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<document id="CCBBD3F1307004B7F4F1CA892E882BE4" ID-CLB-Dataset="28049" ID-DOI="10.11646/zootaxa.4508.2.5" ID-GBIF-Dataset="dbbdad6a-4219-463c-a554-98a619774f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607034" ID-ZooBank="75A8DB8B-CEB1-4D53-AE15-ACE397634479" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585821187" checkinUser="plazi" docAuthor="Yu, Daoyuan, Yang, Xiaodong &amp; Liu, Manqiang" docDate="2018" docId="03F087A17A061056FF5CADF2FD5FFD37" docLanguage="en" docName="zootaxa.4508.2.5.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomocerus nan Yu &amp; Yang &amp; Liu 2018, sp. nov." docType="treatment" docVersion="10" lastPageNumber="232" masterDocId="FFC9FFD97A031051FFCBAD6AFFF8FFFA" masterDocTitle="Three new species of Tomocerus from tropical zone of China (Collembola, Tomoceridae)" masterLastPageNumber="236" masterPageNumber="225" pageNumber="230" updateTime="1698713004284" updateUser="ExternalLinkService">
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<mods:title id="080824A7C07875EE13E5648113C38614">Three new species of Tomocerus from tropical zone of China (Collembola, Tomoceridae)</mods:title>
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<mods:namePart id="B6917F491AFAFE363975F4B9155B20C0">Yu, Daoyuan</mods:namePart>
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<mods:namePart id="9A2C8021B760138A1D5EC13DB2899EF1">Yang, Xiaodong</mods:namePart>
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<mods:namePart id="C660AE6B324BBC73B72D48214C61F471">Liu, Manqiang</mods:namePart>
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<mods:date id="95D752D947F3195C59DD33585C3B370E">2018</mods:date>
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<mods:number id="A10CD2430F0471481066B0512FFF2090">2018-10-31</mods:number>
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<treatment id="03F087A17A061056FF5CADF2FD5FFD37" ID-DOI="http://doi.org/10.5281/zenodo.7042570" ID-GBIF-Taxon="154495769" ID-Zenodo-Dep="7042570" LSID="urn:lsid:plazi:treatment:03F087A17A061056FF5CADF2FD5FFD37" httpUri="http://treatment.plazi.org/id/03F087A17A061056FF5CADF2FD5FFD37" lastPageId="7" lastPageNumber="232" pageId="5" pageNumber="230">
<subSubSection id="C343653C7A061054FF5CADF2FE4FFF48" box="[151,439,152,178]" pageId="5" pageNumber="230" type="nomenclature">
<paragraph id="8BE636B77A061054FF5CADF2FE4FFF48" blockId="5.[151,439,152,211]" box="[151,439,152,178]" pageId="5" pageNumber="230">
<heading id="D0AE81DB7A061054FF5CADF2FE4FFF48" bold="true" box="[151,439,152,178]" fontSize="11" level="1" pageId="5" pageNumber="230" reason="1">
<emphasis id="B92DEAA57A061054FF5CADF2FE4FFF48" bold="true" box="[151,439,152,178]" pageId="5" pageNumber="230">
<taxonomicName id="4C594D347A061054FF5CADF2FEAAFF48" authority="Yu &amp; Yang &amp; Liu, 2018" authorityName="Yu &amp; Yang &amp; Liu" authorityYear="2018" box="[151,338,152,178]" class="Collembola" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="5" pageNumber="230" phylum="Arthropoda" rank="species" species="nan" status="sp. nov.">
<emphasis id="B92DEAA57A061054FF5CADF2FEAAFF48" bold="true" box="[151,338,152,178]" italics="true" pageId="5" pageNumber="230">Tomocerus nan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E57DE7A061054FE92ADF2FE4FFF48" box="[345,439,152,178]" pageId="5" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C343653C7A061054FF5CADD0FEDFFF29" box="[151,295,186,211]" pageId="5" pageNumber="230" type="description">
<paragraph id="8BE636B77A061054FF5CADD0FEDFFF29" blockId="5.[151,439,152,211]" box="[151,295,186,211]" pageId="5" pageNumber="230">
