
|
||
<emphasis id="B92DEACCFFCFFF8462D8FB646D32924B" box="[1005,1047,1269,1292]" italics="true" pageId="8" pageNumber="195">i.e.,</emphasis>
|
||
<taxonomicName id="4C594D5DFFCFFF846517FB646C5C924A" authority="Mechler 1968" authorityName="Mechler" authorityYear="1968" box="[1058,1401,1268,1293]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="colombianus">
|
||
<emphasis id="B92DEACCFFCFFF846517FB646DE8924B" box="[1058,1229,1268,1292]" italics="true" pageId="8" pageNumber="195">L colombianus</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF8465EDFB656C5C924A" author="Mechler" box="[1240,1401,1268,1293]" pageId="8" pageNumber="195" refString="Mechler, B. (1968) Les Geckonides de la Colombie. Revue Suisse de Zoologie, 75, 305 - 371." type="journal article" year="1968">Mechler 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF8464B9FB6468ED9277" authority="Avila-Pires 2001" authorityName="Avila-Pires" authorityYear="2001" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="conolepis">
|
||
<emphasis id="B92DEACCFFCFFF8464B9FB6468269277" italics="true" pageId="8" pageNumber="195">L conolepis</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF84603FFA8668ED9277" author="Avila-Pires" box="[266,456,1303,1328]" pageId="8" pageNumber="195" refString="Avila-Pires, T. C. S. (2001) A new species of Lepidoblepharis (Reptilia: Squamata: Gekkonidae) from Ecuador, with a redescription of Lepidoblepharis grandis Miyata, 1985. Occasional Papers of the Sam Noble Oklahoma Museum of Natural History, 11, 1 - 11." type="journal article" year="2001">Avila-Pires 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF8460E2FA886A689277" authority="Ayala & Castro 1983" authorityName="Ayala & Castro" authorityYear="1983" box="[471,845,1303,1328]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="duolepis">
|
||
<emphasis id="B92DEACCFFCFFF8460E2FA886B719277" box="[471,596,1304,1328]" italics="true" pageId="8" pageNumber="195">L. duolepis</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF846368FA866A689277" author="Ayala" box="[605,845,1303,1328]" pageId="8" pageNumber="195" refString="Ayala, S. C. & Castro, F. (1983) Dos nuevos gecos (Sauria: Gekkonidae, Sphaerodactylinae) para Colombia: Lepidoblepharis xanthostigma (Noble) y descripcion de una nueva especie. Caldasia, 13, 743 - 753." type="journal article" year="1983">Ayala & Castro 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF846269FA886D7E9277" authority="Peracca 1897" authorityName="Peracca" authorityYear="1897" box="[860,1115,1304,1328]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="festae">
|
||
<emphasis id="B92DEACCFFCFFF846269FA886A999277" box="[860,956,1304,1328]" italics="true" pageId="8" pageNumber="195">L. festae</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF8462F1FA886D7E9277" author="Peracca" box="[964,1115,1304,1328]" pageId="8" pageNumber="195" refString="Peracca, M. G. (1897) Viaggio del Dr. Enrico Festa nell'Ecuador e regioni vicine. IV. Rettili. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Universita di Torino, 12, 1 - 20." type="journal article" year="1897">Peracca 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF84655FFA886C539277" authority="Miyata 1985" authorityName="Miyata" authorityYear="1985" box="[1130,1398,1303,1328]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="grandis">
|
||
<emphasis id="B92DEACCFFCFFF84655FFA886DFB9277" box="[1130,1246,1304,1328]" italics="true" pageId="8" pageNumber="195">L. grandis</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF8465D2FA866C539277" author="Miyata" box="[1255,1398,1303,1328]" pageId="8" pageNumber="195" refString="Miyata, K. (1985) A new Lepidoblepharis from the Pacific slope of the Ecuadorian Andes (Sauria: Gekkonidae). Herpetologica, 41, 121 - 126." type="journal article" year="1985">Miyata 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF8464B0FA8868E59213" authority="Vanzolini 1978" authorityName="Vanzolini" authorityYear="1978" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="heyerorum">
