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<document id="39FFE640BE1AB75EBA999ECB6B6FF3DF" ID-DOI="10.5281/zenodo.13643902" ID-ISSN="1732-2421" ID-Zenodo-Dep="13643902" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142865276" checkinUser="felipe" docAuthor="Mutter, Raoul J. &amp; Neuman, Andrew G." docDate="2006" docId="03C987BAFFD0FFC18242FC6AFE00FE7B" docLanguage="en" docName="ActaPalaeontolPol.51.2.271-282.pdf" docOrigin="Acta Palaeontologica Polonica 51 (2)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Listracanthus pectenatus Mutter &amp; Neuman, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="277" masterDocId="FFF0FFC2FFD3FFC6816EFF85FFDEFFDB" masterDocTitle="An enigmatic chondrichthyan with Paleozoic affinities from the Lower Triassic of western Canada" masterLastPageNumber="282" masterPageNumber="271" pageNumber="274" updateTime="1725365894861" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="423E2094A162203D5171F71089F19FD5">An enigmatic chondrichthyan with Paleozoic affinities from the Lower Triassic of western Canada</mods:title>
</mods:titleInfo>
<mods:name id="CCBC6E1B5654C13EEF79D3297256ACA0" type="personal">
<mods:role id="CEE6400AC5EF05AFC7EFF4B555AC97E0">
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<mods:namePart id="5682AD15D77BBC910CA066A8217843F8">Mutter, Raoul J.</mods:namePart>
</mods:name>
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<mods:namePart id="86A24B31E2CABD0ADFF8290C6FBD5D28">Neuman, Andrew G.</mods:namePart>
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<mods:typeOfResource id="600F1321D4A0333B7E89E56456D422E9">text</mods:typeOfResource>
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<mods:title id="545AB402645E23C423CA9936146E834B">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="B4FA499100E72A74850A9E2BD9F21F22">
<mods:date id="C23F1D2232068FCB1D14E676ED8FCD32">2006</mods:date>
<mods:detail id="FA80BA23BE177C0B8992E960413FD913" type="volume">
<mods:number id="C33076709BF93127EA6A2E3BA45366C4">51</mods:number>
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<mods:classification id="8060BBAF318DD24513BA3D8FB14363AB">journal article</mods:classification>
<mods:identifier id="D621DDFCF8B65F2DCCC8725FB507A18B" type="DOI">10.5281/zenodo.13643902</mods:identifier>
<mods:identifier id="1CCCE162604EC114F9C68E0E7789ED71" type="ISSN">1732-2421</mods:identifier>
<mods:identifier id="160720FAEA334C93001D063CAEC352DE" type="Zenodo-Dep">13643902</mods:identifier>
</mods:mods>
<treatment id="03C987BAFFD0FFC18242FC6AFE00FE7B" LSID="urn:lsid:plazi:treatment:03C987BAFFD0FFC18242FC6AFE00FE7B" httpUri="http://treatment.plazi.org/id/03C987BAFFD0FFC18242FC6AFE00FE7B" lastPageId="7" lastPageNumber="277" pageId="3" pageNumber="274">
<subSubSection id="C37A6527FFD0FFC58242FC6AFA10FB93" pageId="3" pageNumber="274" type="nomenclature">
<paragraph id="8BDF36ACFFD0FFC58242FC6AFB07FBD1" blockId="3.[812,1486,1006,1333]" box="[812,1241,1006,1035]" pageId="3" pageNumber="274">
<heading id="D09781C0FFD0FFC58242FC6AFB07FBD1" box="[812,1241,1006,1035]" fontSize="12" level="2" pageId="3" pageNumber="274" reason="9">
<taxonomicName id="4C604D2FFFD0FFC58242FC6AFBB3FBD0" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[812,1133,1007,1035]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="3" pageNumber="274" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD0FFC58242FC6AFBB3FBD0" box="[812,1133,1007,1035]" italics="true" pageId="3" pageNumber="274">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD0FFC5851BFC6BFB07FBD1" box="[1141,1241,1006,1034]" pageId="3" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BDF36ACFFD0FFC58242FB92FA10FB93" blockId="3.[812,1486,1006,1333]" pageId="3" pageNumber="274">
<emphasis id="B914EABEFFD0FFC58242FB92FBDAFBF7" box="[812,1028,1047,1068]" italics="true" pageId="3" pageNumber="274">Derivation of the name</emphasis>
: Latin
<emphasis id="B914EABEFFD0FFC58528FB92FB5AFBF7" box="[1094,1156,1047,1068]" italics="true" pageId="3" pageNumber="274">pecten</emphasis>
, a comb, referring to the comblike arrangement of processes at the posterior border in the large denticles.
