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<document id="3A41094D9710E0AE4FD78510708C6645" ID-CLB-Dataset="102846" ID-DOI="10.11646/zootaxa.5166.1.1" ID-GBIF-Dataset="7e35a6a2-7ddc-453b-99ec-20142ba284f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6876209" ID-ZooBank="17952A48-902C-47A0-A344-8B07490F3B28" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658417435455" checkinUser="plazi" docAuthor="Cadena-Castañeda, Oscar J., Braun, Holger &amp; García, Alexander García" docDate="2022" docId="03C0C519CF2E6144D4A28D5D4F82FEFB" docLanguage="en" docName="zootaxa.5166.1.1.pdf" docOrigin="Zootaxa 5166 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5166.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dysoniini Rehn 1950" docType="treatment" docVersion="6" lastPageNumber="22" masterDocId="FFF9BD61CF3D6151D43589634C49FF9E" masterDocTitle="The tribe Dysoniini part VI: Phylogeny, biogeography and evolutionary trends of the lichen katydid genera (Orthoptera: Tettigoniidae: Phaneropterinae). Eleventh contribution to the suprageneric organization of Neotropical phaneropterines" masterLastPageNumber="93" masterPageNumber="1" pageNumber="20" updateTime="1699361192069" updateUser="ExternalLinkService">
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<mods:title id="990CCDE9EDCC3A7CEDEAA8C5E04E5929">The tribe Dysoniini part VI: Phylogeny, biogeography and evolutionary trends of the lichen katydid genera (Orthoptera: Tettigoniidae: Phaneropterinae). Eleventh contribution to the suprageneric organization of Neotropical phaneropterines</mods:title>
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<mods:namePart id="CB34CDA818BD046DC9EF9DC9657CF032">Cadena-Castañeda, Oscar J.</mods:namePart>
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<mods:namePart id="9228F1C9FBA9B2F5819028F20664B39B">Braun, Holger</mods:namePart>
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<mods:namePart id="5AFB9E0732673CB1686FF761460EE433">García, Alexander García</mods:namePart>
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<treatment id="03C0C519CF2E6144D4A28D5D4F82FEFB" ID-DOI="http://doi.org/10.5281/zenodo.6885770" ID-GBIF-Taxon="197841194" ID-Zenodo-Dep="6885770" LSID="urn:lsid:plazi:treatment:03C0C519CF2E6144D4A28D5D4F82FEFB" httpUri="http://treatment.plazi.org/id/03C0C519CF2E6144D4A28D5D4F82FEFB" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="C3732784CF2E6142D4A28D5D4DA4FBC6" box="[151,493,1086,1112]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BD6740FCF2E6142D4A28D5D4DA4FBC6" blockId="19.[151,493,1086,1112]" box="[151,493,1086,1112]" pageId="19" pageNumber="20">
<heading id="D09EC363CF2E6142D4A28D5D4DA4FBC6" bold="true" box="[151,493,1086,1112]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="3">
<emphasis id="B91DA81DCF2E6142D4A28D5D4DA4FBC6" bold="true" box="[151,493,1086,1112]" pageId="19" pageNumber="20">
Tribe
<taxonomicName id="4C690F8CCF2E6142D4D78D5D4DA4FBC6" ID-CoL="8PT46" authority="Rehn, 1950" authorityName="Rehn" authorityYear="1950" box="[226,493,1086,1112]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="tribe" tribe="Dysoniini">
Dysoniini
<bibRefCitation id="EFF809FECF2E6142D5548D5D4DA4FBC6" author="Rehn, J. A. G." box="[353,493,1086,1112]" pageId="19" pageNumber="20" pagination="271 - 319" refId="ref42693" refString="Rehn, J. A. G. (1950 [1949]) Studies in the Group Dysoniae (Aphidniae of authors) (Orthoptera: Tettigoniidae: Phaneropterinae) Part I. Transactions of the American Entomological Society, 75 (3 - 4), 271 - 319." type="journal article" year="1950">Rehn, 1950</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3732784CF2E6142D4A28DE54E9DF9CF" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BD6740FCF2E6142D4A28DE54E9DF9CF" blockId="19.[151,1437,1158,2013]" pageId="19" pageNumber="20">
<emphasis id="B91DA81DCF2E6142D4A28DE54D5BFB3E" bold="true" box="[151,274,1158,1184]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Small to medium-sized (
