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<document id="C54F4E370E037C18B0C1F51F6BCFBDA9" ID-CLB-Dataset="36804" ID-DOI="10.11646/zootaxa.3852.1.1" ID-GBIF-Dataset="2fb83006-f93c-4492-aed0-f5f0707f553e" ID-ISSN="1175-5326" ID-Zenodo-Dep="286956" ID-ZooBank="7DE3BCBA-E5F0-4F0D-B2FD-B5B59E4DAE51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460883007773" checkinUser="plazi" docAuthor="Galea, Horia R., Schories, Dirk, Försterra, Günter &amp; Häussermann, Verena" docDate="2014" docId="03D92A2C4770FFC5FF51868358E0A1C1" docLanguage="en" docName="zt03852p050.pdf" docOrigin="Zootaxa 3852 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrodendron chilense Galea &amp; Schories, sp. nov." docType="treatment" docVersion="15" lastPageNumber="30" masterDocId="FFE052544768FFD8FFC68437590FA657" masterDocTitle="New species and new records of hydroids (Cnidaria: Hydrozoa) from Chile" masterLastPageNumber="50" masterPageNumber="1" pageNumber="25" updateTime="1698380691636" updateUser="ExternalLinkService">
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<mods:title id="D14AB74AEE955672A5BDFB7BB04F90A2">New species and new records of hydroids (Cnidaria: Hydrozoa) from Chile</mods:title>
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<mods:namePart id="03480978E7D8646A1D4E08B33BEBFEC8">Galea, Horia R.</mods:namePart>
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<mods:namePart id="CABB9C2613F258FCFD2FAA678AC6DA77">Häussermann, Verena</mods:namePart>
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<treatment id="03D92A2C4770FFC5FF51868358E0A1C1" ID-DOI="http://doi.org/10.5281/zenodo.6143187" ID-GBIF-Taxon="127669456" ID-Zenodo-Dep="6143187" LSID="urn:lsid:plazi:treatment:03D92A2C4770FFC5FF51868358E0A1C1" httpUri="http://treatment.plazi.org/id/03D92A2C4770FFC5FF51868358E0A1C1" lastPageId="29" lastPageNumber="30" pageId="24" pageNumber="25">
<subSubSection id="C36AC8B14770FFC0FF5186835870A4B8" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BCF9B3A4770FFC0FF5186835A0EA499" blockId="24.[151,769,692,751]" box="[151,769,692,718]" pageId="24" pageNumber="25">
<heading id="D0872C564770FFC0FF5186835A0EA499" bold="true" box="[151,769,692,718]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B90447284770FFC0FF5186835A0EA499" bold="true" box="[151,769,692,718]" pageId="24" pageNumber="25">
<taxonomicName id="4C70E0B94770FFC0FF5186835B99A499" authority="Galea &amp; Schories" authorityName="Galea &amp; Schories" box="[151,662,692,718]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="chilense" status="sp. nov.">
<emphasis id="B90447284770FFC0FF51868358BCA499" bold="true" box="[151,435,692,718]" italics="true" pageId="24" pageNumber="25">Hydrodendron chilense</emphasis>
Galea &amp; Schories
</taxonomicName>
,
<taxonomicNameLabel id="A237FA534770FFC0FD6586835A0EA499" box="[675,769,692,718]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF9B3A4770FFC0FF5186E15870A4B8" blockId="24.[151,769,692,751]" box="[151,383,726,751]" pageId="24" pageNumber="25">
(
<figureCitation id="134B87BF4770FFC0FF5986E15818A4B8" box="[159,279,726,751]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="24" pageNumber="25">Fig. 5AE</figureCitation>
,
<tableCitation id="C6F2AE814770FFC0FEE586E15879A4B8" box="[291,374,726,751]" captionStart="TABLE 5" captionStartId="26.[151,239,152,175]" captionTargetBox="[159,1431,203,414]" captionTargetPageId="26" captionText="TABLE 5. Morphometric comparison of Hydrodendron species (dimensions in µm)." httpUri="http://table.plazi.org/id/DF0FCBB24772FFC2FF5184AF5D10A6F8" pageId="24" pageNumber="25" tableUuid="DF0FCBB24772FFC2FF5184AF5D10A6F8">Table 5</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36AC8B14770FFC0FF5187295A62A5BC" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BCF9B3A4770FFC0FF5187295A62A5BC" blockId="24.[151,1437,798,1903]" pageId="24" pageNumber="25">
<emphasis id="B90447284770FFC0FF5187295873A560" bold="true" box="[151,380,798,823]" pageId="24" pageNumber="25">Material examined</emphasis>
. Corral, Chaihuin/Huiro, lat. -39.95000, long. -73.61667,
<date id="FFCEBDFA4770FFC0FBD987285D9FA560" box="[1055,1168,799,823]" pageId="24" pageNumber="25" value="2011-10-17">17.x.2011</date>
, 8
<quantity id="4C8836DF4770FFC0FB7D87285DF6A561" box="[1211,1273,799,823]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="m" value="12.0">12 m</quantity>
, sample 36: a male colony on seaweed (
<typeStatus id="54CB25984770FFC0FE1587755B34A50C" box="[467,571,834,859]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
: MHNG-INVE-86227).
<collectingCountry id="F367DBAA4770FFC0FC9D87745AA5A50D" box="[859,938,835,858]" name="France" pageId="24" pageNumber="25">France</collectingCountry>
, Port dAlon creek, lat. 43.146369, long. 5.707545,
<date id="FFCEBDFA4770FFC0FED58750588CA528" box="[275,387,871,895]" pageId="24" pageNumber="25" value="2009-05-20">20.v.2009</date>
,
<quantity id="4C8836DF4770FFC0FE52875158D4A529" box="[404,475,870,895]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="0.5">0.5 m</quantity>
, sample HRG-0020: a female colony of
<taxonomicName id="4C70E0B94770FFC0FC0587505C0DA528" authority="Hincks, 1866" authorityName="Hincks" authorityYear="1866" box="[963,1282,870,895]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284770FFC0FC0587505D46A529" box="[963,1097,870,894]" italics="true" pageId="24" pageNumber="25">H. mirabile</emphasis>
(
<bibRefCitation id="EFE1E6CB4770FFC0FB9B87515DF4A528" author="Hincks" box="[1117,1275,870,895]" pageId="24" pageNumber="25" refString="Hincks, T. (1866) On Ophiodes, a new genus of Hydroida. Annals and Magazine of Natural History, Series 3, 18, 421 - 423." type="journal article" year="1866">Hincks, 1866</bibRefCitation>
)
</taxonomicName>
on seaweed.
<collectingCountry id="F367DBAA4770FFC0FF5187BC59E9A5F5" box="[151,230,907,930]" name="France" pageId="24" pageNumber="25">France</collectingCountry>
, Cap Taillat, lat. 43.174040, long. 6.640635,
<date id="FFCEBDFA4770FFC0FD3587BD5A63A5F4" box="[755,876,906,931]" pageId="24" pageNumber="25" value="2011-06-09">09.vi.2011</date>
,
<quantity id="4C8836DF4770FFC0FCBC87BD5AB0A5F5" box="[890,959,906,931]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="0.5">0.5 m</quantity>
, sample HRG-0567: a male colony of
<taxonomicName id="4C70E0B94770FFC0FA4787BC59F9A591" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284770FFC0FA4787BC59F9A591" italics="true" pageId="24" pageNumber="25">H. mirabile</emphasis>
</taxonomicName>
on seaweed.
<collectingCountry id="F367DBAA4770FFC0FE4D879858D5A591" box="[395,474,943,966]" name="France" pageId="24" pageNumber="25">France</collectingCountry>
,
<collectingCountry id="F367DBAA4770FFC0FE2387995B6AA590" box="[485,613,942,967]" name="Martinique" pageId="24" pageNumber="25">Martinique</collectingCountry>
, Anse Dufour, lat. 14.527162, long. -61.090033,
<date id="FFCEBDFA4770FFC0FB4F87995DFBA590" box="[1161,1268,942,967]" pageId="24" pageNumber="25" value="2014-01-12">12.i.2014</date>
,
<quantity id="4C8836DF4770FFC0FAC687995C4DA591" box="[1280,1346,942,967]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="0.5">0.5 m</quantity>
, sample M277: several colonies, some fertile, of
<taxonomicName id="4C70E0B94770FFC0FD9D87E45BD3A5BC" box="[603,732,979,1003]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284770FFC0FD9D87E45BD3A5BC" box="[603,732,979,1003]" italics="true" pageId="24" pageNumber="25">H. mirabile</emphasis>
</taxonomicName>
on seaweed.
</paragraph>
</subSubSection>
<subSubSection id="C36AC8B14770FFC0FF0187C158C9A07C" pageId="24" pageNumber="25" type="description">
<paragraph id="8BCF9B3A4770FFC0FF0187C158C9A07C" blockId="24.[151,1437,798,1903]" pageId="24" pageNumber="25">
<emphasis id="B90447284770FFC0FF0187C1585EA258" bold="true" box="[199,337,1014,1039]" pageId="24" pageNumber="25">Description</emphasis>
. Colony arising from a creeping, branching, anastomosing hydrorhiza; perisarc with irregular, thick, internal projections. Hydrocauli upright but small, not surpassing
<emphasis id="B90447284770FFC0FC03802C5AEFA265" box="[965,992,1051,1074]" italics="true" pageId="24" pageNumber="25">ca</emphasis>
.
<quantity id="4C8836DF4770FFC0FC29802C5D3FA265" box="[1007,1072,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="25" unit="mm" value="3.0">3 mm</quantity>
in height. Stems monosiphonic, divided into internodes by transverse constrictions of the perisarc. Internodes broadly cylindrical, of varied length, each with a proximal bulge, with or without a lateral apophysis supporting the following internode or a hydrophore, and with or without a nematotheca; perisarc moderately thick, surface smooth to locally wrinkled. Branching pattern clearly sympodial, with additional internodes or hydrophores given off at acute angles from the apophyses of preceding internodes; apophyses not in one plane, but arising from any side of the stems. Hydrophores of varied length, with a basal bulge, widening gradually distally to merge into hydrotheca; both separated by a thin diaphragm projecting into the lumen of hydrophore. Hydrotheca tronconical, obviously deep, rim flaring, margin conspicuously everted. Ring of desmocytes irregular, placed at a fair distance above diaphragm. Hydranths reduced to mere buds, possibly due to reproductive exhaustion. Nematothecae irregular and scarce, given off from both the stolon and some stem internodes or hydrophores; beaker shaped, rather large and deep, with a distinctly everted rim; nematophores long (up to
<emphasis id="B90447284770FFC0FE3D81905B19A3E9" box="[507,534,1447,1470]" italics="true" pageId="24" pageNumber="25">ca</emphasis>
.
<quantity id="4C8836DF4770FFC0FDE381905B66A3E8" box="[549,617,1447,1471]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="mm" value="2.0">2 mm</quantity>
) and slender, with solid core and a large, distal capitulum. Gonothecae, male in the present material, borne abundantly on the stolon; elongated tubular for most of their length, tapering abruptly proximally into a short, indistinct pedicel; truncate distally above a narrowed neck; walls provided with 78 distant, spiral grooves.