</paragraph>
</subSubSection>
<subSubSection id="C37A6527FFD0FFC18242FBD3FE00FE7B" lastPageId="7" lastPageNumber="278" pageId="3" pageNumber="274" type="materials_examined">
<paragraph id="8BDF36ACFFD0FFC58242FBD3FC6DFB5D" blockId="3.[812,1486,1006,1333]" pageId="3" pageNumber="274">
<materialsCitation id="3B083CF1FFD0FFC58242FBD3FC6DFB5D" collectionCode="UALVP" pageId="3" pageNumber="274" specimenCode="UALVP 47002" specimenCount="1" typeStatus="holotype">
<emphasis id="B914EABEFFD0FFC58242FBD3FC5AFBB0" box="[812,900,1110,1131]" italics="true" pageId="3" pageNumber="274">
<typeStatus id="54DB880EFFD0FFC58242FBD3FC5AFBB0" box="[812,900,1110,1131]" pageId="3" pageNumber="274" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="DBC69ED7FFD0FFC582FFFBD0FBFDFBB1" box="[913,1059,1109,1130]" collectionCode="UALVP" pageId="3" pageNumber="274">UALVP 47002</specimenCode>
(
<figureCitation id="135B2A29FFD0FFC5855FFBD0FBB2FBB1" box="[1073,1132,1109,1130]" captionStart="Fig" captionStartId="3.[102,135,762,782]" captionTargetBox="[102,1485,229,732]" captionTargetId="figure-541@3.[103,1485,229,731]" captionTargetPageId="3" captionText="Fig. 2. Small and large dermal denticles in holotype specimen UALVP 47002, a possible body part of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. A. A large number of denticles is irregularly arranged but an unusually high number is clustered to the left. On the right, denticle types I and II are widely spaced, clearly unrelated to each other. B. Three large denticles (type I) as preserved (closeup).C. Small denticles (type II) as preserved, showing variable crown shape (closeup)." figureDoi="http://doi.org/10.5281/zenodo.13643908" httpUri="https://zenodo.org/record/13643908/files/figure.png" pageId="3" pageNumber="274">Fig. 2</figureCitation>
); assemblage of small and large dermal denticles.
</materialsCitation>
</paragraph>
<paragraph id="8BDF36ACFFD0FFC58242FB11FBF2FAEE" blockId="3.[812,1486,1006,1333]" pageId="3" pageNumber="274">
<emphasis id="B914EABEFFD0FFC58242FB11FC29FB72" box="[812,1015,1172,1193]" italics="true" pageId="3" pageNumber="274">Locality and horizon</emphasis>
:
<materialsCitation id="3B083CF1FFD0FFC5856BFB11FBF6FAEE" county="lower Smithian" latitude="54.51" location="Wapiti Lake Provincial Park" longLatPrecision="642" longitude="-120.71" municipality="Lake Provincial Park" pageId="3" pageNumber="274" specimenCount="1" typeStatus="holotype">
<location id="8EBF6077FFD0FFC5856BFB11FAC3FB72" LSID="urn:lsid:plazi:treatment:03C987BAFFD0FFC18242FC6AFE00FE7B:8EBF6077FFD0FFC5856BFB11FAC3FB72" box="[1029,1309,1172,1193]" county="lower Smithian" latitude="54.51" longLatPrecision="642" longitude="-120.71" municipality="Lake Provincial Park" name="Wapiti Lake Provincial Park" pageId="3" pageNumber="274">
<location id="8EBF6077FFD0FFC5856BFB11FBA3FB72" LSID="urn:lsid:plazi:treatment:03C987BAFFD0FFC18242FC6AFE00FE7B:8EBF6077FFD0FFC5856BFB11FBA3FB72" box="[1029,1149,1172,1193]" county="lower Smithian" latitude="54.51" longLatPrecision="642" longitude="-120.71" municipality="Lake Provincial Park" name="Wapiti Lake" pageId="3" pageNumber="274">
Wapiti
<collectingMunicipality id="6BBBACD6FFD0FFC58523FB11FBA3FB72" box="[1101,1149,1172,1193]" pageId="3" pageNumber="274">Lake</collectingMunicipality>
</location>
Provincial Park
</location>
, near “Fossil Fish Lake” (
<geoCoordinate id="EE54506BFFD0FFC5821CFB35FC1BFB1E" box="[882,965,1199,1221]" degrees="54.51" direction="north" orientation="latitude" pageId="3" pageNumber="274" precision="555" value="54.51">
54.51
<emphasis id="B914EABEFFD0FFC582C7FB2AFC6DFB64" box="[937,947,1199,1215]" italics="true" pageId="3" pageNumber="274">°</emphasis>
N
</geoCoordinate>
/
<geoCoordinate id="EE54506BFFD0FFC582A2FB35FBF0FB1E" box="[972,1070,1199,1221]" degrees="120.71" direction="west" orientation="longitude" pageId="3" pageNumber="274" precision="555" value="-120.71">
120.71
<emphasis id="B914EABEFFD0FFC58562FB2AFBC8FB64" box="[1036,1046,1199,1215]" italics="true" pageId="3" pageNumber="274">°</emphasis>
W
</geoCoordinate>
) center of C cirque (see