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). Eyes globose, distance of the antennal sockets not greater than the width of the scapus. Vertex elevated in diverse shapes, from a slightly raised tubercle or crest to long spines. Fastigium with a developed ventral tooth, ocellar tubercle present and only slightly or well developed. Pronotal disc also with various modifications (like spines, expansions, denticulations). Tegmina narrow or moderately widened, Rs vein originating at the middle of R, MA vein prolonged to the middle of the tegmina and then turning into the anal margin. Anterior coxae dorsally armed with a fronto-basal spine; tympana exposed on both sides of the fore tibiae; leg spines lamellate, developed as spinule or long, pointed spines. Genicular lobes of fore and middle femora are usually armed with a spine (in
<taxonomicName id="4C690F8CCF2E6142D5DC8CE04E64FA03" authorityName="Hebard" authorityYear="1926" box="[489,557,1411,1437]" class="Insecta" family="Tettigoniidae" genus="Quiva" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D5DC8CE04E64FA03" box="[489,557,1411,1437]" italics="true" pageId="19" pageNumber="20">Quiva</emphasis>
</taxonomicName>
armed with two small spines). Meso- and metasternum never longer than wide, meso- and metafurcal furrow narrow and usually rounded. Tenth tergite unmodified; male cerci forked, usually divided in the horizontal plane (except
<taxonomicName id="4C690F8CCF2E6142D67A8CA94E81FA7A" authorityName="Rehn" authorityYear="1950" box="[591,712,1482,1508]" class="Insecta" family="Tettigoniidae" genus="Apolinaria" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D67A8CA94E81FA7A" box="[591,712,1482,1508]" italics="true" pageId="19" pageNumber="20">Apolinaria</emphasis>
</taxonomicName>
, where the both branches are directed dorsally); male subgenital plate unmodified (except for
<taxonomicName id="4C690F8CCF2E6142D5D58C8D4ED5F996" authorityName="Cadena-Castaneda" authorityYear="2011" box="[480,668,1518,1544]" class="Insecta" family="Tettigoniidae" genus="Lichenomorphus" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D5D58C8D4ED5F996" box="[480,668,1518,1544]" italics="true" pageId="19" pageNumber="20">Lichenomorphus</emphasis>
</taxonomicName>
, where it is elongated and very flexible), and styli developed to different degrees. Ovipositor of females varying in shape, from as long as the pronotum, curved and broadened, to long and slender, surpassing the length of the pronotum.
</paragraph>
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<emphasis id="B91DA81DCF2E6142D4F28F394D1DF9EA" bold="true" box="[199,340,1626,1652]" pageId="19" pageNumber="20">
<typeStatus id="54D2CAADCF2E6142D4F28F394D4BF9EA" box="[199,258,1626,1652]" pageId="19" pageNumber="20">Type</typeStatus>
genus.
</emphasis>
<taxonomicName id="4C690F8CCF2E6142D56E8F384E05F9EA" ID-CoL="8NR58" authority="White, 1862" authorityName="White" authorityYear="1862" box="[347,588,1626,1652]" class="Insecta" family="Tettigoniidae" genus="Dysonia" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D56E8F384DF1F9EA" box="[347,440,1627,1652]" italics="true" pageId="19" pageNumber="20">Dysonia</emphasis>
<bibRefCitation id="EFF809FECF2E6142D58A8F394E05F9EA" author="White, A." box="[447,588,1626,1652]" pageId="19" pageNumber="20" pagination="175 - 284" refId="ref43428" refString="White, A. (1862) Class V - Insects (Insecta). Richardson's Museum of Natural History. 2, 175 - 284." type="journal article" year="1862">White, 1862</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="C3732784CF2E6142D4F28F1D48FAF906" box="[199,1203,1662,1689]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BD6740FCF2E6142D4F28F1D48FAF906" blockId="19.[151,1437,1158,2013]" box="[199,1203,1662,1689]" pageId="19" pageNumber="20">
<emphasis id="B91DA81DCF2E6142D4F28F1D4D28F906" bold="true" box="[199,353,1662,1688]" pageId="19" pageNumber="20">Distribution.</emphasis>
Neotropical, from northeastern
<collectingCountry id="F37E349FCF2E6142D6FD8F1D4F57F906" box="[712,798,1662,1688]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
