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<document id="A975E3C7726CF883AA36B913110C8C1D" ID-CLB-Dataset="21945" ID-DOI="10.5852/ejt.2014.101" ID-GBIF-Dataset="3216f841-dd7a-4f3e-bf08-fee6aa61d454" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838770" ID-ZooBank="C320F2F8-E9BB-46FC-9A0C-56ECF4B1CE2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590096262994" checkinUser="tatiana" docAuthor="Taverne, Louis &amp; Capasso, Luigi" docDate="2014" docId="03C987FADB62FFA450DAF9C9FEBAFE11" docLanguage="en" docName="ejt-2014-101.pdf.imf" docOrigin="European Journal of Taxonomy 101" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Joinvillichthys kriweti Taverne &amp; Capasso 2014, sp. nov." docType="treatment" docUuid="B8C4822F-44CA-4776-8455-452C5DBDCA1C" docUuidSource="ZooBank" docVersion="7" lastPageNumber="19" masterDocId="FFF0FF82DB6EFFB65253FFA2FF86FFEA" masterDocTitle="On the “ Coccodus ” lindstroemi species complex (Pycnodontiformes, Gladiopycnodontidae) from the marine Late Cretaceous of Lebanon, with the description of two new genera" masterLastPageNumber="27" masterPageNumber="1" pageNumber="13" updateTime="1698840789535" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="73418825A0DA050ACEE17B910129DD12">On the “ Coccodus ” lindstroemi species complex (Pycnodontiformes, Gladiopycnodontidae) from the marine Late Cretaceous of Lebanon, with the description of two new genera</mods:title>
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<mods:name id="D95ED5223A874590E8A8835BC6DE1681" type="personal">
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<mods:namePart id="AF6556468BCB69A3DDA3E36252AC3C50">Taverne, Louis</mods:namePart>
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<mods:affiliation id="D1704A16F818758169E7F9344A8F01E3">Royal Belgian Institute of Natural Sciences of Belgium, Directorate Earth and History of Life, Rue Vautier 29, B- 1000 Brussels, Belgium. E-mail: louis. taverne @ gmail. com &amp; urn: lsid: zoobank. org: author: 0 CF 81641 - 1 DD 1 - 4 CBD- 9735 - F 1 FE 7 EB 0 BCF 5</mods:affiliation>
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<mods:namePart id="5D6C49BBD62823F98FB3BBC9BF2C2367">Capasso, Luigi</mods:namePart>
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<mods:affiliation id="908D52FB6A05A215FA5275574BE071D1">Museo Universitario dellUniversitá “ G. dAnnunzio ” di Chieti-Pescara, Piazza Trento e Trieste 1, I- 661000 Chieti, Italy. E-mail: lcapasso @ unich. it &amp; urn: lsid: zoobank. org: author: C 79 C 14 CF-C 1 D 7 - 48 E 3 - 9 BA 9 - CAD 8 AA 0909 F 7</mods:affiliation>
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<mods:title id="C1C11E897C047470BA9478BD16AE3E7A">European Journal of Taxonomy</mods:title>
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<mods:date id="D5BECE95208D87B870AFE3A3523DAE88">2014</mods:date>
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<treatment id="03C987FADB62FFA450DAF9C9FEBAFE11" ID-DOI="http://doi.org/10.5281/zenodo.3852234" ID-GBIF-Taxon="164247853" ID-Zenodo-Dep="3852234" LSID="urn:lsid:zoobank.org:act:B8C4822F-44CA-4776-8455-452C5DBDCA1C" httpUri="http://treatment.plazi.org/id/03C987FADB62FFA450DAF9C9FEBAFE11" lastPageId="18" lastPageNumber="19" pageId="12" pageNumber="13">
<subSubSection id="C37A6567DB62FFBA50DAF9C9FC71F96C" box="[649,1015,1643,1670]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BDF36ECDB62FFBA50DAF9C9FC71F96C" blockId="12.[400,1263,1643,1740]" box="[649,1015,1643,1670]" pageId="12" pageNumber="13">
<heading id="D0978180DB62FFBA50DAF9C9FC71F96C" box="[649,1015,1643,1670]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C604D6FDB62FFBA50DAF9C9FC13F96C" authority="Taverne &amp; Capasso, 2014" authorityName="Taverne &amp; Capasso" authorityYear="2014" box="[649,917,1643,1670]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kriweti" status="sp. nov.">
<emphasis id="B914EAFEDB62FFBA50DAF9C9FC13F96C" bold="true" box="[649,917,1643,1670]" italics="true" pageId="12" pageNumber="13">Joinvillichthys kriweti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2275785DB62FFBA51CFF9CEFC71F96C" box="[924,1015,1644,1670]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<paragraph id="8BDF36ECDB62FFBA5153F910FCF9F921" blockId="12.[400,1263,1643,1740]" box="[768,895,1713,1740]" pageId="12" pageNumber="13">
<figureCitation id="135B2A69DB62FFBA5153F910FCF9F921" box="[768,895,1713,1740]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="13.[225,268,2038,2064]" captionStartId-1="14.[227,270,2038,2064]" captionStartId-2="15.[227,270,2006,2032]" captionStartId-3="16.[227,270,2002,2028]" captionStartId-4="17.[225,268,2000,2026]" captionTargetBox-0="[227,1436,1189,1999]" captionTargetBox-1="[269,1391,1278,1956]" captionTargetBox-2="[281,1384,965,1949]" captionTargetBox-3="[432,1274,999,1916]" captionTargetBox-4="[370,1266,988,1944]" captionTargetId-0="figure@13.[227,1436,1189,1999]" captionTargetId-1="figure@14.[264,1398,1270,1992]" captionTargetId-2="figure@15.[265,1399,948,1965]" captionTargetId-3="figure@16.[397,1305,985,1959]" captionTargetId-4="figure@17.[355,1280,952,1953]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionText-0="Fig. 12. Joinvillichthys kriweti gen. et sp. nov. Holotype, CLC S-137." captionText-1="Fig. 13. Joinvillichthys kriweti gen. et sp. nov. General reconstruction based on holotype, CLC S-137." captionText-2="Fig. 14. Joinvillichthys kriweti gen. et sp. nov. Reconstruction of the skull and the pectoral girdle based on holotype, CLC S-137." captionText-3="Fig. 15. Joinvillichthys kriweti gen. et sp. nov. Caudal skeleton of holotype, CLC S-137. The arrows indicate the positions of the most external dorsal and ventral procurrent rays of the caudal fin." captionText-4="Fig. 16. Joinvillichthys kriweti gen. et sp. nov. Holotype, CLC S-137. A. Dorsal ridge scutes. B. Anterior flake-like body scales. C. Posterior scute-like body scale. D. Ventral keel scutes." figureDoi-0="http://doi.org/10.5281/zenodo.3838796" figureDoi-1="http://doi.org/10.5281/zenodo.3838798" figureDoi-2="http://doi.org/10.5281/zenodo.3838800" figureDoi-3="http://doi.org/10.5281/zenodo.3838802" figureDoi-4="http://doi.org/10.5281/zenodo.3838804" httpUri-0="https://zenodo.org/record/3838796/files/figure.png" httpUri-1="https://zenodo.org/record/3838798/files/figure.png" httpUri-2="https://zenodo.org/record/3838800/files/figure.png" httpUri-3="https://zenodo.org/record/3838802/files/figure.png" httpUri-4="https://zenodo.org/record/3838804/files/figure.png" pageId="12" pageNumber="13">Figs 12-16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A6567DB62FFBA52B0F951FBB0F822" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BDF36ECDB62FFBA52B0F951FEDAF8E7" blockId="12.[227,348,1779,1805]" box="[227,348,1779,1805]" pageId="12" pageNumber="13">
<heading id="D0978180DB62FFBA52B0F951FEDAF8E7" bold="true" box="[227,348,1779,1805]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B914EAFEDB62FFBA52B0F951FEDAF8E7" bold="true" box="[227,348,1779,1805]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36ECDB62FFBA52B0F880FBB0F822" blockId="12.[227,1437,1825,1993]" pageId="12" pageNumber="13">
<taxonomicName id="4C604D6FDB62FFBA52B0F880FE08F8D6" authorityName="Taverne &amp; Capasso" authorityYear="2014" box="[227,398,1826,1852]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B914EAFEDB62FFBA52B0F880FE08F8D6" box="[227,398,1826,1852]" italics="true" pageId="12" pageNumber="13">Joinvillichthys</emphasis>
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with a body depth equal to 46.0 % of the standard length. A dorsal prominence present on the frontal. Maxilla elongated. Large parietal. Dermosupraoccipital sutured to the parietal and not to the dermopterotic. Dermopterotic much longer than deep. Small dermosphenotic. Comma-shaped opercle. Anterior ventral branch of the cleithrum lost. Broad and short pectoral spine articulated on the ventral margin of the cleithral posterior process. Caudal fin double emarginated.
</paragraph>
</subSubSection>
<subSubSection id="C37A6567DB63FFBB52B2FE92FD45FE71" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BDF36ECDB63FFBB52B2FE92FEEEFEA0" blockId="13.[225,360,304,330]" box="[225,360,304,330]" pageId="13" pageNumber="14">
<heading id="D0978180DB63FFBB52B2FE92FEEEFEA0" bold="true" box="[225,360,304,330]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B914EAFEDB63FFBB52B2FE92FEEEFEA0" bold="true" box="[225,360,304,330]" pageId="13" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36ECDB63FFBB52B2FEFCFD45FE71" blockId="13.[225,1436,350,411]" pageId="13" pageNumber="14">The name of the new species is dedicated to Dr Jürgen Kriwet (Vienna) who has greatly improved our knowledge of the pycnodontiform fishes.</paragraph>
</subSubSection>
<subSubSection id="C37A6567DB63FFBB52B2FE61FABBFD75" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BDF36ECDB63FFBB52B2FE61FED5FE37" blockId="13.[225,339,451,477]" box="[225,339,451,477]" pageId="13" pageNumber="14">
<heading id="D0978180DB63FFBB52B2FE61FED5FE37" bold="true" box="[225,339,451,477]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B914EAFEDB63FFBB52B2FE61FED5FE37" bold="true" box="[225,339,451,477]" pageId="13" pageNumber="14">
<typeStatus id="54DB884EDB63FFBB52B2FE61FED5FE37" box="[225,339,451,477]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36ECDB63FFBB52B2FE53FADDFDE6" blockId="13.[225,1371,497,524]" box="[225,1371,497,524]" pageId="13" pageNumber="14">
<materialsCitation id="3B083CB1DB63FFBB52B2FE53FADDFDE6" ID-GBIF-Occurrence="2621773303" box="[225,1371,497,524]" country="Lebanon" location="LEBANON" pageId="13" pageNumber="14" specimenCode="sample CLC S-137" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F377767CDB63FFBB52B2FE53FEF4FDE1" box="[225,370,497,523]" name="Lebanon" pageId="13" pageNumber="14">LEBANON</collectingCountry>
:
<specimenCode id="DBC69E97DB63FFBB532EFE53FDE6FDE1" box="[381,608,497,523]" pageId="13" pageNumber="14">sample CLC S-137</specimenCode>
, a complete specimen from Haqel (
<figureCitation id="135B2A69DB63FFBB5653FE53FBD1FDE1" box="[1024,1111,497,523]" captionStart="Fig" captionStartId="13.[225,268,2038,2064]" captionTargetBox="[227,1436,1189,1999]" captionTargetId="figure@13.[227,1436,1189,1999]" captionTargetPageId="13" captionText="Fig. 12. Joinvillichthys kriweti gen. et sp. nov. Holotype, CLC S-137." figureDoi="http://doi.org/10.5281/zenodo.3838796" httpUri="https://zenodo.org/record/3838796/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
), total length:
<quantity id="4C989B09DB63FFBB5752FE53FAD1FDE6" box="[1281,1367,497,524]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="13" pageNumber="14" unit="mm" value="91.0">91 mm</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDF36ECDB63FFBB52B2FD91FED5FDA7" blockId="13.[225,339,563,589]" box="[225,339,563,589]" pageId="13" pageNumber="14">
<heading id="D0978180DB63FFBB52B2FD91FED5FDA7" bold="true" box="[225,339,563,589]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B914EAFEDB63FFBB52B2FD91FED5FDA7" bold="true" box="[225,339,563,589]" pageId="13" pageNumber="14">
<typeStatus id="54DB884EDB63FFBB52B2FD91FED5FDA7" box="[225,339,563,589]" pageId="13" pageNumber="14" type="paratype">Paratype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36ECDB63FFBB52B2FDC3FABBFD75" blockId="13.[225,1437,609,671]" pageId="13" pageNumber="14">
<collectingCountry id="F377767CDB63FFBB52B2FDC3FEF4FD91" box="[225,370,609,635]" name="Lebanon" pageId="13" pageNumber="14">LEBANON</collectingCountry>
: sample
<collectionCode id="ED71AE29DB63FFBB538BFDC3FDB4FD91" box="[472,562,609,635]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="14">AMNH</collectionCode>
4517a (3698) and counterpart, an incomplete specimen from Hgula (
<bibRefCitation id="EFF14B1DDB63FFBB5739FDC3FEA6FD74" author="Hay O. P." pageId="13" pageNumber="14" pagination="395 - 452" refId="ref9350" refString="Hay O. P. 1903. On a collection of Upper Cretaceous fishes from Mount Lebanon, Syria, with descriptions of four new genera and nineteen new species. Bulletin of the American Museum of Natural History 19 (10): 395 - 452." type="journal article" year="1903">Hay 1903</bibRefCitation>
: pl. 29,
<figureCitation id="135B2A69DB63FFBB532EFD26FE3EFD74" box="[381,440,644,670]" captionStart="Fig" captionStartId="4.[227,270,2036,2062]" captionTargetBox="[227,1436,1194,2000]" captionTargetId="figure@4.[227,1436,1194,2000]" captionText="Fig. 1. Joinvillichthys lindstroemi (Davis, 1890). Holotype, NRM PZ P. 2073." figureDoi="http://doi.org/10.5281/zenodo.3838772" httpUri="https://zenodo.org/record/3838772/files/figure.png" pageId="13" pageNumber="14">fig. 1</figureCitation>
); only the head and the beginning of the body are preserved, length:
<quantity id="4C989B09DB63FFBB56B0FD26FABFFD75" box="[1251,1337,644,671]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="13" pageNumber="14" unit="mm" value="63.0">63 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A6567DB63FFBB52B2FD67FCF3FCE4" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BDF36ECDB63FFBB52B2FD67FD86FD35" blockId="13.[225,512,709,736]" box="[225,512,709,736]" pageId="13" pageNumber="14">
<heading id="D0978180DB63FFBB52B2FD67FD86FD35" bold="true" box="[225,512,709,736]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B914EAFEDB63FFBB52B2FD67FD86FD35" bold="true" box="[225,512,709,736]" pageId="13" pageNumber="14">Formation and locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36ECDB63FFBB52B2FD56FCF3FCE4" blockId="13.[225,885,756,783]" box="[225,885,756,783]" pageId="13" pageNumber="14">
Marine Upper Cenomanian, Haqel and Hgula,
<collectingCountry id="F377767CDB63FFBB5155FD56FCF7FCE4" box="[774,881,756,782]" name="Lebanon" pageId="13" pageNumber="14">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A6567DB63FFA452B2FC97FEBAFE11" lastPageId="18" lastPageNumber="19" pageId="13" pageNumber="14" type="description">
<paragraph id="8BDF36ECDB63FFBB52B2FC97FDC0FCBA" blockId="13.[225,582,821,848]" box="[225,582,821,848]" pageId="13" pageNumber="14">
<heading id="D0978180DB63FFBB52B2FC97FE5EFCBA" box="[225,472,821,848]" pageId="13" pageNumber="14">
<emphasis id="B914EAFEDB63FFBB52B2FC97FE5EFCBA" bold="true" box="[225,472,821,848]" pageId="13" pageNumber="14">Morphometric data</emphasis>
</heading>
(
<figureCitation id="135B2A69DB63FFBB53BBFC94FDBBFCBA" box="[488,573,822,848]" captionStart="Fig" captionStartId="14.[227,270,2038,2064]" captionTargetBox="[269,1391,1278,1956]" captionTargetId="figure@14.[264,1398,1270,1992]" captionText="Fig. 13. Joinvillichthys kriweti gen. et sp. nov. General reconstruction based on holotype, CLC S-137." figureDoi="http://doi.org/10.5281/zenodo.3838798" httpUri="https://zenodo.org/record/3838798/files/figure.png" pageId="13" pageNumber="14">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="8BDF36ECDB63FFBB52B2FCC6FB0EFC95" blockId="13.[225,1160,868,895]" box="[225,1160,868,895]" pageId="13" pageNumber="14">
The morphometric data are given in % of the
<typeStatus id="54DB884EDB63FFBB50ABFCC6FCD9FC94" box="[760,863,868,894]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
standard length (
<quantity id="4C989B09DB63FFBB5679FCC6FB06FC95" box="[1066,1152,868,895]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="13" pageNumber="14" unit="mm" value="76.0">76 mm</quantity>
)
</paragraph>
<paragraph id="8BDF36ECDB63FFB852B2FC04FA1AFE32" blockId="13.[225,1436,933,1101]" lastBlockId="14.[227,1436,305,472]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">Length of the head (dermosupraoccipital included) …………………………………………… 54.2 % Length of the cephalo-thorax (cleithrum included) ………………………………………… 75.3 % Depth of the head (without the nuchal horn) ……………………………………………… 43.8 % Length of the nuchal horn …………………………………………………………………… 27.7 % Maximum depth of the body (just behind the nuchal horn) ………………………………… 46.0 % Predorsal length ………………………………………………………………………………… 76.6 % Basal length of the dorsal fin …………………………………………………………………… 9.8 % Preanal length …………………………………………………………………………………… 80.4 % Basal length of the anal fin ………………………………………………………………………… 6.4 % Depth of the caudal peduncle ……………………………………………………………………… 9.8 %</paragraph>
<caption id="DF1F6664DB63FFBB52B2F854FB97F7FA" ID-DOI="http://doi.org/10.5281/zenodo.3838796" ID-Zenodo-Dep="3838796" box="[225,1041,2038,2065]" httpUri="https://zenodo.org/record/3838796/files/figure.png" pageId="13" pageNumber="14" startId="13.[225,268,2038,2064]" targetBox="[227,1436,1189,1999]" targetPageId="13">
<paragraph id="8BDF36ECDB63FFBB52B2F854FB97F7FA" blockId="13.[225,1041,2038,2065]" box="[225,1041,2038,2065]" pageId="13" pageNumber="14">
<emphasis id="B914EAFEDB63FFBB52B2F854FEB8F7FA" bold="true" box="[225,318,2038,2064]" pageId="13" pageNumber="14">Fig. 12.</emphasis>
<taxonomicName id="4C604D6FDB63FFBB5316F854FDCEF7FA" authority="Taverne &amp; Capasso, 2014" authorityName="Taverne &amp; Capasso" authorityYear="2014" box="[325,584,2038,2064]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="kriweti" status="gen. et sp. nov.">
<emphasis id="B914EAFEDB63FFBB5316F854FDCEF7FA" box="[325,584,2038,2064]" italics="true" pageId="13" pageNumber="14">Joinvillichthys kriweti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2275785DB63FFBB5003F855FC86F7FB" box="[592,768,2039,2065]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Holotype, CLC S-137.
</paragraph>
</caption>
<paragraph id="8BDF36ECDB60FFB852B0FE5DFED9FDF3" blockId="14.[227,351,511,537]" box="[227,351,511,537]" pageId="14" pageNumber="15">
<heading id="D0978180DB60FFB852B0FE5DFED9FDF3" bold="true" box="[227,351,511,537]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B914EAFEDB60FFB852B0FE5DFED9FDF3" bold="true" box="[227,351,511,537]" pageId="14" pageNumber="15">Osteology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36ECDB60FFB85343FD8CFD96FDA2" blockId="14.[227,1436,558,725]" box="[272,528,558,584]" pageId="14" pageNumber="15">
<emphasis id="B914EAFEDB60FFB85343FD8CFE27FDA2" bold="true" box="[272,417,558,584]" pageId="14" pageNumber="15">1. The skull</emphasis>
(
<figureCitation id="135B2A69DB60FFB853E1FD8CFD80FDA2" box="[434,518,558,584]" captionStart="Fig" captionStartId="15.[227,270,2006,2032]" captionTargetBox="[281,1384,965,1949]" captionTargetId="figure@15.[265,1399,948,1965]" captionTargetPageId="15" captionText="Fig. 14. Joinvillichthys kriweti gen. et sp. nov. Reconstruction of the skull and the pectoral girdle based on holotype, CLC S-137." figureDoi="http://doi.org/10.5281/zenodo.3838800" httpUri="https://zenodo.org/record/3838800/files/figure.png" pageId="14" pageNumber="15">Fig. 14</figureCitation>
)
</paragraph>
<paragraph id="8BDF36ECDB60FFB852B0FDF3FEEBFD3F" blockId="14.[227,1436,558,725]" pageId="14" pageNumber="15">
The general morphology of the skull is rather close to that of
<taxonomicName id="4C604D6FDB60FFB851E1FDF3FB6FFD81" baseAuthorityName="Davis" baseAuthorityYear="1890" box="[946,1257,593,619]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lindstroemi">
<emphasis id="B914EAFEDB60FFB851E1FDF3FB6FFD81" box="[946,1257,593,619]" italics="true" pageId="14" pageNumber="15">Joinvillichthys lindstroemi</emphasis>
</taxonomicName>
and the cranial dermal bones also are ornamented with small tubercles. However, there are many small differences in the head skeleton of the two fishes. Thus the description that follows will principally emphasize these differences.
</paragraph>
<paragraph id="8BDF36ECDB60FFB852B0FD5EFCA9FCB6" blockId="14.[227,1436,764,861]" pageId="14" pageNumber="15">
The skull is shorter and deeper than in
<taxonomicName id="4C604D6FDB60FFB850E0FD5EFC6DFCFC" baseAuthorityName="Davis" baseAuthorityYear="1890" box="[691,1003,764,790]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lindstroemi">
<emphasis id="B914EAFEDB60FFB850E0FD5EFC6DFCFC" box="[691,1003,764,790]" italics="true" pageId="14" pageNumber="15">Joinvillichthys lindstroemi</emphasis>
</taxonomicName>
. Its depth, from the upper margin of the dermosupraoccipital to the lower margin of the cleithrum, is equal to 83 to 86 % of its length, from the tip of the snout to the basis of the nuchal horn.
</paragraph>
<paragraph id="8BDF36ECDB60FFB852B0FC26FADFFB9B" blockId="14.[227,1437,900,1138]" pageId="14" pageNumber="15">
The rostrum lengthening is less pronounced. The prefrontal is broader and has a very sinuous suture with the frontal. Its anterior tip also bears very small spines but is less outpacing of the lower jaw level. The frontal is broader but does not outpace posteriorly the level of the orbit. The bone bears a small dorsal prominence. The dermosupraoccipital is longer and is sutured to the parietal and the supratemporal but not with the dermopterotic. The parietal is considerably larger. The dermopterotic is longer but much thinner. The supratemporal is sutured to the parietal and reaches the dermopterotic at only one point. As in
<taxonomicName id="4C604D6FDB60FFB85352FBF5FDB1FB9B" baseAuthorityName="Davis" baseAuthorityYear="1890" box="[257,567,1111,1137]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lindstroemi">
<emphasis id="B914EAFEDB60FFB85352FBF5FDB1FB9B" box="[257,567,1111,1137]" italics="true" pageId="14" pageNumber="15">Joinvillichthys lindstroemi</emphasis>
</taxonomicName>
, the long nuchal horn is supported only by the dermosupraoccipital.
</paragraph>
<caption id="DF1F6664DB60FFB852B0F854FA08F7FA" ID-DOI="http://doi.org/10.5281/zenodo.3838798" ID-Zenodo-Dep="3838798" box="[227,1422,2038,2065]" httpUri="https://zenodo.org/record/3838798/files/figure.png" pageId="14" pageNumber="15" startId="14.[227,270,2038,2064]" targetBox="[269,1391,1278,1956]" targetPageId="14">
<paragraph id="8BDF36ECDB60FFB852B0F854FE95F7FA" blockId="14.[227,275,2038,2064]" box="[227,275,2038,2064]" pageId="14" pageNumber="15">
<emphasis id="B914EAFEDB60FFB852B0F854FE95F7FA" bold="true" box="[227,275,2038,2064]" pageId="14" pageNumber="15">Fig.</emphasis>
</paragraph>
<paragraph id="8BDF36ECDB60FFB85349F854FA08F7FA" blockId="14.[282,1337,2038,2065]" box="[282,1422,2038,2065]" lastBlockId="14.[1344,1422,2038,2064]" pageId="14" pageNumber="15">
<emphasis id="B914EAFEDB60FFB85349F854FEB9F7FA" bold="true" box="[282,319,2038,2064]" pageId="14" pageNumber="15">13.</emphasis>
<taxonomicName id="4C604D6FDB60FFB85315F854FDCCF7FA" authority="Taverne &amp; Capasso, 2014" authorityName="Taverne &amp; Capasso" authorityYear="2014" box="[326,586,2038,2064]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="kriweti" status="gen. et sp. nov.">
<emphasis id="B914EAFEDB60FFB85315F854FDCCF7FA" box="[326,586,2038,2064]" italics="true" pageId="14" pageNumber="15">Joinvillichthys kriweti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2275785DB60FFB85002F855FC84F7FB" box="[593,770,2039,2065]" pageId="14" pageNumber="15" rank="species">gen. et sp. nov.</taxonomicNameLabel>
General reconstruction based on
<typeStatus id="54DB884EDB60FFB856D8F854FB73F7FA" box="[1163,1269,2038,2064]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
, CLC S-137.
</paragraph>
</caption>
<paragraph id="8BDF36ECDB61FFB952B0FE92FDA7FE87" blockId="15.[227,1436,304,366]" pageId="15" pageNumber="16">The orbitosphenoid and the pleurosphenoid are present in the orbit, just below the frontal, but the basisphenoid is not visible.</paragraph>
<paragraph id="8BDF36ECDB61FFB952B0FE37FD3AFDF2" blockId="15.[227,1437,405,536]" pageId="15" pageNumber="16">The toothless premaxilla is longer and narrower. The toothless maxilla also is narrower and more elongate. The lower jaw is composed with the same bones but is longer. The dentary bears two small incisiform teeth and its ventral margin is denticulated. The articulation with the quadrate is located at the level of the posterior border of the orbit.</paragraph>
<paragraph id="8BDF36ECDB61FFB952B0FDE2FD34FD97" blockId="15.[227,1437,576,637]" pageId="15" pageNumber="16">A fragment of a large first infraorbital is preserved on the suture between the prefrontal and the premaxilla. The sclerotic ring is visible in the orbit.</paragraph>
<paragraph id="8BDF36ECDB61FFB952B0FD06FE95FCA1" blockId="15.[227,1437,676,843]" pageId="15" pageNumber="16">
The hyomandibula and the preopercle are sutured together. The exposed part of the hyomandibuladermohyomandibula is larger than in
<taxonomicName id="4C604D6FDB61FFB95090FD6AFB84FD08" baseAuthorityName="Davis" baseAuthorityYear="1890" box="[707,1026,712,738]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lindstroemi">
<emphasis id="B914EAFEDB61FFB95090FD6AFB84FD08" box="[707,1026,712,738]" italics="true" pageId="15" pageNumber="16">Joinvillichthys lindstroemi</emphasis>
</taxonomicName>
but still much smaller than the considerably enlarged preopercle. The opercle is broader and comma-shaped, with the sharp end dorsally located. A part of the anterior ceratohyal and two small branchiostegal rays are visible behind the lower jaw.
</paragraph>
<caption id="DF1F6664DB61FFB952B0F874FD8DF7F9" ID-DOI="http://doi.org/10.5281/zenodo.3838800" ID-Zenodo-Dep="3838800" httpUri="https://zenodo.org/record/3838800/files/figure.png" pageId="15" pageNumber="16" startId="15.[227,270,2006,2032]" targetBox="[281,1384,965,1949]" targetPageId="15">
<paragraph id="8BDF36ECDB61FFB952B0F874FD8DF7F9" blockId="15.[227,1436,2006,2068]" pageId="15" pageNumber="16">
<emphasis id="B914EAFEDB61FFB952B0F874FEB9F81A" bold="true" box="[227,319,2006,2032]" pageId="15" pageNumber="16">Fig. 14.</emphasis>
<taxonomicName id="4C604D6FDB61FFB95315F874FDCFF81A" authority="Taverne &amp; Capasso, 2014" authorityName="Taverne &amp; Capasso" authorityYear="2014" box="[326,585,2006,2032]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="kriweti" status="gen. et sp. nov.">
<emphasis id="B914EAFEDB61FFB95315F874FDCFF81A" box="[326,585,2006,2032]" italics="true" pageId="15" pageNumber="16">Joinvillichthys kriweti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2275785DB61FFB95002F874FC87F81A" box="[593,769,2006,2032]" pageId="15" pageNumber="16" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Reconstruction of the skull and the pectoral girdle based on holotype, CLC S-137.
</paragraph>
</caption>
<paragraph id="8BDF36ECDB7EFFA65343FE93FDDEFEA6" blockId="16.[227,1436,305,472]" box="[272,600,305,332]" pageId="16" pageNumber="17">
<emphasis id="B914EAFEDB7EFFA65343FE93FE3CFEA1" bold="true" box="[272,442,305,331]" pageId="16" pageNumber="17">2. The girdles</emphasis>
(
<figureCitation id="135B2A69DB7EFFA6539AFE93FDA3FEA1" box="[457,549,305,331]" captionStart="Fig" captionStartId="15.[227,270,2006,2032]" captionTargetBox="[281,1384,965,1949]" captionTargetId="figure@15.[265,1399,948,1965]" captionTargetPageId="15" captionText="Fig. 14. Joinvillichthys kriweti gen. et sp. nov. Reconstruction of the skull and the pectoral girdle based on holotype, CLC S-137." figureDoi="http://doi.org/10.5281/zenodo.3838800" httpUri="https://zenodo.org/record/3838800/files/figure.png" pageId="16" pageNumber="17">Figs 14</figureCitation>
,
<figureCitation id="135B2A69DB7EFFA65062FE93FDC9FEA6" box="[561,591,305,332]" captionStart="Fig" captionStartId="17.[225,268,2000,2026]" captionTargetBox="[370,1266,988,1944]" captionTargetId="figure@17.[355,1280,952,1953]" captionTargetPageId="17" captionText="Fig. 16. Joinvillichthys kriweti gen. et sp. nov. Holotype, CLC S-137. A. Dorsal ridge scutes. B. Anterior flake-like body scales. C. Posterior scute-like body scale. D. Ventral keel scutes." figureDoi="http://doi.org/10.5281/zenodo.3838804" httpUri="https://zenodo.org/record/3838804/files/figure.png" pageId="16" pageNumber="17">16</figureCitation>
)
</paragraph>
<paragraph id="8BDF36ECDB7EFFA652B0FEF6FAF2FE32" blockId="16.[227,1436,305,472]" pageId="16" pageNumber="17">
The bones of the gigantic pectoral girdle have the same size and shape as in
<taxonomicName id="4C604D6FDB7EFFA6560DFEF6FA13FE84" baseAuthorityName="Davis" baseAuthorityYear="1890" box="[1118,1429,340,366]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="lindstroemi">
<emphasis id="B914EAFEDB7EFFA6560DFEF6FA13FE84" box="[1118,1429,340,366]" italics="true" pageId="16" pageNumber="17">Joinvillichthys lindstroemi</emphasis>
</taxonomicName>
. However, two important differences exist. The anterior ventral branch of the cleithrum is lost. No postcleithrum is visible, but that is perhaps due to the fossilization. The pectoral spine is shorter, much broader and is not articulated with the rear of the cleithrum but more anteriorly on its lower margin.
</paragraph>
<paragraph id="8BDF36ECDB7EFFA652B0FDA2FDB2FDD6" blockId="16.[227,1436,511,573]" pageId="16" pageNumber="17">A small pelvic girdle is present. Indeed, a part of a vertically oriented pelvic bone is visible under a broken part of the cleithrum.</paragraph>
<paragraph id="8BDF36ECDB7EFFA65343FDC6FDFBFD94" blockId="16.[227,1437,612,849]" box="[272,637,612,638]" pageId="16" pageNumber="17">
<emphasis id="B914EAFEDB7EFFA65343FDC6FD96FD94" bold="true" box="[272,528,612,638]" pageId="16" pageNumber="17">3. The axial skeleton</emphasis>
(
<figureCitation id="135B2A69DB7EFFA65073FDC6FDF2FD94" box="[544,628,612,638]" captionStart="Fig" captionStartId="14.[227,270,2038,2064]" captionTargetBox="[269,1391,1278,1956]" captionTargetId="figure@14.[264,1398,1270,1992]" captionText="Fig. 13. Joinvillichthys kriweti gen. et sp. nov. General reconstruction based on holotype, CLC S-137." figureDoi="http://doi.org/10.5281/zenodo.3838798" httpUri="https://zenodo.org/record/3838798/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="8BDF36ECDB7EFFA652B0FD25FC43FCBB" blockId="16.[227,1437,612,849]" pageId="16" pageNumber="17">
The trunk is fusiform but proportionally deeper than in
<taxonomicName id="4C604D6FDB7EFFA6513BFD25FB19FD4B" baseAuthorityName="Davis" baseAuthorityYear="1890" box="[872,1183,647,673]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="lindstroemi">
<emphasis id="B914EAFEDB7EFFA6513BFD25FB19FD4B" box="[872,1183,647,673]" italics="true" pageId="16" pageNumber="17">Joinvillichthys lindstroemi</emphasis>
</taxonomicName>
. The axial skeleton is incomplete. Three vertebral segments are missing near the caudal peduncle. There are 16 neural spines (the three missing ones included) before the epichordal series. Only 8 haemal spines are preserved. The total number of haemal spines must be around 12 or 13. The neural and haemal spines are short but broad. The neural and haemal arches surround almost completely the notochord. No ribs are visible. The postcoelomic bone is well developed and backwardly oriented.
</paragraph>
<caption id="DF1F6664DB7EFFA652B0F870FAA8F7E5" ID-DOI="http://doi.org/10.5281/zenodo.3838802" ID-Zenodo-Dep="3838802" httpUri="https://zenodo.org/record/3838802/files/figure.png" pageId="16" pageNumber="17" startId="16.[227,270,2002,2028]" targetBox="[432,1274,999,1916]" targetPageId="16">
<paragraph id="8BDF36ECDB7EFFA652B0F870FA1AF807" blockId="16.[227,1436,2002,2029]" box="[227,1436,2002,2029]" pageId="16" pageNumber="17">
<emphasis id="B914EAFEDB7EFFA652B0F870FEC7F806" bold="true" box="[227,321,2002,2028]" pageId="16" pageNumber="17">Fig. 15.</emphasis>
<taxonomicName id="4C604D6FDB7EFFA65318F870FDD6F806" authority="Taverne &amp; Capasso, 2014" authorityName="Taverne &amp; Capasso" authorityYear="2014" box="[331,592,2002,2028]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kriweti" status="gen. et sp. nov.">
<emphasis id="B914EAFEDB7EFFA65318F870FDD6F806" box="[331,592,2002,2028]" italics="true" pageId="16" pageNumber="17">Joinvillichthys kriweti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2275785DB7EFFA65009F871FC97F807" box="[602,785,2003,2029]" pageId="16" pageNumber="17" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Caudal skeleton of holotype, CLC S-137. The arrows
</paragraph>
<paragraph id="8BDF36ECDB7EFFA652B0F857FAA8F7E5" blockId="16.[227,320,2037,2063]" box="[227,1326,2037,2064]" lastBlockId="16.[1322,1326,2037,2063]" pageId="16" pageNumber="17">indicate the positions of the most external dorsal and ventral procurrent rays of the caudal fin.</paragraph>
</caption>
<paragraph id="8BDF36ECDB7FFFA7535CFE93FD41FEA1" blockId="17.[225,1436,305,437]" box="[271,711,305,331]" pageId="17" pageNumber="18">
<emphasis id="B914EAFEDB7FFFA7535CFE93FDDFFEA1" bold="true" box="[271,601,305,331]" pageId="17" pageNumber="18">4. The dorsal and anal fins</emphasis>
(
<figureCitation id="135B2A69DB7FFFA7503AFE93FD38FEA1" box="[617,702,305,331]" captionStart="Fig" captionStartId="14.[227,270,2038,2064]" captionTargetBox="[269,1391,1278,1956]" captionTargetId="figure@14.[264,1398,1270,1992]" captionText="Fig. 13. Joinvillichthys kriweti gen. et sp. nov. General reconstruction based on holotype, CLC S-137." figureDoi="http://doi.org/10.5281/zenodo.3838798" httpUri="https://zenodo.org/record/3838798/files/figure.png" pageId="17" pageNumber="18">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="8BDF36ECDB7FFFA752B2FEF6FB26FE5F" blockId="17.[225,1436,305,437]" pageId="17" pageNumber="18">The short dorsal and anal fins are located at the mid-length of the body. There are 9 pterygiophores and 9 rays in the dorsal fin. The anal fin contains 7 rays, but the number of anal pterygiophores is not determinable. The first dorsal and anal ray is spiny. The other rays are segmented.</paragraph>
<paragraph id="8BDF36ECDB7FFFA7535CFE7EFD14FE1D" blockId="17.[225,1437,476,643]" box="[271,658,476,503]" pageId="17" pageNumber="18">
<emphasis id="B914EAFEDB7FFFA7535CFE7EFDA3FE1C" bold="true" box="[271,549,476,502]" pageId="17" pageNumber="18">5. The caudal skeleton</emphasis>
(
<figureCitation id="135B2A69DB7FFFA75066FE7EFD0CFE1D" box="[565,650,476,503]" captionStart="Fig" captionStartId="16.[227,270,2002,2028]" captionTargetBox="[432,1274,999,1916]" captionTargetId="figure@16.[397,1305,985,1959]" captionTargetPageId="16" captionText="Fig. 15. Joinvillichthys kriweti gen. et sp. nov. Caudal skeleton of holotype, CLC S-137. The arrows indicate the positions of the most external dorsal and ventral procurrent rays of the caudal fin." figureDoi="http://doi.org/10.5281/zenodo.3838802" httpUri="https://zenodo.org/record/3838802/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
)
</paragraph>
<paragraph id="8BDF36ECDB7FFFA752B2FE5DFBF1FD69" blockId="17.[225,1437,476,643]" pageId="17" pageNumber="18">
The caudal skeleton of the
<typeStatus id="54DB884EDB7FFFA75070FE5DFD0CFDF3" box="[547,650,511,537]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
is partly preserved. There are 6 short and broad epichordals and 7 hypochordals. However, one or two anterior hypochordals are missing, so the complete series must be composed of 8 or 9 elements. The fifth preserved hypochordal is strongly enlarged. No urodermal is visible, but the region where theses bones are usually present is not preserved.
</paragraph>
<paragraph id="8BDF36ECDB7FFFA752B2FD08FDDCFCE1" blockId="17.[225,1436,682,779]" pageId="17" pageNumber="18">
The caudal fin is double emarginated (
<bibRefCitation id="EFF14B1DDB7FFFA750F4FD08FC70FD2F" author="Poyato-Ariza F. J. &amp; Wenz S." box="[679,1014,682,709]" pageId="17" pageNumber="18" pagination="139 - 248" refId="ref9514" refString="Poyato-Ariza F. J. &amp; Wenz S. 2002. A new insight into pycnodontiform fishes. Geodiversitas 24 (1): 139 - 248." type="journal article" year="2002">Poyato-Ariza &amp; Wenz 2002</bibRefCitation>
: fig. 36E) and contains 19 principal segmented caudal rays, the 2 external being pointed and the 17 others branched. There are 6 ventral and at least 4 dorsal procurrent rays.
</paragraph>
<caption id="DF1F6664DB7FFFA752B2F872FB09F7E4" ID-DOI="http://doi.org/10.5281/zenodo.3838804" ID-Zenodo-Dep="3838804" httpUri="https://zenodo.org/record/3838804/files/figure.png" pageId="17" pageNumber="18" startId="17.[225,268,2000,2026]" targetBox="[370,1266,988,1944]" targetPageId="17">
<paragraph id="8BDF36ECDB7FFFA752B2F872FB09F7E4" blockId="17.[225,1436,2000,2062]" pageId="17" pageNumber="18">
<emphasis id="B914EAFEDB7FFFA752B2F872FEBAF800" bold="true" box="[225,316,2000,2026]" pageId="17" pageNumber="18">Fig. 16.</emphasis>
<taxonomicName id="4C604D6FDB7FFFA75311F873FDC2F801" authority="Taverne &amp; Capasso, 2014" authorityName="Taverne &amp; Capasso" authorityYear="2014" box="[322,580,2001,2027]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kriweti" status="gen. et sp. nov.">
<emphasis id="B914EAFEDB7FFFA75311F873FDC2F801" box="[322,580,2001,2027]" italics="true" pageId="17" pageNumber="18">Joinvillichthys kriweti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2275785DB7FFFA75018F873FD7EF801" box="[587,760,2001,2027]" pageId="17" pageNumber="18" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Holotype, CLC S-137.
<emphasis id="B914EAFEDB7FFFA75658F872FBA6F800" bold="true" box="[1035,1056,2000,2026]" pageId="17" pageNumber="18">A</emphasis>
. Dorsal ridge scutes.
<emphasis id="B914EAFEDB7FFFA7574AF873FAABF801" bold="true" box="[1305,1325,2001,2027]" pageId="17" pageNumber="18">B</emphasis>
. Anterior flake-like body scales.
<emphasis id="B914EAFEDB7FFFA753BDF856FD85F7E4" bold="true" box="[494,515,2036,2062]" pageId="17" pageNumber="18">C</emphasis>
. Posterior scute-like body scale.
<emphasis id="B914EAFEDB7FFFA751DBF856FC1BF7E4" bold="true" box="[904,925,2036,2062]" pageId="17" pageNumber="18">D</emphasis>
. Ventral keel scutes.
</paragraph>
</caption>
<paragraph id="8BDF36ECDB7CFFA45343FE93FDB5FEA6" blockId="18.[227,1437,305,507]" box="[272,563,305,332]" pageId="18" pageNumber="19">
<emphasis id="B914EAFEDB7CFFA45343FE93FE43FEA1" bold="true" box="[272,453,305,331]" pageId="18" pageNumber="19">6. Squamation</emphasis>
(
<figureCitation id="135B2A69DB7CFFA45386FE93FDACFEA6" box="[469,554,305,332]" captionStart="Fig" captionStartId="17.[225,268,2000,2026]" captionTargetBox="[370,1266,988,1944]" captionTargetId="figure@17.[355,1280,952,1953]" captionTargetPageId="17" captionText="Fig. 16. Joinvillichthys kriweti gen. et sp. nov. Holotype, CLC S-137. A. Dorsal ridge scutes. B. Anterior flake-like body scales. C. Posterior scute-like body scale. D. Ventral keel scutes." figureDoi="http://doi.org/10.5281/zenodo.3838804" httpUri="https://zenodo.org/record/3838804/files/figure.png" pageId="18" pageNumber="19">Fig. 16</figureCitation>
)
</paragraph>
<paragraph id="8BDF36ECDB7CFFA452B0FEF6FEBAFE11" blockId="18.[227,1437,305,507]" pageId="18" pageNumber="19">
The squamation is the same as in
<taxonomicName id="4C604D6FDB7CFFA450D6FEF6FC46FE84" baseAuthorityName="Davis" baseAuthorityYear="1890" box="[645,960,340,366]" class="Actinopterygii" family="Gladiopycnodontidae" genus="Joinvillichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pycnodontiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="lindstroemi">
<emphasis id="B914EAFEDB7CFFA450D6FEF6FC46FE84" box="[645,960,340,366]" italics="true" pageId="18" pageNumber="19">Joinvillichthys lindstroemi</emphasis>
</taxonomicName>
. There are 7 spiny scutes in the dorsal ridge and at least 3 spiny scutes in the ventral keel. The two posterior ventral scutes are associated with the ventral margin of the postcoelomic bone. The body scales are slightly ornamented with tubercles. Anteriorly, they are small and flake-like. Posteriorly, there are much larger, irregular and scute-like shaped.
</paragraph>
</subSubSection>
</treatment>
</document>