<figureCitation id="13622A327A061054FF5CADD0FF08FF29" box="[151,240,186,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,997,1019]" captionTargetBox="[152,1435,663,976]" captionTargetId="figure@2.[152,1435,663,976]" captionTargetPageId="2" captionText="FIGURE 1. Photographs of the species in alcohol (lateral view) (A) Tomocerus tropicus sp. nov.; (B) Tomocerus nan sp. nov.; (C) Tomocerus nabanensis sp. nov. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2607036/files/figure.png" pageId="5" pageNumber="230">Figs 1B</figureCitation>
,
<figureCitation id="13622A327A061054FF36ADD1FEF5FF29" box="[253,269,187,211]" captionStart="FIGURE 4" captionStartId="5.[151,250,1832,1854]" captionTargetBox="[186,1396,471,1789]" captionTargetId="figure@5.[151,1436,443,1810]" captionTargetPageId="5" captionText="FIGURE 4. Tomocerus nan sp. nov. (A) cephalic dorsal chaetotaxy (dorsal view); (B) dorsal chaetotaxy of Th. IIAbd. V (dorsal view); (C) tibiotarsus I, II and III (lateral view); (D) hind claw complex (posterior view); (E) tenaculum (anterior view). Symbols as for Figure 2." httpUri="https://zenodo.org/record/2607042/files/figure.png" pageId="5" pageNumber="230">4</figureCitation>
,
<figureCitation id="13622A327A061054FED2ADD0FEDFFF29" box="[281,295,186,211]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[191,1375,721,1682]" captionTargetId="figure@6.[151,1436,694,1713]" captionTargetPageId="6" captionText="FIGURE 5. Tomocerus nan sp. nov. (A) left side of manubrium (dorsal view); (B) basal part of dens (dorsal view); (C) dorsal and ventral sides of large dental spine; (D) right mucro (outer view). Symbols as for Figures 2 and 3." httpUri="https://zenodo.org/record/2607044/files/figure.png" pageId="5" pageNumber="230">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C343653C7A061054FF5CAC68FD2DFE7D" pageId="5" pageNumber="230" type="materials_examined">
<paragraph id="8BE636B77A061054FF5CAC68FD2DFE7D" blockId="5.[151,1436,258,391]" pageId="5" pageNumber="230">
<emphasis id="B92DEAA57A061054FF5CAC68FEBEFEE1" bold="true" box="[151,326,258,283]" pageId="5" pageNumber="230">Type material.</emphasis>
<materialsCitation id="3B313CEA7A061054FE85AC68FE01FE99" ID-GBIF-Occurrence="2028138847" collectingDate="2015-12-25" collectorName="Daoyuan Yu &amp; Chunyan Qin" country="China" county="Lingshui Autonomous County" elevation="933" latitude="18.727777" location="Mt. Diaoluo National Nature Reserve" longLatPrecision="21" longitude="109.8675" pageId="5" pageNumber="230" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="54E288157A061054FE85AC68FE43FEE1" box="[334,443,258,283]" pageId="5" pageNumber="230" type="holotype">Holotype</typeStatus>
: female on slide,
<location id="8E86606C7A061054FD4EAC69FBCAFEE0" LSID="urn:lsid:plazi:treatment:03F087A17A061056FF5CADF2FD5FFD37:8E86606C7A061054FD4EAC69FBCAFEE0" box="[645,1074,258,283]" country="China" county="Lingshui Autonomous County" latitude="18.727777" longLatPrecision="21" longitude="109.8675" name="Mt. Diaoluo National Nature Reserve" pageId="5" pageNumber="230" stateProvince="Hainan">Mt. Diaoluo National Nature Reserve</location>
,
<collectingCounty id="62874E3B7A061054FBF4AC68FA60FEE1" box="[1087,1432,258,283]" pageId="5" pageNumber="230">Lingshui Autonomous County</collectingCounty>
,
<collectingRegion id="499DF8557A061054FF5CAC4CFEAFFEC5" box="[151,343,294,319]" country="China" name="Hainan" pageId="5" pageNumber="230">Hainan Province</collectingRegion>
,
<collectingCountry id="F34E76277A061054FEA8AC4CFE51FEC5" box="[355,425,294,319]" name="China" pageId="5" pageNumber="230">China</collectingCountry>
,
<geoCoordinate id="EE6D50707A061054FE7EAC4CFDC1FEC5" box="[437,569,294,319]" direction="east" orientation="longitude" pageId="5" pageNumber="230" precision="15" value="109.8675">109°52'3&quot;E</geoCoordinate>
,
<geoCoordinate id="EE6D50707A061054FD8EAC4DFD34FEC5" box="[581,716,295,319]" direction="north" orientation="latitude" pageId="5" pageNumber="230" precision="15" value="18.727777">18°43'40&quot;N</geoCoordinate>
, alt.