<bibRefCitation id="EFF14B5DFFD0FFC58461FB35FC82FB3A" author="Neuman, A. G. &amp; Mutter, R. J." pageId="3" pageNumber="274" pagination="25 - 36" refId="ref7232" refString="Neuman, A. G. and Mutter, R. J. 2005. Helmolepis cyphognathus, sp. nov., a new platysiagid actinopterygian from the Lower Triassic Sulphur Mountain Formation (BC, Canada). Canadian Journal of Earth Sciences 42: 25 - 36." type="journal article" year="2005">Neuman and Mutter 2005</bibRefCitation>
for detailed information on locality). Exact layer is unknown (
<typeStatus id="54DB880EFFD0FFC5825DFB6DFC56FB26" box="[819,904,1256,1277]" pageId="3" pageNumber="274" type="holotype">holotype</typeStatus>
is from talus), but specimens come most likely from between 34 and 42 meters above the PaleozoicMesozoic contact (
<collectingCounty id="62BE4E20FFD0FFC58420FA81FC92FAEE" pageId="3" pageNumber="274">lower Smithian</collectingCounty>
, see discussion above)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF36ACFFD0FFC58242FACDFB2BF9FB" blockId="3.[812,1485,1351,1568]" pageId="3" pageNumber="274">
<emphasis id="B914EABEFFD0FFC58242FACDFC40FA84" box="[812,926,1352,1375]" italics="true" pageId="3" pageNumber="274">Diagnosis</emphasis>
.—
<taxonomicName id="4C604D2FFFD0FFC582AFFAC2FB05FA84" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[961,1243,1351,1375]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="3" pageNumber="274" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD0FFC582AFFAC2FB05FA84" box="[961,1243,1351,1375]" italics="true" pageId="3" pageNumber="274">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD0FFC5858AFACDFAE3FA84" box="[1252,1341,1352,1375]" pageId="3" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
differs from all other species of
<taxonomicName id="4C604D2FFFD0FFC5856EFAE2FB44FAA4" authorityName="Newberry and Worthen" authorityYear="1870" box="[1024,1178,1383,1407]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="3" pageNumber="274" phylum="Chordata" rank="genus">
<emphasis id="B914EABEFFD0FFC5856EFAE2FB44FAA4" box="[1024,1178,1383,1407]" italics="true" pageId="3" pageNumber="274">Listracanthus</emphasis>
</taxonomicName>
in having the posterior processes of
<typeStatus id="54DB880EFFD0FFC582F9FA0DFC19FA7B" box="[919,967,1416,1440]" pageId="3" pageNumber="274">type</typeStatus>
Identicles showing comblike and very regular, parallel arrangement. These processes project perpendicular to the denticles long axis and are not confluent with the ridges in the shaft. The anterior border is ornamented by about one dozen short and curved fulcra.
</paragraph>
<paragraph id="8BDF36ACFFD0FFC58242F9B7FA96F971" blockId="3.[812,1485,1585,2026]" pageId="3" pageNumber="274">
<emphasis id="B914EABEFFD0FFC58242F9B7FC6DF992" box="[812,947,1586,1609]" italics="true" pageId="3" pageNumber="274">Description</emphasis>
.—The denticles recognized as belonging to
<taxonomicName id="4C604D2FFFD0FFC58242F9D4FB93F9B2" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[812,1101,1617,1641]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="3" pageNumber="274" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD0FFC58242F9D4FB93F9B2" box="[812,1101,1617,1641]" italics="true" pageId="3" pageNumber="274">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD0FFC5853BF9D7FB6EF9B2" box="[1109,1200,1618,1641]" pageId="3" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
range in size from
<quantity id="4C989B49FFD0FFC584FCF9D4FCABF951" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="3" pageNumber="274" unit="mm" value="55.0" valueMax="70.0" valueMin="40.0">40 to 70mm</quantity>
in length (denticle
<typeStatus id="54DB880EFFD0FFC58536F9F7FB57F951" box="[1112,1161,1650,1674]" pageId="3" pageNumber="274">type</typeStatus>
I; width about 1/4 to 1/3 of the length) or from
<quantity id="4C989B49FFD0FFC58562F914FBA7F971" box="[1036,1145,1681,1706]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="3" pageNumber="274" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1 to 4mm</quantity>
(denticle
<typeStatus id="54DB880EFFD0FFC58582F917FAC3F971" box="[1260,1309,1682,1706]" pageId="3" pageNumber="274">type</typeStatus>
II).