to northern
<collectingCountry id="F37E349FCF2E6142D7938F1D485CF906" box="[934,1045,1662,1688]" name="Argentina" pageId="19" pageNumber="20">Argentina</collectingCountry>
(
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Distribution map of Hammatofera nodicornis." captionText-1="Map 2. Distribution map of Machima species." captionText-10="Map 11. Distribution map of Dissonulichen (Dissonulichen) simplicipes simplicipes and D. (D.) simplicipes meridionalis in Colombia and Ecuador." captionText-11="Map 12. Distribution map of Dissonulichospinus n. gen. species." captionText-12="Map 13. Distribution map of Paraphidnia species." captionText-13="Map 14. Distribution map of Anaphidna species in Central America." captionText-14="Map 15. Distribution map of Anaphidna species in South America." captionText-15="Map 16. Distribution map of Dysonia species." captionText-16="Map 17. Distribution map of Lichenomorphus species." captionText-17="Map 18. Distribution map of Lichenomorphus species in Southeast Brazil and Argentina." captionText-18="Map 19. Distribution map of Lichenodentix dentatithorax." captionText-2="Map 3. Distribution map of Machimoides species." captionText-3="Map 4. Distribution map of Apolinaria hygracantha and Lichenodraculus species." captionText-4="Map 5. Distribution map of Markia species (Hystrix and Sarriai groups)." captionText-5="Map 6. Distribution map of Markia species (Erinaceus group)." captionText-6="Map 7. Distribution map of Quiva species." captionText-7="Map 8. Distribution map of Yungasacris species." captionText-8="Map 9. Distribution map of Alexanderellus n. gen. species." captionText-9="Map 10. Distribution map of Dissonulichen s.s. species." figureDoi-0="http://doi.org/10.5281/zenodo.6885555" figureDoi-1="http://doi.org/10.5281/zenodo.6885558" figureDoi-10="http://doi.org/10.5281/zenodo.6885587" figureDoi-11="http://doi.org/10.5281/zenodo.6885589" figureDoi-12="http://doi.org/10.5281/zenodo.6885591" figureDoi-13="http://doi.org/10.5281/zenodo.6885593" figureDoi-14="http://doi.org/10.5281/zenodo.6885595" figureDoi-15="http://doi.org/10.5281/zenodo.6885599" figureDoi-16="http://doi.org/10.5281/zenodo.6885601" figureDoi-17="http://doi.org/10.5281/zenodo.6885603" figureDoi-18="http://doi.org/10.5281/zenodo.6885605" figureDoi-2="http://doi.org/10.5281/zenodo.6885560" figureDoi-3="http://doi.org/10.5281/zenodo.6885564" figureDoi-4="http://doi.org/10.5281/zenodo.6885567" figureDoi-5="http://doi.org/10.5281/zenodo.6885571" figureDoi-6="http://doi.org/10.5281/zenodo.6885573" figureDoi-7="http://doi.org/10.5281/zenodo.6885575" figureDoi-8="http://doi.org/10.5281/zenodo.6885577" figureDoi-9="http://doi.org/10.5281/zenodo.6885583" httpUri-0="https://zenodo.org/record/6885555/files/figure.png" httpUri-1="https://zenodo.org/record/6885558/files/figure.png" httpUri-10="https://zenodo.org/record/6885587/files/figure.png" httpUri-11="https://zenodo.org/record/6885589/files/figure.png" httpUri-12="https://zenodo.org/record/6885591/files/figure.png" httpUri-13="https://zenodo.org/record/6885593/files/figure.png" httpUri-14="https://zenodo.org/record/6885595/files/figure.png" httpUri-15="https://zenodo.org/record/6885599/files/figure.png" httpUri-16="https://zenodo.org/record/6885601/files/figure.png" httpUri-17="https://zenodo.org/record/6885603/files/figure.png" httpUri-18="https://zenodo.org/record/6885605/files/figure.png" httpUri-2="https://zenodo.org/record/6885560/files/figure.png" httpUri-3="https://zenodo.org/record/6885564/files/figure.png" httpUri-4="https://zenodo.org/record/6885567/files/figure.png" httpUri-5="https://zenodo.org/record/6885571/files/figure.png" httpUri-6="https://zenodo.org/record/6885573/files/figure.png" httpUri-7="https://zenodo.org/record/6885575/files/figure.png" httpUri-8="https://zenodo.org/record/6885577/files/figure.png" httpUri-9="https://zenodo.org/record/6885583/files/figure.png" pageId="19" pageNumber="20">Maps 1–19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3732784CF2E6144D4F28FC14F82FEFB" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BD6740FCF2E6142D4F28FC14822F842" blockId="19.[151,1437,1158,2013]" pageId="19" pageNumber="20">
<emphasis id="B91DA81DCF2E6142D4F28FC14D1BF922" bold="true" box="[199,338,1698,1724]" pageId="19" pageNumber="20">Comments:</emphasis>
Recently,
<bibRefCitation id="EFF809FECF2E6142D5FC8FC14ECDF922" author="Gorochov, A. V." box="[457,644,1698,1724]" pageId="19" pageNumber="20" pagination="1 - 14" refId="ref40746" refString="Gorochov, A. V. (2014) Taxonomy of the katydids (Orthoptera: Tettigoniidae) from East Asia and adjacent islands. Communication 8. Far Eastern Entomologist, 273, 1 - 14." type="journal article" year="2014">Gorochov (2014</bibRefCitation>
,
<bibRefCitation id="EFF809FECF2E6142D6A58FC14E83F922" author="Gorochov, A. V." box="[656,714,1698,1724]" pageId="19" pageNumber="20" pagination="135 - 157" refId="ref40784" refString="Gorochov, A. V. (2016) Systematics of the American katydids (Orthoptera: Tettigoniidae). Communication 6. Proceedings of the Zoological Institute of the Russian Academy of Sciences, 320 (2), 135 - 157. https: // doi. org / 10.31610 / trudyzin / 2016.320.2.135" type="journal article" year="2016">2016</bibRefCitation>
) considered
<taxonomicName id="4C690F8CCF2E6142D76C8FC148C3F922" authority="Dohrn, 1892" authorityName="Dohrn" authorityYear="1892" box="[857,1162,1698,1724]" class="Insecta" family="Tettigoniidae" genus="Trachyzulpha" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D76C8FC14FB8F922" box="[857,1009,1698,1724]" italics="true" pageId="19" pageNumber="20">Trachyzulpha</emphasis>
Dohrn, 1892
</taxonomicName>
, a genus from Southeast Asia, to be a member of
<taxonomicName id="4C690F8CCF2E6142D5988FA54E57F97E" authorityName="Rehn" authorityYear="1950" box="[429,542,1734,1760]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="tribe" tribe="Dysoniini">Dysoniini</taxonomicName>
, placing it in its own subtribe
<taxonomicName id="4C690F8CCF2E6142D7478FA5486CF97E" authorityName="Gorochov" authorityYear="2014" box="[882,1061,1734,1760]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subTribe" subTribe="Trachyzulphina">Trachyzulphina</taxonomicName>
. The supposed justification being the shared lichen mimicry. However, this
<typeStatus id="54D2CAADCF2E6142D6458F884EE9F89B" box="[624,672,1771,1797]" pageId="19" pageNumber="20">type</typeStatus>
of mimicry has arisen several times in the subfamily
<taxonomicName id="4C690F8CCF2E6142D1318F894C88F8B7" authorityName="Burmeister" authorityYear="1838" class="Insecta" family="Phaneropterinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Phaneropterinae</taxonomicName>
, and also in other katydids like
<taxonomicName id="4C690F8CCF2E6142D6198E6D486AF8B6" authority="Rentz, Su &amp; Ueshima, 2012" authorityName="Rentz, Su &amp; Ueshima" authorityYear="2012" box="[556,1059,1806,1833]" class="Insecta" family="Tettigoniidae" genus="Lichenagraecia" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D6198E6D4E95F8B6" box="[556,732,1806,1832]" italics="true" pageId="19" pageNumber="20">Lichenagraecia</emphasis>
Rentz, Su &amp; Ueshima, 2012
</taxonomicName>
from
<collectingCountry id="F37E349FCF2E6142D05D8E6D4886F8B6" box="[1128,1231,1806,1832]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
(
<taxonomicName id="4C690F8CCF2E6142D0EB8E6D49D0F8B6" authorityName="Burmeister" authorityYear="1838" box="[1246,1433,1806,1832]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Conocephalinae">Conocephalinae</taxonomicName>
:
<taxonomicName id="4C690F8CCF2E6142D4A28E514D25F8D2" authorityName="Rentz, Su &amp; Ueshima" authorityYear="2012" box="[151,364,1842,1868]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="tribe" tribe="Armadillagraeciini">Armadillagraeciini</taxonomicName>
) as well as several groups of the family
<taxonomicName id="4C690F8CCF2E6142D7008E514FB8F8D2" box="[821,1009,1842,1868]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Pseudophyllinae">Pseudophyllinae</taxonomicName>