</paragraph>
</subSubSection>
<subSubSection id="C36AC8B14770FFC1FF0182015A0FA6AF" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8BCF9B3A4770FFC0FF0182015DF8A138" blockId="24.[151,1437,798,1903]" pageId="24" pageNumber="25">
<emphasis id="B90447284770FFC0FF018201583BA018" bold="true" box="[199,308,1590,1615]" pageId="24" pageNumber="25">Remarks</emphasis>
. The new species is characterized, among others, by the presence of spirally twisted gonothecae. Four congeners, namely
<taxonomicName id="4C70E0B94770FFC0FE6C826C5BE4A024" authority="Fraser, 1938" authorityName="Fraser" authorityYear="1938" box="[426,747,1627,1651]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="alternatum">
<emphasis id="B90447284770FFC0FE6C826C5B49A024" box="[426,582,1627,1651]" italics="true" pageId="24" pageNumber="25">H. alternatum</emphasis>
(
<bibRefCitation id="EFE1E6CB4770FFC0FD92826C5BECA024" author="Fraser" box="[596,739,1627,1651]" pageId="24" pageNumber="25" refString="Fraser, C. M. (1938) Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pacific Expeditions, 4 (3), 129 - 152." type="journal article" year="1938">Fraser, 1938</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C70E0B94770FFC0FD30826C5D22A024" authority="Fraser, 1948" authorityName="Fraser" authorityYear="1948" box="[758,1069,1627,1651]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="expansum">
<emphasis id="B90447284770FFC0FD30826C5A87A025" box="[758,904,1627,1650]" italics="true" pageId="24" pageNumber="25">H. expansum</emphasis>
(
<bibRefCitation id="EFE1E6CB4770FFC0FC50826C5D2AA024" author="Fraser" box="[918,1061,1627,1651]" pageId="24" pageNumber="25" refString="Fraser, C. M. (1948) Hydroids of the Allan Hancock Pacific Expeditions since March, 1938. Allan Hancock Pacific Expedition s, 4 (5), 179 - 343." type="journal article" year="1948">Fraser, 1948</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C70E0B94770FFC0FBFE826C5C67A024" authority="Hincks, 1866" authorityName="Hincks" authorityYear="1866" box="[1080,1384,1626,1651]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284770FFC0FBFE826C5DB7A024" box="[1080,1208,1627,1651]" italics="true" pageId="24" pageNumber="25">H. mirabile</emphasis>
(
<bibRefCitation id="EFE1E6CB4770FFC0FB01826D5C6EA024" author="Hincks" box="[1223,1377,1626,1651]" pageId="24" pageNumber="25" refString="Hincks, T. (1866) On Ophiodes, a new genus of Hydroida. Annals and Magazine of Natural History, Series 3, 18, 421 - 423." type="journal article" year="1866">Hincks, 1866</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C70E0B94770FFC0FF5182485B84A0C0" authority="Millard &amp; Bouillon, 1974" authorityName="Millard &amp; Bouillon" authorityYear="1974" box="[151,651,1662,1687]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="sympodiformis">
<emphasis id="B90447284770FFC0FF5182485850A0C1" box="[151,351,1662,1686]" italics="true" pageId="24" pageNumber="25">H. sympodiformis</emphasis>
Millard &amp; Bouillon, 1974
</taxonomicName>
, share the same character. Both
<taxonomicName id="4C70E0B94770FFC0FC3082485D9BA0C1" box="[1014,1172,1662,1686]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="alternatum">
<emphasis id="B90447284770FFC0FC3082485D9BA0C1" box="[1014,1172,1662,1686]" italics="true" pageId="24" pageNumber="25">H. alternatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C70E0B94770FFC0FB0D82485C52A0C1" box="[1227,1373,1663,1686]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="expansum">
<emphasis id="B90447284770FFC0FB0D82485C52A0C1" box="[1227,1373,1663,1686]" italics="true" pageId="24" pageNumber="25">H. expansum</emphasis>
</taxonomicName>
were described from the eastern Pacific. Similarly to
<taxonomicName id="4C70E0B94770FFC0FD6982945A22A0EC" box="[687,813,1699,1723]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="chilense">
<emphasis id="B90447284770FFC0FD6982945A22A0EC" box="[687,813,1699,1723]" italics="true" pageId="24" pageNumber="25">H. chilense</emphasis>
</taxonomicName>
, the internodes of the former are given off sympodially at sharp angles from the preceding internodes, but there is reportedly a characteristic alternate arrangement of the hydrothecae along both sides of the stems and, when present, side branches (
<bibRefCitation id="EFE1E6CB4770FFC0FC2D82DC5D7AA154" author="Fraser" box="[1003,1141,1771,1795]" pageId="24" pageNumber="25" refString="Fraser, C. M. (1938) Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pacific Expeditions, 4 (3), 129 - 152." type="journal article" year="1938">Fraser 1938</bibRefCitation>
), a situation not met in the new species. In addition,
<taxonomicName id="4C70E0B94770FFC0FE0783385B6DA171" box="[449,610,1806,1830]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="alternatum">
<emphasis id="B90447284770FFC0FE0783385B6DA171" box="[449,610,1806,1830]" italics="true" pageId="24" pageNumber="25">H. alternatum</emphasis>
</taxonomicName>
is further distinguished through the apparently exclusive and abundant occurrence of nematothecae on its stems [“commonly one to each pedicel” (
<emphasis id="B90447284770FFC0FC2D83045D05A11D" box="[1003,1034,1843,1866]" italics="true" pageId="24" pageNumber="25">sic</emphasis>
)], and its uncommon ability to give rise to gonothecae, not only from the stolon, but also from the hydrophores and the stem internodes.
</paragraph>
<paragraph id="8BCF9B3A4771FFC1FF0184A05A0FA6AF" blockId="25.[151,1437,151,1304]" pageId="25" pageNumber="26">
Unlike
<taxonomicName id="4C70E0B94771FFC1FED984AE58ADA6E7" box="[287,418,152,176]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="chilense">
<emphasis id="B90447284771FFC1FED984AE58ADA6E7" box="[287,418,152,176]" italics="true" pageId="25" pageNumber="26">H. chilense</emphasis>
</taxonomicName>
,
<taxonomicName id="4C70E0B94771FFC1FE7384AE5B42A6E7" box="[437,589,153,176]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="expansum">
<emphasis id="B90447284771FFC1FE7384AE5B42A6E7" box="[437,589,153,176]" italics="true" pageId="25" pageNumber="26">H. expansum</emphasis>
</taxonomicName>
is a species with polysiphonic stems, collinear internodes delimited by oblique nodes, and large gonothecae with strongly ringed walls, which are given off “from the fascicled portion of the stem, projecting at right angles to it” (
<bibRefCitation id="EFE1E6CB4771FFC1FDAE84D75BFCA6AF" author="Fraser" box="[616,755,224,248]" pageId="25" pageNumber="26" refString="Fraser, C. M. (1948) Hydroids of the Allan Hancock Pacific Expeditions since March, 1938. Allan Hancock Pacific Expedition s, 4 (5), 179 - 343." type="journal article" year="1948">Fraser 1948</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36AC8B14771FFC1FF01853359EEA287" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8BCF9B3A4771FFC1FF0185335AFBA4FA" blockId="25.[151,1437,151,1304]" pageId="25" pageNumber="26">
<taxonomicName id="4C70E0B94771FFC1FF01853358C3A74B" box="[199,460,260,284]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="chilense">
<emphasis id="B90447284771FFC1FF01853358C3A74B" box="[199,460,260,284]" italics="true" pageId="25" pageNumber="26">Hydrodendron chilense</emphasis>
</taxonomicName>
shares several features with
<taxonomicName id="4C70E0B94771FFC1FCCA85325A83A74B" box="[780,908,260,284]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FCCA85325A83A74B" box="[780,908,260,284]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
, among which the sympodial growth pattern of the stems15. The material in hands was compared with three samples that we assign to
<taxonomicName id="4C70E0B94771FFC1FBB0851C5DF6A714" box="[1142,1273,299,323]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FBB0851C5DF6A714" box="[1142,1273,299,323]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
: a male and a female16 colony from the Mediterranean (
<figureCitation id="134B87BF4771FFC1FDB885665BCBA73D" box="[638,708,337,362]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5</figureCitation>
FJ), as well as a male colony from
<collectingCountry id="F367DBAA4771FFC1FB4785665DF2A73D" box="[1153,1277,337,362]" name="Martinique" pageId="25" pageNumber="26">Martinique</collectingCountry>
(
<figureCitation id="134B87BF4771FFC1FAD785665C58A73D" box="[1297,1367,337,362]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5</figureCitation>
L
<collectingCountry id="F367DBAA4771FFC1FABD85665C9FA73D" box="[1403,1424,337,362]" name="Iceland" pageId="25" pageNumber="26">O</collectingCountry>
). Although all exhibit a sympodial growth, in
<taxonomicName id="4C70E0B94771FFC1FD6785415A2DA7DA" box="[673,802,373,397]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="chilense">
<emphasis id="B90447284771FFC1FD6785415A2DA7DA" box="[673,802,373,397]" italics="true" pageId="25" pageNumber="26">H. chilense</emphasis>
</taxonomicName>
the hydrothecae are much deeper and have a strongly everted rim, the nematothecae are bigger and their rim is equally everted, and the gonothecae are more elongated and reach as much as two times the length of their Caribbean and Mediterranean counterparts (see measurements in
<tableCitation id="C6F2AE814771FFC1FF5185D659E4A7AD" box="[151,235,481,506]" captionStart="TABLE 5" captionStartId="26.[151,239,152,175]" captionTargetBox="[159,1431,203,414]" captionTargetPageId="26" captionText="TABLE 5. Morphometric comparison of Hydrodendron species (dimensions in µm)." httpUri="http://table.plazi.org/id/DF0FCBB24772FFC2FF5184AF5D10A6F8" pageId="25" pageNumber="26" tableUuid="DF0FCBB24772FFC2FF5184AF5D10A6F8">Table 5</tableCitation>
). In all available accounts dealing with fertile material [notably
<bibRefCitation id="EFE1E6CB4771FFC1FC7E85D65D9EA7AD" author="Rho" box="[952,1169,481,506]" pageId="25" pageNumber="26" refString="Rho, B. J. &amp; Park, J. H. (1980) A systematic study on the marine hydroids in Korea 6. Thecata. Journal of Korean Research Institute for Better Living, 25, 15 - 43." type="journal article" year="1980">Rho &amp; Park (1980)</bibRefCitation>
; Hirohito (1974, 1995)] attributable, with little doubt, to
<taxonomicName id="4C70E0B94771FFC1FDC586315B8AA44A" box="[515,645,517,541]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FDC586315B8AA44A" box="[515,645,517,541]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
, the gonothecae never reach the huge size seen in the Chilean species. The Vema material studied by
<bibRefCitation id="EFE1E6CB4771FFC1FE31861E5BA0A415" author="Millard" box="[503,687,553,578]" pageId="25" pageNumber="26" refString="Millard, N. A. H. (1966 b) Hydroids of the Vema Seamount. Annals of the South African Museum, 48 (19), 489 - 496." type="journal article" year="1966" yearSuffix="b">Millard (1966b)</bibRefCitation>
, however, possesses gonothecae whose size is comparable to that observed in
<taxonomicName id="4C70E0B94771FFC1FEE7867958AFA432" box="[289,416,589,613]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="chilense">
<emphasis id="B90447284771FFC1FEE7867958AFA432" box="[289,416,589,613]" italics="true" pageId="25" pageNumber="26">H. chilense</emphasis>
</taxonomicName>
, but their walls are only “lightly annulated”. In addition, their trophosome is said “similar to the South African material” that she described earlier as having “the hydrophores alternately to right and left” (
<bibRefCitation id="EFE1E6CB4771FFC1FF5886A25838A4F9" author="Millard" box="[158,311,661,686]" pageId="25" pageNumber="26" refString="Millard, N. A. H. (1957) The Hydrozoa of False Bay, South Africa. Annals of the South African Museum, 43 (4) (6), 173 - 243." type="journal article" year="1957">Millard 1957</bibRefCitation>
), thus differing from the typical growth pattern of
<taxonomicName id="4C70E0B94771FFC1FCAA86A15AE2A4FA" box="[876,1005,661,685]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FCAA86A15AE2A4FA" box="[876,1005,661,685]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCF9B3A4771FFC1FF01868E5B2AA569" blockId="25.[151,1437,151,1304]" pageId="25" pageNumber="26">
<taxonomicName id="4C70E0B94771FFC1FF01868E5B18A486" box="[199,535,697,721]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="sympodiformis">
<emphasis id="B90447284771FFC1FF01868E5B18A486" box="[199,535,697,721]" italics="true" pageId="25" pageNumber="26">Hydrodendron sympodiformis</emphasis>
</taxonomicName>
is almost indistinguishable morphologically from the specimens of
<taxonomicName id="4C70E0B94771FFC1FADC868D5C93A486" box="[1306,1436,697,721]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FADC868D5C93A486" box="[1306,1436,697,721]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
examined by us [compare Millard &amp; Bouillons (1974)
<figureCitation id="134B87BF4771FFC1FCC886EA5A5EA4A1" box="[782,849,733,758]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5</figureCitation>
with our
<figureCitation id="134B87BF4771FFC1FC0786EA5D0CA4A1" box="[961,1027,733,758]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5</figureCitation>
F, I, L,
<collectingCountry id="F367DBAA4771FFC1FB9C86E95D7DA4A1" box="[1114,1138,734,758]" name="Iceland" pageId="25" pageNumber="26">O</collectingCountry>
], and its measurements fit well those of the Mediterranean and Martinican samples (see
<tableCitation id="C6F2AE814771FFC1FC8087365A95A54D" box="[838,922,769,794]" captionStart="TABLE 5" captionStartId="26.[151,239,152,175]" captionTargetBox="[159,1431,203,414]" captionTargetPageId="26" captionText="TABLE 5. Morphometric comparison of Hydrodendron species (dimensions in µm)." httpUri="http://table.plazi.org/id/DF0FCBB24772FFC2FF5184AF5D10A6F8" pageId="25" pageNumber="26" tableUuid="DF0FCBB24772FFC2FF5184AF5D10A6F8">Table 5</tableCitation>
). For this reason, we suspect that it could be a junior synonym of Hincks species.