</paragraph>
<paragraph id="8BDF36ACFFD0FFC5823CF934FA12F831" blockId="3.[812,1485,1585,2026]" pageId="3" pageNumber="274">
<materialsCitation id="3B083CF1FFD0FFC5823CF934FA12F831" collectionCode="TMP, UALVP" pageId="3" pageNumber="274" specimenCode="TMP 89.131, UALVP 47004, 47005, 47015" specimenCount="1" typeStatus="holotype">
The arrangement of type Identicles on several specimens suggest that actual parts of the body of the fish may be preserved (e.g.,
<typeStatus id="54DB880EFFD0FFC582D2F974FBC0F8D1" box="[956,1054,1777,1802]" pageId="3" pageNumber="274" type="holotype">holotype</typeStatus>
specimen
<figureCitation id="135B2A29FFD0FFC585FBF974FB07F8D1" box="[1173,1241,1777,1802]" captionStart="Fig" captionStartId="3.[102,135,762,782]" captionTargetBox="[102,1485,229,732]" captionTargetId="figure-541@3.[103,1485,229,731]" captionTargetPageId="3" captionText="Fig. 2. Small and large dermal denticles in holotype specimen UALVP 47002, a possible body part of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. A. A large number of denticles is irregularly arranged but an unusually high number is clustered to the left. On the right, denticle types I and II are widely spaced, clearly unrelated to each other. B. Three large denticles (type I) as preserved (closeup).C. Small denticles (type II) as preserved, showing variable crown shape (closeup)." figureDoi="http://doi.org/10.5281/zenodo.13643908" httpUri="https://zenodo.org/record/13643908/files/figure.png" pageId="3" pageNumber="274">Fig. 2</figureCitation>
; also specimens
<specimenCode id="DBC69ED7FFD0FFC584FDF977FC5DF8F1" collectionCode="TMP" country="South Africa" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="3" pageNumber="274">TMP 89.131</specimenCode>
.1,
<specimenCode id="DBC69ED7FFD0FFC582F1F894FB90F8F1" box="[927,1102,1809,1834]" collectionCode="UALVP" pageId="3" pageNumber="274">UALVP 47004</specimenCode>
,
<specimenCode id="DBC69ED7FFD0FFC58535F894FB7AF8F1" box="[1115,1188,1809,1834]" collectionCode="UALVP" pageId="3" pageNumber="274">47005</specimenCode>
,
<specimenCode id="DBC69ED7FFD0FFC585DFF894FB22F8F1" box="[1201,1276,1809,1834]" collectionCode="UALVP" pageId="3" pageNumber="274">47015</specimenCode>
). The smaller dermal denticles (type II) have various conical shapes, some are recurved at their tips, are straightconical or are almost completely flat (
<figureCitation id="135B2A29FFD0FFC582D6F8F4FC20F851" box="[952,1022,1905,1930]" captionStart="Fig" captionStartId="4.[102,135,1312,1332]" captionText="Fig. 3. A. Variation of type IIdenticles of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. A shred of fairly regularly arranged in situdenticles as preserved in specimen UALVP 17938. B. Specimen UALVP 46540. B1. A single small denticle, recurved at the tip. B2. A single, small, seemingly straight denticle, possibly anteroposteriorly embedded. B3. Sketch of a small denticle showing the poorly developed base." pageId="3" pageNumber="274">Fig. 3</figureCitation>
). The bases are not conspicuously separated from the crowns (there is no neck), and the crowns were probably circular in crosssection (not laterally compressed), bearing on average eight to ten ridges running basad from the
</materialsCitation>
</paragraph>
<paragraph id="8BDF36ACFFD7FFC28107FF2EFA17FF1B" blockId="4.[105,1481,171,192]" box="[105,1481,171,192]" pageId="4" pageNumber="275">
MUTTER AND NEUMAN—ENIGMATIC CHONDRICHTHYAN FROM LOWER TRIASSIC OF
<collectingCountry id="F377763CFFD7FFC2853AFF2EFB65FF1B" box="[1108,1211,171,192]" name="Canada" pageId="4" pageNumber="275">CANADA</collectingCountry>
275
</paragraph>
<paragraph id="8BDF36ACFFD7FFC28204FD44FA8EFB32" pageId="4" pageNumber="275">
<quantity id="4C989B49FFD7FFC28204FD44FD0AFB30" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="275" unit="mm" value="1.0">1 mm 1 mm</quantity>
<emphasis id="B914EABEFFD7FFC28515FC29FB1EFB1E" bold="true" box="[1147,1267,940,1221]" pageId="4" pageNumber="275">crownlshaft base</emphasis>
<quantity id="4C989B49FFD7FFC28474FB53FA8EFB32" box="[1306,1360,1238,1257]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="275" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
<caption id="DF1F6624FFD7FFC28108FAA5FA85FAB7" pageId="4" pageNumber="275" startId="4.[102,135,1312,1332]" targetType="figure">
<paragraph id="8BDF36ACFFD7FFC28108FAA5FA85FAB7" blockId="4.[102,1486,1312,1392]" pageId="4" pageNumber="275">
Fig. 3.
<emphasis id="B914EABEFFD7FFC281CAFAA5FF6AFAEF" bold="true" box="[164,180,1312,1332]" pageId="4" pageNumber="275">A</emphasis>
. Variation of type IIdenticles of
<taxonomicName id="4C604D2FFFD7FFC280B0FAA4FD1EFAEF" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[478,704,1313,1333]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="4" pageNumber="275" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD7FFC280B0FAA4FD1EFAEF" box="[478,704,1313,1333]" italics="true" pageId="4" pageNumber="275">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD7FFC283ABFAA4FCD0FAEF" box="[709,782,1313,1332]" pageId="4" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
, Lower Triassic, Wapiti Lake, British Columbia. A shred of fairly regularly arranged in situdenticles as preserved in specimen UALVP 17938.
<emphasis id="B914EABEFFD7FFC283DEFAB8FD61FA8B" bold="true" box="[688,703,1341,1360]" pageId="4" pageNumber="275">B</emphasis>
. Specimen UALVP 46540. B
<subScript id="17E434E9FFD7FFC282A7FAC3FC0FFA8F" attach="left" box="[969,977,1350,1364]" fontSize="6" pageId="4" pageNumber="275">1</subScript>
. A single small denticle, recurved at the tip. B
<subScript id="17E434E9FFD7FFC28404FAC3FAACFA8F" attach="left" box="[1386,1394,1350,1364]" fontSize="6" pageId="4" pageNumber="275">2</subScript>
. A single, small, seemingly straight denticle, possibly anteroposteriorly embedded. B
<subScript id="17E434E9FFD7FFC2827CFAE7FCC4FAAB" attach="left" box="[786,794,1378,1392]" fontSize="6" pageId="4" pageNumber="275">3</subScript>
. Sketch of a small denticle showing the poorly developed base.
</paragraph>
</caption>
<paragraph id="8BDF36ACFFD7FFC28108FA37FDA3F9F1" blockId="4.[102,775,1457,2026]" pageId="4" pageNumber="275">
apex (
<figureCitation id="135B2A29FFD7FFC281C1FA34FF26FA11" box="[175,248,1457,1482]" captionStart="Fig" captionStartId="4.[102,135,1312,1332]" captionText="Fig. 3. A. Variation of type IIdenticles of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. A shred of fairly regularly arranged in situdenticles as preserved in specimen UALVP 17938. B. Specimen UALVP 46540. B1. A single small denticle, recurved at the tip. B2. A single, small, seemingly straight denticle, possibly anteroposteriorly embedded. B3. Sketch of a small denticle showing the poorly developed base." pageId="4" pageNumber="275">Fig. 3</figureCitation>
).
<typeStatus id="54DB880EFFD7FFC2807DFA37FE93FA11" box="[275,333,1458,1482]" pageId="4" pageNumber="275">Type</typeStatus>
IIdenticles possess a short, compact crown with usually converging ridges. The bases of the small denticles are rarely clearly visible, were probably weakly developed and had no visible foramina (
<figureCitation id="135B2A29FFD7FFC28363F994FDB2F9F6" box="[525,620,1553,1581]" captionStart="Fig" captionStartId="4.[102,135,1312,1332]" captionText="Fig. 3. A. Variation of type IIdenticles of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. A shred of fairly regularly arranged in situdenticles as preserved in specimen UALVP 17938. B. Specimen UALVP 46540. B1. A single small denticle, recurved at the tip. B2. A single, small, seemingly straight denticle, possibly anteroposteriorly embedded. B3. Sketch of a small denticle showing the poorly developed base." pageId="4" pageNumber="275">
Fig. 3B
<subScript id="17E434E9FFD7FFC2830DF998FDB2F9F6" attach="left" box="[611,620,1565,1581]" fontSize="7" pageId="4" pageNumber="275">3</subScript>
</figureCitation>
).
</paragraph>
<paragraph id="8BDF36ACFFD7FFC281E2F9B4FB42F9B2" blockId="4.[102,775,1457,2026]" lastBlockId="4.[812,1485,1457,2026]" pageId="4" pageNumber="275">
<materialsCitation id="3B083CF1FFD7FFC281E2F9B4FB46F9B2" collectionCode="UALVP" pageId="4" pageNumber="275" specimenCode="UALVP 46575" specimenCount="2">
The larger denticles (
<typeStatus id="54DB880EFFD7FFC280EFF9B4FE6DF991" box="[385,435,1585,1610]" pageId="4" pageNumber="275">type</typeStatus>
I) consist of a long, laterally flattened shaft (~crown) that tapers towards the apex and has longitudinal ridges. The shafts show distinctive anterior and posterior borders. The ridges in this
<typeStatus id="54DB880EFFD7FFC2837CF917FD9CF971" box="[530,578,1682,1706]" pageId="4" pageNumber="275">type</typeStatus>
vary in number, comprising as a maximum 45 near the base of the shaft and 15 near the apex (
<specimenCode id="DBC69ED7FFD7FFC28046F954FE0DF931" box="[296,467,1745,1770]" collectionCode="UALVP" pageId="4" pageNumber="275">UALVP 46575</specimenCode>
and 46577). The upper half of the convex anterior border is hemmed by about one dozen, one millimeter to one and a half millimeters long, single, short and curved fulcra, that are regularly and increasingly wider spaced towards the base and are absent in some denticles in the basal third of the denticles entire length (as can be seen in
<figureCitation id="135B2A29FFD7FFC2807DF814FE87F871" box="[275,345,1937,1962]" captionStart="Fig" captionStartId="5.[102,135,1777,1797]" captionTargetBox="[181,1405,225,1749]" captionTargetId="figure-67@5.[181,1405,225,1749]" captionTargetPageId="5" captionText="Fig. 4. Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. Specimen UALVP 46551 showing morphological details of a large denticle (type I). A. A complete large denticle. Large denticles may vary in length, width and curvature. B. The apex of a large denticle with the fulcral ridges arranged like a “tuft”. C. The posterior border as preserved in the same specimen. Note that each straight process possesses a striated shaft (1) and a triangular base (2) lateral to the process (see also thin sections in Fig. 7). D. Detail of posterobasal corner of the posterior border." figureDoi="http://doi.org/10.5281/zenodo.13643910" httpUri="https://zenodo.org/record/13643910/files/figure.png" pageId="4" pageNumber="275">Fig. 4</figureCitation>
). The concave posterior border sends out numerous straight and striated processes that give the posterior border of the denticle a comblike appearance. These small processes run perpendicular to the denticles long axis, are oriented more apicad near the base and are not “confluent” with the longitudinal ridges on the lateral wall of the shaft (see
<figureCitation id="135B2A29FFD7FFC282A1F994FBC9F9F1" box="[975,1047,1553,1578]" captionStart="Fig" captionStartId="5.[102,135,1777,1797]" captionTargetBox="[181,1405,225,1749]" captionTargetId="figure-67@5.[181,1405,225,1749]" captionTargetPageId="5" captionText="Fig. 4. Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. Specimen UALVP 46551 showing morphological details of a large denticle (type I). A. A complete large denticle. Large denticles may vary in length, width and curvature. B. The apex of a large denticle with the fulcral ridges arranged like a “tuft”. C. The posterior border as preserved in the same specimen. Note that each straight process possesses a striated shaft (1) and a triangular base (2) lateral to the process (see also thin sections in Fig. 7). D. Detail of posterobasal corner of the posterior border." figureDoi="http://doi.org/10.5281/zenodo.13643910" httpUri="https://zenodo.org/record/13643910/files/figure.png" pageId="4" pageNumber="275">Fig. 4</figureCitation>
). There are smaller, triangular shaped processes intercalated at the posterior border just lateral to the comblike straight processes
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF36ACFFD7FFC2823CF9F4FA51F831" blockId="4.[812,1485,1457,2026]" pageId="4" pageNumber="275">
The base is slightly broader than the maximum width of the crown of the denticle and is usually poorly preserved or was originally weakly developed. As seen from thin sections (see discussion below), the shaft of
<typeStatus id="54DB880EFFD7FFC285D1F957FB31F931" box="[1215,1263,1746,1770]" pageId="4" pageNumber="275">type</typeStatus>
Identicle is probably secondarily ossified near the core centre and is histologically indistinguishable from the base. In ornamentation, the base is clearly delimited from the crown by lacking a superficial striation and by possessing wide cavities. The ventral border of the base is normally convex but may occasionally be straight or even concave (
<figureCitation id="135B2A29FFD7FFC285C8F814FB28F871" box="[1190,1270,1937,1962]" captionStart="Fig" captionStartId="5.[102,135,1777,1797]" captionTargetBox="[181,1405,225,1749]" captionTargetId="figure-67@5.[181,1405,225,1749]" captionTargetPageId="5" captionText="Fig. 4. Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. Specimen UALVP 46551 showing morphological details of a large denticle (type I). A. A complete large denticle. Large denticles may vary in length, width and curvature. B. The apex of a large denticle with the fulcral ridges arranged like a “tuft”. C. The posterior border as preserved in the same specimen. Note that each straight process possesses a striated shaft (1) and a triangular base (2) lateral to the process (see also thin sections in Fig. 7). D. Detail of posterobasal corner of the posterior border." figureDoi="http://doi.org/10.5281/zenodo.13643910" httpUri="https://zenodo.org/record/13643910/files/figure.png" pageId="4" pageNumber="275">Figs. 4</figureCitation>
,
<figureCitation id="135B2A29FFD7FFC2846AF814FACAF871" box="[1284,1300,1937,1962]" captionStart="Fig" captionStartId="6.[102,135,1154,1174]" captionTargetBox="[103,1483,225,1127]" captionTargetId="figure-68@6.[103,1483,225,1127]" captionTargetPageId="6" captionText="Fig. 5. Type Identicles of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia, showing in situ arrangement (A) and aberrant morphology (B). A. As preserved in specimen TMP 95.114.52. B. Aberrant denticle as preserved in specimen TMP 2001.18.01. B1. General view. B2. Closeup of the midportion." figureDoi="http://doi.org/10.5281/zenodo.13643912" httpUri="https://zenodo.org/record/13643912/files/figure.png" pageId="4" pageNumber="275">5</figureCitation>
). The apex consists of fulcral ridges arranged as a “tuft” (
<figureCitation id="135B2A29FFD7FFC2847EF834FAB9F811" box="[1296,1383,1969,1994]" captionStart="Fig" captionStartId="5.[102,135,1777,1797]" captionTargetBox="[181,1405,225,1749]" captionTargetId="figure-67@5.[181,1405,225,1749]" captionTargetPageId="5" captionText="Fig. 4. Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia. Specimen UALVP 46551 showing morphological details of a large denticle (type I). A. A complete large denticle. Large denticles may vary in length, width and curvature. B. The apex of a large denticle with the fulcral ridges arranged like a “tuft”. C. The posterior border as preserved in the same specimen. Note that each straight process possesses a striated shaft (1) and a triangular base (2) lateral to the process (see also thin sections in Fig. 7). D. Detail of posterobasal corner of the posterior border." figureDoi="http://doi.org/10.5281/zenodo.13643910" httpUri="https://zenodo.org/record/13643910/files/figure.png" pageId="4" pageNumber="275">Fig. 4B</figureCitation>
) that exhibits individual variation in structure and complexity.
</paragraph>
<caption id="DF1F6624FFD6FFC38108F974FB20F882" ID-DOI="http://doi.org/10.5281/zenodo.13643910" ID-Zenodo-Dep="13643910" httpUri="https://zenodo.org/record/13643910/files/figure.png" pageId="5" pageNumber="276" startId="5.[102,135,1777,1797]" targetBox="[181,1405,225,1749]" targetPageId="5" targetType="figure">
<paragraph id="8BDF36ACFFD6FFC38108F974FB20F882" blockId="5.[102,1485,1777,1881]" pageId="5" pageNumber="276">
Fig. 4.
<taxonomicName id="4C604D2FFFD6FFC381C9F977FE54F8DF" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[167,394,1778,1798]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="5" pageNumber="276" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD6FFC381C9F977FE54F8DF" box="[167,394,1778,1798]" italics="true" pageId="5" pageNumber="276">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD6FFC380FEF977FE04F8DE" box="[400,474,1778,1797]" pageId="5" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
, Lower Triassic, Wapiti Lake, British Columbia. Specimen UALVP 46551 showing morphological details of a large denticle (type I).
<emphasis id="B914EABEFFD6FFC38041F888FEE1F8FA" bold="true" box="[303,319,1805,1825]" pageId="5" pageNumber="276">A</emphasis>
. A complete large denticle. Large denticles may vary in length, width and curvature.
<emphasis id="B914EABEFFD6FFC3855DF88BFB9CF8FA" bold="true" box="[1075,1090,1806,1825]" pageId="5" pageNumber="276">B</emphasis>
. The apex of a large denticle with the fulcral ridges arranged like a “tuft”.
<emphasis id="B914EABEFFD6FFC38004F8ACFEA4F8E6" bold="true" box="[362,378,1833,1853]" pageId="5" pageNumber="276">C</emphasis>
. The posterior border as preserved in the same specimen. Note that each straight process possesses a striated shaft (1) and a triangular base (2) lateral to the process (see also thin sections in Fig. 7).
<emphasis id="B914EABEFFD6FFC38393F8C3FCD3F882" bold="true" box="[765,781,1862,1881]" pageId="5" pageNumber="276">D</emphasis>
. Detail of posterobasal corner of the posterior border.
</paragraph>
</caption>
<paragraph id="8BDF36ACFFD6FFC081E2F814FD9BF831" blockId="5.[102,775,1937,2026]" lastBlockId="6.[102,775,1937,2026]" lastPageId="6" lastPageNumber="277" pageId="5" pageNumber="276">
<materialsCitation id="3B083CF1FFD6FFC081E2F814FD9FF831" collectionCode="UALVP" lastPageId="6" lastPageNumber="277" pageId="5" pageNumber="276" specimenCode="UALVP 46568, 47004, 47005" specimenCount="3" typeStatus="holotype">
The denticles weather to bluishgrey or white. Most specimens consist of
<specimenCount id="9D66FD25FFD6FFC3804BF837FE5BF811" box="[293,389,1970,1994]" count="1" pageId="5" pageNumber="276" type="generic" typeStatus="type">one type</specimenCount>
of denticle or the respective slabs either contain the great majority of large or small denticles. However, there are several larger slabs, that show an accumulation of largest and smallest denticles (e.g.,
<specimenCode id="DBC69ED7FFD6FFC38401F834FCABF831" collectionCode="UALVP" pageId="5" pageNumber="276">UALVP 46568</specimenCode>
,
<specimenCode id="DBC69ED7FFD6FFC38211F854FC16F831" box="[895,968,2001,2026]" collectionCode="UALVP" pageId="5" pageNumber="276">47004</specimenCode>
,
<specimenCode id="DBC69ED7FFD6FFC382BCF854FBC5F831" box="[978,1051,2001,2026]" collectionCode="UALVP" pageId="5" pageNumber="276">47005</specimenCode>
, and 47015).
<specimenCount id="9D66FD25FFD6FFC385C1F857FA83F831" box="[1199,1373,2001,2026]" count="2" pageId="5" pageNumber="276" type="generic">Two specimens</specimenCount>
show that denticles may become “split” in half during growth and that the largest denticles were probably also fairly regularly arranged and connected to the bases (
<figureCitation id="135B2A29FFD5FFC0809CF854FDE9F831" box="[498,567,2001,2026]" captionStart="Fig" captionStartId="6.[102,135,1154,1174]" captionTargetBox="[103,1483,225,1127]" captionTargetId="figure-68@6.[103,1483,225,1127]" captionTargetPageId="6" captionText="Fig. 5. Type Identicles of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia, showing in situ arrangement (A) and aberrant morphology (B). A. As preserved in specimen TMP 95.114.52. B. Aberrant denticle as preserved in specimen TMP 2001.18.01. B1. General view. B2. Closeup of the midportion." figureDoi="http://doi.org/10.5281/zenodo.13643912" httpUri="https://zenodo.org/record/13643912/files/figure.png" pageId="6" pageNumber="277">Fig. 5</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="DF1F6624FFD5FFC08108FB07FE53FB15" ID-DOI="http://doi.org/10.5281/zenodo.13643912" ID-Zenodo-Dep="13643912" httpUri="https://zenodo.org/record/13643912/files/figure.png" pageId="6" pageNumber="277" startId="6.[102,135,1154,1174]" targetBox="[103,1483,225,1127]" targetPageId="6" targetType="figure">
<paragraph id="8BDF36ACFFD5FFC08108FB07FE53FB15" blockId="6.[102,1486,1154,1234]" pageId="6" pageNumber="277">
Fig. 5. Type Identicles of
<taxonomicName id="4C604D2FFFD5FFC0803CFB06FDEAFB4E" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[338,564,1155,1175]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD5FFC0803CFB06FDEAFB4E" box="[338,564,1155,1175]" italics="true" pageId="6" pageNumber="277">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD5FFC08356FB06FD5FFB4D" box="[568,641,1155,1174]" pageId="6" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
, Lower Triassic, Wapiti Lake, British Columbia, showing in situ arrangement (
<emphasis id="B914EABEFFD5FFC0842FFB07FA8FFB4D" bold="true" box="[1345,1361,1154,1174]" pageId="6" pageNumber="277">A</emphasis>
) and aberrant morphology (
<emphasis id="B914EABEFFD5FFC0818AFB1AFF2DFB69" bold="true" box="[228,243,1183,1202]" pageId="6" pageNumber="277">B</emphasis>
).