|
||
<emphasis id="B92DEACCFFCFFF8464B0FA88682A9213" italics="true" pageId="8" pageNumber="195">L. heyerorum</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF846023FAAD68E59213" author="Vanzolini" box="[278,448,1340,1365]" pageId="8" pageNumber="195" refString="Vanzolini, P. E. (1978) Lepidoblepharis in Amazonia (Sauria, Gekkonidae). Papeis Avulsos de Zoologia (Sao Paulo), 31, 203 - 211." type="journal article" year="1978">Vanzolini 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF8460FBFAAC6A0E9212" authority="Avila-Pires 1995" authorityName="Avila-Pires" authorityYear="1995" box="[462,811,1340,1365]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="B92DEACCFFCFFF8460FBFAAC6B4D9213" box="[462,616,1340,1364]" italics="true" pageId="8" pageNumber="195">L. hoogmoedi</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF84635BFAAD6A0E9212" author="Avila-Pires" box="[622,811,1340,1365]" pageId="8" pageNumber="195" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia. (Reptilia: Squamata). Zoologische Verhandelingen Leiden, 299, 1 - 706." type="journal article" year="1995">Avila-Pires 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C594D5DFFCFFF84620DFAAC6DB19213" authority="Boulenger 1914" authorityName="Boulenger" authorityYear="1914" box="[824,1172,1340,1365]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="B92DEACCFFCFFF84620DFAAC6AFF9213" box="[824,986,1340,1364]" italics="true" pageId="8" pageNumber="195">L. intermedius</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF8462D4FAAD6DB19213" author="Boulenger" box="[993,1172,1340,1365]" pageId="8" pageNumber="195" refString="Boulenger, G. A. (1914) On a second collection of batrachians and reptiles made by Dr. H. G. F. Spurrell, F. Z. S. in the Choco, Colombia. Proceedings of the Zoological Society London, 1914, 813 - 817." type="proceedings paper" year="1914">Boulenger 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C594D5DFFCFFF8465E4FAAC69EA923F" authority="Parker 1926" authorityName="Parker" authorityYear="1926" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="ruthveni">
|
||
<emphasis id="B92DEACCFFCFFF8465E4FAAC6C699213" box="[1233,1356,1340,1364]" italics="true" pageId="8" pageNumber="195">L. ruthveni</emphasis>
|
||
<bibRefCitation id="EFC84B2FFFCFFF846466FAAD69EA923F" author="Parker" pageId="8" pageNumber="195" refString="Parker, H. W. (1926) The neotropical lizards of the genera Lepidoblepharis, Pseudogonatodes, Lanthrogecko, and Sphaerodactylus, with the description of a new genus. The Annals and Magazine of Natural History, 17, 291 - 301. http: // dx. doi. org / 10.1080 / 00222932608633413" type="journal article" year="1926">Parker 1926</bibRefCitation>
|
||
</taxonomicName>
|
||
are long-toed with eleven or more lamellae under the fourth toe (
|
||
<quantity id="4CA19B3BFFCFFF846299FACE6AD1923F" box="[940,1012,1375,1400]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="8" pageNumber="195" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9 in</quantity>
|
||
<taxonomicName id="4C594D5DFFCFFF8462CEFAF06DFC923F" box="[1019,1241,1376,1400]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="195" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFCFFF8462CEFAF06DFC923F" box="[1019,1241,1376,1400]" italics="true" pageId="8" pageNumber="195">L. emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3436555FFCFFF8561F2FA156B299117" lastPageId="9" lastPageNumber="196" pageId="8" pageNumber="195" type="description">
|
||
<paragraph id="8BE636DEFFCFFF8561F2FA1568D694C7" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1437,151,1941]" lastPageId="9" lastPageNumber="196" pageId="8" pageNumber="195">
|
||