</paragraph>
<paragraph id="8BCF9B3A4771FFC1FF01877E589AA268" blockId="25.[151,1437,151,1304]" pageId="25" pageNumber="26">
The cnidome composition of all specimens studied here, as well as that of a sterile congener from
<collectingCountry id="F367DBAA4771FFC1FAD3877E5C93A535" box="[1301,1436,841,866]" name="Guadeloupe" pageId="25" pageNumber="26">Guadeloupe</collectingCountry>
(see
<bibRefCitation id="EFE1E6CB4771FFC1FF0A87475852A5DE" author="Galea" box="[204,349,880,905]" pageId="25" pageNumber="26" refString="Galea, H. R. (2010 a) Additional shallow-water thecate hydroids (Cnidaria: Hydrozoa) from Guadeloupe and Les Saintes, French Lesser Antilles. Zootaxa, 2570, 1 - 40." type="journal article" year="2010" yearSuffix="a">Galea 2010a</bibRefCitation>
, p. 13,
<figureCitation id="134B87BF4771FFC1FE7587475B3FA5DE" box="[435,560,880,905]" captionStart="FIGURE 4. A F, M" captionStartId="20.[151,250,1819,1841]" captionTargetBox="[154,1431,248,1754]" captionTargetId="figure@20.[151,1436,243,1755]" captionTargetPageId="20" captionText="FIGURE 4. A F, M: Cnidomes of some Halecium species discussed herein— H. flexile Allman, 1888 (A); H. balei (B); H. humeriformis sp. nov. (C); H. maximum sp. nov. (D); H. tristaniensis sp. nov. (E); H. modestum sp. nov. (F). G: Halecium pallens Jäderholm, 1904 — female gonotheca. H M: Halecium tehuelchum (dOrbigny, 1842) — fragments of colonies (H), primary hydrophore and renovated hydrotheca (I); female gonotheca (J) and shape variation (K); male gonothecae (L); cnidome (M). Scale bars: 10 µm (A F, M); 300 µm (I, J, L), 500 µm (G, K), 1 mm (H)." httpUri="https://zenodo.org/record/286959/files/figure.png" pageId="25" pageNumber="26">Fig. 4AD</figureCitation>
), comprises six capsules: a small (
<figureCitation id="134B87BF4771FFC1FC7C87475D00A5DE" box="[954,1039,880,905]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5E</figureCitation>
3, K3, P3, Q3) and a very large (
<figureCitation id="134B87BF4771FFC1FAA8874759BAA5E7" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5E</figureCitation>
<figureCitation id="134B87BF4771FFC1FF7087A359B1A5F5" box="[182,190,916,930]" captionStart="FIGURE 1. A, B" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[151,1430,198,1707]" captionTargetId="figure@6.[151,1436,195,1707]" captionTargetPageId="6" captionText="FIGURE 1. A, B: Hebella cf. scandens (Bale, 1888) — hydrotheca (A); gonotheca (B). C D: Hebella striata Allman, 1888 — hydrotheca (C); gonotheca (D). E, G, H: Phialella? falklandica Browne, 1902 — portion of erect stem (E) compared with that of P. quadrata (Forbes, 1848) (F); gonotheca seen laterally (G) and apically (H), compared with those of P. quadrata (I, stolonal; J, borne of stem). K, L: Silicularia? bilabiata (Coughtrey, 1876) — two hydrothecae (K); female gonotheca (L). M O: Silicularia rosea Meyen, 1834 — two hydrothecae (M) compared with specimens from Tristan da Cunha assigned to S. hemisphaerica (Allman, 1888) (P); female (N) and male (O) gonothecae, compared to homologous gonothecae (Q and R, respectively) from Tristan. S: Halecium corrugatissimum Trebilcock, 1828 — small stem with female gonotheca. Scale bars: 300 µm (A, B, E, F, S), 500 µm (C, D, G J), and 1 mm (K R)." httpUri="https://zenodo.org/record/286957/files/figure.png" pageId="25" pageNumber="26">1</figureCitation>
, K1, P1, Q1) pseudostenotele, as well as four microbasic mastigophores (
<figureCitation id="134B87BF4771FFC1FC3A87A05D5EA5E7" box="[1020,1105,919,944]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5E</figureCitation>
<figureCitation id="134B87BF4771FFC1FB9787A35D55A5F5" box="[1105,1114,916,930]" captionStart="FIGURE 2. A, B" captionStartId="15.[151,250,1739,1761]" captionTargetBox="[154,1435,200,1713]" captionTargetId="figure@15.[151,1436,195,1713]" captionTargetPageId="15" captionText="FIGURE 2. A, B: Halecium flexile Allman, 1888 — portion of stem (A) compared with H. balei Fraser, 1911 (C); detail of hydrophore and renovated hydrotheca (B), and comparison with H. balei (D). E, F: Halecium humeriformis sp. nov. — stem fragments (E); hydrophore and renovated hydrotheca (F). G, H: Halecium maximum sp. nov. — stem fragments (G); hydrophore and renovated hydrotheca (H). I L: Halecium modestum sp. nov. — stem fragments (I K); hydrophore with hydrotheca (L). M, N: Halecium platythecum sp. nov. — stem fragment (M); hydrophore and renovated hydrotheca (N). Scale bars: 300 µm (B, D, F, H, L, N) and 1 mm (A, C, E, G, I K, M)." httpUri="https://zenodo.org/record/286958/files/figure.png" pageId="25" pageNumber="26">2</figureCitation>
,
<figureCitation id="134B87BF4771FFC1FBA787A35D66A5F5" box="[1121,1129,916,930]" captionStart="FIGURE 4. A F, M" captionStartId="20.[151,250,1819,1841]" captionTargetBox="[154,1431,248,1754]" captionTargetId="figure@20.[151,1436,243,1755]" captionTargetPageId="20" captionText="FIGURE 4. A F, M: Cnidomes of some Halecium species discussed herein— H. flexile Allman, 1888 (A); H. balei (B); H. humeriformis sp. nov. (C); H. maximum sp. nov. (D); H. tristaniensis sp. nov. (E); H. modestum sp. nov. (F). G: Halecium pallens Jäderholm, 1904 — female gonotheca. H M: Halecium tehuelchum (dOrbigny, 1842) — fragments of colonies (H), primary hydrophore and renovated hydrotheca (I); female gonotheca (J) and shape variation (K); male gonothecae (L); cnidome (M). Scale bars: 10 µm (A F, M); 300 µm (I, J, L), 500 µm (G, K), 1 mm (H)." httpUri="https://zenodo.org/record/286959/files/figure.png" pageId="25" pageNumber="26">4</figureCitation>
6, K2, 46, P2, 46, Q2, 46). The shape and size of the capsules appears uniform in all specimens with sympodial colonies (compare
<figureCitation id="134B87BF4771FFC1FB20878C5C34A583" box="[1254,1339,955,980]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5E</figureCitation>
, K &amp; P), while a difference in the size of the large pseudostenoteles (
<figureCitation id="134B87BF4771FFC1FCBD87E85ACBA5AF" box="[891,964,991,1016]" captionStart="FIGURE 5. A E" captionStartId="27.[151,250,1780,1802]" captionTargetBox="[153,1431,197,1758]" captionTargetId="figure@27.[151,1436,194,1759]" captionTargetPageId="27" captionText="FIGURE 5. A E: Hydrodendron chilense sp. nov. — colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). F K: H. mirabile (Hincks, 1866) from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). L P: H. mirabile from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q: Hydrodendron sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L)." httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="25" pageNumber="26">Fig. 5</figureCitation>
Q) could be noted in the species from
<collectingCountry id="F367DBAA4771FFC1FF518034582EA24B" box="[151,289,1027,1052]" name="Guadeloupe" pageId="25" pageNumber="26">Guadeloupe</collectingCountry>
, whose structure of stems is radically different, suggesting that, in some cases, species-specific differences may occur.
</paragraph>
<paragraph id="8BCF9B3A4771FFC1FF01807B59EEA287" blockId="25.[151,1437,151,1304]" pageId="25" pageNumber="26">
It has been shown earlier (Galea
<emphasis id="B90447284771FFC1FDF1807B5B67A234" box="[567,616,1099,1123]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2012) that the
<typeStatus id="54CB25984771FFC1FCDF807B5A5BA233" box="[793,852,1100,1124]" pageId="25" pageNumber="26">types</typeStatus>
of capsules within the genus
<taxonomicName id="4C70E0B94771FFC1FB66807C59C0A2DF" ID-CoL="7PPVL" authority="Bouillon, 1986" authorityName="Bouillon" authorityYear="1986" class="Hydrozoa" family="Haleciidae" genus="Nemalecium" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="genus">
<emphasis id="B90447284771FFC1FB66807C5C23A234" box="[1184,1324,1099,1123]" italics="true" pageId="25" pageNumber="26">Nemalecium</emphasis>
Bouillon, 1986
</taxonomicName>
do not change, but their sizes vary to a certain extent among species. Conversely, in
<taxonomicName id="4C70E0B94771FFC1FB6480585C93A2DF" ID-CoL="7P55M" authority="Oken, 1815" authorityName="Oken" authorityYear="1815" box="[1186,1436,1135,1160]" class="Hydrozoa" family="Haleciidae" genus="Halecium" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="genus">
<emphasis id="B90447284771FFC1FB6480585C00A2D0" box="[1186,1295,1135,1159]" italics="true" pageId="25" pageNumber="26">Halecium</emphasis>
Oken, 1815
</taxonomicName>
there is a significant variation in the shape,
<typeStatus id="54CB25984771FFC1FD4180A35BB8A2FB" box="[647,695,1172,1196]" pageId="25" pageNumber="26">type</typeStatus>
and size of the capsules, as illustrated with the species discussed above.