<quantity id="4CA19B527A061054FCCFAC4DFCB6FEC4" box="[772,846,295,319]" metricMagnitude="2" metricUnit="m" metricValue="9.33" pageId="5" pageNumber="230" unit="m" value="933.0">
<elevation id="0074D1847A061054FCCFAC4DFCB6FEC4" box="[772,846,295,319]" metricMagnitude="2" metricUnit="m" metricValue="9.33" pageId="5" pageNumber="230" unit="m" value="933.0">933 m</elevation>
</quantity>
,
<date id="FFE710777A061054FC90AC4CFC25FEC5" box="[859,989,294,319]" pageId="5" pageNumber="230" value="2015-12-25">
<collectingDate id="EFA3E99F7A061054FC90AC4CFC25FEC5" box="[859,989,294,319]" pageId="5" pageNumber="230" value="2015-12-25">25.xii.2015</collectingDate>
</date>
, litters and rotten woods, leg.
<collectorName id="26AC53617A061054FAFCAC4DFF4EFE98" pageId="5" pageNumber="230">Daoyuan Yu</collectorName>
and
<collectorName id="26AC53617A061054FF3BAC20FE70FE99" box="[240,392,330,355]" pageId="5" pageNumber="230">Chunyan Qin</collectorName>
(15HN2)
</materialsCitation>
.
<typeStatus id="54E288157A061054FDCEAC21FD8FFE99" box="[517,631,331,355]" pageId="5" pageNumber="230" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5FFD3E7A061054FD48AC21FD0BFE99" box="[643,755,330,355]" pageId="5" pageNumber="230" type="female">2 females</specimenCount>
,
<specimenCount id="9D5FFD3E7A061054FCCBAC21FCA1FE99" box="[768,857,330,355]" pageId="5" pageNumber="230" type="male">3 males</specimenCount>
and
<specimenCount id="9D5FFD3E7A061054FC59AC21FBF0FE99" box="[914,1032,330,355]" pageId="5" pageNumber="230" type="subadult">1 subadult</specimenCount>
on slides,
<materialsCitation id="3B313CEA7A061054FB4EAC21FD29FE7D" ID-GBIF-Occurrence="2028138845" collectingDate="2015-12-25" collectionCode="NJAU" collectorName="Daoyuan Yu &amp; Chunyan Qin" country="China" county="Lingshui Autonomous County" elevation="933" latitude="18.727777" location="Mt. Diaoluo National Nature Reserve" longLatPrecision="21" longitude="109.8675" pageId="5" pageNumber="230" specimenCount="9" specimenCount-adult="9" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9D5FFD3E7A061054FB4EAC21FB27FE99" box="[1157,1247,330,355]" pageId="5" pageNumber="230" type="adult">9 adults</specimenCount>
and juveniles in alcohol, same data as holotype. All types in
<collectionCode id="ED48AE727A061054FD4DAC05FD29FE7D" box="[646,721,367,391]" pageId="5" pageNumber="230">NJAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF26663F7A061054FF5CAA42FE69F881" httpUri="https://zenodo.org/record/2607042/files/figure.png" pageId="5" pageNumber="230" startId="5.[151,250,1832,1854]" targetBox="[186,1396,471,1789]" targetPageId="5">
<paragraph id="8BE636B77A061054FF5CAA42FE69F881" blockId="5.[151,1437,1832,1915]" pageId="5" pageNumber="230">
<emphasis id="B92DEAA57A061054FF5CAA42FEEDF8C7" bold="true" box="[151,277,1832,1854]" pageId="5" pageNumber="230">FIGURE 4.</emphasis>
<taxonomicName id="4C594D347A061054FED5AA40FE43F8C5" authority="Yu &amp; Yang &amp; Liu, 2018" authorityName="Yu &amp; Yang &amp; Liu" authorityYear="2018" box="[286,443,1834,1855]" class="Collembola" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="5" pageNumber="230" phylum="Arthropoda" rank="species" species="nan" status="sp. nov.">
<emphasis id="B92DEAA57A061054FED5AA40FE43F8C5" box="[286,443,1834,1855]" italics="true" pageId="5" pageNumber="230">Tomocerus nan</emphasis>
</taxonomicName>
<emphasis id="B92DEAA57A061054FE0FAA43FDEFF8C4" bold="true" box="[452,535,1833,1854]" pageId="5" pageNumber="230">
<taxonomicNameLabel id="A21E57DE7A061054FE0FAA43FDEFF8C4" box="[452,535,1833,1854]" pageId="5" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) cephalic dorsal chaetotaxy (dorsal view); (B) dorsal chaetotaxy of Th. IIAbd. V (dorsal view); (C) tibiotarsus I, II and III (lateral view); (D) hind claw complex (posterior view); (E) tenaculum (anterior view). Symbols as for Figure 2.