, and even in Caelifera such as pygmy grasshoppers of the genera
<taxonomicName id="4C690F8CCF2E6142D5E58E354F4EF8EE" authority="Serville, 1838" authorityName="Serville" authorityYear="1838" box="[464,775,1878,1904]" class="Insecta" family="Tetrigidae" genus="Amorphopus" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D5E58E354E16F8EE" box="[464,607,1878,1904]" italics="true" pageId="19" pageNumber="20">Amorphopus</emphasis>
<bibRefCitation id="EFF809FECF2E6142D65D8E354F4EF8EE" author="Serville, J. G. A." box="[616,775,1878,1904]" pageId="19" pageNumber="20" pagination="1 - 776" refId="ref42963" refString="Serville, J. G. A. (1838 [1839]) Histoire naturelle des insectes. Orthopteres, 1839, i - xviii, 1 - 776." type="journal article" year="1838">Serville, 1838</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C690F8CCF2E6142D7748E3548DBF8EE" authority="Hancock, 1907" authorityName="Hancock" authorityYear="1907" box="[833,1170,1878,1904]" class="Insecta" family="Tetrigidae" genus="Eomorphopus" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA81DCF2E6142D7748E354F97F8EE" box="[833,990,1878,1904]" italics="true" pageId="19" pageNumber="20">Eomorphopus</emphasis>
Hancock, 1907
</taxonomicName>
from the Amazon (
<bibRefCitation id="EFF809FECF2E6142D1478E354D9FF80A" author="Cadena-Castaneda, O. J. &amp; Silva, D. S. M. &amp; Mendes, D. M. D. &amp; Pereira, M. &amp; Domenico, F. &amp; Sperber, C. F." pageId="19" pageNumber="20" pagination="45 - 62" refId="ref39767" refString="Cadena-Castaneda, O. J., Silva, D. S. M., Mendes, D. M. D., Pereira, M., Domenico, F. &amp; Sperber, C. F. (2019 a) Review of the tribe Amorphopini (Orthoptera: Tetrigidae: Metrodorinae): pygmy moss-lichen tetrigids from the Amazon rainforest. Journal of Orthoptera Research, 29 (1), 45 - 62. https: // doi. org / 10.3897 / jor. 29.33717" type="journal article" year="2019">
Cadena-Castañeda
<emphasis id="B91DA81DCF2E6142D5658E184DC8F80A" box="[336,385,1914,1940]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 2019a
</bibRefCitation>
,
<bibRefCitation id="EFF809FECF2E6142D5D78E194E67F80A" author="Cadena-Castaneda, O. J. &amp; Mendes, D. M. D. &amp; Silva, D. S. M. &amp; Cardona-Granda, J. M. &amp; Garcia Garcia, A. &amp; Tumbrinck, J." box="[482,558,1914,1940]" pageId="19" pageNumber="20" pagination="1 - 65" refId="ref39859" refString="Cadena-Castaneda, O. J., Mendes, D. M. D., Silva, D. S. M., Cardona-Granda, J. M., Garcia Garcia, A. &amp; Tumbrinck, J. (2019 b) Systematics and biogeography of the genus Scaria Bolivar, 1887 (Orthoptera: Tetrigidae: Batrachideinae). Zootaxa, 4675 (1), 1 - 65. https: // doi. org / 10.11646 / zootaxa. 4675.1.1" type="journal article" year="2019">2019b</bibRefCitation>
). An adaptation to a particular
<typeStatus id="54D2CAADCF2E6142D7BD8E184FF1F80B" box="[904,952,1915,1941]" pageId="19" pageNumber="20">type</typeStatus>
of environment or microhabitat is not necessarily associated with a particular taxonomic group. Therefore, the Trachyzulphini
<emphasis id="B91DA81DCF2E6142D0618EFD48ECF826" bold="true" box="[1108,1189,1950,1976]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A22E1566CF2E6142D0618EFD48ECF826" box="[1108,1189,1950,1976]" pageId="19" pageNumber="20">n. stat.</taxonomicNameLabel>
</emphasis>
are here elevated to a separate tribe of the
<taxonomicName id="4C690F8CCF2E6142D5498EA14E7CF842" authorityName="Burmeister" authorityYear="1838" box="[380,565,1986,2012]" class="Insecta" family="Phaneropterinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Phaneropterinae</taxonomicName>
, based on the original diagnosis (
<bibRefCitation id="EFF809FECF2E6142D79C8EA14816F842" author="Gorochov, A. V." box="[937,1119,1986,2012]" pageId="19" pageNumber="20" pagination="1 - 14" refId="ref40746" refString="Gorochov, A. V. (2014) Taxonomy of the katydids (Orthoptera: Tettigoniidae) from East Asia and adjacent islands. Communication 8. Far Eastern Entomologist, 273, 1 - 14." type="journal article" year="2014">Gorochov 2014</bibRefCitation>
).