</paragraph>
</subSubSection>
<subSubSection id="C36AC8B14771FFC1FF0180EC5AC5A2A3" box="[199,970,1243,1268]" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BCF9B3A4771FFC1FF0180EC5AC5A2A3" blockId="25.[151,1437,151,1304]" box="[199,970,1243,1268]" pageId="25" pageNumber="26">
<emphasis id="B90447284771FFC1FF0180EC5849A2A3" bold="true" box="[199,326,1243,1268]" pageId="25" pageNumber="26">Etymology</emphasis>
. The new species is named after the country of its origin.
</paragraph>
</subSubSection>
<subSubSection id="C36AC8B14771FFC5FF0180C858E0A1C1" lastPageId="29" lastPageNumber="30" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BCF9B3A4771FFC1FF0180C85BE4A34F" blockId="25.[151,1437,151,1304]" box="[199,747,1279,1304]" pageId="25" pageNumber="26">
<emphasis id="B90447284771FFC1FF0180C858B1A34F" bold="true" box="[199,446,1279,1304]" pageId="25" pageNumber="26">
Distribution in
<collectingCountry id="F367DBAA4771FFC1FEB880C858B1A34F" box="[382,446,1279,1304]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
</emphasis>
. Only known from Corral.
</paragraph>
<paragraph id="8BCF9B3A4771FFC1FF5181845C94A1E4" blockId="25.[151,1436,1459,2033]" pageId="25" pageNumber="26">
15. The stems of
<taxonomicName id="4C70E0B94771FFC1FE94818358C7A39E" box="[338,456,1459,1481]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FE94818358C7A39E" box="[338,456,1459,1481]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
were originally described as “rudely annulated at the base, and jointed at intervals throughout”, with the “branching […] usually consisting of a single division of the stem”, at most two (
<bibRefCitation id="EFE1E6CB4771FFC1FB5581E65C11A3B0" author="Hincks" box="[1171,1310,1489,1511]" pageId="25" pageNumber="26" refString="Hincks, T. (1866) On Ophiodes, a new genus of Hydroida. Annals and Magazine of Natural History, Series 3, 18, 421 - 423." type="journal article" year="1866">Hincks 1866</bibRefCitation>
). A modern illustration of the
<typeStatus id="54CB25984771FFC1FEBB81C658C2A051" box="[381,461,1521,1542]" pageId="25" pageNumber="26" type="neotype">neotype</typeStatus>
(BMNH 1899.5.1.169) is given by
<bibRefCitation id="EFE1E6CB4771FFC1FCFA81C75D52A051" author="Cornelius" box="[828,1117,1520,1542]" pageId="25" pageNumber="26" refString="Cornelius, P. F. S. (1995 a) North-west European thecate hydroids and their medusae. Part 1. Introduction, Laodiceidae to Haleciidae. Synopses of the British fauna, 50, 1 - 347." type="journal article" year="1995" yearSuffix="a">Cornelius (1995a, Fig. 73C)</bibRefCitation>
, who showed the characteristic sympodial growth pattern of the colonies. The description and illustration of the gonotheca goes back to
<bibRefCitation id="EFE1E6CB4771FFC1FB2882385C8CA072" author="Hincks" box="[1262,1411,1551,1573]" pageId="25" pageNumber="26" refString="Hincks, T. (1868) A history of British hydroid zoophytes. John van Voorst, London, 338 pp." type="book" year="1868">Hincks (1868)</bibRefCitation>
. It is worth underlining that the description of
<taxonomicName id="4C70E0B94771FFC1FD4F82185BF0A014" box="[649,767,1581,1604]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FD4F82185BF0A014" box="[649,767,1581,1604]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFE1E6CB4771FFC1FC4C821A5D38A014" author="Cornelius" box="[906,1079,1581,1603]" pageId="25" pageNumber="26" refString="Cornelius, P. F. S. (1975) A revision of the species of Lafoeidae and Haleciidae (Coelenterata: Hydroida) recorded from Britain and nearby seas. Bulletin of the British Museum (Natural History), Zoology, 28 (8), 373 - 426." type="journal article" year="1975">Cornelius (1975)</bibRefCitation>
resulted undoubtedly from pooled data based, among others, on material dealt with earlier by
<bibRefCitation id="EFE1E6CB4771FFC1FCE7827B5DD9A035" author="Ritchie" box="[801,1238,1612,1634]" pageId="25" pageNumber="26" refString="Ritchie, J. (1908) On collections of the Cape Verde Islands marine fauna, made by Cyril Crossland, MA. (Cantab.), B. Sc. (Lond.), F. Z. S., of St. Andrews University, July to September, 1904. - The Hydroids. Procedings of the Zoological Society of London, 1907, 488 - 514." type="journal article" year="1908">
Ritchie (1908, as
<taxonomicName id="4C70E0B94771FFC1FC13827B5DC2A035" box="[981,1229,1612,1634]" class="Hydrozoa" family="Phylactothecidae" genus="Ophiodissa" higherTaxonomySource="CoL" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="caciniformis">
<emphasis id="B90447284771FFC1FC13827B5DC2A035" box="[981,1229,1612,1634]" italics="true" pageId="25" pageNumber="26">Ophiodissa caciniformis</emphasis>
</taxonomicName>
)
</bibRefCitation>
and
<bibRefCitation id="EFE1E6CB4771FFC1FACE827B5C97A035" author="Millard" box="[1288,1432,1612,1634]" pageId="25" pageNumber="26" refString="Millard, N. A. H. (1957) The Hydrozoa of False Bay, South Africa. Annals of the South African Museum, 43 (4) (6), 173 - 243." type="journal article" year="1957">Millard (1957</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FF01825C5804A0D6" author="Millard" box="[199,267,1643,1665]" pageId="25" pageNumber="26" refString="Millard, N. A. H. (1966 b) Hydroids of the Vema Seamount. Annals of the South African Museum, 48 (19), 489 - 496." type="journal article" year="1966" yearSuffix="b">1966b</bibRefCitation>
). In Cornelius (1975, 1995a) view, widely held by subsequent authors,
<taxonomicName id="4C70E0B94771FFC1FC2F825B5D87A0D6" box="[1001,1160,1643,1665]" class="Hydrozoa" family="Phylactothecidae" genus="Ophiodissa" higherTaxonomySource="CoL" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="caciniformis">
<emphasis id="B90447284771FFC1FC2F825B5D87A0D6" box="[1001,1160,1643,1665]" italics="true" pageId="25" pageNumber="26">H. caciniformis</emphasis>
</taxonomicName>
represents not more than a junior synonym of
<taxonomicName id="4C70E0B94771FFC1FE4F82BD58F1A0C8" box="[393,510,1673,1695]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FE4F82BD58F1A0C8" box="[393,510,1673,1695]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
, and it appears that he mainly focused his attention to the similarities between them, rather than to certain, though not less important differences that are related to their respective growth patterns. Indeed, his illustration (
<bibRefCitation id="EFE1E6CB4771FFC1FE8082F05B59A08A" author="Cornelius" box="[326,598,1735,1757]" pageId="25" pageNumber="26" refString="Cornelius, P. F. S. (1975) A revision of the species of Lafoeidae and Haleciidae (Coelenterata: Hydroida) recorded from Britain and nearby seas. Bulletin of the British Museum (Natural History), Zoology, 28 (8), 373 - 426." type="journal article" year="1975">Cornelius 1975, Fig. 14A</bibRefCitation>
) of the
<typeStatus id="54CB25984771FFC1FD6982F05A38A08A" box="[687,823,1735,1757]" pageId="25" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C70E0B94771FFC1FCA782FF5D0CA08A" box="[865,1027,1735,1757]" class="Hydrozoa" family="Phylactothecidae" genus="Ophiodissa" higherTaxonomySource="CoL" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="caciniformis">
<emphasis id="B90447284771FFC1FCA782FF5D0CA08A" box="[865,1027,1735,1757]" italics="true" pageId="25" pageNumber="26">
<collectingCountry id="F367DBAA4771FFC1FCA782FF5A79A08A" box="[865,886,1736,1757]" name="Iceland" pageId="25" pageNumber="26">O</collectingCountry>
. caciniformis
</emphasis>
</taxonomicName>
shows a species with almost collinear internodes delimited by slightly oblique nodes, thus differing from the sympodial growth pattern characteristic of Hincks species. In addition, the
<typeStatus id="54CB25984771FFC1FE03833258FEA14D" box="[453,497,1797,1818]" pageId="25" pageNumber="26">type</typeStatus>
material of
<taxonomicName id="4C70E0B94771FFC1FDB483325A1DA14D" box="[626,786,1796,1818]" class="Hydrozoa" family="Phylactothecidae" genus="Ophiodissa" higherTaxonomySource="CoL" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="caciniformis">
<emphasis id="B90447284771FFC1FDB483325A1DA14D" box="[626,786,1796,1818]" italics="true" pageId="25" pageNumber="26">H. caciniformis</emphasis>
</taxonomicName>
is sterile. A careful check of the relevant literature shows that specimens with both sympodial [
<emphasis id="B90447284771FFC1FDE683135B4AA16E" box="[544,581,1828,1849]" italics="true" pageId="25" pageNumber="26">e.g.</emphasis>
<bibRefCitation id="EFE1E6CB4771FFC1FD8883145BD7A16E" author="Babic" box="[590,728,1827,1849]" pageId="25" pageNumber="26" refString="Babic, K. (1913) Uber einige Haleciiden. Zoologischer Anzeiger, 41, 468 - 474." type="journal article" year="1913">Babić (1913)</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FD2383145D1AA16E" box="[741,1045,1827,1849]" pageId="25" pageNumber="26" refString="Garcia Corrales, P., Aguirre Inchaurbe, A. &amp; Gonzales Mora, D. (1978) Contribucion al conocimiento de los hidrozoos de las costas espanolas. Parte I: Halecidos, Campanularidos y Plumularidos. Boletin del Instituto Espanol de Oceanografia, 4 (253), 5 - 73." type="journal article">
García Corrales
<emphasis id="B90447284771FFC1FC5683135AB0A16E" box="[912,959,1827,1849]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. (1978)
</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FBE783145DE1A16E" author="Rho" box="[1057,1262,1827,1849]" pageId="25" pageNumber="26" refString="Rho, B. J. &amp; Park, J. H. (1980) A systematic study on the marine hydroids in Korea 6. Thecata. Journal of Korean Research Institute for Better Living, 25, 15 - 43." type="journal article" year="1980">Rho &amp; Park (1980)</bibRefCitation>
, Hirohito (1974, 1995),
<bibRefCitation id="EFE1E6CB4771FFC1FEC883765B85A100" box="[270,650,1857,1879]" pageId="25" pageNumber="26" refString="Ramil Blanco, F. &amp; Iglesias Diaz, A. (1988) La Familia Haleciidae (Cnidaria, Hydrozoa) en las costas de la Galicia. Thalassas, 6, 71 - 78." type="journal article">Ramil Blanco &amp; Iglesias Diaz (1988)</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FD5583765A2AA100" author="Altuna" box="[659,805,1857,1879]" pageId="25" pageNumber="26" refString="Altuna, A. (1994) Estudio faunistico, ecologico y biogeografico de los cnidarios bentonicos de la costa Vasca. PhD thesis. Universidad de Navarra, Pamplona, 769 pp." type="book" year="1994">Altuna (1994)</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FCE883755DD5A100" box="[814,1242,1857,1879]" pageId="25" pageNumber="26" refString="Pena Cantero, A. L. &amp; Garcia Carrascosa, A. M. (2002) The benthic hydroid fauna of the Chafarinas islands (Alboran Sea, western Mediterranean). Zoologische Verhandelingen, Leiden, 337, 1 - 180." type="journal article">Peña Cantero &amp; García Carrascosa (2002)</bibRefCitation>
] and alternate [
<emphasis id="B90447284771FFC1FAB183755C93A100" box="[1399,1436,1858,1879]" italics="true" pageId="25" pageNumber="26">e.g.</emphasis>
<bibRefCitation id="EFE1E6CB4771FFC1FF0183575854A121" author="Millard" box="[199,347,1888,1910]" pageId="25" pageNumber="26" refString="Millard, N. A. H. (1957) The Hydrozoa of False Bay, South Africa. Annals of the South African Museum, 43 (4) (6), 173 - 243." type="journal article" year="1957">Millard (1957</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FEAF835758A4A121" author="Millard" box="[361,427,1888,1910]" pageId="25" pageNumber="26" refString="Millard, N. A. H. (1966 b) Hydroids of the Vema Seamount. Annals of the South African Museum, 48 (19), 489 - 496." type="journal article" year="1966" yearSuffix="b">1966b</bibRefCitation>
),
<bibRefCitation id="EFE1E6CB4771FFC1FE0783575B41A121" author="Ralph" box="[449,590,1888,1910]" pageId="25" pageNumber="26" refString="Ralph, P. M. (1958) New Zealand thecate hydroids. Part II. Families Lafoeidae, Lineolariidae, Haleciidae and Syntheciidae. Transactions of the Royal Society of New Zealand, 85 (2), 301 - 356." type="journal article" year="1958">Ralph (1958)</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FD9A83565A0AA121" author="Vervoort" box="[604,773,1888,1910]" pageId="25" pageNumber="26" refString="Vervoort, W. (1959) The Hydroida of the tropical west coast of Africa. Atlantide Report, 5, 211 - 332." type="journal article" year="1959">Vervoort (1959)</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FCD583575A85A121" author="Gili" box="[787,906,1888,1910]" pageId="25" pageNumber="26" refString="Gili, J. M. (1986) Estudio sistematico y faunistico de los cnidarios de la costa Catalana. PhD thesis. Universitat Autonoma de Barcelona, Barcelona, 565 pp." type="book" year="1986">Gili (1986)</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FC5F83575D64A121" box="[921,1131,1888,1910]" pageId="25" pageNumber="26" refString="Medel, M. D., Garcia, F. G. &amp; Vervoort, W. (1998) The family Haleciidae (Cnidaria: Hydrozoa) from the Strait of Gibraltar and nearby areas. Zoologische Meddedelingen, Leiden, 72 (3), 29 - 50." type="journal article">
Medel
<emphasis id="B90447284771FFC1FC2283565D1AA121" box="[996,1045,1888,1910]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. (1998)
</bibRefCitation>
,
<bibRefCitation id="EFE1E6CB4771FFC1FBBF83565C9AA121" author="Vervoort" box="[1145,1429,1888,1910]" pageId="25" pageNumber="26" refString="Vervoort, W. &amp; Watson, J. E. (2003) The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir, 119, 1 - 538." type="journal article" year="2003">Vervoort &amp; Watson (2003)</bibRefCitation>
] growth patterns were equally included in either
<taxonomicName id="4C70E0B94771FFC1FD6B83B75A2CA1C2" box="[685,803,1919,1941]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284771FFC1FD6B83B75A2CA1C2" box="[685,803,1919,1941]" italics="true" pageId="25" pageNumber="26">H. mirabile</emphasis>
</taxonomicName>
or
<taxonomicName id="4C70E0B94771FFC1FC8383B75AECA1C2" box="[837,995,1919,1941]" class="Hydrozoa" family="Phylactothecidae" genus="Ophiodissa" higherTaxonomySource="CoL" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="26" phylum="Cnidaria" rank="species" species="caciniformis">
<emphasis id="B90447284771FFC1FC8383B75AECA1C2" box="[837,995,1919,1941]" italics="true" pageId="25" pageNumber="26">H. caciniformis</emphasis>
</taxonomicName>
. It therefore becomes obvious that some of the records cited above, especially certain based on sterile material, are either incorrect or belong to unidentifiable species.
</paragraph>
<footnote id="E86B87344771FFC1FF51838B5AF5A1A6" pageId="25" pageNumber="26">
<paragraph id="8BCF9B3A4771FFC1FF51838B5AF5A1A6" blockId="25.[151,1436,1459,2033]" pageId="25" pageNumber="26">16. A sexual dimorphism of the gonothecae could be noted in specimens from the Mediterranean: the females are more slender and more elongate compared to the males (compare Fig. 5J and 5I, respectively).</paragraph>
</footnote>
<caption id="DF0FCBB24772FFC2FF5184AF5D10A6F8" ID-Table-UUID="DF0FCBB24772FFC2FF5184AF5D10A6F8" box="[151,1055,151,175]" httpUri="http://table.plazi.org/id/DF0FCBB24772FFC2FF5184AF5D10A6F8" pageId="26" pageNumber="27" targetBox="[159,1431,203,414]" targetIsTable="true" targetPageId="26">
<paragraph id="8BCF9B3A4772FFC2FF5184AF5D10A6F8" blockId="26.[151,1055,151,175]" box="[151,1055,151,175]" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF5184AF580CA6F8" bold="true" box="[151,259,152,175]" pageId="26" pageNumber="27">TABLE 5</emphasis>
. Morphometric comparison of
<taxonomicName id="4C70E0B94772FFC2FD9784A05BE3A6F8" box="[593,748,151,175]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="genus">
<emphasis id="B90447284772FFC2FD9784A05BE3A6F8" box="[593,748,151,175]" italics="true" pageId="26" pageNumber="27">Hydrodendron</emphasis>
</taxonomicName>
species (dimensions in µm).
</paragraph>
</caption>
<paragraph id="8BCF9B3A4772FFC2FF5984FC5DD3A7C9" pageId="26" pageNumber="27">
<table id="F970699A47720027FF5984FC5C98A7C9" box="[159,1431,203,414]" gridcols="5" gridrows="5" pageId="26" pageNumber="27">
<tr id="3540997847720027FF5984FC5C98A6B6" box="[159,1431,203,225]" gridrow="0" pageId="26" pageNumber="27">
<th id="7691F00447720027FF5984FC5811A6B6" box="[159,286,203,225]" gridcol="0" gridrow="0" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF5984FC59E5A6B7" bold="true" box="[159,234,203,224]" pageId="26" pageNumber="27">Species</emphasis>
</th>
<th id="7691F00447720027FEB384FC58F9A6B6" box="[373,502,203,225]" gridcol="1" gridrow="0" pageId="26" pageNumber="27">
<taxonomicName id="4C70E0B94772FFC2FEB384FC58EAA6B6" box="[373,485,203,225]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="chilense">
<emphasis id="B90447284772FFC2FEB384FC58EAA6B6" bold="true" box="[373,485,203,225]" italics="true" pageId="26" pageNumber="27">H. chilense</emphasis>
</taxonomicName>
</th>
<th id="7691F00447720027FD8A84FC5BD7A6B6" box="[588,728,203,225]" gridcol="2" gridrow="0" pageId="26" pageNumber="27">
<taxonomicName id="4C70E0B94772FFC2FD8A84FC5BB2A6B6" box="[588,701,203,225]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284772FFC2FD8A84FC5BB2A6B6" bold="true" box="[588,701,203,225]" italics="true" pageId="26" pageNumber="27">H. mirabile</emphasis>
</taxonomicName>
</th>
<th id="7691F00447720027FCE484FC5AACA6B6" box="[802,931,203,225]" gridcol="3" gridrow="0" pageId="26" pageNumber="27">
<taxonomicName id="4C70E0B94772FFC2FCE484FC5A9CA6B6" box="[802,915,203,225]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284772FFC2FCE484FC5A9CA6B6" bold="true" box="[802,915,203,225]" italics="true" pageId="26" pageNumber="27">H. mirabile</emphasis>
</taxonomicName>
</th>
<th id="7691F00447720027FC3F84FC5C98A6B6" box="[1017,1431,203,225]" gridcol="4" gridrow="0" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FC3F84FC5C8CA6B6" bold="true" box="[1017,1411,203,225]" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FC3F84FC5C6EA6B6" bold="true" box="[1017,1377,203,225]" italics="true" pageId="26" pageNumber="27">
<taxonomicName id="4C70E0B94772FFC2FC3F84FC5DA6A6B6" box="[1017,1193,203,225]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="sympodiformis">H. sympodiformis</taxonomicName>
<taxonomicName id="4C70E0B94772FFC2FB1684FC5C6EA6B6" box="[1232,1377,203,225]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="genus">Hydrodendron</taxonomicName>
</emphasis>
sp.
</emphasis>
</th>
</tr>
<tr id="3540997847720027FF5984C45C98A771" box="[159,1431,243,294]" gridrow="1" pageId="26" pageNumber="27">
<th id="7691F00447720027FF5984C45811A771" box="[159,286,243,294]" gridcol="0" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF5984C45811A75F" bold="true" box="[159,286,243,264]" pageId="26" pageNumber="27">Reference(s)</emphasis>
</th>
<td id="7691F00447720027FEB384C458F9A771" box="[373,502,243,294]" gridcol="1" gridrow="1" pageId="26" pageNumber="27">Present study</td>
<td id="7691F00447720027FD8A84C45BD7A771" box="[588,728,243,294]" gridcol="2" gridrow="1" pageId="26" pageNumber="27">Present study</td>
<td id="7691F00447720027FCE484C45AACA771" box="[802,931,243,294]" gridcol="3" gridrow="1" pageId="26" pageNumber="27">Present study</td>
<td id="7691F00447720027FC3F84C45C98A771" box="[1017,1431,243,294]" gridcol="4" gridrow="1" pageId="26" pageNumber="27">Millard &amp; Bouillon Galea (2010a), (1974) present study</td>
</tr>
<tr id="3540997847720027FF59850E5C98A719" box="[159,1431,313,334]" gridrow="2" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59850E5811A719" box="[159,286,313,334]" gridcol="0" gridrow="2" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF59850E5813A719" bold="true" box="[159,284,313,334]" pageId="26" pageNumber="27">Distribution</emphasis>
</th>
<td id="7691F00447720027FEB3850E58F9A719" box="[373,502,313,334]" gridcol="1" gridrow="2" pageId="26" pageNumber="27">Chile</td>
<td id="7691F00447720027FD8A850E5BD7A719" box="[588,728,313,334]" gridcol="2" gridrow="2" pageId="26" pageNumber="27">Mediterranean</td>
<td id="7691F00447720027FCE4850E5AACA719" box="[802,931,313,334]" gridcol="3" gridrow="2" pageId="26" pageNumber="27">Martinique</td>
<td id="7691F00447720027FC3F850E5C98A719" box="[1017,1431,313,334]" gridcol="4" gridrow="2" pageId="26" pageNumber="27">Mozambique Guadeloupe</td>
</tr>
<tr id="3540997847720027FF5985565C98A721" box="[159,1431,353,374]" gridrow="3" pageId="26" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7691F00447720027FF5985565811A721" box="[159,286,353,374]" gridcol="0" gridrow="3" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF5985565802A721" bold="true" box="[159,269,353,374]" pageId="26" pageNumber="27">Internodes</emphasis>
</th>
</tr>
<tr id="3540997847720027FF5985BE5C98A7C9" box="[159,1431,393,414]" gridrow="4" pageId="26" pageNumber="27">
<th id="7691F00447720027FF5985BE5811A7C9" box="[159,286,393,414]" gridcol="0" gridrow="4" pageId="26" pageNumber="27">- length</th>
<td id="7691F00447720027FEB385BE58F9A7C9" box="[373,502,393,414]" gridcol="1" gridrow="4" pageId="26" pageNumber="27">100315</td>
<td id="7691F00447720027FD8A85BE5BD7A7C9" box="[588,728,393,414]" gridcol="2" gridrow="4" pageId="26" pageNumber="27">120405</td>
<td id="7691F00447720027FCE485BE5AACA7C9" box="[802,931,393,414]" gridcol="3" gridrow="4" pageId="26" pageNumber="27">115270</td>
<td id="7691F00447720027FC3F85BE5C98A7C9" box="[1017,1431,393,414]" gridcol="4" gridrow="4" pageId="26" pageNumber="27"> </td>
</tr>
</table>
</paragraph>
<paragraph id="8BCF9B3A4772FFC2FF5985865802A25E" pageId="26" pageNumber="27">
<table id="F970699A47720027FF5985865C98A25E" box="[159,1431,433,1033]" gridcols="5" gridrows="14" pageId="26" pageNumber="27">
<tr id="3540997847720027FF5985865C98A791" box="[159,1431,433,454]" gridrow="0" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59858658C6A791" box="[159,457,433,454]" gridcol="0" gridrow="0" pageId="26" pageNumber="27">- diameter at node 75105</th>
<th id="7691F00447720027FD8A85865BAEA791" box="[588,673,433,454]" gridcol="1" gridrow="0" pageId="26" pageNumber="27">7090</th>
<th id="7691F00447720027FCE485865A78A791" box="[802,887,433,454]" gridcol="2" gridrow="0" pageId="26" pageNumber="27">7090</th>
<th id="7691F00447720027FC3F85865D43A791" box="[1017,1100,433,454]" gridcol="3" gridrow="0" pageId="26" pageNumber="27"></th>
<th id="7691F00447720027FB1685865C98A791" box="[1232,1431,433,454]" gridcol="4" gridrow="0" pageId="26" pageNumber="27"></th>
</tr>
<tr id="3540997847720027FF5985EE5C98A7B9" box="[159,1431,473,494]" gridrow="1" pageId="26" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7691F00447720027FF5985EE58C6A7B9" box="[159,457,473,494]" gridcol="0" gridrow="1" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF5985EE5805A7B9" bold="true" box="[159,266,473,494]" pageId="26" pageNumber="27">Apophysis</emphasis>
</th>
</tr>
<tr id="3540997847720027FF5986365C98A441" box="[159,1431,513,534]" gridrow="2" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59863658C6A441" box="[159,457,513,534]" gridcol="0" gridrow="2" pageId="26" pageNumber="27">- length 110195</th>
<td id="7691F00447720027FD8A86365BAEA441" box="[588,673,513,534]" gridcol="1" gridrow="2" pageId="26" pageNumber="27">75125</td>
<td id="7691F00447720027FCE486365A78A441" box="[802,887,513,534]" gridcol="2" gridrow="2" pageId="26" pageNumber="27">65120</td>
<td id="7691F00447720027FC3F86365D43A441" box="[1017,1100,513,534]" gridcol="3" gridrow="2" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB1686365C98A441" box="[1232,1431,513,534]" gridcol="4" gridrow="2" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF59861E5C98A469" box="[159,1431,553,574]" gridrow="3" pageId="26" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7691F00447720027FF59861E58C6A469" box="[159,457,553,574]" gridcol="0" gridrow="3" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF59861E5812A469" bold="true" box="[159,285,553,574]" pageId="26" pageNumber="27">Hydrophore</emphasis>
</th>
</tr>
<tr id="3540997847720027FF5986665C98A431" box="[159,1431,593,614]" gridrow="4" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59866658C6A431" box="[159,457,593,614]" gridcol="0" gridrow="4" pageId="26" pageNumber="27">- length 85415</th>
<td id="7691F00447720027FD8A86665BAEA431" box="[588,673,593,614]" gridcol="1" gridrow="4" pageId="26" pageNumber="27">95145</td>
<td id="7691F00447720027FCE486665A78A431" box="[802,887,593,614]" gridcol="2" gridrow="4" pageId="26" pageNumber="27">205560</td>
<td id="7691F00447720027FC3F86665D43A431" box="[1017,1100,593,614]" gridcol="3" gridrow="4" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB1686665C98A431" box="[1232,1431,593,614]" gridcol="4" gridrow="4" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF59864E5C98A4D9" box="[159,1431,633,654]" gridrow="5" pageId="26" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7691F00447720027FF59864E58C6A4D9" box="[159,457,633,654]" gridcol="0" gridrow="5" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF59864E5817A4D9" bold="true" box="[159,280,633,654]" pageId="26" pageNumber="27">Hydrotheca</emphasis>
</th>
</tr>
<tr id="3540997847720027FF5986965C98A4E1" box="[159,1431,673,694]" gridrow="6" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59869658C6A4E1" box="[159,457,673,694]" gridcol="0" gridrow="6" pageId="26" pageNumber="27">- height 90130</th>
<td id="7691F00447720027FD8A86965BAEA4E1" box="[588,673,673,694]" gridcol="1" gridrow="6" pageId="26" pageNumber="27">6085</td>
<td id="7691F00447720027FCE486965A78A4E1" box="[802,887,673,694]" gridcol="2" gridrow="6" pageId="26" pageNumber="27">3040</td>
<td id="7691F00447720027FC3F86965D43A4E1" box="[1017,1100,673,694]" gridcol="3" gridrow="6" pageId="26" pageNumber="27">60100</td>
<td id="7691F00447720027FB1686965C98A4E1" box="[1232,1431,673,694]" gridcol="4" gridrow="6" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF5986FE5C98A489" box="[159,1431,713,734]" gridrow="7" pageId="26" pageNumber="27">
<th id="7691F00447720027FF5986FE58C6A489" box="[159,457,713,734]" gridcol="0" gridrow="7" pageId="26" pageNumber="27">- diameter at rim 225280</th>
<td id="7691F00447720027FD8A86FE5BAEA489" box="[588,673,713,734]" gridcol="1" gridrow="7" pageId="26" pageNumber="27">195220</td>
<td id="7691F00447720027FCE486FE5A78A489" box="[802,887,713,734]" gridcol="2" gridrow="7" pageId="26" pageNumber="27">180205</td>
<td id="7691F00447720027FC3F86FE5D43A489" box="[1017,1100,713,734]" gridcol="3" gridrow="7" pageId="26" pageNumber="27">170220</td>
<td id="7691F00447720027FB1686FE5C98A489" box="[1232,1431,713,734]" gridcol="4" gridrow="7" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF5986C65C98A574" box="[159,1431,753,803]" gridrow="8" pageId="26" pageNumber="27">
<th id="7691F00447720027FF5986C658C6A574" box="[159,457,753,803]" gridcol="0" gridrow="8" pageId="26" pageNumber="27">- diameter at 110160 diaphragm</th>
<td id="7691F00447720027FD8A86C65BAEA574" box="[588,673,753,803]" gridcol="1" gridrow="8" pageId="26" pageNumber="27">120130</td>
<td id="7691F00447720027FCE486C65A78A574" box="[802,887,753,803]" gridcol="2" gridrow="8" pageId="26" pageNumber="27">135160</td>
<td id="7691F00447720027FC3F86C65D43A574" box="[1017,1100,753,803]" gridcol="3" gridrow="8" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB1686C65C98A574" box="[1232,1431,753,803]" gridcol="4" gridrow="8" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF5987015C98A51C" box="[159,1431,822,843]" gridrow="9" pageId="26" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7691F00447720027FF59870158C6A51C" box="[159,457,822,843]" gridcol="0" gridrow="9" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF598701582AA51C" bold="true" box="[159,293,822,843]" pageId="26" pageNumber="27">Nematotheca</emphasis>
</th>
</tr>
<tr id="3540997847720027FF5987695C98A524" box="[159,1431,862,883]" gridrow="10" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59876958C6A524" box="[159,457,862,883]" gridcol="0" gridrow="10" pageId="26" pageNumber="27">- length 165200</th>
<td id="7691F00447720027FD8A87695BAEA524" box="[588,673,862,883]" gridcol="1" gridrow="10" pageId="26" pageNumber="27">105145</td>
<td id="7691F00447720027FCE487695A78A524" box="[802,887,862,883]" gridcol="2" gridrow="10" pageId="26" pageNumber="27">110150</td>
<td id="7691F00447720027FC3F87695D43A524" box="[1017,1100,862,883]" gridcol="3" gridrow="10" pageId="26" pageNumber="27">140200</td>
<td id="7691F00447720027FB1687695C98A524" box="[1232,1431,862,883]" gridcol="4" gridrow="10" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF5987B15C98A5EE" box="[159,1431,902,953]" gridrow="11" pageId="26" pageNumber="27">
<th id="7691F00447720027FF5987B158C6A5EE" box="[159,457,902,953]" gridcol="0" gridrow="11" pageId="26" pageNumber="27">- diameter at 105135 aperture</th>
<td id="7691F00447720027FD8A87B15BAEA5EE" box="[588,673,902,953]" gridcol="1" gridrow="11" pageId="26" pageNumber="27">7585</td>
<td id="7691F00447720027FCE487B15A78A5EE" box="[802,887,902,953]" gridcol="2" gridrow="11" pageId="26" pageNumber="27">6580</td>
<td id="7691F00447720027FC3F87B15D43A5EE" box="[1017,1100,902,953]" gridcol="3" gridrow="11" pageId="26" pageNumber="27">90110</td>
<td id="7691F00447720027FB1687B15C98A5EE" box="[1232,1431,902,953]" gridcol="4" gridrow="11" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF5987FB5C98A5B6" box="[159,1431,972,993]" gridrow="12" pageId="26" pageNumber="27" rowspan-3="1">
<th id="7691F00447720027FF5987FB58C6A5B6" box="[159,457,972,993]" gridcol="0" gridrow="12" pageId="26" pageNumber="27">
-
<taxonomicName id="4C70E0B94772FFC2FF6B87FB5801A5B6" box="[173,270,972,993]" class="Hydrozoa" family="Haleciidae" genus="Halecium" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="maximum">maximum</taxonomicName>
width 85105
</th>
<td id="7691F00447720027FD8A87FB5BAEA5B6" box="[588,673,972,993]" gridcol="1" gridrow="12" pageId="26" pageNumber="27">5565</td>
<td id="7691F00447720027FCE487FB5A78A5B6" box="[802,887,972,993]" gridcol="2" gridrow="12" pageId="26" pageNumber="27">6065</td>
<td id="7691F00447720027FB1687FB5C98A5B6" box="[1232,1431,972,993]" gridcol="4" gridrow="12" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF5987C35C98A25E" box="[159,1431,1012,1033]" gridrow="13" pageId="26" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7691F00447720027FF5987C358C6A25E" box="[159,457,1012,1033]" gridcol="0" gridrow="13" pageId="26" pageNumber="27">Gonotheca</th>