<emphasis id="B92DEACCFFCFFF8461F2FA156B0492DA" bold="true" box="[199,545,1412,1437]" pageId="8" pageNumber="195">
|
||
Description of the
|
||
<typeStatus id="54E2887CFFCFFF846086FA156B3992DA" box="[435,540,1412,1437]" pageId="8" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Variation among the entire
|
||
<typeStatus id="54E2887CFFCFFF846247FA146A8792DA" box="[882,930,1413,1437]" pageId="8" pageNumber="195">type</typeStatus>
|
||
series is given in parentheses for selected characters (see
|
||
<tableCitation id="C6DB0365FFCFFF846070FA3668BD9287" box="[325,408,1447,1472]" captionStart="TABLE 1" captionStartId="5.[151,239,1362,1385]" captionTargetBox="[159,1427,1452,1887]" captionTargetPageId="5" captionText="TABLE 1. Selected morphological characters among our examined specimens of Central American Lepidoblepharis." httpUri="http://table.plazi.org/id/DF266656FFC2FF8961A2FAC36C59922E" pageId="8" pageNumber="195" tableUuid="DF266656FFC2FF8961A2FAC36C59922E">Table 1</tableCitation>
|
||
for details and variation among all examined specimens). Adult male as judged by everted hemipenes; SVL
|
||
<quantity id="4CA19B3BFFCFFF84606AFA5D689492A3" box="[351,433,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="195" unit="mm" value="26.0">26 mm</quantity>
|
||
(
|
||
<quantity id="4CA19B3BFFCFFF8460F7FA5D6B6692A3" box="[450,579,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.7" metricValueMin="2.1" pageId="8" pageNumber="195" unit="mm" value="24.0" valueMax="27.0" valueMin="21.0">21–27 mm</quantity>
|
||
), TL
|
||
<quantity id="4CA19B3BFFCFFF8463B0FA5D6BF292A3" box="[645,727,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="195" unit="mm" value="27.0">27 mm</quantity>
|
||
(measured while the now broken tail was still intact) (
|
||
<quantity id="4CA19B3BFFCFFF846460FA5D69E2914F" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.7" metricValueMin="1.8" pageId="8" pageNumber="195" unit="mm" value="22.5" valueMax="27.0" valueMin="18.0">18–27 mm</quantity>
|
||
), HL
|
||
<quantity id="4CA19B3BFFCFFF846030FA7E687E9140" box="[261,347,1519,1544]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="8" pageNumber="195" unit="mm" value="5.6">5.6 mm</quantity>
|
||
(5.0–
|
||
<quantity id="4CA19B3BFFCFFF8460A9FA7E68D3914F" box="[412,502,1519,1544]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="8" pageNumber="195" unit="mm" value="5.6">5.6 mm</quantity>
|
||
), HW
|
||
<quantity id="4CA19B3BFFCFFF84630BFA616BB19140" box="[574,660,1520,1544]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="195" unit="mm" value="3.8">3.8 mm</quantity>
|
||
(
|
||
<quantity id="4CA19B3BFFCFFF846397FA616A0B914F" box="[674,814,1520,1544]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="8" pageNumber="195" unit="mm" value="3.8" valueMax="4.4" valueMin="3.2">3.2–4.4 mm</quantity>
|
||
), forelimbs
|
||
<quantity id="4CA19B3BFFCFFF846286FA7E6D2E9140" box="[947,1035,1519,1544]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="8" pageNumber="195" unit="mm" value="5.2">5.2 mm</quantity>
|
||
, hind limbs
|
||
<quantity id="4CA19B3BFFCFFF8465A1FA7E6DC99140" box="[1172,1260,1519,1544]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="195" unit="mm" value="8.6">8.6 mm</quantity>
|
||
, shank
|
||
<quantity id="4CA19B3BFFCFFF84640AFA616CBD9140" box="[1343,1432,1520,1544]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="195" unit="mm" value="3.7">3.7 mm</quantity>
|
||
; rostral large, clearly visible from above, with a shallow, horseshoe-shaped posterior depression and a long median cleft; postrostrals including supranasals three (3–4), the median ones smaller than supranasals, and indenting the rostral, the median postrostral about the same size as the posterior scales on snout; postnasals two (1–2), both about the same size as posteriorly adjacent loreal scales; scales on snout small and smooth, 20 (18–21) scales across snout between anterior sutures of second SPLs; loreal scales juxtaposed, elevated, and rounded, 7 (6–7) on a longitudinal line between postnasals and orbit; scales on top of head small, granular, juxtaposed, generally pointing upward, about as half as large as those on the middle area of snout; superciliary flap with two enlarged scales on anterior border, of which the first is slightly longer than the second, followed by four small scales; supralabials three (3–4), posteriormost one below center of eye; ear-opening small, oval, in oblique orientation; mental large, posterior margin slightly convex in the middle, with two small clefts bordering this convexity, resulting M-shaped; postmentals 7 (4–7), median scales slightly larger than the others, postmentals larger than the posteriorly adjacent chin scales (
|
||
<figureCitation id="13622A5BFFCFFF84601CF831685990FF" box="[297,380,1952,1977]" captionStart="FIGURE 3" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[244,1340,193,1892]" captionTargetId="figure@11.[244,1340,193,1893]" captionTargetPageId="11" captionText="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." httpUri="https://zenodo.org/record/243283/files/figure.png" pageId="8" pageNumber="195">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13622A5BFFCFFF8460B9F83168B990FF" box="[396,412,1952,1976]" captionStart="FIGURE 7" captionStartId="18.[151,250,1802,1824]" captionTargetBox="[335,1251,193,1780]" captionTargetId="figure@18.[335,1251,193,1780]" captionTargetPageId="18" captionText="FIGURE 7. Comparison between Lepidoblepharis emberawoundule, L. microlepis (holotype), and L. peraccae (holotype). Mental plate and postmental scales in (A) L. microlepis and (B) L. emerawondule; (C) right foot of L. microlepis; (D) right foot of L. emberawoundule; (E) subcaudal scales in L. microlepis, each of the slightly enlarged median subcaudal scales is bordered laterodistally by only one scale, leading to a regular tail sequence of 1 ' 1 '; (F) subcaudal scales in L. emberawoundule, the larger of the enlarged subcaudals are bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1 ' 1 ''; (G) right foot and (H) right hand of the holotype of L. peraccae; scale bars equal 1 mm." httpUri="https://zenodo.org/record/243287/files/figure.png" pageId="8" pageNumber="195">7</figureCitation>
|
||
); scales on chin small and juxtaposed, most of them flat but some slightly pointed, on posterior region granular, approximately vertical in position, with a slight reduction in size towards posterior portion of chin; scales near posterior infralabials flat, subimbricate, and larger than scales in median area of chin; infralabials four, first largest, fourth below center of eye; throat with small granular scales, the posterior region with larger, granular, and pointed scales directed upward (some directed posteriorly); dorsal scales on neck and body small, granular, and juxtaposed; dorsals around midbody and on posterior portion of trunk pointed, granular, or, in frontal view, triangular, mostly directed posteriorly, some scales on flanks and in lumbar region slightly flattened; 40 (36–50) middorsal scales in one HL, 89 between levels of axilla and groin; ventrals flat, smooth, imbricate, with an ovoid posterior margin, increasing moderately in size from gular region to belly, posterior ventral scales longer than wide; 12 (11–15) midventral scales in one HL, 27 between levels of axilla and groin, 32 to border of cloaca; ventral escutcheon patch with 53 (38–46) scales, some of which have slightly pointed posterior margins, 7 (6–7) scales long and 13 (10–13) wide, escutcheon long/wide ratio 54% (50–70%); subfemoral escutcheon scales five (4–5) per thigh, arranged in a single row; transition between ventrals and scales on flanks abrupt; scales around midbody 92 (69–92), of which 18 are ventrals (16–19); scales on precloacal plate similar to ventrals, except for those on border of cloaca, which are smaller; tail dorsally and laterally with flattened, smooth, and imbricate scales, less elongated than ventrals (with a transitional zone at base of tail); underside of tail with a median row of moderately enlarged scales, mostly with a repeated series of one median scale bordered laterodistally by one scale, followed by a slightly larger median scale that is in contact laterodistally with two scales (