</paragraph>
<caption id="DF162487CF296145D4A28E1E4EB9F80B" ID-DOI="http://doi.org/10.5281/zenodo.6876239" ID-Zenodo-Dep="6876239" box="[151,752,1917,1942]" httpUri="https://zenodo.org/record/6876239/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1917,1941]" targetBox="[151,1436,182,1892]" targetPageId="20">
<paragraph id="8BD6740FCF296145D4A28E1E4EB9F80B" blockId="20.[151,752,1917,1942]" box="[151,752,1917,1942]" pageId="20" pageNumber="21">
<emphasis id="B91DA81DCF296145D4A28E1E4D57F80B" bold="true" box="[151,286,1917,1941]" pageId="20" pageNumber="21">FIGURE 11.</emphasis>
Pictorial key to the genera of tribe
<taxonomicName id="4C690F8CCF296145D6B08E1E4EA5F80B" authorityName="Rehn" authorityYear="1950" box="[645,748,1917,1941]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="tribe" tribe="Dysoniini">Dysoniini</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD6740FCF286144D4F289F44F9FFEDF" blockId="21.[151,1437,151,357]" pageId="21" pageNumber="22">
Moreover,
<bibRefCitation id="EFF809FECF286144D57689F44E4CFF2F" author="Gorochov, A. V." box="[323,517,151,177]" pageId="21" pageNumber="22" pagination="135 - 157" refId="ref40784" refString="Gorochov, A. V. (2016) Systematics of the American katydids (Orthoptera: Tettigoniidae). Communication 6. Proceedings of the Zoological Institute of the Russian Academy of Sciences, 320 (2), 135 - 157. https: // doi. org / 10.31610 / trudyzin / 2016.320.2.135" type="journal article" year="2016">Gorochov (2016)</bibRefCitation>
proposed to reduce
<taxonomicName id="4C690F8CCF286144D6DF89F44F37FF2F" authority="Cadena-Castañeda &amp; Braun &amp; García, 2022" authorityName="Cadena-Castañeda &amp; Braun &amp; García" authorityYear="2022" box="[746,894,151,177]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="78" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Pycnopalpini">Pycnopalpini</taxonomicName>
to a subtribe of
<taxonomicName id="4C690F8CCF286144D00289F448E1FF2F" authorityName="Rehn" authorityYear="1950" box="[1079,1192,151,177]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Dysoniini">Dysoniini</taxonomicName>
, arguing that the posteromedial denticle or spine on the upper rostral tubercle is also present in some representatives of that group. However,
<taxonomicName id="4C690F8CCF286144D53989BC4DE9FF67" authority="Cadena-Castañeda &amp; Braun &amp; García, 2022" authorityName="Cadena-Castañeda &amp; Braun &amp; García" authorityYear="2022" box="[268,416,223,249]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="78" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Pycnopalpini">Pycnopalpini</taxonomicName>
+
<taxonomicName id="4C690F8CCF286144D58A89BC4E64FF67" authorityName="Rehn" authorityYear="1950" box="[447,557,223,249]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Dysoniini">Dysoniini</taxonomicName>
do not belong to a monophyletic group according to the most comprehensive phylogeny of
<taxonomicName id="4C690F8CCF286144D50388604EACFE83" authority="(Mugleston et al. 2018)" baseAuthorityName="Mugleston" baseAuthorityYear="2018" box="[310,741,259,285]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">
Tettigoniidae (
<bibRefCitation id="EFF809FECF286144D5EF88604E94FE83" author="Mugleston, J. D. &amp; Naegle, M. &amp; Song, H. &amp; Whiting, M. F." box="[474,733,259,285]" pageId="21" pageNumber="22" pagination="1 - 27" refId="ref41921" refString="Mugleston, J. D., Naegle, M., Song, H. &amp; Whiting, M. F. (2018) A comprehensive phylogeny of Tettigoniidae (Orthoptera: Ensifera) reveals extensive ecomorph convergence and widespread taxonomic incongruence. Insect Systematics and Diversity, 2 (4), 1 - 27. https: // doi. org / 10.1093 / isd / ixy 010" type="journal article" year="2018">
