437 lines
42 KiB
XML
437 lines
42 KiB
XML
<document ID-DOI="10.1206/3990.1" ID-GBIF-Dataset="5e5fa3ea-10cd-4646-abd6-58aa6c664b31" ID-ISSN="0003-0082" checkinTime="1665198941087" checkinUser="felipe" docAuthor="Bernt, Maxwell J. & Stiassny, Melanie L. J." docDate="2022" docId="03D0F929FFD3FFE7FE008D23FD75FB0C" docLanguage="en" docName="AmMusNovit.2022.3990.1-20.pdf" docOrigin="American Museum Novitates 2022 (3990)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2022/issue-3990/3990.1/A-New-Species-of-Air-Breathing-Catfish-Clariidae--Clarias/10.1206/3990.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.4:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="4" docTitle="Clarias monsembulai Bernt & Stiassny 2022, new species" docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="FFE98151FFD0FFEFFFC5882AFFDDFFB8" masterDocTitle="A New Species of Air-breathing Catfish (Clariidae: Clarias) from Salonga National Park, Democratic Republic of the Congo" masterLastPageNumber="20" masterPageNumber="1" pageNumber="4" updateTime="1665199951882" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A New Species of Air-breathing Catfish (Clariidae: Clarias) from Salonga National Park, Democratic Republic of the Congo</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bernt, Maxwell J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stiassny, Melanie L. J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>American Museum Novitates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2022</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3990</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>20</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://bioone.org/journals/american-museum-novitates/volume-2022/issue-3990/3990.1/A-New-Species-of-Air-Breathing-Catfish-Clariidae--Clarias/10.1206/3990.1.full</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/3990.1</mods:identifier>
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<mods:identifier type="GBIF-Dataset">5e5fa3ea-10cd-4646-abd6-58aa6c664b31</mods:identifier>
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<mods:identifier type="ISSN">0003-0082</mods:identifier>
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</mods:mods>
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||
<treatment LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C" httpUri="http://treatment.plazi.org/id/03D0F929FFD3FFE7FE008D23FD75FB0C" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4">
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||
<subSubSection box="[453,866,1289,1316]" pageId="3" pageNumber="4" type="nomenclature">
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||
<paragraph blockId="3.[453,866,1289,1316]" box="[453,866,1289,1316]" pageId="3" pageNumber="4">
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||
<heading box="[453,866,1289,1316]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
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<emphasis bold="true" box="[453,719,1289,1316]" pageId="3" pageNumber="4">
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<taxonomicName authority="Bernt & Stiassny, 2022" authorityName="Bernt & Stiassny" authorityYear="2022" box="[453,713,1289,1316]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="monsembulai" status="new species">
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<emphasis bold="true" box="[453,713,1289,1316]" italics="true" pageId="3" pageNumber="4">Clarias monsembulai</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel box="[728,866,1289,1316]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="3" pageNumber="4" type="description">
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<paragraph blockId="3.[545,775,1345,1372]" box="[545,775,1345,1372]" pageId="3" pageNumber="4">
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<heading box="[545,775,1345,1372]" centered="true" fontSize="36" level="3" pageId="3" pageNumber="4" reason="7">
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<figureCitation box="[545,685,1345,1372]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[108,194,666,687]" captionStartId-1="6.[108,192,889,910]" captionStartId-2="7.[109,196,889,910]" captionStartId-3="8.[108,194,1019,1040]" captionTargetBox-0="[184,1156,282,613]" captionTargetBox-1="[108,1212,224,860]" captionTargetBox-2="[108,1212,224,858]" captionTargetBox-3="[408,934,232,992]" captionTargetId-0="figure-404@4.[108,1212,224,640]" captionTargetId-1="figure-193@6.[108,1212,224,860]" captionTargetId-2="figure-269@7.[108,1212,224,861]" captionTargetId-3="figure-233@8.[408,936,227,992]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 2. Lateral view of Clarias monsembulai new species, holotype AMNH 244176, 226 mm SL. Luilaka River at Ilenge, Salonga National Park. Scale bar = 1 cm." captionText-1="FIGURE 3. Computed tomography scan reconstruction of the skull and pectoral girdle of Clarias monsembulai (at left and above), AMNH 252267, 179 mm SL in A, dorsal; B, ventral; and C, lateral views. Abbreviations: ang, anguloarticular; apal, autopalatine; br, branchiostegal rays; ch-a, anterior ceratohyal; ch-p, posterior ceratohyal; cl, cleithrum; cor, coracoid; den, dentary; fr, frontal; hhv, ventral hypohyal; hm, hyomandibula; io-ii, infraorbital ii; io-iii, infraorbital iii; io-iv, infraorbital iv; iop, interopercle; lac, lacrimal; leth, lateral ethmoid; meth, mesethmoid; mx, maxilla; na, nasal; ns, neural spines; op, opercle; o-susp, os suspensorium; pmx, premaxilla; pop, preopercle; pp-v4, parapophysis of 4th vertebra; pp-v5, parapophysis of 5th vertebra; ps, pectoral spine; pt, pterotic; pt-scl, posttemporo-supracleithrum; q, quadrate; rad, radials; soc, supraoccipital; sph, sphenotic; spop, suprapreopercle; trsc, transcapular process; uh, urohyal; vo, vomer. Scale bars = 1 cm." captionText-2="FIGURE 4. Suprabranchial organ of Clarias monsembulai, AMNH 244162, 165 mm SL (suprapreopercle removed). Scale bar = 1 cm." captionText-3="FIGURE 5. Exposed cleithrum (demarcated by dashed line) of A, Clarias buthupogon, AMNH 227571, 162 mm SL and B, Clarias monsembulai, AMNH 244162, 183 mm SL. Scale bar = 1 cm." pageId="3" pageNumber="4">Figures 2–5</figureCitation>
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, table 1
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</heading>
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</paragraph>