<emphasis id="B914EABEFFD5FFC08067FB1BFEC7FB69" bold="true" box="[265,281,1182,1202]" pageId="6" pageNumber="277">A</emphasis>
. As preserved in specimen TMP 95.114.52.
<emphasis id="B914EABEFFD5FFC083AFFB1AFD0EFB69" bold="true" box="[705,720,1183,1202]" pageId="6" pageNumber="277">B</emphasis>
. Aberrant denticle as preserved in specimen TMP 2001.18.01. B
<subScript id="17E434E9FFD5FFC0845BFB2DFAE3FB6D" attach="left" box="[1333,1341,1192,1206]" fontSize="6" pageId="6" pageNumber="277">1</subScript>
. General view. B
<subScript id="17E434E9FFD5FFC0811BFB41FFA3FB09" attach="left" box="[117,125,1220,1234]" fontSize="6" pageId="6" pageNumber="277">2</subScript>
. Closeup of the midportion.
</paragraph>
</caption>
<caption id="DF1F6624FFD5FFC08108F8A0FB82F88E" ID-DOI="http://doi.org/10.5281/zenodo.13643914" ID-Zenodo-Dep="13643914" httpUri="https://zenodo.org/record/13643914/files/figure.png" pageId="6" pageNumber="277" startId="6.[102,135,1829,1849]" targetBox="[228,1341,1295,1802]" targetPageId="6" targetType="figure">
<paragraph id="8BDF36ACFFD5FFC08108F8A0FB82F88E" blockId="6.[102,1485,1829,1877]" pageId="6" pageNumber="277">
Fig. 6. Associated fragment (
<emphasis id="B914EABEFFD5FFC08006F8A0FEA6F8E2" bold="true" box="[360,376,1829,1849]" pageId="6" pageNumber="277">A</emphasis>
) and sketch (
<emphasis id="B914EABEFFD5FFC0809EF8A3FE21F8E2" bold="true" box="[496,511,1830,1849]" pageId="6" pageNumber="277">B</emphasis>
) of a?toothlike fragment or skeletal fragment preserved on a slab with both types of denticles (not shown) of
<taxonomicName id="4C604D2FFFD5FFC08108F8C4FE97F88F" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[102,329,1857,1877]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD5FFC08108F8C4FE97F88F" box="[102,329,1857,1877]" italics="true" pageId="6" pageNumber="277">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD5FFC08020F8C7FE46F88E" box="[334,408,1858,1877]" pageId="6" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
, Lower Triassic, Wapiti Lake, British Columbia, in specimen UALVP 38562.
</paragraph>
</caption>
<paragraph id="8BDF36ACFFD5FFC1823CF814FE00FE7B" blockId="6.[812,1485,1937,2026]" lastBlockId="7.[102,775,232,416]" lastPageId="7" lastPageNumber="278" pageId="6" pageNumber="277">
No head or fin region or articulated remains of jaws or a dentition have been detected in any specimen. The internal skeleton was undoubtedly largely uncalcified. However, a?toothlike fragment or skeletal element is preserved with large and small denticles in specimen
<collectionCode id="ED71AE69FFD4FFC18375FE8DFDA6FEFA" box="[539,632,264,289]" pageId="7" pageNumber="278">UALVP</collectionCode>
38562 (
<figureCitation id="135B2A29FFD4FFC183B7FE8DFFA8FE9A" captionStart="Fig" captionStartId="6.[102,135,1829,1849]" captionTargetBox="[228,1341,1295,1802]" captionTargetId="figure-191@6.[228,1358,1295,1802]" captionTargetPageId="6" captionText="Fig. 6. Associated fragment (A) and sketch (B) of a?toothlike fragment or skeletal fragment preserved on a slab with both types of denticles (not shown) of Listracanthus pectenatus sp. nov., Lower Triassic, Wapiti Lake, British Columbia, in specimen UALVP 38562." figureDoi="http://doi.org/10.5281/zenodo.13643914" httpUri="https://zenodo.org/record/13643914/files/figure.png" pageId="7" pageNumber="278">Fig. 6</figureCitation>
). The questionable skeletal element is only partly preserved, oval in crosssection and apparently hollow. It is not clear whether this remain at all represents a skeletal part of
<taxonomicName id="4C604D2FFFD4FFC18108FE0DFEA1FE7B" authorityName="Mutter &amp; Neuman" authorityYear="2006" box="[102,383,392,416]" class="Holocephali" family="Menaspidae" genus="Listracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chimaeriformes" pageId="7" pageNumber="278" phylum="Chordata" rank="species" species="pectenatus" status="sp. nov.">
<emphasis id="B914EABEFFD4FFC18108FE0DFEA1FE7B" box="[102,383,392,416]" italics="true" pageId="7" pageNumber="278">Listracanthus pectenatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757C5FFD4FFC180E8FE0CFE00FE7B" box="[390,478,393,416]" pageId="7" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>