|
||
<figureCitation id="13622A5BFFCEFF8565B0FD256DF1958B" box="[1157,1236,692,717]" captionStart="FIGURE 3" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[244,1340,193,1892]" captionTargetId="figure@11.[244,1340,193,1893]" captionTargetPageId="11" captionText="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." httpUri="https://zenodo.org/record/243283/files/figure.png" pageId="9" pageNumber="196">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13622A5BFFCEFF8565D5FD256DD5958B" box="[1248,1264,692,716]" captionStart="FIGURE 7" captionStartId="18.[151,250,1802,1824]" captionTargetBox="[335,1251,193,1780]" captionTargetId="figure@18.[335,1251,193,1780]" captionTargetPageId="18" captionText="FIGURE 7. Comparison between Lepidoblepharis emberawoundule, L. microlepis (holotype), and L. peraccae (holotype). Mental plate and postmental scales in (A) L. microlepis and (B) L. emerawondule; (C) right foot of L. microlepis; (D) right foot of L. emberawoundule; (E) subcaudal scales in L. microlepis, each of the slightly enlarged median subcaudal scales is bordered laterodistally by only one scale, leading to a regular tail sequence of 1 ' 1 '; (F) subcaudal scales in L. emberawoundule, the larger of the enlarged subcaudals are bordered laterodistally by two scales, the smaller ones by one, forming a regular tail sequence of 1 ' 1 ''; (G) right foot and (H) right hand of the holotype of L. peraccae; scale bars equal 1 mm." httpUri="https://zenodo.org/record/243287/files/figure.png" pageId="9" pageNumber="196">7</figureCitation>
|
||
), constituting a regular tail sequence of 1'1''; dorsal scales on forelimbs granular; scales on hind limbs flat, smooth, imbricate on anteroventral thigh and shank surfaces, granular elsewhere; fingers, from longest to shortest, IV-III-II-V-I; toes IV- III-II-V-I, fourth and third toes about the same length; lamellae under fourth finger six (5–8), under fourth toe eight (8–9), proximal lamellae slightly larger than distal ones; claws enclosed by an ungual sheath composed of six scales, as typical for the genus.
|
||
</paragraph>
|
||
<paragraph id="8BE636DEFFCEFF8561F2FC1D687D9372" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="196">
|
||
<emphasis id="B92DEACCFFCEFF8561F2FC1D68CD94E2" bold="true" box="[199,488,908,933]" pageId="9" pageNumber="196">Hemipenis morphology.</emphasis>
|
||
The everted hemipenis of SMF 50968 (
|
||
<figureCitation id="13622A5BFFCEFF8562FEFC1D6D3694E2" box="[971,1043,908,933]" captionStart="FIGURE 6" captionStartId="14.[151,250,944,966]" captionTargetBox="[216,1371,176,923]" captionTargetId="figure@14.[216,1371,169,923]" captionTargetPageId="14" captionText="FIGURE 6. Hemipenis morphology of four Lepidoblepharis species from Panama. Sulcate-asulcate view, (A – B) L. emberawoundule SMF 50968; (C – D) L. sanctaemartae SMF 97419 (E – F) L. victormartinezi SMF 50950; (G – H) L. xanthostigma SMF 91558. All scale bars equal 1 mm." httpUri="https://zenodo.org/record/243286/files/figure.png" pageId="9" pageNumber="196">Fig. 6</figureCitation>
|
||
A–B) is a small, bilobate organ, divided for around one third of its length, with a naked base; sulcus spermaticus bordered by well-developed, smooth sulcal lips; ornamentation of papillate calyces present on each lobe, asulcate area of the truncus covered by small spines; a third lobule-like rising from the pedicel, not connected to the sulcus spermaticus, and covered with papillate calyces.
|
||
</paragraph>
|
||
<paragraph id="8BE636DEFFCEFF8561F2FBAE69D6924A" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="196">
|
||
<emphasis id="B92DEACCFFCEFF8561F2FBAE6B25931F" bold="true" box="[199,512,1087,1112]" pageId="9" pageNumber="196">Coloration in preservative</emphasis>
|
||
(alcohol 70%; variation among the
|
||
<typeStatus id="54E2887CFFCEFF8562A7FBD16AD8931F" box="[914,1021,1088,1112]" pageId="9" pageNumber="196" type="paratype">paratypes</typeStatus>
|
||
in parentheses). Dorsal ground color Hair Brown (277); occipital marks Beige (254) (Gray Horn Color (268) in females), posterior margin of orbit bordered with Sepia (279); dorsum of head with small Beige (254) blotches; infra- and supralabials with alternating Sepia (279) and white bars; chin and throat with Sepia (286) marks on a dirty white background; venter slightly pigmented with Vandyke Brown (281); escutcheon scales unpigmented in the center, with Olive Brown (278) borders.
|
||
</paragraph>
|
||
<paragraph id="8BE636DEFFCEFF8561F2FA866B299117" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="196">
|
||
<emphasis id="B92DEACCFFCEFF8561F2FA8668BF9277" bold="true" box="[199,410,1303,1328]" pageId="9" pageNumber="196">Coloration in life</emphasis>
|
||
(
|
||
<figureCitation id="13622A5BFFCEFF85609BFA866B209277" box="[430,517,1303,1328]" captionStart="FIGURE 10" captionStartId="22.[151,250,1971,1993]" captionTargetBox="[188,1398,1299,1948]" captionTargetId="figure@22.[188,1398,1299,1949]" captionTargetPageId="22" captionText="FIGURE 10. Holotype of Lepidoblepharis emberawoundule (male SMF 50968) in life. (A) entire specimen; (B) dorsal view of the head; (C) lateral view of the head." httpUri="https://zenodo.org/record/243290/files/figure.png" pageId="9" pageNumber="196">Fig. 10</figureCitation>
|
||
; variation among the
|
||
<typeStatus id="54E2887CFFCEFF85623DFA886A569276" box="[776,883,1305,1329]" pageId="9" pageNumber="196" type="paratype">paratypes</typeStatus>
|
||
in parentheses). Dorsal ground color Glaucous (272), with small scattered Lavender Blue (195) and Sepia (286) dots; neck region Olive Brown (278); an indistinct Sepia (286) line from tip of snout to anterior border of eye; two diffuse postorbital Sepia (286) lines, one directed towards the occipital region and the other towards the ear; infra- and supralabials with alternating Sepia (279) and white bars; a vaguely M-shaped dirty white (Smoke Gray (267) in females) occipital mark bordered with Burnt Sienna (38); top of head suffused with Fawn Color (258) and Lavender Blue (195); chin and throat with Sepia (286) reticulations on a Chamois (84) background; venter Lavender Blue (195) suffused with Cinnamon-Rufous (31); an indistinct dorsolateral pale line from behind the ear to mid tail; tail Cinnamon-Rufous (31); a Sepia (286) line from above groin to mid tail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3436555FFCEFF8561F2F9CD6D8B91A7" pageId="9" pageNumber="196" type="distribution">
|
||
<paragraph id="8BE636DEFFCEFF8561F2F9CD6D8B91A7" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="196">
|
||
<emphasis id="B92DEACCFFCEFF8561F2F9CD68D39132" bold="true" box="[199,502,1628,1653]" pageId="9" pageNumber="196">Distribution and habitat.</emphasis>
|
||
<taxonomicName id="4C594D5DFFCEFF8560CAF9CD6AA49133" box="[511,897,1628,1652]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="196" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFCEFF8560CAF9CD6AA49133" box="[511,897,1628,1652]" italics="true" pageId="9" pageNumber="196">Lepidoblepharis emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
is currently known from a few sites in eastern Panamanian montane forests and Chocó-Darién moist forests (
|
||
<bibRefCitation id="EFC84B2FFFCEFF856262F9EE6AF091DF" author="Fund" box="[855,981,1663,1688]" pageId="9" pageNumber="196" refString="Fund, W. (2011) Isthmian-Atlantic moist forests. Available from http: // www. eoearth. org / view / article / 153927 (accessed 24 October 2013)" type="journal article" year="2011">Fund 2011</bibRefCitation>
|
||
), from
|
||
<quantity id="4CA19B3BFFCEFF856512F9116D9891DF" box="[1063,1213,1664,1688]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="7.7299999999999995" metricValueMin="2.27" pageId="9" pageNumber="196" unit="m" value="500.0" valueMax="773.0" valueMin="227.0">227 to 773 m</quantity>
|
||
elevation in Darién and
|
||
<collectingCountry id="F34E764EFFCEFF8561F3F935683B91FB" box="[198,286,1700,1724]" name="Panama" pageId="9" pageNumber="196">Panamá</collectingCountry>
|
||
provinces as well as in the Comarcas Emberá and Guna Yala. Most probably,
|
||
<taxonomicName id="4C594D5DFFCEFF8565B6F9346C4491FB" box="[1155,1377,1700,1724]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="196" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFCEFF8565B6F9346C4491FB" box="[1155,1377,1700,1724]" italics="true" pageId="9" pageNumber="196">L. emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
lives in the leaf-litter and feeds on small invertebrates like other
|
||
<taxonomicName id="4C594D5DFFCEFF856218F9596D8F91A7" authority="Vitt et al. 2005" authorityName="Vitt et al." authorityYear="2005" box="[813,1194,1735,1760]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="196" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92DEACCFFCEFF856218F9596AC091A7" box="[813,997,1736,1760]" italics="true" pageId="9" pageNumber="196">Lepidoblepharis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC84B2FFFCEFF8562C6F9566D8791A7" author="Vitt" box="[1011,1186,1735,1760]" pageId="9" pageNumber="196" refString="Vitt, L. J., Sartorius, S. S., Avila-Pires, T. C., Zani, P. A. & Esposito M. C. (2005) Small in a big world: Ecology of leaf-litter geckos in new world tropical forests. Herpetological Monographs, 19, 137 - 152. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2005) 019 [0137: SIABWE] 2.0. CO; 2" type="journal article" year="2005">
|
||
Vitt
|
||
<emphasis id="B92DEACCFFCEFF856513F9586D7391A7" box="[1062,1110,1736,1760]" italics="true" pageId="9" pageNumber="196">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3436555FFCEFF8561F2F97D6B6090D2" pageId="9" pageNumber="196" type="etymology">
|
||
<paragraph id="8BE636DEFFCEFF8561F2F97D6B6090D2" blockId="9.[151,1437,151,1941]" pageId="9" pageNumber="196">
|
||
<emphasis id="B92DEACCFFCEFF8561F2F97D686B9042" bold="true" box="[199,334,1772,1797]" pageId="9" pageNumber="196">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C594D5DFFCEFF8560E3F97D6BB29043" box="[470,663,1772,1796]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="196" phylum="Chordata" rank="species" species="emberawoundule">
|
||
<emphasis id="B92DEACCFFCEFF8560E3F97D6BB29043" box="[470,663,1772,1796]" italics="true" pageId="9" pageNumber="196">emberawoundule</emphasis>
|
||
</taxonomicName>
|
||
is a compound word in honor to “the forest guardians”, the three indigenous peoples inhabiting eastern
|
||
<collectingCountry id="F34E764EFFCEFF85637AF8806B89906F" box="[591,684,1809,1832]" name="Panama" pageId="9" pageNumber="196">Panama</collectingCountry>
|
||
;
|
||
<emphasis id="B92DEACCFFCEFF85638FF8816A2B906F" box="[698,782,1808,1832]" italics="true" pageId="9" pageNumber="196">embera</emphasis>
|
||
: Emberá Indians from the foothills of Jingurudó, Bagre, Sapo, Darién, and Pirre mountain ranges;
|
||
<emphasis id="B92DEACCFFCEFF85635FF8A46B82900B" box="[618,679,1845,1868]" italics="true" pageId="9" pageNumber="196">woun</emphasis>
|
||
: Wounaan Indians, mainly from the Tuira basin and Majé mountain range;
|
||
<emphasis id="B92DEACCFFCEFF8561D0F8C968309037" box="[229,277,1880,1904]" italics="true" pageId="9" pageNumber="196">dule</emphasis>
|
||
: meaning people in the language of the Guna Indians from the Caribbean and Pacific versants of the San Blas and Darién mountain ranges.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |