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<document ID-DOI="http://doi.org/10.5281/zenodo.6485244" ID-GBIF-Dataset="3dfc64e7-8c23-456d-9678-1bb0858ac470" ID-GBIF-Taxon="150645363" ID-Zenodo-Dep="6485244" checkinTime="1545431109525" checkinUser="PlaziZenodoSync" docAuthor="Jan van Tol &amp; André Günther" docDate="2018" docId="0395A91CAC56FFB49536FC68FE7CFCF4" docLanguage="en" docName="10.5281zenodo.1481114_vanTol_Guenther_2018.pdf" docOrigin="Odonatologica 47 (3)" docStyle="DocumentStyle{}" docTitle="Rhinocypha monochroa Selys 1873" docType="treatment" docVersion="4" lastPageId="50" lastPageNumber="349" masterDocId="FFACD164AC71FF869443FFAFFF05FFBC" masterDocTitle="The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)" masterLastPageNumber="386" masterPageNumber="299" originalUpdateDomain="Boston" originalUpdateTime="1544520335069" originalUpdateUser="PlaziZenodoSync" pageId="39" pageNumber="338" updateTime="1650928538772" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jan van Tol</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>André Günther</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Odonatologica</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-12-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
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<mods:start>299</mods:start>
<mods:end>386</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.1481114</mods:identifier>
<mods:identifier type="GBIF-Dataset">3dfc64e7-8c23-456d-9678-1bb0858ac470</mods:identifier>
<mods:identifier type="Zenodo-Dep">1481114</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6485244" ID-GBIF-Taxon="150645363" ID-Zenodo-Dep="6485244" LSID="urn:lsid:plazi:treatment:0395A91CAC56FFB49536FC68FE7CFCF4" httpUri="http://treatment.plazi.org/id/0395A91CAC56FFB49536FC68FE7CFCF4" lastPageId="50" lastPageNumber="349" pageId="39" pageNumber="338">
<subSubSection box="[373,821,967,995]" pageId="39" pageNumber="338" type="nomenclature">
<paragraph blockId="39.[159,1037,967,1250]" box="[373,821,967,995]" pageId="39" pageNumber="338">
<heading bold="true" box="[373,821,967,995]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="338" reason="4">
<taxonomicName authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[373,821,967,995]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="338" phylum="Arthropoda" rank="species" species="monochroa">
<emphasis bold="true" box="[373,821,967,995]" pageId="39" pageNumber="338">
<emphasis bold="true" box="[373,661,967,995]" italics="true" pageId="39" pageNumber="338">Rhinocypha monochroa</emphasis>
(Selys, 1873)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[496,698,1001,1029]" pageId="39" pageNumber="338" type="description">
<paragraph blockId="39.[159,1037,967,1250]" box="[496,698,1001,1029]" pageId="39" pageNumber="338">
<heading box="[496,698,1001,1029]" centered="true" fontSize="11" level="4" pageId="39" pageNumber="338" reason="4">
(
<figureCitation box="[502,575,1001,1029]" captionStart="Figure 5" captionStartId="15.[828,896,320,345]" captionText="Figure 5. Rhinocypha monochroa, male. Stream Sg. Pattunuang Asue, Southwest Sulawesi. Photo: AG (01- -viii-2011)" pageId="39" pageNumber="338">Figs 5</figureCitation>
,
<figureCitation box="[586,604,1001,1029]" captionStart="Figure 6" captionStartId="15.[828,896,675,700]" captionText="Figure 6. Rhinocypha monochroa, female. Stream Sg. Pattunuang Asue, Southwest Sulawesi. Photo: AG (01- -viii-2011)" pageId="39" pageNumber="338">6</figureCitation>
,
<figureCitation box="[615,692,1001,1029]" captionStart-0="Figure 38" captionStart-1="Figures 3944" captionStart-2="Figures 4552" captionStart-3="Figure 53" captionStartId-0="40.[159,227,1495,1520]" captionStartId-1="43.[159,238,1348,1373]" captionStartId-2="47.[765,844,588,613]" captionStartId-3="48.[159,227,1498,1523]" captionTargetBox-0="[239,960,915,1456]" captionTargetBox-1="[168,1033,254,1310]" captionTargetBox-2="[156,1039,694,1459]" captionTargetBox-3="[156,1039,694,1459]" captionTargetId-0="figure@40.[235,960,906,1456]" captionTargetId-1="figure@43.[161,601,741,1037]" captionTargetId-2="figure@47.[198,727,885,1049]" captionTargetId-3="figure@48.[156,1039,694,1459]" captionTargetPageId-0="40" captionTargetPageId-1="43" captionTargetPageId-2="47" captionTargetPageId-3="48" captionText-0="Figure 38. Rhinocypha monochroa. Male, lectotype [NHMUK 013324013]. Photo: B. Price, Natural History Museum, London" captionText-1="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" captionText-2="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" captionText-3="Figure 53. Rhinocypha monochroa. Simplified scheme of coloration of underside of hind wing. Artwork by AG" httpUri-0="https://zenodo.org/record/2200481/files/figure.png" httpUri-1="https://zenodo.org/record/2200483/files/figure.png" httpUri-2="https://zenodo.org/record/2200487/files/figure.png" httpUri-3="https://zenodo.org/record/2200495/files/figure.png" pageId="39" pageNumber="338">3853</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="338" type="reference_group">
<paragraph blockId="39.[159,1037,967,1250]" pageId="39" pageNumber="338">
<taxonomicName authorityName="Selys" authorityPageNumber="614" authorityYear="1873" box="[159,823,1042,1066]" class="Insecta" family="Chlorocyphidae" genus="Libellago" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis box="[159,252,1042,1066]" italics="true" pageId="39" pageNumber="338">Libellago</emphasis>
(
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[268,392,1042,1066]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis box="[268,392,1042,1066]" italics="true" pageId="39" pageNumber="338">Rhinocypha</emphasis>
</taxonomicName>
)
<emphasis box="[408,532,1042,1066]" italics="true" pageId="39" pageNumber="338">monochroa.</emphasis>
— SELYS (1873b): 614615
</taxonomicName>
(sep. 6162) (origi- nal description, type locality Celebes).
</paragraph>
<paragraph blockId="39.[159,1037,967,1250]" pageId="39" pageNumber="338">
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[159,411,1103,1127]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="338" phylum="Arthropoda" rank="species" species="monochroa">
<emphasis box="[159,411,1103,1127]" italics="true" pageId="39" pageNumber="338">Rhinocypha monochroa</emphasis>
</taxonomicName>
. — KIRBY (1890): 114 (catalogue, distribution Celebes); SELYS (1891): 215 (distribution Ceylon [
<emphasis box="[612,676,1134,1158]" italics="true" pageId="39" pageNumber="338">lapsus</emphasis>
], relationships); RIS (1916): 311 312 (pro parte) (key species Celebes, material); LIEFTINCK (1935): 177 (compared with
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[237,354,1195,1219]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="338" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[237,354,1195,1219]" italics="true" pageId="39" pageNumber="338">
R.
<taxonomicName box="[268,354,1195,1219]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="338" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
</emphasis>
</taxonomicName>
, distribution); COWLEY (1937): 6 (penis structure in group A); KIMMINS (1969): 309 (lectotype designated, in BMNH).
</paragraph>
</subSubSection>
<subSubSection lastPageId="40" lastPageNumber="339" pageId="39" pageNumber="338" type="diagnosis">
<paragraph blockId="39.[159,1036,1299,1550]" box="[159,281,1299,1325]" pageId="39" pageNumber="338">
<heading bold="true" box="[159,281,1299,1325]" fontSize="11" level="3" pageId="39" pageNumber="338" reason="6">
<emphasis bold="true" box="[159,281,1299,1325]" pageId="39" pageNumber="338">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="39.[159,1036,1299,1550]" pageId="39" pageNumber="338">
Stripe over synthorax with a ventral projection on metepimeron against metakatepisternum (
<figureCitation box="[403,540,1374,1400]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="39" pageNumber="338">Figs 4142</figureCitation>
), which is diagnostic among the Sulawe- si
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[187,324,1411,1437]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis box="[187,324,1411,1437]" italics="true" pageId="39" pageNumber="338">Rhinocypha</emphasis>
</taxonomicName>
species. Males have more extensive brownish black opaque markings on the wings than other species on Sulawesi, leaving only the area anterior to the Arculus, or just distal to the Arculus, transparent. Frons black without blue markings. Mesopreepisternum black, not blue or pale yellow
</paragraph>
<paragraph blockId="40.[159,1037,254,429]" pageId="40" pageNumber="339">
as in other species on Sulawesi (except
<taxonomicName authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[634,800,254,279]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="339" phylum="Arthropoda" rank="species" species="togeanensis">
<emphasis box="[634,800,254,279]" italics="true" pageId="40" pageNumber="339">
R.
<taxonomicName box="[669,800,254,279]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="339" phylum="Arthropoda" rank="species" species="togeanensis">togeanensis</taxonomicName>
</emphasis>
</taxonomicName>
), although in some females this plate has a very small and indistinct pale marking. Synthorax (
<figureCitation box="[166,250,328,354]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="40" pageNumber="339">Fig. 41</figureCitation>
) with upperside of transverse blue stripe more or less straight, usu- ally without conspicuous projection. Some specimens, however, may have such a projection (
<figureCitation box="[373,460,403,429]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="40" pageNumber="339">Fig. 42</figureCitation>
). Abdomen black on dorsum, without pruinosity.
</paragraph>
</subSubSection>
<subSubSection lastPageId="44" lastPageNumber="343" pageId="40" pageNumber="339" type="description">
<paragraph blockId="40.[159,1036,477,877]" box="[159,305,477,503]" pageId="40" pageNumber="339">
<heading bold="true" box="[159,305,477,503]" fontSize="11" level="3" pageId="40" pageNumber="339" reason="6">
<emphasis bold="true" box="[159,305,477,503]" pageId="40" pageNumber="339">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="40.[159,1036,477,877]" box="[159,220,515,541]" pageId="40" pageNumber="339">
<heading bold="true" box="[159,220,515,541]" fontSize="11" level="3" pageId="40" pageNumber="339" reason="6">
<emphasis bold="true" box="[159,220,515,541]" pageId="40" pageNumber="339">Male</emphasis>
</heading>
</paragraph>
<paragraph blockId="40.[159,1036,477,877]" pageId="40" pageNumber="339">
Head (
<figureCitation box="[235,318,552,578]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="40" pageNumber="339">Fig. 39</figureCitation>
) — Labium brownish black, median lobe black; labrum swol- len, black; mandibles black, with a large blue mark, its base distinctly wider than near labrum, its lower side running parallel to border of mandible, leaving a narrow black margin; clypeus projected as in other Chlorocyphi- dae; anteclypeus shining black; postclypeus, frons, vertex, occiput and post- ocular lobe velvet-black, a paired cream spot behind lateral ocelli, and small cream post-ocular spots close to occiput; genae with large blue spot just above mandibles, extending dorsally with a narrow blue line along the eyes up to the level of the median ocellus.
</paragraph>
<caption httpUri="https://zenodo.org/record/2200481/files/figure.png" pageId="40" pageNumber="339" startId="40.[159,227,1495,1520]" targetBox="[239,960,915,1456]" targetPageId="40">
<paragraph blockId="40.[159,1037,1494,1549]" pageId="40" pageNumber="339">
<emphasis bold="true" box="[159,267,1495,1520]" pageId="40" pageNumber="339">Figure 38.</emphasis>
<emphasis box="[274,529,1494,1518]" italics="true" pageId="40" pageNumber="339">
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[274,400,1494,1518]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="339" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
monochroa
</emphasis>
. Male, lectotype [NHMUK 013324013]. Photo: B. Price, Natural History Museum, London
</paragraph>
</caption>
<paragraph blockId="41.[159,1037,253,877]" pageId="41" pageNumber="340">
Thorax — Pronotum (
<figureCitation box="[413,496,254,280]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="41" pageNumber="340">Fig. 40</figureCitation>
) black, with pale-blue markings as follows:an- terior lobe with small spot in latero-posterior corner, continuing onto me- dian lobe, where it is wider anteriorly and tapers in a sharp point, reaching just the posterior half of the median lobe; spots on lateral lobe very variable, some large as in
<figureCitation box="[363,447,403,429]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="41" pageNumber="340">Fig. 40</figureCitation>
, in other specimens small or even hardly visible; posterior lobe black, unmarked.
</paragraph>
<paragraph blockId="41.[159,1037,253,877]" pageId="41" pageNumber="340">
Synthorax (
<figureCitation box="[321,414,478,504]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="41" pageNumber="340">Fig. 41</figureCitation>
). Mesopreepisternum black, Mesepisternum black, umarked; a wide, rather straight, blue stripe in lower half of synthorax, run- ning anteriorly from ventro-posterior corner of mesokatepisternum, continuing on ventro-anterior part of mesepimeron; dorsal side of stripe irregular, but usually without conspicuous projection, ending well before posterior border of mesepimeron; lower side of stripe runs from mesokatepisternum just under metathoracic spiracle to metepimeron, where it begins sharply in ventro-anterior corner, continuing at some distance from lower margin of metepimeron, reaching its posterior margin; dorsal side of stripe poste- riorly leaving metapleural suture and meeting the posterior margin of met- epimeron in the middle; metakatepisternum black.
</paragraph>
<paragraph blockId="41.[159,968,923,952]" box="[159,968,923,952]" pageId="41" pageNumber="340">Legs — Black, but distal ¾ on innerside of mid and hind tibiae white.</paragraph>
<paragraph blockId="41.[159,1036,1000,1549]" pageId="41" pageNumber="340">Wings — Black with metallic sheen; basal part to the level of Arculus transparent, posteriorly also to level of distal side of quadrangle; metallic blue iridescence on underside of hind wing to level of Px10, distal part of wing with deep, golden copper iridescence; pterostigma of fore and hind wings dark brown, nearly black against veins.</paragraph>
<paragraph blockId="41.[159,1036,1000,1549]" pageId="41" pageNumber="340">
There are remarkable differences in the coloration of the wings between the males of different populations of
<taxonomicName box="[602,765,1224,1250]" pageId="41" pageNumber="340">
<emphasis box="[602,765,1224,1250]" italics="true" pageId="41" pageNumber="340">R. monochroa</emphasis>
</taxonomicName>
(
<figureCitation box="[780,914,1224,1251]" captionStart-0="Figures 4552" captionStart-1="Figure 53" captionStartId-0="47.[765,844,588,613]" captionStartId-1="48.[159,227,1498,1523]" captionTargetBox-1="[156,1039,694,1459]" captionTargetId-0="figure@47.[198,727,885,1049]" captionTargetId-1="figure@48.[156,1039,694,1459]" captionTargetPageId-0="47" captionTargetPageId-1="48" captionText-0="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" captionText-1="Figure 53. Rhinocypha monochroa. Simplified scheme of coloration of underside of hind wing. Artwork by AG" httpUri-0="https://zenodo.org/record/2200487/files/figure.png" httpUri-1="https://zenodo.org/record/2200495/files/figure.png" pageId="41" pageNumber="340">Figs 4553</figureCitation>
). This applies particularly to the iridescent colours of the undersides of the hind wings. Also, being strongly iridescent structural colour, both the colours and the extent of the coloured zones perceived are affected by the angle of reflection, with shallow viewing angles resulting in shorter wavelength reflected light (cf.
<figureCitation box="[350,524,1411,1437]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="41" pageNumber="340">Figs 46 and 47</figureCitation>
). The best overall colour impression is seen on photographs of living individuals. Reflective patterns of newly preserved specimens are fairly stable. However, the colours on the wings of pinned specimens can change or fade with time, especially in younger specimens.
</paragraph>
<paragraph blockId="42.[159,1037,253,1027]" pageId="42" pageNumber="341">
The populations in the southern part of south-western Sulawesi and on Sa- layar Island show a metallic light blue to deep blue sheen in the proximal half of the underside of the hind wing (blue triangle), distally followed by a narrow yellow band and a golden brown distal part (
<figureCitation box="[794,867,365,392]" captionStart="Figure 5" captionStartId="15.[828,896,320,345]" captionText="Figure 5. Rhinocypha monochroa, male. Stream Sg. Pattunuang Asue, Southwest Sulawesi. Photo: AG (01- -viii-2011)" pageId="42" pageNumber="341">Figs 5</figureCitation>
and
<figureCitation box="[926,958,365,392]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">45</figureCitation>
). Such coloration is also shown in the lectotype (
<figureCitation box="[643,728,403,429]" captionStart="Figure 38" captionStartId="40.[159,227,1495,1520]" captionTargetBox="[239,960,915,1456]" captionTargetId="figure@40.[235,960,906,1456]" captionTargetPageId="40" captionText="Figure 38. Rhinocypha monochroa. Male, lectotype [NHMUK 013324013]. Photo: B. Price, Natural History Museum, London" httpUri="https://zenodo.org/record/2200481/files/figure.png" pageId="42" pageNumber="341">Fig. 38</figureCitation>
). In specimens from Palopo (
<figureCitation box="[204,288,440,466]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">Fig. 47</figureCitation>
) and Madjene (
<figureCitation box="[472,556,440,466]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">Fig. 48</figureCitation>
) the reflection of the basal blue triangle is, depending on the angle of examination, more deep blue to blue violet, while the reflection of the distal parts is golden coppery brown, at slightly altered angle of incidence changing to green. Specimens from Malino and Watampone (
<figureCitation box="[314,398,589,616]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">Fig. 46</figureCitation>
) have an intermediate coloration. The tripartite reflec- tion pattern with a basal blue triangle is characteristic of populations from the south-western peninsula. Specimens of the south-eastern peninsula lack this clear zoning in the reflection patterns; the underside of the hind wings reflect bronzy brown with slightly golden sheen (Kolaka,
<figureCitation box="[842,926,739,765]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">Fig. 49</figureCitation>
), purple- brown to purple (Kendari,
<figureCitation box="[475,556,776,803]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">Fig. 50</figureCitation>
) or green to bluish with a greenish distal part of the wing (Moramo (
<figureCitation box="[495,584,813,840]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">Fig. 51</figureCitation>
). A distinctly different reflection pat- tern is shown by specimens from Kabaena Island, off south-eastern Sulawesi (
<figureCitation box="[166,253,888,915]" captionStart="Figures 4552" captionStartId="47.[765,844,588,613]" captionTargetId="figure@47.[198,727,885,1049]" captionTargetPageId="47" captionText="Figures 4552. Rhinocypha monochroa. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG" httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="42" pageNumber="341">Fig. 52</figureCitation>
). The main area of the wings reflects green, and towards the distal end one finds a colour gradient from gold to blue violet. The space between C and R1 reflects, depending on the view angle, steel blue to purple, and the space between R1 and R2 purple (see also the conclusions below).
</paragraph>
<paragraph blockId="42.[159,1036,1075,1288]" pageId="42" pageNumber="341">
Abdomen (
<figureCitation box="[293,380,1075,1101]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="42" pageNumber="341">Fig. 43</figureCitation>
) — Dorsum black; side of tergites with extensive blue markings as follows: S1 large, sub-quadrangular, S2 rectangular, but ventro- anterior corner emarginate, S3S8 with progressively smaller markings, that on S3 large and rectangular, those on S6S7 tapering dorso-posteriorly, S8 with oblong triangular marking, pointed posteriorly, ending before poste- rior margin of segment; S9S10 and anal appendages black.
</paragraph>
<paragraph blockId="42.[159,1037,1336,1549]" box="[159,247,1336,1362]" pageId="42" pageNumber="341">
<heading bold="true" box="[159,247,1336,1362]" fontSize="11" level="3" pageId="42" pageNumber="341" reason="6">
<emphasis bold="true" box="[159,247,1336,1362]" pageId="42" pageNumber="341">Female</emphasis>
</heading>
</paragraph>
<paragraph blockId="42.[159,1037,1336,1549]" pageId="42" pageNumber="341">
Head (
<figureCitation box="[235,319,1374,1400]" captionStart="Figures 3944" captionStartId="43.[159,238,1348,1373]" captionTargetBox="[168,1033,254,1310]" captionTargetId="figure@43.[161,601,741,1037]" captionTargetPageId="43" captionText="Figures 3944. Rhinocypha monochroa. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT" httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="42" pageNumber="341">Fig. 44</figureCitation>
) — Mandible with squarish pale bluish-white marking in ba- sal part, distal part shining black; gena with similar marking to
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[903,1029,1411,1437]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="341" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[903,1029,1411,1437]" italics="true" pageId="42" pageNumber="341">R. frontalis</emphasis>
</taxonomicName>
, extending along the eyes as a very narrow stripe up to the level of vertex; frons black without pale marking; a paired spot beyond the lateral ocelli, about the size of the ocelli; post-ocular spots of similar size.
</paragraph>
<caption httpUri="https://zenodo.org/record/2200483/files/figure.png" pageId="43" pageNumber="342" startId="43.[159,238,1348,1373]" targetBox="[168,1033,254,1310]" targetPageId="43">
<paragraph blockId="43.[159,1037,1348,1549]" pageId="43" pageNumber="342">
<emphasis bold="true" box="[159,324,1348,1373]" pageId="43" pageNumber="342">Figures 3944.</emphasis>
<emphasis box="[335,596,1348,1372]" italics="true" pageId="43" pageNumber="342">
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[335,461,1348,1372]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="43" pageNumber="342" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
monochroa
</emphasis>
. All specimens from Sulawesi Selatan. 39 Male, head, frontal view. Bantimurung, 19­iv­1991 [JvT 05698]; 40 Male, pronotum, lef lateral view. Bantimurung, 1923­iv­1991 [JvT 05708]; 41 Male, synthorax, lef lateral view. Bantimurung, 19­iv­1991 [JvT 05698]; 42 Male, syn- thorax, lef lateral view. Madjene, 810­xi­1939 [JvT 05343]; 43 Male, abdomen, lef lateral view. Bantimurung, 19­iv­1991 [JvT 06630]; 44 Female, head, dorsal view. Bantimurung, 19­iv­1991 [JvT 05702]. Artwork by JvT
</paragraph>
</caption>
<paragraph blockId="44.[159,1036,253,504]" pageId="44" pageNumber="343">Thorax — Pronotum velvet black with just a small irregular bluish white marking from the extreme lateral corner of anterior lobe to middle of me- dian lobe; marking widest in the middle.</paragraph>
<paragraph blockId="44.[159,1036,253,504]" pageId="44" pageNumber="343">Synthorax. Mesopreepisternum black.Synthorax velvet black with a longi- tudinal bluish white stripe from ventro-posterior corner of mesokatepister- num, over ventro-anterior part of mesepimeron, continuing over meta- thoracic spiracle on metepisternum, and over middle of metepimeron.</paragraph>
<paragraph blockId="44.[159,325,552,579]" box="[159,325,552,579]" pageId="44" pageNumber="343">Legs — Black.</paragraph>
<paragraph blockId="44.[159,1036,627,840]" pageId="44" pageNumber="343">Wings — Basal part of fore wing opaque dark brown up to level of Px24; posterior part transparent white, the extreme tip opaque white; pterostigma white, basal corner brown; hind wing dark brown, with tip posterior to posterior side of pterostigma opaque white; pterostigma with significant variation in both fore and hind wings, from basal half dark brown and rest creamy white, to nearly fully dirty white.</paragraph>
<paragraph blockId="44.[159,1036,888,1138]" pageId="44" pageNumber="343">
Abdomen — Black, with small bluish white markings as follows: S1 with small oblong spot mediolaterally; S2 with longitudinal marking about length of segment mediolaterally, with sub-cylindrical spot in same line against posterior margin, ventrally a spot of
<emphasis box="[676,701,1001,1026]" italics="true" pageId="44" pageNumber="343">ca</emphasis>
¼ of segment length, half as high as long; a series of similar spots at same level on S3S7, large on S3S5, smaller on S6S7; anterior margin of S3S6 with narrow stripe, very small on S7; rest of abdomen black.
</paragraph>
<paragraph blockId="44.[159,1036,1187,1549]" box="[159,338,1187,1213]" pageId="44" pageNumber="343">
<heading bold="true" box="[159,338,1187,1213]" fontSize="11" level="3" pageId="44" pageNumber="343" reason="6">
<emphasis bold="true" box="[159,338,1187,1213]" pageId="44" pageNumber="343">Measurements</emphasis>
</heading>
</paragraph>
<paragraph blockId="44.[159,1036,1187,1549]" pageId="44" pageNumber="343">Males: Measurements of specimens from south-western Sulawesi [mm] — Hind wing length (n=5) x=24.5 (23.026.0); hind wing width (n=5) x= 6.0 (5.56.5); abdomen length (including appendages) (n=5) x=20.5 (19.522.0). Females: Measurements of specimens from south-western Sulawesi [mm] — Hind wing length (n =5) x= 26.0 (25.529.5); hind wing width (n =5) x=6.5 (6.07.0); abdomen length (including appendages) (n =5) x= 20.5 (19.021.5). Specimens from Salayar [mm] — Hind wing length (n =3) 24.025.0; hind wing width (n= 3) 5.56.0; abdomen length (including appendages) (n= 3) 18.020.0.</paragraph>
</subSubSection>
<subSubSection lastPageId="46" lastPageNumber="345" pageId="45" pageNumber="344" type="materials_examined">
<paragraph blockId="45.[159,1037,253,1544]" box="[159,522,253,280]" pageId="45" pageNumber="344">
<heading bold="true" box="[159,522,253,280]" fontSize="11" level="3" pageId="45" pageNumber="344" reason="6">
<emphasis bold="true" box="[159,360,253,279]" pageId="45" pageNumber="344">Material studied</emphasis>
(290♂ 196♀)
</heading>
</paragraph>
<paragraph blockId="45.[159,1037,253,1544]" box="[159,621,286,310]" pageId="45" pageNumber="344">
<emphasis bold="true" box="[159,255,286,310]" pageId="45" pageNumber="344">Sulawesi</emphasis>
(Celebes, without further locality)
</paragraph>
<paragraph blockId="45.[159,1037,253,1544]" pageId="45" pageNumber="344">
<materialsCitation ID-GBIF-Occurrence="1949138306" box="[159,700,316,341]" collectingDate="1873" location="Celebes. Before" municipality="Celebes. Before" pageId="45" pageNumber="344" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<specimenCount box="[159,192,316,340]" pageId="45" pageNumber="344" type="male">1♂</specimenCount>
(
<typeStatus box="[209,311,317,341]" pageId="45" pageNumber="344">
<emphasis bold="true" box="[209,311,317,341]" pageId="45" pageNumber="344">lectotype</emphasis>
</typeStatus>
),
<collectingMunicipality box="[333,499,316,340]" pageId="45" pageNumber="344">Celebes. Before</collectingMunicipality>
<collectingDate box="[507,558,316,340]" pageId="45" pageNumber="344" value="1873">1873</collectingDate>
(in BMNH);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="1949138304" pageId="45" pageNumber="344">
<specimenCount box="[708,740,316,340]" pageId="45" pageNumber="344" type="male">1♂</specimenCount>
<specimenCount box="[748,781,316,340]" pageId="45" pageNumber="344" type="female">3♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:0395A91CAC56FFB49536FC68FE7CFCF4:8EE34ED1AC5CFFAB9758FE93FC74FEE8" box="[795,881,316,340]" pageId="45" pageNumber="344">Celebes</location>
, [before
<date box="[975,1029,316,340]" pageId="45" pageNumber="344" value="1916">1916</date>
]
<botanicCollectionCodeGroup box="[159,438,347,371]" pageId="45" pageNumber="344">
(ded.
<collectorName box="[218,323,347,371]" pageId="45" pageNumber="344">R. Martin</collectorName>
; in
<collectionCode box="[357,431,347,371]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="45" pageNumber="344">SMFD</collectionCode>
)
</botanicCollectionCodeGroup>
;
</materialsCitation>
1♂ 1♀, Celebes, [before 1916]. Leg. C. Ribbe (in SMFD).
<emphasis bold="true" box="[159,342,385,409]" pageId="45" pageNumber="344">Sulawesi Selatan</emphasis>
(South Sulawesi, including present-day Sulawesi Barat, West Su- lawesi)
</paragraph>
<paragraph blockId="45.[159,1037,253,1544]" lastBlockId="46.[159,1038,253,1343]" lastPageId="46" lastPageNumber="345" pageId="45" pageNumber="344">28♂ 13♀, Celebes mer[idional]. 1896. Leg. H. Fruhstorfer, (in SMFD); 1♂, Lompa-Battau. 3000. Mar. 1896. Leg. H. Fruhstorfer; 6♂ 1♀, Bonthain, 1897. Leg. H. Fruhstorfer; 1♀, Maros, 34 Sep. 1904. Leg. E. Lorenz Meyer; 2♂ 1♀, Maros. 28 Apr. 1914. Leg. L. Martin; 4♀, Surroundings Makassar, June 1929. Leg. G. Overdijkink; 1♂ 5♀, Bantimoeroeng, 3 Oct. 1930. Leg. J. van der Vecht; 1♂ 1♀, Bantimoe- roeng. Aug. 1931. Leg. G. Heinrich; 7♂ 6♀, Watampone. Lonrong. 150 m asl. 25 June 1936. Leg. L.J. Toxopeus; Maros; 1♂, Bantimoerong. 200 m. 27 June 1936. Leg. L.J. Toxopeus; 1♂ 1♀, Bonthain. Makassar-Malino Rd. pl. 47 and pl. 46. 200 m. 24 July 1936. Leg. L.J. Toxopeus; 2♂, Maros. Bantimoerong. 24 July 1936. Leg. L.J. Toxopeus; 1♀, Bonthain, July 1938. J.P. Kalis; 23♂ 15♀, Eiland Saleier [Salayer Is- land]. Pariangan. 100 m. OctNov 1938. Leg. J.P.A. Kalis; 6♂, Paloppo. 1939. Leg. M.E. Walsh-Held; 3♂ 2♀, Madjene. 810 Nov. 1939. Leg. J.J. van der Starre; 9♂ 2♀, Maros n/b Msar [near Makassar]. 1319 Nov. 1939. Leg. J.J. van der Starre; 1♂ 2♀, Madjene. 30 Nov. 1939. Leg. J.J. van der Starre; 1♂, Masimboe [Masimbu]. 18 Mar. 1940. 500 m. Leg. J.J. van der Starre; 1♀, Madjene, 20 Mar.1940. Leg. J.J. van der Starre; 1♂ 1♀, Loewoe. Masamba. S. [Sungai] Baeboenta [appr. 8 km W of Masamba]. 2223 June 1940. Leg. L.L.A. Maurenbrecher; 1♀, Masimboe [Ma- simbu]. 20 Aug. 1940. Leg. J.J. van der Starre; 1♂ 1♀, Bantimoeroeng. CdR XXX. 11 Apr. 1941. Leg. L. Coomans de Ruiter; 2♀, Maros. Bantimoeroeng. Omgeving waterval [Surroundings waterfall] CdR XXX. 22 May 1941. Leg. L. Coomans de Ruiter; 1♂, Loewoe. Masamba. S. Baebunta. 25 May 1941. Leg. L.L.A. Maurenbre- cher; 2♂ 3♀, Maros. Bantimoeroeng. Omgeving waterval. CdR XXX. 1112 June 1941. Leg. L. Coomans de Ruiter; 1♀, Luwu distr. Masamba. Lamasi. 12 June 1941. Leg. L.L.A. Maurenbrecher; 1♂, Maros. Bantimoeroeng. CdR XXX. 17 Aug. 1941. Leg. L. Coomans de Ruiter; 5♂ 1♀, km 47 road to Malino [from Makassar], CdR 36. 23 Aug. 1941. Leg. L. Coomans de Ruiter; 2♀, Maros. Bantimurung. 6 July 1949. Leg. W.A. van der Noordaa; 4♂ 2♀, Bantimurung. 1018 Aug. 1949. Leg. A. Diakonoff; 6♂, Bantimurung area. Pattunuang Asue. 200 m. 2931 May 1982. Leg. M.A. Lieftinck; 1♂, Mt. Lompobatang area. Malino. 1100 m. 5°15S, 119°51E. 210 Jun 1982. Leg. M.A. Lieftinck; 3♂ 2♀, Mt. Lompobatang area. Affl. Berang river. 400 m. 37 June 1982. Leg. M.A. Lieftinck; 8♂ 2♀, E of Maros. Bantimu- rung area. Surroundings of Bantimurung waterfall. QK94. 18 Sep14 Oct 1983. Leg. S.S. Pariwono; 2♂ 1♀, E of Maros. Bantimurung area. QK94. Biseang Labboro. 275 m. 411 October 1983. Leg. S.S. Pariwono; 13♂ 4♀, E of Maros. Bantimurung area. QK 94. Pattunuang Asue. 250 m. 21 Sep10 Oct 1983. Leg. S.S. Pariwono; 3♂ 3♀, E of Maros: rivulet in Bantimurung area. 5°02S, 119°40E. 1623 June 1985. Leg. J. van Tol; 2♂ 3♀, E of Maros. 5°02S, 119°40E. Rivulet in Bantimurung area. Semi-cultivated. 23 June 1985. Sample A. Leg. J. van Tol; 4♂ 4♀, E of Maros: Ban- timurung area, Sg. Pattunuang Asue. 5°03S, 119°41E. Fast flowing stream through disturbed forest on limestone. Large boulders, ponded areas, clear water. Sample 91JvT07. 19 Apr. 1991. Leg. J. van Tol; 11♂ 2♀, Circa 10 km E of Maros: surround- ings of Bantimurung waterfall. River through disturbed forest on limestone, and ponds in semi-cultivated area, c. 200 m asl. 1923 Apr. 1991 (91JvT09). Leg. J. van Tol; 6♂ 2♀, Bantimurung, Sungai Bantimurung (5°01S, 119°41E), 2930 Mar. 1993. Leg. A. Günther (in AGPC); 2♀, Bantimurung, Sungai Bantimurung (5°01S, 119°41E), 21 Aug. 1993. Leg. A. Günther (in AGPC); 9♂ 2♀, Pattunuang Asue, Sungai Pattunuang Asue (5°04S, 119°43E), 23 Aug. 1994. Leg. A. Günther &amp; F. Randow (in AGPC); 3♂ 3♀, Pattunuang Asue, Sungai Pattunuang Asue (5°04S, 119°43E), 12 Mar. 1997. Leg. A. Günther (in AGPC); 1♂, Pattunuang Asue, Sungai Pattunuang Asue (5°04S, 119°43E), 14 Sep. 2010. Leg. A. Günther (in AGPC); 1♂ 1♀, Sulawesi Selatan, Pattunuang Asue, Sungai Pattunuang Asue (5°04S, 119°43E), 31 July1 Aug. 2011. Leg. A. Günther (in AGPC).</paragraph>
<paragraph blockId="46.[159,1038,253,1343]" box="[159,543,751,776]" pageId="46" pageNumber="345">
<emphasis bold="true" box="[159,341,751,776]" pageId="46" pageNumber="345">Sulawesi Tengah</emphasis>
(Central Sulawesi)
</paragraph>
<paragraph blockId="46.[159,1038,253,1343]" pageId="46" pageNumber="345">1♂, Palu: Pekawa, 15 Sep. 1912. Leg. L. Martin; 1♂, Palu: Soeroemana, Feb. 1913. Leg. L. Martin.</paragraph>
<paragraph blockId="46.[159,1038,253,1343]" box="[159,510,851,875]" pageId="46" pageNumber="345">
<emphasis bold="true" box="[159,324,851,875]" pageId="46" pageNumber="345">Sulawesi Utara</emphasis>
(North Sulawesi)
</paragraph>
<paragraph blockId="46.[159,1038,253,1343]" pageId="46" pageNumber="345">9♂ 17♀, Toli Toli [locality considered a lapsus, not on the map], [ca. 1897]. Leg. H. Fruhstorfer (in SMFD).</paragraph>
<paragraph blockId="46.[159,1038,253,1343]" box="[159,633,950,975]" pageId="46" pageNumber="345">
<emphasis bold="true" box="[159,363,950,975]" pageId="46" pageNumber="345">Sulawesi Tenggara</emphasis>
(South-eastern Sulawesi)
</paragraph>
<paragraph blockId="46.[159,1038,253,1343]" pageId="46" pageNumber="345">1♂, Kolaka. T. Elbert. Ovt. 1909. [Partly handwritten] (in SMFD); 11♂ 2♀, 30 km WSW Kendari: Sg Amoito. Small stream and trickles through disturbed forest N of Pegunungan Boroboro. Clear water. Alt. 100 m. 5 Feb. 1989. Sample 89JvT09. Leg. J. van Tol; 5♂ 2♀, 18 km ENE Kolaka: Sg. Koloimba near crossing with road. Black- water stream with vegetation. 4°02S, 121°47E. Sample 89JvT11. 7 Feb. 1989. Leg. J. van Tol; 2♂, Pulau Kabaena. Tangkeno. 23 Nov. 1989. Leg. C. Lepelaar; 1♀, Pulau Kabaena. 4 km S of Tangkeno. Cave camp. S8928R. Open riverine forest along Sg. Lakambula. 58 Nov. 1989. Leg. R. de Jong &amp; J. Huisman; 1♀, Pulau Kabaena. 1 km S of Tangkeno. Riverine forest, bamboo, grassy slopes (S8931). 550 m. 89 Nov. 1989. Leg. R. de Jong; 7♂, Pulau Kabaena. Batuawu. Cultivations, mainly coconut (S8933). Sea level. 11 Nov. 1989. Leg. R. de Jong; 1♀, Moramo. Sg. Sena. Forest with dense undergrowth along stream. 50 m. S 8938. Leg. R. de Jong.</paragraph>
</subSubSection>
<subSubSection lastPageId="49" lastPageNumber="348" pageId="46" pageNumber="345" type="distribution">
<paragraph blockId="46.[159,1036,1406,1545]" box="[159,459,1406,1433]" pageId="46" pageNumber="345">
<heading bold="true" box="[159,459,1406,1433]" fontSize="11" level="3" pageId="46" pageNumber="345" reason="6">
<emphasis bold="true" box="[159,459,1406,1433]" pageId="46" pageNumber="345">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="46.[159,1036,1406,1545]" pageId="46" pageNumber="345">Widespread in south-western Sulawesi below about 400 m, northward up to Palopo. Some specimens were collected along the coast near Palu by L. Mar- tin (in SMFD); these records need confirmation.More local in other parts of</paragraph>
<caption httpUri="https://zenodo.org/record/2200487/files/figure.png" pageId="47" pageNumber="346" targetBox="[159,727,249,1547]" targetPageId="47">
<paragraph blockId="47.[765,1037,588,1552]" pageId="47" pageNumber="346">
<emphasis bold="true" box="[765,942,588,613]" pageId="47" pageNumber="346">Figures 4552.</emphasis>
<emphasis italics="true" pageId="47" pageNumber="346">
<taxonomicName authorityName="Rambur" authorityYear="1842" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="47" pageNumber="346" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
monochroa
</emphasis>
. Male, hind wing, ventral view. 45 Sulawesi Selatan. Bantimurung, Sg. Pat- tunuangAsue,05-ix-1999 (in life); 46 Sulawesi Se- latan, Watampone, Lonrong, 25-vi-1936. Leg.L.J. Toxopeus [JvT 05452]; 47 Sulawesi Selatan, Palopo, 1939. Leg. M.E. Walsh [JvT 05342]; 48 Sulawesi Selatan, Madjene, 810­xi­1939. Leg. J.J. van der Starre [JvT 05345]; 49 Sulawesi Tenggara, 18 km ENE of Kolaka: Sg. Koloimba. Leg. J. van Tol, 07-ii-1989 [JvT 06622]; 50 Su- lawesi Tenggara, 30 km WSW of Kendari, Sg. Amoito, 05-ii-1989. Leg. J. van Tol [JvT 05503]; 51 Sulawesi Tenggara, Moramo, Sg. Moramo, 1617-xi-1989. Leg. R. de Jong [JvT 05507]; 52 Sulawesi Tenggara, Pulau Kabaena, Batuawu, 11- -xi-1989. Leg. R. de Jong [JvT 05330]. Photos: AG
</paragraph>
</caption>
<paragraph blockId="48.[159,1037,253,653]" pageId="48" pageNumber="347">
Sulawesi (
<figureCitation box="[272,357,253,280]" captionStart="Figure 85" captionStartId="81.[711,779,1497,1522]" captionTargetBox="[161,665,912,1547]" captionTargetId="figure@81.[159,665,911,1547]" captionTargetPageId="81" captionText="Figure 85. Distribution Rhinocypha monochroa." httpUri="https://zenodo.org/record/2200561/files/figure.png" pageId="48" pageNumber="347">Fig. 85</figureCitation>
). The locality of specimens in RMNH labelled Gorontalo / Minahasa (northern arm of Sulawesi) is considered doubtful and as such in- cluded on the map. In south-eastern Sulawesi it is much rarer than
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="347" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis italics="true" pageId="48" pageNumber="347">R. frontalis</emphasis>
</taxonomicName>
. Although they occur in the same regions,
<taxonomicName box="[717,878,365,391]" pageId="48" pageNumber="347">
<emphasis box="[717,878,365,391]" italics="true" pageId="48" pageNumber="347">R. monochroa</emphasis>
</taxonomicName>
has not been observed at the same sites as
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[497,624,403,429]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="347" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[497,624,403,429]" italics="true" pageId="48" pageNumber="347">R. frontalis</emphasis>
</taxonomicName>
or
<emphasis box="[662,829,403,429]" italics="true" pageId="48" pageNumber="347">
<taxonomicName box="[662,825,403,429]" pageId="48" pageNumber="347">R. phantasma</taxonomicName>
.
</emphasis>
It is the only spe- cies of
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[238,375,440,466]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis box="[238,375,440,466]" italics="true" pageId="48" pageNumber="347">Rhinocypha</emphasis>
</taxonomicName>
recorded from the islands of Salayar and Kabaena, while it is not known from Buton, where both
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[621,746,477,503]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="347" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[621,746,477,503]" italics="true" pageId="48" pageNumber="347">R. frontalis</emphasis>
</taxonomicName>
and
<taxonomicName box="[801,959,477,503]" pageId="48" pageNumber="347">
<emphasis box="[801,959,477,503]" italics="true" pageId="48" pageNumber="347">R. phantasma</emphasis>
</taxonomicName>
occur.
</paragraph>
<paragraph blockId="48.[159,1037,253,653]" lastBlockId="49.[159,1036,253,429]" lastPageId="49" lastPageNumber="348" pageId="48" pageNumber="347">
Little has been recorded about the habitat requirements of this species, with the exception of the frequently visited streams at Bantimurung and Pattunuang Asue near Maros in SW Sulawesi (
<figureCitation box="[718,805,589,616]" captionStart="Figure 15" captionStartId="18.[159,227,405,430]" captionTargetBox="[509,1035,249,600]" captionTargetId="figure@18.[509,1035,249,600]" captionTargetPageId="18" captionText="Figure 15. Habitat of Rhinocypha monochroa, Libellago rufescens and L. asclepiades. Stream Sg. Pattunuang Asue in Bulusaraung National Park, Southwest Sulawesi. Photo: AG (31-vii-2011)" httpUri="https://zenodo.org/record/2200415/files/figure.png" pageId="48" pageNumber="347">Fig. 15</figureCitation>
). Our observations indicate that
<taxonomicName box="[315,477,627,653]" pageId="48" pageNumber="347">
<emphasis box="[315,477,627,653]" italics="true" pageId="48" pageNumber="347">R. monochroa</emphasis>
</taxonomicName>
frequents more open streams than
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[901,1029,627,653]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="347" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[901,1029,627,653]" italics="true" pageId="48" pageNumber="347">
R.
<taxonomicName box="[934,1029,627,653]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="347" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
</emphasis>
</taxonomicName>
. The species was found in clear and fast flowing streams, mostly in areas of lowland primary or secondary forests. The species seems to be absent from muddy watercourses,
<emphasis box="[413,444,329,354]" italics="true" pageId="49" pageNumber="348">e.g</emphasis>
., near larger forest clearings and below paddy field or settlements. The streams in south-western Sulawesi have partially very large fluctuations in flow, caused by seasonal influences.
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/2200495/files/figure.png" pageId="48" pageNumber="347" startId="48.[159,227,1498,1523]" targetBox="[156,1039,694,1459]" targetPageId="48">
<paragraph blockId="48.[159,1036,1498,1552]" pageId="48" pageNumber="347">
<emphasis bold="true" box="[159,268,1498,1523]" pageId="48" pageNumber="347">Figure 53.</emphasis>
<emphasis box="[276,540,1498,1522]" italics="true" pageId="48" pageNumber="347">
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[276,402,1498,1522]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="48" pageNumber="347" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
monochroa.
</emphasis>
Simplified scheme of coloration of underside of hind wing. Artwork by AG
</paragraph>
</caption>
<subSubSection pageId="49" pageNumber="348" type="biology_ecology">
<paragraph blockId="49.[159,1037,477,1101]" box="[159,286,477,503]" pageId="49" pageNumber="348">
<heading bold="true" box="[159,286,477,503]" fontSize="11" level="3" pageId="49" pageNumber="348" reason="6">
<emphasis bold="true" box="[159,286,477,503]" pageId="49" pageNumber="348">Behaviour</emphasis>
</heading>
</paragraph>
<paragraph blockId="49.[159,1037,477,1101]" pageId="49" pageNumber="348">
The behaviour was studied by GÜNTHER (2008a) in south-western Sulawesi. Oviposition sites were dead rotting logs, driftwood or other plant substrates lying in the water. Males defend territories around these sites with long- lasting threat displays. The patterns of these threat flights are unique among all
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[192,329,664,690]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="49" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis box="[192,329,664,690]" italics="true" pageId="49" pageNumber="348">Rhinocypha</emphasis>
</taxonomicName>
species described here. The synchronous wing beat is signifi- cantly modified. Single hind wings are presented forward and laterally to the rival male. During this display the wing beat of these wings pauses over 23 strokes. The strongly banded, contrasting colours of the hind wings, described above for the males, probably intensifies the signal effect in this style of presentation.
</paragraph>
<paragraph blockId="49.[159,1037,477,1101]" pageId="49" pageNumber="348">
The presentation of the hind wings is of obvious importance during male courtship displays to females.It is conceivable that what we describe as
<emphasis italics="true" pageId="49" pageNumber="348">
<taxonomicName authorityName="Rambur" authorityYear="1842" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="49" pageNumber="348" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
monochroa
</emphasis>
, actually consists of several genetically distinct populations that do not interbreed since they are isolated by their behaviour and male colour patterns, and should therefore be treated as separate species. Further behavioural and genetic studies are needed to help clarify this matter.
</paragraph>
</subSubSection>
<subSubSection lastPageId="50" lastPageNumber="349" pageId="49" pageNumber="348" type="discussion">
<paragraph blockId="49.[159,1036,1150,1549]" box="[159,267,1150,1176]" pageId="49" pageNumber="348">
<heading bold="true" box="[159,267,1150,1176]" fontSize="11" level="3" pageId="49" pageNumber="348" reason="6">
<emphasis bold="true" box="[159,267,1150,1176]" pageId="49" pageNumber="348">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="49.[159,1036,1150,1549]" lastBlockId="50.[159,1037,253,840]" lastPageId="50" lastPageNumber="349" pageId="49" pageNumber="348">
As already mentioned above, RIS (1916: 310 ff.) treated the
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[899,1036,1187,1213]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="49" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis box="[899,1036,1187,1213]" italics="true" pageId="49" pageNumber="348">Rhinocypha</emphasis>
</taxonomicName>
specimens available at the time, now preserved in SMFD. Ris distinguished two forms of
<taxonomicName box="[312,472,1261,1287]" pageId="49" pageNumber="348">
<emphasis box="[312,472,1261,1287]" italics="true" pageId="49" pageNumber="348">R. monochroa</emphasis>
</taxonomicName>
. Form a (»typisch«) was mentioned from »Süd- Celebes«, Toli-Toli, Lombok [considered erroneous], and Maros. Form b was mentioned from the Takala Mts and Suramana and Pekawa, both near Palu. From Suramana and Pekawa, also
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[694,822,1373,1399]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="49" pageNumber="348" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[694,822,1373,1399]" italics="true" pageId="49" pageNumber="348">
R.
<taxonomicName box="[727,822,1373,1399]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="49" pageNumber="348" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
</emphasis>
</taxonomicName>
was reported. We have examined these specimens in SMFD, and conclude the following: all specimens of
<taxonomicName box="[325,489,1448,1474]" pageId="49" pageNumber="348">
<emphasis box="[325,489,1448,1474]" italics="true" pageId="49" pageNumber="348">R. monochroa</emphasis>
</taxonomicName>
Form a indeed belong to
<taxonomicName box="[823,988,1448,1474]" pageId="49" pageNumber="348">
<emphasis box="[823,988,1448,1474]" italics="true" pageId="49" pageNumber="348">R. monochroa</emphasis>
</taxonomicName>
, al- though we have strong doubts whether the locality Toli-Toli is correct (not included on the map). There is a male specimen of a
<taxonomicName authorityName="Selys" authorityYear="1840" box="[783,883,1523,1549]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="49" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis box="[783,883,1523,1549]" italics="true" pageId="49" pageNumber="348">Euphaea</emphasis>
</taxonomicName>
(ex UMMZ) with a similar label in RMNH. It was identified as
<taxonomicName authority="Kruger" authorityName="Kruger" box="[755,1000,253,280]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="lara">
<taxonomicName box="[755,909,253,279]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="lara">
<emphasis box="[755,909,253,279]" italics="true" pageId="50" pageNumber="349">
<taxonomicName box="[755,855,253,279]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="genus">Euphaea</taxonomicName>
<taxonomicName authority="Kruger" authorityName="Kruger" box="[863,1000,253,280]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="lara">
<taxonomicName box="[863,909,253,279]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="lara">lara</taxonomicName>
Krüger
</taxonomicName>
</emphasis>
</taxonomicName>
</taxonomicName>
by M. Hämäläinen (with question mark), and previously as
<emphasis box="[831,1036,291,317]" italics="true" pageId="50" pageNumber="349">
<taxonomicName authorityName="Selys" authorityYear="1840" box="[831,931,291,317]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="genus">Euphaea</taxonomicName>
brunnea
</emphasis>
Selys by F. Förster. It may actually represent a specimen of the type series of
<taxonomicName authority="McLachlan. No" authorityName="McLachlan. No" box="[159,699,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName authority="McLachlan." authorityName="McLachlan." box="[159,659,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName authority="McLachlan" authorityName="McLachlan" box="[159,654,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName box="[159,513,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName box="[159,311,365,391]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="lara">
<emphasis box="[159,311,365,391]" italics="true" pageId="50" pageNumber="349">
<taxonomicName box="[159,259,365,391]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="genus">Euphaea</taxonomicName>
<taxonomicName authority="McLachlan. No" authorityName="McLachlan. No" box="[265,699,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName authority="McLachlan." authorityName="McLachlan." box="[265,659,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
lara
<taxonomicName authority="McLachlan. No" authorityName="McLachlan. No" box="[316,699,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName authority="McLachlan." authorityName="McLachlan." box="[316,659,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName authority="McLachlan" authorityName="McLachlan" box="[316,654,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName box="[316,513,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
var.
<taxonomicName authority="McLachlan. No" authorityName="McLachlan. No" box="[363,699,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName authority="McLachlan." authorityName="McLachlan." box="[363,659,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName authority="McLachlan" authorityName="McLachlan" box="[363,654,365,392]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<taxonomicName box="[363,513,365,391]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="variety" species="lara" variety="lombockensis">
<emphasis box="[363,513,365,391]" italics="true" pageId="50" pageNumber="349">lombockensis</emphasis>
</taxonomicName>
McLachlan
</taxonomicName>
.
</taxonomicName>
No
</taxonomicName>
</taxonomicName>
</taxonomicName>
</taxonomicName>
</taxonomicName>
</taxonomicName>
</taxonomicName>
</emphasis>
</taxonomicName>
</taxonomicName>
</taxonomicName>
</taxonomicName>
</taxonomicName>
other specimen of the family
<taxonomicName authorityName="Jacobson &amp; Bianchi" authorityYear="1905" box="[159,295,403,429]" class="Insecta" family="Euphaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="family">Euphaeidae</taxonomicName>
is known from Sulawesi. We conclude that labels of specimens collected by H. Fruhstorfer from »Toli Toli« are unreliable.
</paragraph>
<paragraph blockId="50.[159,1037,253,840]" pageId="50" pageNumber="349">
The specimens characterised as
<taxonomicName box="[567,731,477,503]" pageId="50" pageNumber="349">
<emphasis box="[567,731,477,503]" italics="true" pageId="50" pageNumber="349">R. monochroa</emphasis>
</taxonomicName>
“Form b” are labelled by F. Ris as
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[268,398,515,541]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[268,398,515,541]" italics="true" pageId="50" pageNumber="349">R. frontalis</emphasis>
</taxonomicName>
[sic!] “Form b” in the collection. We have identified these specimens as follows: the 3♂ 2♀ from the Takala Mts are
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="virgulata" status="sp. nov.">
<emphasis italics="true" pageId="50" pageNumber="349">
R.
<taxonomicName class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="virgulata" status="sp. nov.">virgulata</taxonomicName>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[211,301,590,616]" pageId="50" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
(Ris no. 1441, 1442, 1446 and 1447), and
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[799,939,589,615]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="flavipoda" status="sp. nov.">
<emphasis box="[799,939,589,615]" italics="true" pageId="50" pageNumber="349">R. flavipoda</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[947,1037,590,616]" pageId="50" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
(Ris no. 1443, ♂). The specimens from Pekawa and Soeroemana / Suramana had been correctly identified: Pekawa:
<taxonomicName box="[614,775,664,690]" pageId="50" pageNumber="349">
<emphasis box="[614,775,664,690]" italics="true" pageId="50" pageNumber="349">R. monochroa</emphasis>
</taxonomicName>
(Ris no. 1437, ♂) and
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[159,287,701,727]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[159,287,701,727]" italics="true" pageId="50" pageNumber="349">R. frontalis</emphasis>
</taxonomicName>
(Ris no. 14381439, ♂ and Ris no. 1440, ♀), and Soeroemana:
<taxonomicName box="[159,322,739,765]" pageId="50" pageNumber="349">
<emphasis box="[159,322,739,765]" italics="true" pageId="50" pageNumber="349">R. monochroa</emphasis>
</taxonomicName>
(Ris no. 1445), and
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[572,702,739,765]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[572,702,739,765]" italics="true" pageId="50" pageNumber="349">R. frontalis</emphasis>
</taxonomicName>
(Ris no. 1444), both males. These specimens represent the only observations of sympatry of
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[910,1036,776,802]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="50" pageNumber="349" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[910,1036,776,802]" italics="true" pageId="50" pageNumber="349">R. frontalis</emphasis>
</taxonomicName>
and
<taxonomicName box="[209,370,813,839]" pageId="50" pageNumber="349">
<emphasis box="[209,370,813,839]" italics="true" pageId="50" pageNumber="349">R. monochroa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>