</tr>
</table>
</paragraph>
<paragraph id="8BCF9B3A4772FFC2FF59802B5BCAA219" blockId="26.[159,1244,1052,1102]" pageId="26" pageNumber="27">
- total length 13501720 (♂) 830855 (♂) 670830 (♂) 630800 (♀)
<emphasis id="B90447284772FFC2FD8A800E5B6DA219" box="[588,610,1081,1102]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 1120 (♀)
</paragraph>
<paragraph id="8BCF9B3A4772FFC2FF5980565C27A168" pageId="26" pageNumber="27">
<table id="F970699A47720027FF5980565C98A108" box="[159,1431,1121,1887]" gridcols="5" gridrows="9" pageId="26" pageNumber="27">
<tr id="3540997847720027FF5980565C98A2C4" box="[159,1431,1121,1171]" gridrow="0" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59805658F7A2C4" box="[159,504,1121,1171]" gridcol="0" gridrow="0" pageId="26" pageNumber="27">
-
<taxonomicName id="4C70E0B94772FFC2FF6B80565801A221" box="[173,270,1121,1142]" class="Hydrozoa" family="Haleciidae" genus="Halecium" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="maximum">maximum</taxonomicName>
width 505565 (♂)
</th>
<th id="7691F00447720027FD8A80565BC0A2C4" box="[588,719,1121,1171]" gridcol="1" gridrow="0" pageId="26" pageNumber="27">
405430 (♂)
<emphasis id="B90447284772FFC2FD8A80495B6DA2C4" box="[588,610,1150,1171]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 365 (♀)
</th>
<th id="7691F00447720027FCE580565AAAA2C4" box="[803,933,1121,1171]" gridcol="2" gridrow="0" pageId="26" pageNumber="27">350415 (♂)</th>
<th id="7691F00447720027FC3F80565D78A2C4" box="[1017,1143,1121,1171]" gridcol="3" gridrow="0" pageId="26" pageNumber="27">390440 (♀)</th>
<th id="7691F00447720027FB1680565C98A2C4" box="[1232,1431,1121,1171]" gridcol="4" gridrow="0" pageId="26" pageNumber="27"></th>
</tr>
<tr id="3540997847720027FF5980915C98A28E" box="[159,1431,1190,1241]" gridrow="1" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59809158F7A28E" box="[159,504,1190,1241]" gridcol="0" gridrow="1" pageId="26" pageNumber="27">- width at aperture 395465 (♂)</th>
<td id="7691F00447720027FD8A80915BC0A28E" box="[588,719,1190,1241]" gridcol="1" gridrow="1" pageId="26" pageNumber="27">
310335 (♂)
<emphasis id="B90447284772FFC2FD8A80F35B6DA28E" box="[588,610,1220,1241]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 250 (♀)
</td>
<td id="7691F00447720027FCE580915AAAA28E" box="[803,933,1190,1241]" gridcol="2" gridrow="1" pageId="26" pageNumber="27">220280 (♂)</td>
<td id="7691F00447720027FC3F80915D78A28E" box="[1017,1143,1190,1241]" gridcol="3" gridrow="1" pageId="26" pageNumber="27">200260 (♀)</td>
<td id="7691F00447720027FB1680915C98A28E" box="[1232,1431,1190,1241]" gridcol="4" gridrow="1" pageId="26" pageNumber="27"></td>
</tr>
<tr id="3540997847720027FF5980DB5C98A356" box="[159,1431,1260,1281]" gridrow="2" pageId="26" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="7691F00447720027FF5980DB58F7A356" box="[159,504,1260,1281]" gridcol="0" gridrow="2" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FF5980DB59F3A356" bold="true" box="[159,252,1260,1281]" pageId="26" pageNumber="27">Cnidome</emphasis>
</th>
</tr>
<tr id="3540997847720027FF5981235C98A332" box="[159,1431,1300,1381]" gridrow="3" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59812358F7A332" box="[159,504,1300,1381]" gridcol="0" gridrow="3" pageId="26" pageNumber="27">- large (29.030.1) × pseudostenoteles (12.412.9) (Fig. 5E1, K1, P1, Q1)</th>
<td id="7691F00447720027FD8A81235BC0A332" box="[588,719,1300,1381]" gridcol="1" gridrow="3" pageId="26" pageNumber="27">(26.031.7) × (11.813.4)</td>
<td id="7691F00447720027FCE581235AAAA332" box="[803,933,1300,1381]" gridcol="2" gridrow="3" pageId="26" pageNumber="27">(25.527.9) × (11.512.9)</td>
<td id="7691F00447720027FC3F81235D78A332" box="[1017,1143,1300,1381]" gridcol="3" gridrow="3" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FC3F81235D00A37E" box="[1017,1039,1300,1321]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 22.5 × 10
</td>
<td id="7691F00447720027FB1681235C98A332" box="[1232,1431,1300,1381]" gridcol="4" gridrow="3" pageId="26" pageNumber="27">(18.320.4) × (8.69.1)</td>
</tr>
<tr id="3540997847720027FF59814F5C98A39D" box="[159,1431,1400,1482]" gridrow="4" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59814F58F7A39D" box="[159,504,1400,1482]" gridcol="0" gridrow="4" pageId="26" pageNumber="27">- small (5.66.2) × pseudostenoteles (3.23.8) (Fig. 5E3, K3, P3, Q3)</th>
<td id="7691F00447720027FD8A814F5BC0A39D" box="[588,719,1400,1482]" gridcol="1" gridrow="4" pageId="26" pageNumber="27">(7.28.0) × (4.04.6)</td>
<td id="7691F00447720027FCE5814F5AAAA39D" box="[803,933,1400,1482]" gridcol="2" gridrow="4" pageId="26" pageNumber="27">(6.26.4) × (3.53.8)</td>
<td id="7691F00447720027FC3F814F5D78A39D" box="[1017,1143,1400,1482]" gridcol="3" gridrow="4" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB16814F5C98A39D" box="[1232,1431,1400,1482]" gridcol="4" gridrow="4" pageId="26" pageNumber="27">(5.15.4) × (2.93.2)</td>
</tr>
<tr id="3540997847720027FF5981EA5C98A067" box="[159,1431,1501,1584]" gridrow="5" pageId="26" pageNumber="27">
<th id="7691F00447720027FF5981EA58F7A067" box="[159,504,1501,1584]" gridcol="0" gridrow="5" pageId="26" pageNumber="27">- microbasic (4.35.4) × mastigophores (Fig. (2.12.4) 5E2, K2, P2, Q2)</th>
<td id="7691F00447720027FD8A81EA5BC0A067" box="[588,719,1501,1584]" gridcol="1" gridrow="5" pageId="26" pageNumber="27">
(4.64.8) ×
<emphasis id="B90447284772FFC2FD8A81CC5B6DA047" box="[588,610,1531,1552]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 1.9
</td>
<td id="7691F00447720027FCE581EA5AAAA067" box="[803,933,1501,1584]" gridcol="2" gridrow="5" pageId="26" pageNumber="27">(4.64.8) × (1.92.1)</td>
<td id="7691F00447720027FC3F81EA5D78A067" box="[1017,1143,1501,1584]" gridcol="3" gridrow="5" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB1681EA5C98A067" box="[1232,1431,1501,1584]" gridcol="4" gridrow="5" pageId="26" pageNumber="27">
<emphasis id="B90447284772FFC2FB1681EA5DE8A3A5" box="[1232,1255,1501,1522]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 3.2 × 1.9
</td>
</tr>
<tr id="3540997847720027FF5982745C98A0C2" box="[159,1431,1603,1685]" gridrow="6" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59827458F7A0C2" box="[159,504,1603,1685]" gridcol="0" gridrow="6" pageId="26" pageNumber="27">
- microbasic (6.26.4) × mastigophores (Fig.
<emphasis id="B90447284772FFC2FEB382575883A022" box="[373,396,1632,1653]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 1.6 5E4, K4, P4, Q4)
</th>
<td id="7691F00447720027FD8A82745BC0A0C2" box="[588,719,1603,1685]" gridcol="1" gridrow="6" pageId="26" pageNumber="27">
(5.96.2) ×
<emphasis id="B90447284772FFC2FD8A82575B6DA022" box="[588,610,1632,1653]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 1.6
</td>
<td id="7691F00447720027FCE582745AAAA0C2" box="[803,933,1603,1685]" gridcol="2" gridrow="6" pageId="26" pageNumber="27">
(5.96.2) ×
<emphasis id="B90447284772FFC2FCE582575A36A022" box="[803,825,1632,1653]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 1.6
</td>
<td id="7691F00447720027FC3F82745D78A0C2" box="[1017,1143,1603,1685]" gridcol="3" gridrow="6" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB1682745C98A0C2" box="[1232,1431,1603,1685]" gridcol="4" gridrow="6" pageId="26" pageNumber="27">
(5.65.9) ×
<emphasis id="B90447284772FFC2FB1682575DE8A022" box="[1232,1255,1632,1653]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 1.6
</td>
</tr>
<tr id="3540997847720027FF59829F5C98A0AD" box="[159,1431,1704,1786]" gridrow="7" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59829F58F7A0AD" box="[159,504,1704,1786]" gridcol="0" gridrow="7" pageId="26" pageNumber="27">- microbasic (7.57.8) × mastigophores (Fig. (2.12.4) 5E5, K5, P5, Q5)</th>
<td id="7691F00447720027FD8A829F5BC0A0AD" box="[588,719,1704,1786]" gridcol="1" gridrow="7" pageId="26" pageNumber="27">
(7.88.3) ×
<emphasis id="B90447284772FFC2FD8A82F25B6DA08D" box="[588,610,1733,1754]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 2.1
</td>
<td id="7691F00447720027FCE5829F5AAAA0AD" box="[803,933,1704,1786]" gridcol="2" gridrow="7" pageId="26" pageNumber="27">
(7.07.5) ×
<emphasis id="B90447284772FFC2FCE582F25A36A08D" box="[803,825,1733,1754]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 2.1
</td>
<td id="7691F00447720027FC3F829F5D78A0AD" box="[1017,1143,1704,1786]" gridcol="3" gridrow="7" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB16829F5C98A0AD" box="[1232,1431,1704,1786]" gridcol="4" gridrow="7" pageId="26" pageNumber="27">
(6.77.0) ×
<emphasis id="B90447284772FFC2FB1682F25DE8A08D" box="[1232,1255,1733,1754]" italics="true" pageId="26" pageNumber="27">ca</emphasis>
. 2.4
</td>
</tr>
<tr id="3540997847720027FF59833A5C98A108" box="[159,1431,1805,1887]" gridrow="8" pageId="26" pageNumber="27">
<th id="7691F00447720027FF59833A58F7A108" box="[159,504,1805,1887]" gridcol="0" gridrow="8" pageId="26" pageNumber="27">- microbasic (8.99.9) × mastigophores (Fig. (3.23.5) 5E6, K6, P6, Q6)</th>
<td id="7691F00447720027FD8A833A5BC0A108" box="[588,719,1805,1887]" gridcol="1" gridrow="8" pageId="26" pageNumber="27">(8.99.7) × (2.93.2)</td>
<td id="7691F00447720027FCE5833A5AAAA108" box="[803,933,1805,1887]" gridcol="2" gridrow="8" pageId="26" pageNumber="27">(7.88.6) × (2.73.2)</td>
<td id="7691F00447720027FC3F833A5D78A108" box="[1017,1143,1805,1887]" gridcol="3" gridrow="8" pageId="26" pageNumber="27"></td>
<td id="7691F00447720027FB16833A5C98A108" box="[1232,1431,1805,1887]" gridcol="4" gridrow="8" pageId="26" pageNumber="27">(8.99.4) × (2.93.5)</td>
</tr>
</table>
</paragraph>
<caption id="DF0FCBB24773FFC3FF5182C35D78A1D2" httpUri="https://zenodo.org/record/286960/files/figure.png" pageId="27" pageNumber="28" targetBox="[153,1431,197,1758]" targetPageId="27">
<paragraph id="8BCF9B3A4773FFC3FF5182C35D78A1D2" blockId="27.[151,1437,1780,1925]" pageId="27" pageNumber="28">
<emphasis id="B90447284773FFC3FF5182C3581FA15D" bold="true" box="[151,272,1780,1802]" pageId="27" pageNumber="28">FIGURE 5</emphasis>
. AE:
<taxonomicName id="4C70E0B94773FFC3FEA682C35B5FA15D" box="[352,592,1780,1802]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="chilense" status="sp. nov.">
<emphasis id="B90447284773FFC3FEA682C35B5FA15D" box="[352,592,1780,1802]" italics="true" pageId="27" pageNumber="28">Hydrodendron chilense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534773FFC3FD9C82C25BA4A15D" box="[602,683,1780,1802]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
—colonies and fragments (A); hydrotheca (B); nematothecae (C); male gonotheca (D); cnidome (E). FK:
<taxonomicName id="4C70E0B94773FFC3FDCB83235A25A17E" authority="Hincks, 1866" authorityName="Hincks" authorityYear="1866" box="[525,810,1811,1833]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284773FFC3FDCB83235B8AA17E" box="[525,645,1811,1833]" italics="true" pageId="27" pageNumber="28">H. mirabile</emphasis>
(Hincks, 1866)
</taxonomicName>
from the Mediterranean—colony (F); hydrothecae (G) and nematothecae (H); male (I) and female (J) gonothecae; cnidome (K). LP:
<taxonomicName id="4C70E0B94773FFC3FC6583045D15A11F" box="[931,1050,1842,1864]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="mirabile">
<emphasis id="B90447284773FFC3FC6583045D15A11F" box="[931,1050,1842,1864]" italics="true" pageId="27" pageNumber="28">H. mirabile</emphasis>
</taxonomicName>
from Martinique—four colonies (L); hydrothecae (M) and nematothecae (N); male gonotheca (O); cnidome (P). Q:
<taxonomicName id="4C70E0B94773FFC3FC7083675D46A131" box="[950,1097,1872,1894]" class="Hydrozoa" family="Haleciidae" genus="Hydrodendron" kingdom="Animalia" order="Leptothecata" pageId="27" pageNumber="28" phylum="Cnidaria" rank="genus">
<emphasis id="B90447284773FFC3FC7083675D46A131" box="[950,1097,1872,1894]" italics="true" pageId="27" pageNumber="28">Hydrodendron</emphasis>
</taxonomicName>
sp. from Guadeloupe—cnidome. Scale bars: 10 µm (E, K, P, Q), 200 µm (B, C, G, H, M, N), 300 µm (D, I, J, O), 500 µm (A, F, L).
</paragraph>
</caption>
<paragraph id="8BCF9B3A4774FFC4FF51831A5C22A1F5" blockId="28.[151,1436,1834,1954]" pageId="28" pageNumber="29">
<emphasis id="B90447284774FFC4FF51831A5802A114" bold="true" box="[151,269,1837,1859]" pageId="28" pageNumber="29">FIGURE 6</emphasis>
. A, B:
<taxonomicName id="4C70E0B94774FFC4FE9F831A5B48A113" box="[345,583,1837,1860]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="28" pageNumber="29" phylum="Cnidaria" rank="species" species="asymmetra" status="sp. nov.">
<emphasis id="B90447284774FFC4FE9F831A5B48A113" box="[345,583,1837,1860]" italics="true" pageId="28" pageNumber="29">Sertularella asymmetra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534774FFC4FD8883195B93A114" box="[590,668,1837,1859]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
—colony fragment (A); hydrotheca in lateral (B1) and apical (B2) views. C:
<taxonomicName id="4C70E0B94774FFC4FF51837B5B63A135" authority="El Beshbeeshy, 2011" authorityName="El Beshbeeshy" authorityYear="2011" box="[151,620,1868,1890]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="28" pageNumber="29" phylum="Cnidaria" rank="species" species="blanconae">
<emphasis id="B90447284774FFC4FF51837B588BA135" box="[151,388,1868,1890]" italics="true" pageId="28" pageNumber="29">Sertularella blanconae</emphasis>
<bibRefCitation id="EFE1E6CB4774FFC4FE49837B5B63A135" author="El" box="[399,620,1868,1890]" pageId="28" pageNumber="29" refString="El Beshbeeshy, M. (2011) Thecate hydroids from the Patagonian shelf (Coelenterata, Hydrozoa, Thecata). G. Jarms (Ed.)." type="book" year="2011">El Beshbeeshy, 2011</bibRefCitation>
</taxonomicName>
—colony fragment. D, E:
<taxonomicName id="4C70E0B94774FFC4FC4F837B5D4DA135" box="[905,1090,1868,1890]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="28" pageNumber="29" phylum="Cnidaria" rank="species" species="curta" status="sp. nov.">
<emphasis id="B90447284774FFC4FC4F837B5D4DA135" box="[905,1090,1868,1890]" italics="true" pageId="28" pageNumber="29">Sertularella curta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534774FFC4FB88837A5DAFA135" box="[1102,1184,1868,1890]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
—colony fragment (D); hydrotheca in lateral (E1) and apical (E2) views. F, H:
<taxonomicName id="4C70E0B94774FFC4FD72835A5AB8A1D4" box="[692,951,1901,1923]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="28" pageNumber="29" phylum="Cnidaria" rank="species" species="pauciramosa" status="sp. nov.">
<emphasis id="B90447284774FFC4FD72835A5AB8A1D4" box="[692,951,1901,1923]" italics="true" pageId="28" pageNumber="29">Sertularella pauciramosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534774FFC4FC7A83595D06A1D4" box="[956,1033,1901,1923]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
—colony fragment (F), and comparison with
<taxonomicName id="4C70E0B94774FFC4FF0C83BA5B0CA1F5" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[202,515,1932,1954]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="28" pageNumber="29" phylum="Cnidaria" rank="species" species="polyzonias">
<emphasis id="B90447284774FFC4FF0C83BA5840A1F5" box="[202,335,1932,1954]" italics="true" pageId="28" pageNumber="29">S. polyzonias</emphasis>
(Linnaeus, 1758)
</taxonomicName>
(G); four hydrothecae (H). Scale bars: 200 µm (B, E, H),
<quantity id="4C8836DF4774FFC4FB8983BB5D85A1F5" box="[1103,1162,1932,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="mm" value="1.0">1 mm</quantity>
(A, C, D, F, G).
</paragraph>
<paragraph id="8BCF9B3A4775FFC5FF5182F658E0A1C1" blockId="29.[151,1437,1726,1942]" pageId="29" pageNumber="30">
<emphasis id="B90447284775FFC5FF5182F6581CA080" bold="true" box="[151,275,1729,1751]" pageId="29" pageNumber="30">FIGURE 7</emphasis>
. A:
<taxonomicName id="4C70E0B94775FFC5FE8A82F65B4FA080" box="[332,576,1729,1751]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="species" species="asymmetra" status="sp. nov.">
<emphasis id="B90447284775FFC5FE8A82F65B4FA080" box="[332,576,1729,1751]" italics="true" pageId="29" pageNumber="30">Sertularella asymmetra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534775FFC5FD8B82F55BAFA080" box="[589,672,1729,1751]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
—female (A1) and male (A2) gonotheca; B:
<taxonomicName id="4C70E0B94775FFC5FB4C82F6585CA0A2" authority="El Beshbeeshy, 2011" authorityName="El Beshbeeshy" authorityYear="2011" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="species" species="blanconae">
<emphasis id="B90447284775FFC5FB4C82F65C76A080" box="[1162,1401,1729,1751]" italics="true" pageId="29" pageNumber="30">Sertularella blanconae</emphasis>
<bibRefCitation id="EFE1E6CB4775FFC5FA4382F6585CA0A2" author="El" pageId="29" pageNumber="30" refString="El Beshbeeshy, M. (2011) Thecate hydroids from the Patagonian shelf (Coelenterata, Hydrozoa, Thecata). G. Jarms (Ed.)." type="book" year="2011">El Beshbeeshy, 2011</bibRefCitation>
</taxonomicName>
—male gonotheca. C:
<taxonomicName id="4C70E0B94775FFC5FD8A82E85A09A0A2" box="[588,774,1759,1781]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="species" species="curta" status="sp. nov.">
<emphasis id="B90447284775FFC5FD8A82E85A09A0A2" box="[588,774,1759,1781]" italics="true" pageId="29" pageNumber="30">Sertularella curta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534775FFC5FCD282D75A67A0A2" box="[788,872,1759,1781]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
—male gonotheca. D:
<taxonomicName id="4C70E0B94775FFC5FBAE82E85C7DA0A2" box="[1128,1394,1759,1781]" class="Hydrozoa" family="Sertulariidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="species" species="pauciramosa" status="sp. nov.">
<emphasis id="B90447284775FFC5FBAE82E85C7DA0A2" box="[1128,1394,1759,1781]" italics="true" pageId="29" pageNumber="30">Sertularella pauciramosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534775FFC5FAB982D759CEA141" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
—male (D1) and female (D2) gonotheca. E, F:
<taxonomicName id="4C70E0B94775FFC5FD0483375D7FA141" authority="Hartlaub, 1901" authorityName="Hartlaub" authorityYear="1901" box="[706,1136,1792,1814]" class="Hydrozoa" family="Sertulariidae" genus="Symplectoscyphus" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="species" species="flexilis">
<emphasis id="B90447284775FFC5FD0483375AC5A141" box="[706,970,1792,1814]" italics="true" pageId="29" pageNumber="30">Symplectoscyphus flexilis</emphasis>
Hartlaub, 1901
</taxonomicName>
—colony fragment (E1) and hydrotheca (E2); gonotheca (F). GJ:
<taxonomicName id="4C70E0B94775FFC5FDDC83155A2CA16F" box="[538,803,1826,1848]" class="Hydrozoa" family="Sertulariidae" genus="Symplectoscyphus" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="species" species="semper" status="sp. nov.">
<emphasis id="B90447284775FFC5FDDC83155A2CA16F" box="[538,803,1826,1848]" italics="true" pageId="29" pageNumber="30">Symplectoscyphus semper</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FA534775FFC5FCED83145A75A16F" box="[811,890,1827,1848]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
—stem fragment with beginning of side branch (G1), and portion of cladium (G2); hydrotheca (H); gonotheca (I), and details of the distal part (J). KM:
<taxonomicName id="4C70E0B94775FFC5FB06837458E8A12F" authority="Coughtrey, 1875" authorityName="Coughtrey" authorityYear="1875" class="Hydrozoa" family="Sertulariidae" genus="Symplectoscyphus" kingdom="Animalia" order="Leptothecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="species" species="subarticulatus">
<emphasis id="B90447284775FFC5FB0683745C77A10E" box="[1216,1400,1859,1881]" italics="true" pageId="29" pageNumber="30">Symplectoscyphus</emphasis>
cf.
<emphasis id="B90447284775FFC5FF5183555827A12F" box="[151,296,1890,1912]" italics="true" pageId="29" pageNumber="30">subarticulatus</emphasis>
(
<bibRefCitation id="EFE1E6CB4775FFC5FEF0835558EFA12F" author="Coughtrey" box="[310,480,1890,1912]" pageId="29" pageNumber="30" refString="Coughtrey, M. (1875) Notes on the New Zealand Hydroideae. Transactions of the Royal Society of New Zealand, 7, 281 - 293." type="journal article" year="1875">Coughtrey, 1875</bibRefCitation>
)
</taxonomicName>
—colony fragment (K); hydrotheca (L); gonotheca (M). Scale bars: 200 µm (H, L), 500 µm (AD, F, I, J, M),
<quantity id="4C8836DF4775FFC5FE8B83B75888A1C1" box="[333,391,1920,1942]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="mm" value="1.0">1 mm</quantity>
(E, G, K).
</paragraph>
</subSubSection>
</treatment>
</document>