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<paragraph blockId="3.[108,855,1403,1427]" box="[108,855,1403,1427]" pageId="3" pageNumber="4">
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<taxonomicName box="[108,316,1403,1427]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="buthupogon">
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<emphasis box="[108,316,1403,1427]" italics="true" pageId="3" pageNumber="4">Clarias buthupogon</emphasis>
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</taxonomicName>
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(in part):
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<bibRefCitation author="Monsembula Iyaba, R. J. C. & M. L. J. Stiassny" box="[427,851,1403,1427]" pageId="3" pageNumber="4" pagination="246 - 256" refId="ref8905" refString="Monsembula Iyaba, R. J. C., and M. L. J. Stiassny. 2013. Fishes of the Salonga National Park (Congo basin, central Africa): a list of species collected in the Luilaka, Salonga, and Yenge rivers (Equateur Province, Democratic Republic of Congo). Check List 9: 246 - 256." type="journal article" year="2013">Monsembula Iyaba and Stiassny (2013)</bibRefCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph blockId="3.[108,1212,1457,1745]" pageId="3" pageNumber="4">
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<materialsCitation collectingDate="2007-11-11" collectionCode="R" collectorName="J. C. Monsembula Iyaba" country="Democratic Republic of the Congo" county="Ilenge" latitude="-2.5963888" location="Salonga National Park" longLatPrecision="21" longitude="21.576668" municipality="Luilaka River" pageId="3" pageNumber="4" specimenCode="AMNH 244176" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[156,284,1457,1483]" pageId="3" pageNumber="4">HOLOTYPE</typeStatus>
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:
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<specimenCode box="[298,484,1457,1484]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH 244176</specimenCode>
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,
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<quantity box="[496,622,1457,1484]" metricMagnitude="-3" metricUnit="m" metricValue="1.226" pageId="3" pageNumber="4" unit="mm" value="1.226">1, 226 mm</quantity>
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SL,
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<collectingCountry box="[674,1088,1457,1483]" name="Democratic Republic of the Congo" pageId="3" pageNumber="4">Democratic Republic of the Congo</collectingCountry>
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, Equateur,
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<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD3FFECFFA88DFDFEA6FA49" box="[109,379,1495,1521]" country="Democratic Republic of the Congo" county="Ilenge" latitude="-2.5963888" longLatPrecision="21" longitude="21.576668" municipality="Luilaka River" name="Salonga National Park" pageId="3" pageNumber="4">Salonga National Park</location>
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,
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<collectingMunicipality box="[393,553,1495,1521]" pageId="3" pageNumber="4">Luilaka River</collectingMunicipality>
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||
at
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||
<collectingCounty box="[595,669,1495,1521]" pageId="3" pageNumber="4">Ilenge</collectingCounty>
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,
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||
<geoCoordinate box="[683,808,1494,1521]" degrees="2" direction="south" minutes="35" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="47" value="-2.5963888">2°35′ 47″S</geoCoordinate>
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||
,
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||
<geoCoordinate box="[823,968,1494,1521]" degrees="21" direction="east" minutes="34" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="36" value="21.576668">21°34′ 36″E</geoCoordinate>
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||
.
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||
<collectionCode box="[981,1003,1495,1521]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="4">R</collectionCode>
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||
.
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||
<collectorName pageId="3" pageNumber="4">J.C. Monsembula Iyaba</collectorName>
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||
,
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||
<date box="[186,413,1532,1558]" pageId="3" pageNumber="4" value="2007-11-11">
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||
<collectingDate box="[186,413,1532,1558]" pageId="3" pageNumber="4" value="2007-11-11">11 November 2007</collectingDate>
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||
</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph blockId="3.[108,1212,1457,1745]" lastBlockId="4.[108,1214,748,1746]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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||
<materialsCitation box="[156,1208,1569,1596]" collectionCode="AMNH" latitude="-1.0638888" location="Yenge River" longLatPrecision="21" longitude="20.745556" municipality="Boyenga" pageId="3" pageNumber="4" specimenCode="AMNH 244163" specimenCount="1" typeStatus="paratype">
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||
<typeStatus box="[156,285,1569,1595]" pageId="3" pageNumber="4">PARATYPES</typeStatus>
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||
:
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||
<specimenCode box="[298,480,1569,1596]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH 244163</specimenCode>
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||
,
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||
<quantity box="[490,612,1569,1596]" metricMagnitude="-3" metricUnit="m" metricValue="1.244" pageId="3" pageNumber="4" unit="mm" value="1.244">1, 244 mm</quantity>
|
||
SL;
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD3FFECFD518E08FCC1F983" box="[660,796,1569,1596]" latitude="-1.0638888" longLatPrecision="21" longitude="20.745556" municipality="Boyenga" name="Yenge River" pageId="3" pageNumber="4">Yenge River</location>
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||
ca.