Mugleston
<emphasis id="B91DA81DCF286144D66888604ED9FE83" box="[605,656,259,285]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. 2018
</bibRefCitation>
)
</taxonomicName>
. The
<taxonomicName id="4C690F8CCF286144D71388604FF3FE83" authority="Cadena-Castañeda &amp; Braun &amp; García, 2022" authorityName="Cadena-Castañeda &amp; Braun &amp; García" authorityYear="2022" box="[806,954,259,285]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="21" pageNumber="78" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Pycnopalpini">Pycnopalpini</taxonomicName>
are considered a tribe with two subtribes, Pycopalpina and Theiina (
<bibRefCitation id="EFF809FECF286144D58E88444E95FEDF" author="Cadena-Castaneda, O. J." box="[443,732,295,321]" pageId="21" pageNumber="22" pagination="149 - 163" refId="ref38888" refString="Cadena-Castaneda, O. J. (2014 a) La tribu Pycnopalpini trib. n. (Orthoptera, Tettigonioidea, Phaneropteridae): segunda contribucion a la organizacion supragenerica de los faneropterinos neotropicales. Animal Biodiversity and Conservation, 37 (2), 149 - 163." type="journal article" year="2014">Cadena-Castañeda 2014a</bibRefCitation>
, Cigliano
<emphasis id="B91DA81DCF286144D77A88444FCEFEDF" box="[847,903,295,321]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2022).
</paragraph>
<paragraph id="8BD6740FCF286144D4F288284F82FEFB" blockId="21.[151,1437,151,357]" box="[199,971,331,357]" pageId="21" pageNumber="22">
<figureCitation id="1352688ACF286144D4F288284D2EFEFB" box="[199,359,331,357]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStartId-0="20.[151,250,1917,1941]" captionStartId-1="21.[151,250,1986,2010]" captionStartId-2="22.[151,250,1872,1896]" captionStartId-3="23.[151,250,1928,1952]" captionTargetBox-0="[151,1436,182,1892]" captionTargetBox-1="[155,1435,381,1962]" captionTargetBox-2="[151,1436,181,1847]" captionTargetBox-3="[151,1425,181,1904]" captionTargetId-0="figure-21@20.[151,1436,181,1893]" captionTargetId-1="figure-110@21.[151,1435,381,1963]" captionTargetId-2="figure-21@22.[151,1436,181,1848]" captionTargetId-3="figure-22@23.[151,1436,181,1905]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="23" captionText-0="FIGURE 11. Pictorial key to the genera of tribe Dysoniini." captionText-1="FIGURE 12. Pictorial key to the genera of Dysoniini (Markiina n. subtr.)." captionText-2="FIGURE 13. Pictorial key to the genera of Dysoniini (Dysoniina n. subtr.)." captionText-3="FIGURE 14. Pictorial key to the genera of Dysoniini (Dysoniina n. subtr.)." figureDoi-0="http://doi.org/10.5281/zenodo.6876239" figureDoi-1="http://doi.org/10.5281/zenodo.6876241" figureDoi-2="http://doi.org/10.5281/zenodo.6876243" figureDoi-3="http://doi.org/10.5281/zenodo.6876247" httpUri-0="https://zenodo.org/record/6876239/files/figure.png" httpUri-1="https://zenodo.org/record/6876241/files/figure.png" httpUri-2="https://zenodo.org/record/6876243/files/figure.png" httpUri-3="https://zenodo.org/record/6876247/files/figure.png" pageId="21" pageNumber="22">Figures 11–14</figureCitation>
present a pictorial key for the identification of genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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