</paragraph>
</caption>
<subSubSection id="C343653C7A051056FF0CADFDFA64FEBA" lastPageId="7" lastPageNumber="232" pageId="6" pageNumber="231" type="description">
<paragraph id="8BE636B77A051057FF0CADFDFE65FF02" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="231">
<emphasis id="B92DEAA57A051057FF0CADFDFEA1FF4A" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="231">Description.</emphasis>
Body length
<quantity id="4CA19B527A051057FE38ADF2FD85FF55" box="[499,637,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.1" metricValueMin="2.2" pageId="6" pageNumber="231" unit="mm" value="2.65" valueMax="3.1" valueMin="2.2">2.23.1 mm</quantity>
. General background colouration of body light yellow. Ant. I and Ant. II distally with light purple pigment, Ant. III and Ant. IV purple. Eye patch black. Tibiotarsi and distal half of hind femur purple (
<figureCitation id="13622A327A051057FEFCADB5FE76FF02" box="[311,398,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,997,1019]" captionTargetBox="[152,1435,663,976]" captionTargetId="figure@2.[152,1435,663,976]" captionTargetPageId="2" captionText="FIGURE 1. Photographs of the species in alcohol (lateral view) (A) Tomocerus tropicus sp. nov.; (B) Tomocerus nan sp. nov.; (C) Tomocerus nabanensis sp. nov. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2607036/files/figure.png" pageId="6" pageNumber="231">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="8BE636B77A051057FF0CAC6EFAC5FE9E" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="231">
Antenna 0.81.0× length of body. Length ratio of Ant I:II:III:IV = 1.0:1.21.3:8.89.8:1.11.3. Cephalic dorsal macrochaetotaxy: anterior area: 2 (A2), 4 (A3, A5); interocular area: 2 (M2), 6 (S2, S5, S
<date id="FFE710777A051057FB42AC4DFB59FEBA" box="[1161,1185,295,320]" pageId="6" pageNumber="231">5i</date>
); postocular area: 2+2 (Pa5, Pa6); posterior area: 1+1 (Pa2). Posterior margin of head with 2035 chaetae on each side (
<figureCitation id="13622A327A051057FB1DAC26FAC9FE9E" box="[1238,1329,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1832,1854]" captionTargetBox="[186,1396,471,1789]" captionTargetId="figure@5.[151,1436,443,1810]" captionTargetPageId="5" captionText="FIGURE 4. Tomocerus nan sp. nov. (A) cephalic dorsal chaetotaxy (dorsal view); (B) dorsal chaetotaxy of Th. IIAbd. V (dorsal view); (C) tibiotarsus I, II and III (lateral view); (D) hind claw complex (posterior view); (E) tenaculum (anterior view). Symbols as for Figure 2." httpUri="https://zenodo.org/record/2607042/files/figure.png" pageId="6" pageNumber="231">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="8BE636B77A051057FF0CAC1AFCA4FE56" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="231">
Pattern of body chaetotaxy as in
<figureCitation id="13622A327A051057FDF9AC05FD70FE72" box="[562,648,367,392]" captionStart="FIGURE 4" captionStartId="5.[151,250,1832,1854]" captionTargetBox="[186,1396,471,1789]" captionTargetId="figure@5.[151,1436,443,1810]" captionTargetPageId="5" captionText="FIGURE 4. Tomocerus nan sp. nov. (A) cephalic dorsal chaetotaxy (dorsal view); (B) dorsal chaetotaxy of Th. IIAbd. V (dorsal view); (C) tibiotarsus I, II and III (lateral view); (D) hind claw complex (posterior view); (E) tenaculum (anterior view). Symbols as for Figure 2." httpUri="https://zenodo.org/record/2607042/files/figure.png" pageId="6" pageNumber="231">Fig. 4B</figureCitation>
. Th. II with macrochaetae a3, a4, a4a and a5a behind anterior marginal macrochaetae cluster; central area with single macrochaeta m3.
</paragraph>
<paragraph id="8BE636B77A051057FF0CACDDFB1DFD7E" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="231">
Trochantero-femoral organ with 1, 1 slender chaetae. Tibiotarsi I, II, III ventrally with 6, 57, 67 strong chaetae, 34, 45, 46 of them blunt (
<figureCitation id="13622A327A051057FD9FACB6FD48FE0E" box="[596,688,476,501]" captionStart="FIGURE 4" captionStartId="5.[151,250,1832,1854]" captionTargetBox="[186,1396,471,1789]" captionTargetId="figure@5.[151,1436,443,1810]" captionTargetPageId="5" captionText="FIGURE 4. Tomocerus nan sp. nov. (A) cephalic dorsal chaetotaxy (dorsal view); (B) dorsal chaetotaxy of Th. IIAbd. V (dorsal view); (C) tibiotarsus I, II and III (lateral view); (D) hind claw complex (posterior view); (E) tenaculum (anterior view). Symbols as for Figure 2." httpUri="https://zenodo.org/record/2607042/files/figure.png" pageId="6" pageNumber="231">Fig. 4C</figureCitation>
). Tenent hair 1.01.1× length of inner edge of unguis; accessory chaetae shorter and thinner than pretarsal chaetae; guard chaetae subequal to tenent hair in length. Unguis slender, with baso-internal ridges about 1/41/3 distance from base; lateral teeth pointed, of moderate size. Inner edge of unguis with basal tooth and 35 more distal teeth, sub-basal tooth slightly stronger than more distal teeth. Unguiculus lanceolate, about 0.600.85× length of unguis, its inner edge with one tooth (
<figureCitation id="13622A327A051057FBB6AF06FB20FD7E" box="[1149,1240,620,645]" captionStart="FIGURE 4" captionStartId="5.[151,250,1832,1854]" captionTargetBox="[186,1396,471,1789]" captionTargetId="figure@5.[151,1436,443,1810]" captionTargetPageId="5" captionText="FIGURE 4. Tomocerus nan sp. nov. (A) cephalic dorsal chaetotaxy (dorsal view); (B) dorsal chaetotaxy of Th. IIAbd. V (dorsal view); (C) tibiotarsus I, II and III (lateral view); (D) hind claw complex (posterior view); (E) tenaculum (anterior view). Symbols as for Figure 2." httpUri="https://zenodo.org/record/2607042/files/figure.png" pageId="6" pageNumber="231">Fig. 4D</figureCitation>
).
</paragraph>
<caption id="DF26663F7A051057FF5CABD1FB6BF915" httpUri="https://zenodo.org/record/2607044/files/figure.png" pageId="6" pageNumber="231" startId="6.[151,250,1723,1745]" targetBox="[191,1375,721,1682]" targetPageId="6">
<paragraph id="8BE636B77A051057FF5CABD1FB6BF915" blockId="6.[151,1436,1723,1775]" pageId="6" pageNumber="231">
<emphasis id="B92DEAA57A051057FF5CABD1FEEBF92A" bold="true" box="[151,275,1723,1745]" pageId="6" pageNumber="231">FIGURE 5.</emphasis>
<taxonomicName id="4C594D347A051057FED2ABD6FE4CF92B" authority="Yu &amp; Yang &amp; Liu, 2018" authorityName="Yu &amp; Yang &amp; Liu" authorityYear="2018" box="[281,436,1724,1745]" class="Collembola" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="231" phylum="Arthropoda" rank="species" species="nan" status="sp. nov.">
<emphasis id="B92DEAA57A051057FED2ABD6FE4CF92B" box="[281,436,1724,1745]" italics="true" pageId="6" pageNumber="231">Tomocerus nan</emphasis>
</taxonomicName>
<emphasis id="B92DEAA57A051057FE70ABD1FDF4F92A" bold="true" box="[443,524,1723,1744]" pageId="6" pageNumber="231">
<taxonomicNameLabel id="A21E57DE7A051057FE70ABD1FDF4F92A" box="[443,524,1723,1744]" pageId="6" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) left side of manubrium (dorsal view); (B) basal part of dens (dorsal view); (C) dorsal and ventral sides of large dental spine; (D) right mucro (outer view). Symbols as for Figures 2 and 3.
</paragraph>
</caption>
<paragraph id="8BE636B77A051056FF0CAA2AFA64FEBA" blockId="6.[151,1436,1856,2025]" lastBlockId="7.[151,1437,151,717]" lastPageId="7" lastPageNumber="232" pageId="6" pageNumber="231">
Ventral tube scaled on both faces. Anterior face with 1420 chaetae on each side, posterior face with 5070 chaetae, each lateral flap unscaled, with 2545 chaetae. Anterior face of tenaculum with 36 chaetae and without scales (
<figureCitation id="13622A327A051057FF25AAE2FEB4F85B" box="[238,332,1928,1953]" captionStart="FIGURE 4" captionStartId="5.[151,250,1832,1854]" captionTargetBox="[186,1396,471,1789]" captionTargetId="figure@5.[151,1436,443,1810]" captionTargetPageId="5" captionText="FIGURE 4. Tomocerus nan sp. nov. (A) cephalic dorsal chaetotaxy (dorsal view); (B) dorsal chaetotaxy of Th. IIAbd. V (dorsal view); (C) tibiotarsus I, II and III (lateral view); (D) hind claw complex (posterior view); (E) tenaculum (anterior view). Symbols as for Figure 2." httpUri="https://zenodo.org/record/2607042/files/figure.png" pageId="6" pageNumber="231">Fig. 4E</figureCitation>
). Ratio of manubrium:dens:mucro = 2.42.9:3.23.7:1.0. Manubrium ventrally scaled without chaetae; laterally with large round scales and 911 chaetae, proximal 12 chaetae slender and almost smooth, distal chaetae strong and slightly serrated; each dorsal chaetal strip with 95120 chaetae of different sizes, an irregular row of scales from base to 1/23/5 length of manubrium along inner edge, and 813 pseudopores on lateral side; without distinct prominent chaetae (
<figureCitation id="13622A327A041056FDF8ADD6FD77FF2F" box="[563,655,188,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[191,1375,721,1682]" captionTargetId="figure@6.[151,1436,694,1713]" captionTargetPageId="6" captionText="FIGURE 5. Tomocerus nan sp. nov. (A) left side of manubrium (dorsal view); (B) basal part of dens (dorsal view); (C) dorsal and ventral sides of large dental spine; (D) right mucro (outer view). Symbols as for Figures 2 and 3." httpUri="https://zenodo.org/record/2607044/files/figure.png" pageId="7" pageNumber="232">Fig. 5A</figureCitation>
); distal corner chaeta as large as small mesochaetae in chaetal strip. Dens basally without prominent dorsal chaeta. Dental spine formula as 36/25,
<emphasis id="B92DEAA57A041056FBFEADB5FBBBFF02" bold="true" box="[1077,1091,223,248]" pageId="7" pageNumber="232">2</emphasis>
(
<figureCitation id="13622A327A041056FB9FADB5FB56FF02" box="[1108,1198,223,248]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[191,1375,721,1682]" captionTargetId="figure@6.[151,1436,694,1713]" captionTargetPageId="6" captionText="FIGURE 5. Tomocerus nan sp. nov. (A) left side of manubrium (dorsal view); (B) basal part of dens (dorsal view); (C) dorsal and ventral sides of large dental spine; (D) right mucro (outer view). Symbols as for Figures 2 and 3." httpUri="https://zenodo.org/record/2607044/files/figure.png" pageId="7" pageNumber="232">Fig. 5B</figureCitation>
); denticles on spines unevenly distributed, all spines with mainly basal moderate to large and a few small denticles, large spines ventrally with additional distal small to moderate denticles (
<figureCitation id="13622A327A041056FCFDAC4DFC77FEBA" box="[822,911,295,320]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[191,1375,721,1682]" captionTargetId="figure@6.[151,1436,694,1713]" captionTargetPageId="6" captionText="FIGURE 5. Tomocerus nan sp. nov. (A) left side of manubrium (dorsal view); (B) basal part of dens (dorsal view); (C) dorsal and ventral sides of large dental spine; (D) right mucro (outer view). Symbols as for Figures 2 and 3." httpUri="https://zenodo.org/record/2607044/files/figure.png" pageId="7" pageNumber="232">Fig. 5C</figureCitation>
). Mucro with 48 intermediate teeth (
<figureCitation id="13622A327A041056FAFDAC4DFA77FEBA" box="[1334,1423,295,320]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[191,1375,721,1682]" captionTargetId="figure@6.[151,1436,694,1713]" captionTargetPageId="6" captionText="FIGURE 5. Tomocerus nan sp. nov. (A) left side of manubrium (dorsal view); (B) basal part of dens (dorsal view); (C) dorsal and ventral sides of large dental spine; (D) right mucro (outer view). Symbols as for Figures 2 and 3." httpUri="https://zenodo.org/record/2607044/files/figure.png" pageId="7" pageNumber="232">Fig. 5D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C343653C7A041056FF0CAC26FB0EFE72" pageId="7" pageNumber="232" type="etymology">
<paragraph id="8BE636B77A041056FF0CAC26FD8BFE9F" blockId="7.[151,1437,151,717]" box="[199,627,332,357]" pageId="7" pageNumber="232">
<emphasis id="B92DEAA57A041056FF0CAC26FEB6FE9F" bold="true" box="[199,334,332,357]" pageId="7" pageNumber="232">Etymology.</emphasis>
Chinese word
<taxonomicName id="4C594D347A041056FE3DAC27FDD8FE9E" authorityName="Yu &amp; Yang &amp; Liu" authorityYear="2018" box="[502,544,333,356]" class="Collembola" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="232" phylum="Arthropoda" rank="species" species="nan">
<emphasis id="B92DEAA57A041056FE3DAC27FDD8FE9E" box="[502,544,333,356]" italics="true" pageId="7" pageNumber="232">nan</emphasis>
</taxonomicName>
: south.
</paragraph>
<paragraph id="8BE636B77A041056FF0CAC05FB0EFE72" blockId="7.[151,1437,151,717]" box="[199,1270,367,392]" pageId="7" pageNumber="232">
<emphasis id="B92DEAA57A041056FF0CAC05FED2FE72" bold="true" box="[199,298,367,392]" pageId="7" pageNumber="232">Habitat.</emphasis>
Mixed forest of
<taxonomicName id="4C594D347A041056FE2DAC05FD4EFE72" authorityName="Hickel" authorityYear="1930" box="[486,694,367,392]" class="Bivalvia" family="Mytilidae" genus="Dacrydium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="232" phylum="Mollusca" rank="species" species="pierrei">
<emphasis id="B92DEAA57A041056FE2DAC05FD4EFE72" box="[486,694,367,392]" italics="true" pageId="7" pageNumber="232">Dacrydium pierrei</emphasis>
</taxonomicName>
and broad leaf trees, in litters and on rotten woods.
</paragraph>
</subSubSection>
<subSubSection id="C343653C7A041056FF0CACFEFD5FFD37" pageId="7" pageNumber="232" type="discussion">
<paragraph id="8BE636B77A041056FF0CACFEFD5FFD37" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="232">
<emphasis id="B92DEAA57A041056FF0CACFEFEC3FE57" bold="true" box="[199,315,404,429]" pageId="7" pageNumber="232">Remarks.</emphasis>
<taxonomicName id="4C594D347A041056FE8FACFFFE0AFE56" authority="Yu &amp; Yang &amp; Liu, 2018" authorityName="Yu &amp; Yang &amp; Liu" authorityYear="2018" box="[324,498,405,428]" class="Collembola" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="232" phylum="Arthropoda" rank="species" species="nan" status="sp. nov.">
<emphasis id="B92DEAA57A041056FE8FACFFFE0AFE56" box="[324,498,405,428]" italics="true" pageId="7" pageNumber="232">Tomocerus nan</emphasis>
</taxonomicName>
<emphasis id="B92DEAA57A041056FE37ACFEFDA0FE57" bold="true" box="[508,600,404,429]" pageId="7" pageNumber="232">
<taxonomicNameLabel id="A21E57DE7A041056FE37ACFEFDA0FE57" box="[508,600,404,429]" pageId="7" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be diagnosed by absence of macrochaetae S0 and Pp3 on the head, presence of only one central macrochaeta m3 on Th. II, absence of distinct prominent chaetae on the furca, presence of dorsal scales on the manubrium and uneven distribution of denticles on large dental spines. The new species resembles most
<taxonomicName id="4C594D347A041056FE61AF6BFD00FDE2" authority="Yu, 2018" authorityName="Yu" authorityYear="2018" box="[426,760,512,536]" class="Collembola" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="232" phylum="Arthropoda" rank="species" species="virgatus">
<emphasis id="B92DEAA57A041056FE61AF6BFD70FDE2" box="[426,648,513,536]" italics="true" pageId="7" pageNumber="232">Tomocerus virgatus</emphasis>
<bibRefCitation id="EFC84B467A041056FD5AAF6AFD00FDE2" author="Yu, D. Y. &amp; Qin, C. Y. &amp; Ding, Y. H. &amp; Hu, F. &amp; Zhang, F. &amp; Liu, M. Q." box="[657,760,512,536]" pageId="7" pageNumber="232" pagination="147 - 172" refId="ref6035" refString="Yu, D. Y., Qin, C. Y., Ding, Y. H., Hu, F., Zhang, F. &amp; Liu, M. Q. (2018) Revealing species diversity of Tomocerus ocreatus complex (Collembola: Tomoceridae): integrative species delimitation and evaluation of taxonomic characters. Arthropod Systematics &amp; Phylogeny, 76 (1), 147 - 172." type="journal article" year="2018">
Yu,
<quantity id="4CA19B527A041056FD0BAF6AFD00FDE2" box="[704,760,512,536]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="7" pageNumber="232" unit="in" value="2018.0">2018</quantity>
</bibRefCitation>
</taxonomicName>
in absence of cephalic macrochaeta S0, presence of single mesothoracic central macrochaeta m3 and absence of prominent chaetae on the furca, but differs from the latter in colour pattern on the body and legs, absence of cephalic macrochaeta Pp3, presence of manubrial dorsal scales and pattern of denticles on large dental spines. Critical characters discriminating
<taxonomicName id="4C594D347A041056FBC9AF07FBB1FD7E" authority="Yu &amp; Yang &amp; Liu, 2018" authorityName="Yu &amp; Yang &amp; Liu" authorityYear="2018" box="[1026,1097,621,644]" class="Collembola" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="232" phylum="Arthropoda" rank="species" species="nan" status="sp. nov.">
<emphasis id="B92DEAA57A041056FBC9AF07FBB1FD7E" box="[1026,1097,621,644]" italics="true" pageId="7" pageNumber="232">T. nan</emphasis>
</taxonomicName>
<emphasis id="B92DEAA57A041056FB99AF06FB55FD7F" bold="true" box="[1106,1197,620,645]" pageId="7" pageNumber="232">
<taxonomicNameLabel id="A21E57DE7A041056FB99AF06FB55FD7F" box="[1106,1197,620,645]" pageId="7" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from similar species are listed in
<tableCitation id="C6DB030C7A041056FED4AFE5FE6BFD52" box="[287,403,655,680]" captionStart="TABLE 1" captionStartId="10.[151,239,152,175]" captionText="TABLE 1. Critical characters that distinguish three new species and the most similar species. *: characters after Denis" pageId="7" pageNumber="232">Table 1. A</tableCitation>
malformed dental spine formula is found on only one side of dens in two individuals as 5/3 4,
<emphasis id="B92DEAA57A041056FF78AFDEFF39FD37" bold="true" box="[179,193,692,717]" pageId="7" pageNumber="232">1</emphasis>
, with proximal spines irregularly arranged.
</paragraph>
</subSubSection>
</treatment>
</document>