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||
<collectingMunicipality box="[840,942,1570,1596]" pageId="3" pageNumber="4">Boyenga</collectingMunicipality>
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||
,
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||
<geoCoordinate box="[951,1066,1569,1596]" degrees="1" direction="south" minutes="03" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="50" value="-1.0638888">1°03′50″S</geoCoordinate>
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||
,
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||
<geoCoordinate box="[1076,1208,1569,1596]" degrees="20" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="44" value="20.745556">20°44′44″E</geoCoordinate>
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||
</materialsCitation>
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||
;
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||
<materialsCitation collectionCode="AMNH" latitude="-2.1363056" location="Katanyongo" longLatPrecision="1" longitude="21.119307" municipality="Salonga National Park" pageId="3" pageNumber="4" specimenCode="AMNH 252236, 3" specimenCount="1" typeStatus="paratype">
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||
<specimenCode box="[108,294,1606,1633]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH 252236</specimenCode>
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||
,
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||
<specimenCode box="[306,321,1607,1633]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">3</specimenCode>
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||
,
|
||
<quantity box="[334,490,1607,1633]" metricMagnitude="-1" metricUnit="m" metricValue="1.785" metricValueMax="1.93" metricValueMin="1.64" pageId="3" pageNumber="4" unit="mm" value="178.5" valueMax="193.0" valueMin="164.0">164–193 mm</quantity>
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||
SL; small stream at
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD3FFECFD1C8E6DFCB5F9D9" box="[729,872,1607,1633]" latitude="-2.1363056" longLatPrecision="1" longitude="21.119307" municipality="Salonga National Park" name="Katanyongo" pageId="3" pageNumber="4">Katanyongo</location>
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||
, near
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||
<collectingMunicipality box="[943,1208,1607,1633]" pageId="3" pageNumber="4">Salonga National Park</collectingMunicipality>
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||
,
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||
<geoCoordinate box="[109,245,1643,1670]" degrees="2" direction="south" minutes="08" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="10.7" value="-2.1363056">2°08′10.7″S</geoCoordinate>
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||
,
|
||
<geoCoordinate box="[256,396,1643,1671]" degrees="21" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="9.5" value="21.119307">21°07′9.5″E</geoCoordinate>
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||
</materialsCitation>
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||
;
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||
<materialsCitation box="[407,1110,1644,1671]" collectionCode="AMNH" location="Salonga River" municipality="Watsikengo" pageId="3" pageNumber="4" specimenCode="AMNH 244162" specimenCount="1" typeStatus="paratype">
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||
<specimenCode box="[407,591,1644,1670]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH 244162</specimenCode>
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||
,
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||
<quantity box="[602,726,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="1.183" pageId="3" pageNumber="4" unit="mm" value="1.183">1, 183 mm</quantity>
|
||
SL;
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD3FFECFCCD8E46FC74F93E" box="[776,937,1644,1670]" municipality="Watsikengo" name="Salonga River" pageId="3" pageNumber="4">Salonga River</location>
|
||
at
|
||
<collectingMunicipality box="[971,1110,1644,1670]" pageId="3" pageNumber="4">Watsikengo</collectingMunicipality>
|
||
</materialsCitation>
|
||
;
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||
<materialsCitation collectionCode="AMNH" latitude="-1.9118721" location="Salonga National Park" longLatPrecision="1" longitude="20.853153" municipality="Bekongo" pageId="3" pageNumber="4" specimenCode="AMNH 252219, 2" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH 252219</specimenCode>
|
||
,
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||
<specimenCode box="[209,225,1681,1707]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">2</specimenCode>
|
||
,
|
||
<quantity box="[239,396,1681,1708]" metricMagnitude="-1" metricUnit="m" metricValue="2.09" metricValueMax="2.31" metricValueMin="1.8699999999999999" pageId="3" pageNumber="4" unit="mm" value="209.0" valueMax="231.0" valueMin="187.0">187–231 mm</quantity>
|
||
SL;
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD3FFECFE068EBBFD11F913" box="[451,716,1681,1707]" latitude="-1.9118721" longLatPrecision="1" longitude="20.853153" municipality="Bekongo" name="Salonga National Park" pageId="3" pageNumber="4">Salonga National Park</location>
|
||
at
|
||
<collectingMunicipality box="[756,861,1681,1707]" pageId="3" pageNumber="4">Bekongo</collectingMunicipality>
|
||
,
|
||
<geoCoordinate box="[875,1026,1681,1708]" degrees="1" direction="south" minutes="54" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="42.74" value="-1.9118721">1°54′42.74″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1040,1208,1681,1708]" degrees="20" direction="east" minutes="51" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="11.35" value="20.853153">20°51′11.35″E</geoCoordinate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation box="[108,874,1719,1745]" collectionCode="AMNH" location="Luilaka River" municipality="Monkoto" pageId="3" pageNumber="4" specimenCode="AMNH 244164" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode box="[108,294,1719,1745]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH 244164</specimenCode>
|
||
,
|
||
<quantity box="[306,434,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.127" pageId="3" pageNumber="4" unit="mm" value="1.127">1, 127 mm</quantity>
|
||
SL,
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD3FFECFE238E9DFD59F969" box="[486,644,1719,1745]" municipality="Monkoto" name="Luilaka River" pageId="3" pageNumber="4">Luilaka River</location>
|
||
, beach at
|
||
<collectingMunicipality box="[763,874,1719,1745]" pageId="3" pageNumber="4">Monkoto</collectingMunicipality>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectionCode="AMNH" county="Salonga National Park" lastPageId="4" lastPageNumber="5" latitude="-1.9221945" location="Mpongo" longLatPrecision="1" longitude="20.863834" municipality="Luilaka River" pageId="3" pageNumber="4" specimenCode="AMNH 252267" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode box="[887,1072,1718,1745]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH 252267</specimenCode>
|
||
,
|
||
<quantity box="[1085,1212,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.179" pageId="3" pageNumber="4" unit="mm" value="1.179">1, 179 mm</quantity>
|
||
SL,
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD4FFEBFF5F8AC7FEDDFCBF" box="[154,256,749,775]" county="Salonga National Park" latitude="-1.9221945" longLatPrecision="1" longitude="20.863834" municipality="Luilaka River" name="Mpongo" pageId="4" pageNumber="5">Mpongo</location>
|
||
, large stream ca. 2 kilometers from
|
||
<collectingMunicipality box="[698,860,748,774]" pageId="4" pageNumber="5">Luilaka River</collectingMunicipality>
|
||
, near
|
||
<collectingCounty box="[937,1208,748,774]" pageId="4" pageNumber="5">Salonga National Park</collectingCounty>
|
||
,
|
||
<geoCoordinate box="[108,252,785,813]" degrees="1" direction="south" minutes="55" orientation="latitude" pageId="4" pageNumber="5" precision="1" seconds="19.9" value="-1.9221945">1°55′19.9″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[264,418,785,813]" degrees="20" direction="east" minutes="51" orientation="longitude" pageId="4" pageNumber="5" precision="1" seconds="49.8" value="20.863834">20°51′49.8″E</geoCoordinate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectionCode="AMNH" latitude="-2.0069168" location="Luilaka River" longLatPrecision="1" longitude="20.96839" municipality="Lokaka in Salonga National Park" pageId="4" pageNumber="5" specimenCode="AMNH 252222" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode box="[430,615,786,813]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="5">AMNH 252222</specimenCode>
|
||
,
|
||
<quantity box="[626,752,786,812]" metricMagnitude="-3" metricUnit="m" metricValue="1.167" pageId="4" pageNumber="5" unit="mm" value="1.167">1, 167 mm</quantity>
|
||
SL;
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD4FFEBFCE68B38FC1DFC94" box="[803,960,786,812]" latitude="-2.0069168" longLatPrecision="1" longitude="20.96839" municipality="Lokaka in Salonga National Park" name="Luilaka River" pageId="4" pageNumber="5">Luilaka River</location>
|
||
at
|
||
<collectingMunicipality pageId="4" pageNumber="5">Lokaka in Salonga National Park</collectingMunicipality>
|
||
,
|
||
<geoCoordinate box="[294,417,822,849]" degrees="2" direction="south" minutes="0" orientation="latitude" pageId="4" pageNumber="5" precision="1" seconds="24.9" value="-2.0069168">2°0′24.9″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[433,574,822,850]" degrees="20" direction="east" minutes="58" orientation="longitude" pageId="4" pageNumber="5" precision="1" seconds="6.2" value="20.96839">20°58′6.2″E</geoCoordinate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectionCode="RMCA" location="Salonga River" municipality="Watsikengo" pageId="4" pageNumber="5" specimenCode="RMCA 2022-019" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode box="[590,798,823,849]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="4" pageNumber="5" type="Museum">RMCA 2022-019</specimenCode>
|
||
-P-0001-0002,
|
||
<quantity box="[974,1165,823,850]" metricMagnitude="-2" metricUnit="m" metricValue="8.70715" metricValueMax="17.2" metricValueMin="0.2143" pageId="4" pageNumber="5" unit="mm" value="87.0715" valueMax="172.0" valueMin="2.143">2, 143–172 mm</quantity>
|
||
SL,
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD4FFEBFFA98B76FED3FCCE" box="[108,270,860,886]" municipality="Watsikengo" name="Salonga River" pageId="4" pageNumber="5">Salonga River</location>
|
||
at
|
||
<collectingMunicipality box="[307,446,860,886]" pageId="4" pageNumber="5">Watsikengo</collectingMunicipality>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption pageId="4" pageNumber="5" startId="4.[108,194,666,687]" targetBox="[184,1156,282,613]" targetPageId="4">
|
||
<paragraph blockId="4.[108,1214,664,718]" pageId="4" pageNumber="5">
|
||
FIGURE 2. Lateral view of
|
||
<taxonomicName authority="Bernt & Stiassny, 2022" authorityName="Bernt & Stiassny" authorityYear="2022" box="[387,600,664,688]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="monsembulai" status="new species">
|
||
<emphasis box="[387,600,664,688]" italics="true" pageId="4" pageNumber="5">Clarias monsembulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[607,729,664,688]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
|
||
, holotype AMNH 244176, 226 mm SL. Luilaka River at Ilenge, Salonga National Park. Scale bar = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="4" pageNumber="5" type="description">
|
||
<paragraph blockId="4.[108,1214,748,1746]" pageId="4" pageNumber="5">
|
||
<materialsCitation collectionCode="AMNH" latitude="-1.75" location="Luilaka River" longLatPrecision="15" longitude="20.680527" municipality="Monkoto in Salonga National Park" pageId="4" pageNumber="5" specimenCode="AMNH 242721, 14" specimenCount="1">
|
||
NONTYPE SPECIMENS:
|
||
<specimenCode box="[420,606,898,924]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="5">AMNH 242721</specimenCode>
|
||
,
|
||
<specimenCode box="[618,648,898,924]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="5">14</specimenCode>
|
||
,
|
||
<quantity box="[661,788,898,925]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="8.1" metricValueMin="5.4" pageId="4" pageNumber="5" unit="mm" value="67.5" valueMax="81.0" valueMin="54.0">54–81 mm</quantity>
|
||
SL, small tributary of
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD4FFEBFBDA8BA8FB61FC24" box="[1055,1212,898,924]" latitude="-1.75" longLatPrecision="15" longitude="20.680527" municipality="Monkoto in Salonga National Park" name="Luilaka River" pageId="4" pageNumber="5">Luilaka River</location>
|
||
near
|
||
<collectingMunicipality box="[169,583,935,961]" pageId="4" pageNumber="5">Monkoto in Salonga National Park</collectingMunicipality>
|
||
,
|
||
<geoCoordinate box="[595,696,934,962]" degrees="1" direction="south" minutes="45" orientation="latitude" pageId="4" pageNumber="5" precision="15" seconds="0" value="-1.75">1°45′0″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[708,863,934,962]" degrees="20" direction="east" minutes="40" orientation="longitude" pageId="4" pageNumber="5" precision="1" seconds="49.9" value="20.680527">20°40′49.9″E</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="5" type="diagnosis">
|
||
<paragraph blockId="4.[108,1214,748,1746]" pageId="4" pageNumber="5">
|
||
DIAGNOSIS:
|
||
<taxonomicName authorityName="Bernt & Stiassny" authorityYear="2022" box="[299,544,972,998]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="monsembulai">
|
||
<emphasis box="[299,544,972,998]" italics="true" pageId="4" pageNumber="5">Clarias monsembulai</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all congeners, with the exception of
|
||
<taxonomicName box="[198,367,1010,1036]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="buthupogon">
|
||
<emphasis box="[198,367,1010,1036]" italics="true" pageId="4" pageNumber="5">C. buthupogon</emphasis>
|
||
</taxonomicName>
|
||
, by its exceptionally long maxillary barbels (60% of standard length or greater) vs. maxillary barbels less than 60% SL (usually considerably less). It differs from
|
||
<taxonomicName class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="buthupogon">
|
||
<emphasis italics="true" pageId="4" pageNumber="5">C. buthupogon</emphasis>
|
||
</taxonomicName>
|
||
in the absence of fine, pale spots over the surface of the body and by an exposed bony surface of the cleithrum reaching 14%–20% of head length (vs. cleithrum deeply imbedded in soft tissue with only a narrow bony ridge visible externally).
|
||
<taxonomicName authorityName="Bernt & Stiassny" authorityYear="2022" box="[918,1164,1159,1185]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="monsembulai">
|
||
<emphasis box="[918,1164,1159,1185]" italics="true" pageId="4" pageNumber="5">Clarias monsembulai</emphasis>
|
||
</taxonomicName>
|
||
can be further differentiated from
|
||
<emphasis box="[461,617,1196,1223]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName authorityName="Steindachner" authorityYear="1866" box="[461,613,1196,1223]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="angolensis">C. angolensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the species with which it shares closest phenotypic similarity, by longer nasal barbels (37%–51% SL vs. 22%–34% SL), longer internal mandibular barbels (29–37% SL vs. 16–27% SL), longer external mandibular barbels (45%–57% SL vs. 24%–40% SL); and by the coloration of maxillary barbels which are white or cream-colored distally over more than half of their length (vs. brown or gray over more than half their length).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="description">
|
||
<paragraph blockId="4.[108,1214,748,1746]" pageId="4" pageNumber="5">
|
||
DESCRIPTION: Morphometrics summarized in table 1. Maximum observed total length
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="5" unit="mm" value="270.0">270 mm</quantity>
|
||
(standard length
|
||
<quantity box="[376,482,1421,1447]" metricMagnitude="-1" metricUnit="m" metricValue="2.44" pageId="4" pageNumber="5" unit="mm" value="244.0">244 mm</quantity>
|
||
). Head broad, depressed with slightly convex dorsal profile, snout rounded. Anterior nares tubular, medial to posterior nares and nasal barbels. Mouth subterminal, lips and bases of barbels papillose. Four pairs of barbels, all extending beyond dorsal fin origin. Barbels rounded at base, becoming flat, ellipsoid in cross section. Body cylindrical between head and anal fin origin, becoming laterally compressed posteriorly.
|
||
</paragraph>
|
||
<paragraph blockId="4.[108,1214,748,1746]" lastBlockId="6.[108,1215,1174,1723]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">Dorsal fin with 80–86 rays, originating a short distance (less than 10% SL) behind supraoccipital process. Anal fin with 57–61 rays, origin nearly equidistant from snout tip and caudal fin base. Dorsal and anal fins not confluent with caudal fin. Pectoral fin with robust spine and 10–12 rays. Spine sharply pointed with slight posterior curve. Strong serration present on both anterior and posterior margins. Serrae angled proximally, larger on posterior margin, increasing in number with body size. Pelvic fins present on all specimens examined, with six rays. Caudal fin rounded, with 18–22 rays.</paragraph>
|
||
<caption pageId="6" pageNumber="7" startId="6.[108,192,889,910]" targetBox="[108,1212,224,860]" targetPageId="6">
|
||
<paragraph blockId="6.[108,1213,887,1145]" pageId="6" pageNumber="7">
|
||
FIGURE 3. Computed tomography scan reconstruction of the skull and pectoral girdle of
|
||
<taxonomicName authorityName="Bernt & Stiassny" authorityYear="2022" box="[1005,1213,887,911]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="monsembulai">
|
||
<emphasis box="[1005,1213,887,911]" italics="true" pageId="6" pageNumber="7">Clarias monsembulai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[117,276,916,940]" italics="true" pageId="6" pageNumber="7">at left and above</emphasis>
|
||
), AMNH 252267, 179 mm SL in
|
||
<emphasis bold="true" box="[606,630,916,940]" pageId="6" pageNumber="7">A,</emphasis>
|
||
dorsal;
|
||
<emphasis bold="true" box="[707,729,916,940]" pageId="6" pageNumber="7">B,</emphasis>
|
||
ventral; and
|
||
<emphasis bold="true" box="[858,875,916,940]" pageId="6" pageNumber="7">C</emphasis>
|
||
, lateral views. Abbreviations:
|
||
<emphasis bold="true" box="[1168,1212,916,940]" pageId="6" pageNumber="7">ang,</emphasis>
|
||
anguloarticular;
|
||
<emphasis bold="true" box="[272,322,945,969]" pageId="6" pageNumber="7">apal,</emphasis>
|
||
autopalatine;
|
||
<emphasis bold="true" box="[462,490,945,969]" pageId="6" pageNumber="7">br,</emphasis>
|
||
branchiostegal rays;
|
||
<emphasis bold="true" box="[697,748,945,969]" pageId="6" pageNumber="7">ch-a,</emphasis>
|
||
anterior ceratohyal;
|
||
<emphasis bold="true" box="[951,1003,945,969]" pageId="6" pageNumber="7">ch-p,</emphasis>
|
||
posterior ceratohyal;
|
||
<emphasis bold="true" box="[108,132,975,999]" pageId="6" pageNumber="7">cl,</emphasis>
|
||
cleithrum;
|
||
<emphasis bold="true" box="[243,282,975,999]" pageId="6" pageNumber="7">cor,</emphasis>
|
||
coracoid;
|
||
<emphasis bold="true" box="[382,427,975,999]" pageId="6" pageNumber="7">den,</emphasis>
|
||
dentary;
|
||
<emphasis bold="true" box="[519,541,974,998]" pageId="6" pageNumber="7">fr,</emphasis>
|
||
frontal;
|
||
<emphasis bold="true" box="[624,667,975,999]" pageId="6" pageNumber="7">hhv,</emphasis>
|
||
ventral hypohyal;
|
||
<emphasis bold="true" box="[847,888,975,999]" pageId="6" pageNumber="7">hm,</emphasis>
|
||
hyomandibula;
|
||
<emphasis bold="true" box="[1046,1095,974,998]" pageId="6" pageNumber="7">io-ii,</emphasis>
|
||
infraorbital ii;
|
||
<emphasis bold="true" box="[134,191,1004,1028]" pageId="6" pageNumber="7">io-iii,</emphasis>
|
||
infraorbital iii;
|
||
<emphasis bold="true" box="[353,405,1004,1028]" pageId="6" pageNumber="7">io-iv,</emphasis>
|
||
infraorbital iv;
|
||
<emphasis bold="true" box="[564,603,1004,1028]" pageId="6" pageNumber="7">iop,</emphasis>
|
||
interopercle;
|
||
<emphasis bold="true" box="[742,777,1004,1028]" pageId="6" pageNumber="7">lac,</emphasis>
|
||
lacrimal;
|
||
<emphasis bold="true" box="[878,925,1004,1028]" pageId="6" pageNumber="7">leth,</emphasis>
|
||
lateral ethmoid;
|
||
<emphasis bold="true" box="[1097,1158,1004,1028]" pageId="6" pageNumber="7">meth,</emphasis>
|
||
mesethmoid;
|
||
<emphasis bold="true" box="[203,243,1033,1057]" pageId="6" pageNumber="7">mx,</emphasis>
|
||
maxilla;
|
||
<emphasis bold="true" box="[334,366,1033,1057]" pageId="6" pageNumber="7">na,</emphasis>
|
||
nasal;
|
||
<emphasis bold="true" box="[434,464,1033,1057]" pageId="6" pageNumber="7">ns,</emphasis>
|
||
neural spines;
|
||
<emphasis bold="true" box="[612,644,1033,1057]" pageId="6" pageNumber="7">op,</emphasis>
|
||
opercle;
|
||
<emphasis bold="true" box="[733,807,1033,1057]" pageId="6" pageNumber="7">o-susp,</emphasis>
|
||
os suspensorium;
|
||
<emphasis bold="true" box="[991,1044,1033,1057]" pageId="6" pageNumber="7">pmx,</emphasis>
|
||
premaxilla;
|
||
<emphasis bold="true" box="[1166,1212,1033,1057]" pageId="6" pageNumber="7">pop,</emphasis>
|
||
preopercle;
|
||
<emphasis bold="true" box="[227,294,1062,1086]" pageId="6" pageNumber="7">pp-v4,</emphasis>
|
||
parapophysis of 4th vertebra;
|
||
<emphasis bold="true" box="[608,674,1063,1087]" pageId="6" pageNumber="7">pp-v5,</emphasis>
|
||
parapophysis of 5th vertebra;
|
||
<emphasis bold="true" box="[989,1018,1063,1087]" pageId="6" pageNumber="7">ps,</emphasis>
|
||
pectoral spine;
|
||
<emphasis bold="true" box="[1183,1212,1063,1087]" pageId="6" pageNumber="7">pt,</emphasis>
|
||
pterotic;
|
||
<emphasis bold="true" box="[195,259,1092,1116]" pageId="6" pageNumber="7">pt-scl,</emphasis>
|
||
posttemporo-supracleithrum;
|
||
<emphasis bold="true" box="[561,581,1092,1116]" pageId="6" pageNumber="7">q,</emphasis>
|
||
quadrate;
|
||
<emphasis bold="true" box="[684,726,1092,1116]" pageId="6" pageNumber="7">rad,</emphasis>
|
||
radials;
|
||
<emphasis bold="true" box="[807,847,1092,1116]" pageId="6" pageNumber="7">soc,</emphasis>
|
||
supraoccipital;
|
||
<emphasis bold="true" box="[1001,1045,1092,1116]" pageId="6" pageNumber="7">sph,</emphasis>
|
||
sphenotic;
|
||
<emphasis bold="true" box="[1156,1212,1092,1116]" pageId="6" pageNumber="7">spop,</emphasis>
|
||
suprapreopercle;
|
||
<emphasis bold="true" box="[278,324,1121,1145]" pageId="6" pageNumber="7">trsc,</emphasis>
|
||
transcapular process;
|
||
<emphasis bold="true" box="[546,580,1121,1145]" pageId="6" pageNumber="7">uh,</emphasis>
|
||
urohyal;
|
||
<emphasis bold="true" box="[674,705,1121,1145]" pageId="6" pageNumber="7">vo,</emphasis>
|
||
vomer. Scale bars = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[108,1215,1174,1723]" pageId="6" pageNumber="7">
|
||
Elements of skull and pectoral girdle shown in figure 3. Neurocranium robustly ossified. Fourth infraorbital contacting suprapreopercle in all specimens examined (
|
||
<quantity box="[1012,1158,1324,1350]" metricMagnitude="-1" metricUnit="m" metricValue="1.6199999999999999" metricValueMax="2.44" metricValueMin="0.8" pageId="6" pageNumber="7" unit="mm" value="162.0" valueMax="244.0" valueMin="80.0">80–244 mm</quantity>
|
||
SL). Frontal fontanel long and narrow (knife shaped). Supraoccipital fontanel narrowly oblong, supraoccipital process sharply pointed in both juveniles and adults.
|
||
</paragraph>
|
||
<paragraph blockId="6.[108,1215,1174,1723]" lastBlockId="7.[108,1214,991,1764]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">Premaxillary and vomerine tooth plates similar in width, about 30% of head length. Premaxillary teeth villiform. Vomerine toothplate with short posterior median process, teeth subgranular. Dentary teeth villiform, extending over half of dorsal surface, nearly reaching coronoid process. First branchial arch with 16–18 slender gill rakers. Second and fourth arches diverticulate, forming arborescent suprabranchial organ (fig. 4). Rosette on second arch with few branches, not overlapping larger, more extensively, branched rosette on fourth arch. Respiratory organ occupies approximately one fourth of suprabranchial chamber volume. Seven or eight branchiostegal rays. Urohyal trifurcate, lateral processes longer than posteromedial process. Cleithrum superficial, visible externally as striated bony band at isthmus (fig. 5). Coracoid with single circular foramen at margin with cleithrum near articulation with pectoral spine.</paragraph>
|
||
<caption pageId="7" pageNumber="8" startId="7.[109,196,889,910]" targetBox="[108,1212,224,858]" targetPageId="7">
|
||
<paragraph blockId="7.[108,1212,886,940]" pageId="7" pageNumber="8">
|
||
FIGURE 4. Suprabranchial organ of
|
||
<taxonomicName authority=", AMNH" authorityName="AMNH" box="[496,808,887,911]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="monsembulai">
|
||
<emphasis box="[496,713,887,911]" italics="true" pageId="7" pageNumber="8">Clarias monsembulai</emphasis>
|
||
, AMNH
|
||
</taxonomicName>
|
||
244162, 165 mm SL (suprapreopercle removed). Scale bar = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[108,1214,991,1764]" pageId="7" pageNumber="8">Parapophyses of fourth and fifth vertebrae expanded, forming hourglass-shaped Weberian apparatus encapsulating bilobed gas bladder. Posterolateral processes of os suspensorium forming ventral floor of Weberian capsule. Fifty-eight to 61 (mode = 60) vertebrae posterior to first four vertebrae comprising Weberian apparatus. Two free neural spines between skull and first dorsal-fin pterygiophore. First rib articulated with seventh vertebra, 12 ribs present. No fusion of caudal fin hypurals.</paragraph>
|
||
<paragraph blockId="7.[108,1214,991,1764]" pageId="7" pageNumber="8">COLORATION: Preserved specimens are dark brown dorsally, fading to light brown, gray, or cream colored ventrally. Some lighter colored specimens exhibit a faint mottling of irregular dark brown markings over a slightly lighter background. Fins uniformly dark brown. Barbels brown at base, becoming creamy white over most of their length. Underside of head pale, but with band of dark pigment above fleshy furrow between mandibular barbels and isthmus. Lateral line visible as a series of small regularly spaced white pores extending from the base of the head to the base of the caudal peduncle. Pores of the secondary canals of the lateral line system form a regular pattern over the flanks, but these pores are not readily visible against the darkly pigmented skin.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph blockId="7.[108,1214,991,1764]" pageId="7" pageNumber="8">
|
||
DISTRIBUTION: The species is currently known only from the Momboyo, Luilaka, Salonga, and Yenge river systems within the Cuvette Centrale of the
|
||
<collectingCountry box="[808,978,1663,1689]" name="Democratic Republic of the Congo" pageId="7" pageNumber="8">middle Congo</collectingCountry>
|
||
River Basin (fig. 6). However additional collecting throughout the region will likely extend this range (E. Decru, personal commun.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption pageId="8" pageNumber="9" startId="8.[108,194,1019,1040]" targetBox="[408,934,232,992]" targetPageId="8">
|
||
<paragraph blockId="8.[108,1212,1017,1071]" pageId="8" pageNumber="9">
|
||
FIGURE 5. Exposed cleithrum (demarcated by dashed line) of
|
||
<emphasis bold="true" box="[759,783,1017,1041]" pageId="8" pageNumber="9">A,</emphasis>
|
||
<taxonomicName authority=", AMNH" authorityName="AMNH" box="[791,1081,1017,1042]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="buthupogon">
|
||
<emphasis box="[791,989,1017,1041]" italics="true" pageId="8" pageNumber="9">Clarias buthupogon</emphasis>
|
||
, AMNH
|
||
</taxonomicName>
|
||
227571, 162 mm SL and
|
||
<emphasis bold="true" box="[235,257,1047,1071]" pageId="8" pageNumber="9">B,</emphasis>
|
||
<taxonomicName authority=", AMNH" authorityName="AMNH" box="[263,566,1047,1071]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="monsembulai">
|
||
<emphasis box="[263,475,1047,1071]" italics="true" pageId="8" pageNumber="9">Clarias monsembulai</emphasis>
|
||
, AMNH
|
||
</taxonomicName>
|
||
244162, 183 mm SL. Scale bar = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph blockId="8.[108,1212,1103,1204]" pageId="8" pageNumber="9">
|
||
<materialsCitation location="central African" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Kinshasa">
|
||
ETYMOLOGY: Named in honor of Raoul Monsembula Iyaba (professor of biology, University of
|
||
<collectingRegion box="[185,293,1140,1166]" country="Republic of the Congo" name="Kinshasa" pageId="8" pageNumber="9">Kinshasa</collectingRegion>
|
||
) for collecting the
|
||
<typeStatus box="[513,564,1141,1167]" pageId="8" pageNumber="9">type</typeStatus>
|
||
series of this species, and in recognition of his substantial contributions to
|
||
<location LSID="urn:lsid:plazi:treatment:03D0F929FFD3FFE7FE008D23FD75FB0C:8EA61EE4FFD8FFE7FE9A8CB0FDCCFB0C" box="[351,529,1178,1204]" name="central African" pageId="8" pageNumber="9" stateProvince="Kinshasa">central African</location>
|
||
ichthyology
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |