466 lines
66 KiB
XML
466 lines
66 KiB
XML
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<mods:title id="333EE2D64C940A55892616648E8A953B">Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<taxonomicName id="658FC8F84B16FFDAE7B8FDEBFDADFDFF" ID-CoL="85LM2" authority="Banner & Banner, 1985" authorityName="Banner & Banner" authorityYear="1985" box="[151,614,582,609]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nipa">
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<emphasis id="90FB6F694B16FFDAE7B8FDEBFDADFDFF" bold="true" box="[151,614,582,609]" pageId="82" pageNumber="83">
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<emphasis id="90FB6F694B16FFDAE7B8FDEBFEF3FDFE" bold="true" box="[151,312,582,609]" italics="true" pageId="82" pageNumber="83">Alpheus nipa</emphasis>
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<bibRefCitation id="C61ECE8A4B16FFDAE610FDEAFDADFDFF" author="Banner, D. M. & Banner, A. H." box="[319,614,582,609]" pageId="82" pageNumber="83" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner, 1985</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="EA95E0F04B16FFDAE7B8FDC6FEA7FD1A" box="[151,364,618,645]" pageId="82" pageNumber="83" type="description">
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<paragraph id="A230B37B4B16FFDAE7B8FDC6FEA7FD1A" blockId="82.[151,614,582,645]" box="[151,364,618,645]" pageId="82" pageNumber="83">
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(
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<figureCitation id="3AB4AFFE4B16FFDAE7B0FDC6FF35FD1A" box="[159,254,619,645]" captionStart="FIGURE 44" captionStartId="82.[151,250,1710,1734]" captionTargetBox="[151,1435,987,1687]" captionTargetId="figure-336@82.[151,1435,987,1687]" captionTargetPageId="82" captionText="FIGURE 44. Alpheus nipa Banner & Banner, 1985: paratypes, males (cl 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526), detached chelipeds; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (right) cheliped, lateral view; D—same, dorsolateral view; E—same, mesial view." figureDoi="http://doi.org/10.5281/zenodo.7912431" httpUri="https://zenodo.org/record/7912431/files/figure.png" pageId="82" pageNumber="83">Figs. 44</figureCitation>
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,
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<figureCitation id="3AB4AFFE4B16FFDAE626FDC7FEECFD1A" box="[265,295,618,645]" captionStart="FIGURE 45" captionStartId="83.[151,250,1527,1551]" captionTargetBox="[170,1409,197,1483]" captionTargetId="figure-15@83.[151,1436,181,1503]" captionTargetPageId="83" captionText="FIGURE 45. Alpheus nipa Banner & Banner, 1985: paratype, male (cl indet., range: 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526); A—rostro-orbital region of carapace, dorsal view; B—frontal region, lateral view; C— ventromesial carina of first article of antennular peduncle, lateral view; D—antennal scaphocerite, dorsal view; E—third maxilliped, lateral view [arthrobranch not drawn]; F—same, detail of branchial structures on exopod, mesial view; G—major (right) cheliped, chela, mesial view; H—same, distal part of chela, fingers open, lateral view; I—second pleopod, appendices masculina and interna; J—right uropod, detail of distolateral angle, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912433" httpUri="https://zenodo.org/record/7912433/files/figure.png" pageId="82" pageNumber="83">45</figureCitation>
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,
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<figureCitation id="3AB4AFFE4B16FFDAE61CFDC7FEAEFD1A" box="[307,357,618,645]" captionStart="FIGURE 52" captionStartId="106.[151,250,1909,1933]" captionTargetBox="[156,1430,181,1885]" captionTargetId="figure-21@106.[156,1430,181,1885]" captionTargetPageId="106" captionText="FIGURE 52. Distributional maps for species of Alpheus treated in this study:A—A. takla sp. nov.; B—A. mangalis sp. nov. and A. cf. mangalis sp. nov.; C—A. songkla Banner & Banner, 1966 and A. cf. songkla (see text); D—A. paludicola Kemp, 1915, A. nipa Banner & Banner, 1985 and A. bunburius Banner & Banner, 1982; E—A. pontederiae de Rochebrune, 1883; F—A. firmus Kim & Abele, 1988 and A. cf. firmus (see text)." figureDoi="http://doi.org/10.5281/zenodo.7912457" httpUri="https://zenodo.org/record/7912457/files/figure.png" pageId="82" pageNumber="83">52D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="EA95E0F04B16FFDAE7B8FD02FDB5FD58" box="[151,638,687,711]" pageId="82" pageNumber="83" type="reference_group">
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<paragraph id="A230B37B4B16FFDAE7B8FD02FDB5FD58" blockId="82.[151,638,687,711]" box="[151,638,687,711]" pageId="82" pageNumber="83">
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<treatmentCitationGroup id="829F94554B16FFDAE7B8FD02FDB5FD58" box="[151,638,687,711]" pageId="82" pageNumber="83">
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<treatmentCitation id="232E956A4B16FFDAE7B8FD02FDFCFD58" author="Banner, D. M. & Banner, A. H." box="[151,567,687,711]" page="21" pageId="82" pageNumber="83" year="1985">
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<taxonomicName id="658FC8F84B16FFDAE7B8FD02FDFCFD58" ID-CoL="85LM2" authority="Banner & Banner 1985: 21" authorityName="Banner & Banner" authorityPageNumber="21" authorityYear="1985" box="[151,567,687,711]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nipa">
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<emphasis id="90FB6F694B16FFDAE7B8FD02FED7FD58" box="[151,284,687,711]" italics="true" pageId="82" pageNumber="83">Alpheus nipa</emphasis>
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<bibRefCitation id="C61ECE8A4B16FFDAE60CFD1DFDFCFD58" author="Banner, D. M. & Banner, A. H." box="[291,567,687,711]" pageId="82" pageNumber="83" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner 1985: 21</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 2.
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</paragraph>
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<paragraph id="A230B37B4B16FFDAE7B8FD58FC25FCE3" blockId="82.[151,1436,757,964]" pageId="82" pageNumber="83">
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<emphasis id="90FB6F694B16FFDAE7B8FD58FE8BFC90" bold="true" box="[151,320,757,783]" pageId="82" pageNumber="83">Type material</emphasis>
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.
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<materialsCitation id="12E7B9264B16FFDAE67FFD5BFC21FCE3" ID-GBIF-Occurrence="4104078314" collectingDate="1951-10-05" collectionCode="ZMUC" country="Indonesia" county="Galathea Expedition" elevation="40" latitude="4.3333335" location="Strait" longLatPrecision="1306" longitude="98.88333" municipality="Medan" pageId="82" pageNumber="83" specimenCode="CRU-7526" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
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<typeStatus id="7D340DD94B16FFDAE67FFD5BFE0BFC8F" box="[336,448,758,784]" pageId="82" pageNumber="83" type="paratype">Paratypes</typeStatus>
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,
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<specimenCount id="B48978F24B16FFDAE6E2FD58FDEEFC90" box="[461,549,757,784]" pageId="82" pageNumber="83" type="male">5 males</specimenCount>
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(cl
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<quantity id="65771E9E4B16FFDAE57BFD58FD14FC8F" box="[596,735,757,784]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.5" metricValueMin="6.2" pageId="82" pageNumber="83" unit="mm" value="6.85" valueMax="7.5" valueMin="6.2">6.2–7.5 mm</quantity>
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, tl 16.0 –20.0 mm, all missing chelipeds, but detached major and minor chelipeds present in the vial), 1 ov. female (cl
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<quantity id="65771E9E4B16FFDAE40AFCB4FCB4FCAB" box="[805,895,793,820]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="82" pageNumber="83" unit="mm" value="9.5">9.5 mm</quantity>
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, tl
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<quantity id="65771E9E4B16FFDAE48BFCB4FBC0FCAB" box="[932,1035,793,820]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="82" pageNumber="83" unit="mm" value="23.5">23.5 mm</quantity>
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, chl 12.0 mm),
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<collectionCode id="C49E2BBE4B16FFDAE3EEFCB4FAD9FCAC" box="[1217,1298,793,819]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="82" pageNumber="83">ZMUC</collectionCode>
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<specimenCode id="F2291B004B16FFDAE235FCB4FA53FCAB" box="[1306,1432,793,820]" collectionCode="CRU-" pageId="82" pageNumber="83">CRU-7526</specimenCode>
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,
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<collectingCountry id="DA98F3EB4B16FFDAE7B8FC90FECCFCC8" box="[151,263,829,855]" name="Indonesia" pageId="82" pageNumber="83">Indonesia</collectingCountry>
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,
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<location id="A750E5A04B16FFDAE63CFC90FE9BFCC8" LSID="urn:lsid:plazi:treatment:2A26026D4B16FFDCE7B8FDEBFE00FB76:A750E5A04B16FFDAE63CFC90FE9BFCC8" box="[275,336,829,855]" country="Indonesia" county="Galathea Expedition" latitude="4.3333335" longLatPrecision="1306" longitude="98.88333" municipality="Medan" name="Strait" pageId="82" pageNumber="83">Strait</location>
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of Malacca off
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<location id="A750E5A04B16FFDAE52BFC90FDAFFCC8" LSID="urn:lsid:plazi:treatment:2A26026D4B16FFDCE7B8FDEBFE00FB76:A750E5A04B16FFDAE52BFC90FDAFFCC8" box="[516,612,829,855]" country="Indonesia" county="Galathea Expedition" latitude="4.3333335" longLatPrecision="1306" longitude="98.88333" municipality="Medan" name="Sumatra" pageId="82" pageNumber="83">Sumatra</location>
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, vicinity of
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<collectingMunicipality id="425429014B16FFDAE5C2FC90FCF5FCC8" box="[749,830,829,855]" pageId="82" pageNumber="83">Medan</collectingMunicipality>
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,
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<collectingCounty id="4B51CBF74B16FFDAE466FC90FBE4FCC8" box="[841,1071,829,855]" pageId="82" pageNumber="83">Galathea Expedition</collectingCounty>
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sta. 325,
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<geoCoordinate id="C7BBD5BC4B16FFDAE3B2FC90FB3BFCC8" box="[1181,1264,829,855]" degrees="4" direction="north" minutes="20" orientation="latitude" pageId="82" pageNumber="83" precision="925" value="4.3333335">4°20’N</geoCoordinate>
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<geoCoordinate id="C7BBD5BC4B16FFDAE3D7FC90FA93FCC7" box="[1272,1368,829,856]" degrees="98" direction="east" minutes="53" orientation="longitude" pageId="82" pageNumber="83" precision="925" value="98.88333">98°53’E</geoCoordinate>
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, palm fronds floating on water surface (depth at collection site:
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<quantity id="65771E9E4B16FFDAE436FCCCFC9EFCE4" box="[793,853,865,891]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="82" pageNumber="83" unit="m" value="40.0">
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<elevation id="29A254484B16FFDAE436FCCCFC9EFCE4" box="[793,853,865,891]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="82" pageNumber="83" unit="m" value="40.0">40 m</elevation>
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</quantity>
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),
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<date id="D63195BB4B16FFDAE446FCCCFC21FCE3" box="[873,1002,865,892]" pageId="82" pageNumber="83" value="1951-10-05">
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<collectingDate id="C6756C534B16FFDAE446FCCCFC21FCE3" box="[873,1002,865,892]" pageId="82" pageNumber="83" value="1951-10-05">10.05.1951</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="EA95E0F04B16FFDAE7E8FC28FE35F8C8" pageId="82" pageNumber="83" type="description">
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<paragraph id="A230B37B4B16FFDAE7E8FC28FD10FC5B" blockId="82.[151,1436,757,964]" pageId="82" pageNumber="83">
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<emphasis id="90FB6F694B16FFDAE7E8FC28FE9AFC00" bold="true" box="[199,337,901,927]" pageId="82" pageNumber="83">Description</emphasis>
|
||
. See
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||
<bibRefCitation id="C61ECE8A4B16FFDAE6A4FC2BFD57FC3F" author="Banner, D. M. & Banner, A. H." box="[395,668,901,928]" pageId="82" pageNumber="83" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner (1985)</bibRefCitation>
|
||
for detailed description and illustrations; complementary illustrations of one of the
|
||
<typeStatus id="7D340DD94B16FFDAE603FC07FE5CFC5B" box="[300,407,938,964]" pageId="82" pageNumber="83" type="paratype">paratypes</typeStatus>
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||
are provided in
|
||
<figureCitation id="3AB4AFFE4B16FFDAE560FC04FD66FC5C" box="[591,685,937,963]" captionStart="FIGURE 44" captionStartId="82.[151,250,1710,1734]" captionTargetBox="[151,1435,987,1687]" captionTargetId="figure-336@82.[151,1435,987,1687]" captionTargetPageId="82" captionText="FIGURE 44. Alpheus nipa Banner & Banner, 1985: paratypes, males (cl 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526), detached chelipeds; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (right) cheliped, lateral view; D—same, dorsolateral view; E—same, mesial view." figureDoi="http://doi.org/10.5281/zenodo.7912431" httpUri="https://zenodo.org/record/7912431/files/figure.png" pageId="82" pageNumber="83">Figs. 44</figureCitation>
|
||
,
|
||
<figureCitation id="3AB4AFFE4B16FFDAE597FC04FD1CFC5B" box="[696,727,937,964]" captionStart="FIGURE 45" captionStartId="83.[151,250,1527,1551]" captionTargetBox="[170,1409,197,1483]" captionTargetId="figure-15@83.[151,1436,181,1503]" captionTargetPageId="83" captionText="FIGURE 45. Alpheus nipa Banner & Banner, 1985: paratype, male (cl indet., range: 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526); A—rostro-orbital region of carapace, dorsal view; B—frontal region, lateral view; C— ventromesial carina of first article of antennular peduncle, lateral view; D—antennal scaphocerite, dorsal view; E—third maxilliped, lateral view [arthrobranch not drawn]; F—same, detail of branchial structures on exopod, mesial view; G—major (right) cheliped, chela, mesial view; H—same, distal part of chela, fingers open, lateral view; I—second pleopod, appendices masculina and interna; J—right uropod, detail of distolateral angle, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912433" httpUri="https://zenodo.org/record/7912433/files/figure.png" pageId="82" pageNumber="83">45</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="F6F0E3F34B16FFDAE7B8F903FC5BF890" ID-DOI="http://doi.org/10.5281/zenodo.7912431" ID-Zenodo-Dep="7912431" httpUri="https://zenodo.org/record/7912431/files/figure.png" pageId="82" pageNumber="83" startId="82.[151,250,1710,1734]" targetBox="[151,1435,987,1687]" targetPageId="82" targetType="figure">
|
||
<paragraph id="A230B37B4B16FFDAE7B8F903FC5BF890" blockId="82.[151,1436,1710,1807]" pageId="82" pageNumber="83">
|
||
<emphasis id="90FB6F694B16FFDAE7B8F903FED0F959" bold="true" box="[151,283,1710,1734]" pageId="82" pageNumber="83">FIGURE 44</emphasis>
|
||
.
|
||
<taxonomicName id="658FC8F84B16FFDAE605F902FD7FF959" authority="Banner & Banner, 1985" authorityName="Banner & Banner" authorityYear="1985" box="[298,692,1710,1735]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B16FFDAE605F902FE7AF959" box="[298,433,1711,1735]" italics="true" pageId="82" pageNumber="83">Alpheus nipa</emphasis>
|
||
<bibRefCitation id="C61ECE8A4B16FFDAE696F902FD7FF959" author="Banner, D. M. & Banner, A. H." box="[441,692,1710,1735]" pageId="82" pageNumber="83" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
: paratypes, males (cl 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526), detached chelipeds; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (right) cheliped, lateral view; D—same, dorsolateral view; E—same, mesial view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A230B37B4B16FFDAE7E8F890FE35F8C8" blockId="82.[151,1436,1853,2023]" box="[199,510,1853,1879]" pageId="82" pageNumber="83">
|
||
<emphasis id="90FB6F694B16FFDAE7E8F890FEB0F8C8" bold="true" box="[199,379,1853,1879]" pageId="82" pageNumber="83">Colour pattern</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EA95E0F04B16FFDAE7E8F8CCFDB0F8E4" box="[199,635,1889,1915]" pageId="82" pageNumber="83" type="materials_examined">
|
||
<paragraph id="A230B37B4B16FFDAE7E8F8CCFDB0F8E4" blockId="82.[151,1436,1853,2023]" box="[199,635,1889,1915]" pageId="82" pageNumber="83">
|
||
<materialsCitation id="12E7B9264B16FFDAE7E8F8CCFDB3F8E4" ID-GBIF-Occurrence="4104078403" box="[199,632,1889,1915]" country="Indonesia" location="Off" pageId="82" pageNumber="83" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
|
||
<emphasis id="90FB6F694B16FFDAE7E8F8CCFEABF8E4" bold="true" box="[199,352,1889,1915]" pageId="82" pageNumber="83">
|
||
<typeStatus id="7D340DD94B16FFDAE7E8F8CCFEC9F8E4" box="[199,258,1889,1915]" pageId="82" pageNumber="83">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<location id="A750E5A04B16FFDAE641F8CCFE5FF8E4" LSID="urn:lsid:plazi:treatment:2A26026D4B16FFDCE7B8FDEBFE00FB76:A750E5A04B16FFDAE641F8CCFE5FF8E4" box="[366,404,1889,1915]" country="Indonesia" name="Off" pageId="82" pageNumber="83" stateProvince="Sumatera Utara">Off</location>
|
||
<collectingRegion id="604B7D994B16FFDAE6B4F8CCFE30F8E4" box="[411,507,1889,1915]" country="Indonesia" name="Sumatera Utara" pageId="82" pageNumber="83">Sumatra</collectingRegion>
|
||
,
|
||
<collectingCountry id="DA98F3EB4B16FFDAE528F8CCFDB3F8E4" box="[519,632,1889,1915]" name="Indonesia" pageId="82" pageNumber="83">Indonesia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EA95E0F04B16FFDAE7E8F828FD99F85B" pageId="82" pageNumber="83" type="distribution">
|
||
<paragraph id="A230B37B4B16FFDAE7E8F828FD99F85B" blockId="82.[151,1436,1853,2023]" pageId="82" pageNumber="83">
|
||
<emphasis id="90FB6F694B16FFDAE7E8F828FE92F800" bold="true" box="[199,345,1925,1951]" pageId="82" pageNumber="83">Distribution</emphasis>
|
||
. Indo-West Pacific: presently known only from the
|
||
<typeStatus id="7D340DD94B16FFDAE495F828FC21F800" box="[954,1002,1925,1951]" pageId="82" pageNumber="83">type</typeStatus>
|
||
locality in the Straits of
|
||
<collectingCountry id="DA98F3EB4B16FFDAE23CF828FAB9F800" box="[1299,1394,1925,1951]" name="Malaysia" pageId="82" pageNumber="83">Malacca</collectingCountry>
|
||
off
|
||
<collectingRegion id="604B7D994B16FFDAE7B8F804FF3CF85C" box="[151,247,1961,1987]" country="Indonesia" name="Sumatera Utara" pageId="82" pageNumber="83">Sumatra</collectingRegion>
|
||
, western
|
||
<collectingCountry id="DA98F3EB4B16FFDAE64EF804FE05F85C" box="[353,462,1961,1987]" name="Indonesia" pageId="82" pageNumber="83">Indonesia</collectingCountry>
|
||
(
|
||
<figureCitation id="3AB4AFFE4B16FFDAE6F2F804FD8DF85B" box="[477,582,1961,1988]" captionStart="FIGURE 52" captionStartId="106.[151,250,1909,1933]" captionTargetBox="[156,1430,181,1885]" captionTargetId="figure-21@106.[156,1430,181,1885]" captionTargetPageId="106" captionText="FIGURE 52. Distributional maps for species of Alpheus treated in this study:A—A. takla sp. nov.; B—A. mangalis sp. nov. and A. cf. mangalis sp. nov.; C—A. songkla Banner & Banner, 1966 and A. cf. songkla (see text); D—A. paludicola Kemp, 1915, A. nipa Banner & Banner, 1985 and A. bunburius Banner & Banner, 1982; E—A. pontederiae de Rochebrune, 1883; F—A. firmus Kim & Abele, 1988 and A. cf. firmus (see text)." figureDoi="http://doi.org/10.5281/zenodo.7912457" httpUri="https://zenodo.org/record/7912457/files/figure.png" pageId="82" pageNumber="83">Fig. 52D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EA95E0F04B16FFDAE7E8F860FCF4F878" box="[199,831,1997,2023]" pageId="82" pageNumber="83" type="vernacular_names">
|
||
<paragraph id="A230B37B4B16FFDAE7E8F860FCF4F878" blockId="82.[151,1436,1853,2023]" box="[199,831,1997,2023]" pageId="82" pageNumber="83">
|
||
<emphasis id="90FB6F694B16FFDAE7E8F860FE38F878" bold="true" box="[199,499,1997,2023]" pageId="82" pageNumber="83">Common name proposed</emphasis>
|
||
. Nipa palm snapping shrimp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F6F0E3F34B17FFDBE7B8FA5AFC0BF95C" ID-DOI="http://doi.org/10.5281/zenodo.7912433" ID-Zenodo-Dep="7912433" httpUri="https://zenodo.org/record/7912433/files/figure.png" pageId="83" pageNumber="84" startId="83.[151,250,1527,1551]" targetBox="[170,1409,197,1483]" targetPageId="83" targetType="figure">
|
||
<paragraph id="A230B37B4B17FFDBE7B8FA5AFC0BF95C" blockId="83.[151,1437,1527,1731]" pageId="83" pageNumber="84">
|
||
<emphasis id="90FB6F694B17FFDBE7B8FA5AFED1F990" bold="true" box="[151,282,1527,1551]" pageId="83" pageNumber="84">FIGURE 45</emphasis>
|
||
.
|
||
<taxonomicName id="658FC8F84B17FFDBE607FA5AFD66F990" authority="Banner & Banner, 1985" authorityName="Banner & Banner" authorityYear="1985" box="[296,685,1527,1551]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B17FFDBE607FA5AFE65F990" box="[296,430,1527,1551]" italics="true" pageId="83" pageNumber="84">Alpheus nipa</emphasis>
|
||
<bibRefCitation id="C61ECE8A4B17FFDBE699FA55FD66F990" author="Banner, D. M. & Banner, A. H." box="[438,685,1527,1551]" pageId="83" pageNumber="84" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
: paratype, male (cl indet., range: 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526); A—rostro-orbital region of carapace, dorsal view; B—frontal region, lateral view; C— ventromesial carina of first article of antennular peduncle, lateral view; D—antennal scaphocerite, dorsal view; E—third maxilliped, lateral view [arthrobranch not drawn]; F—same, detail of branchial structures on exopod, mesial view; G—major (right) cheliped, chela, mesial view; H—same, distal part of chela, fingers open, lateral view; I—second pleopod, appendices masculina and interna; J—right uropod, detail of distolateral angle, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="EA95E0F04B17FFDBE7E8F95FFE42F8E8" pageId="83" pageNumber="84" type="biology_ecology">
|
||
<paragraph id="A230B37B4B17FFDBE7E8F95FFE42F8E8" blockId="83.[151,1436,1777,2020]" pageId="83" pageNumber="84">
|
||
<emphasis id="90FB6F694B17FFDBE7E8F95FFE74F894" bold="true" box="[199,447,1777,1804]" pageId="83" pageNumber="84">Ecology and biology</emphasis>
|
||
. All known specimens of
|
||
<taxonomicName id="658FC8F84B17FFDBE5DBF95FFC82F894" authorityName="Banner & Banner" authorityYear="1985" box="[756,841,1778,1804]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B17FFDBE5DBF95FFC82F894" box="[756,841,1778,1804]" italics="true" pageId="83" pageNumber="84">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
were found on floating fronds of nipa palm (
|
||
<taxonomicName id="658FC8F84B17FFDBE24EF95FFEACF8B0" authority="Wurmb" authorityName="Wurmb" class="Liliopsida" family="Arecaceae" genus="Nypa" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="fructicans">
|
||
<emphasis id="90FB6F694B17FFDBE24EF95FFECCF8AF" italics="true" pageId="83" pageNumber="84">Nypa fructicans</emphasis>
|
||
Wurmb
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="658FC8F84B17FFDBE65DF8BBFE20F8AF" box="[370,491,1814,1840]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
|
||
), suggesting that they may inhabit estuarine muddy banks fringed by nipa palms, a habitat that is both difficult to access and dangerous to sample due to the presence of saltwater crocodiles [
|
||
<taxonomicName id="658FC8F84B17FFDBE233F894FEB4F8E8" authority="(Schneider)" baseAuthorityName="Schneider" class="Reptilia" family="Crocodylidae" genus="Crocodylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="83" pageNumber="84" phylum="Chordata" rank="species" species="porosus">
|
||
<emphasis id="90FB6F694B17FFDBE233F894FF24F8E8" italics="true" pageId="83" pageNumber="84">Crocodylus porosus</emphasis>
|
||
(Schneider)
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EA95E0F04B17FFDCE7E8F82CFE00FB76" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="discussion">
|
||
<paragraph id="A230B37B4B17FFDCE7E8F82CFB12FE16" blockId="83.[151,1436,1777,2020]" lastBlockId="84.[151,1437,150,1257]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
|
||
<emphasis id="90FB6F694B17FFDBE7E8F82CFE75F803" bold="true" box="[199,446,1921,1948]" pageId="83" pageNumber="84">Taxonomic remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="C61ECE8A4B17FFDBE6FCF82FFCCAF803" author="Banner, D. M. & Banner, A. H." box="[467,769,1921,1948]" pageId="83" pageNumber="84" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner (1985)</bibRefCitation>
|
||
provided a reasonably detailed description of
|
||
<taxonomicName id="658FC8F84B17FFDBE212F82FFA5EF804" authorityName="Banner & Banner" authorityYear="1985" box="[1341,1429,1922,1948]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B17FFDBE212F82FFA5EF804" box="[1341,1429,1922,1948]" italics="true" pageId="83" pageNumber="84">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
, accompanied by illustrations of all major diagnostic features. Two important features of
|
||
<taxonomicName id="658FC8F84B17FFDBE3B5F80BFB25F820" authorityName="Banner & Banner" authorityYear="1985" box="[1178,1262,1958,1984]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B17FFDBE3B5F80BFB25F820" box="[1178,1262,1958,1984]" italics="true" pageId="83" pageNumber="84">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
not mentioned nor illustrated by
|
||
<bibRefCitation id="C61ECE8A4B17FFDBE64BF867FD4AF87B" author="Banner, D. M. & Banner, A. H." box="[356,641,1993,2020]" pageId="83" pageNumber="84" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner (1985)</bibRefCitation>
|
||
are the unarmed antennal basicerite (
|
||
<figureCitation id="3AB4AFFE4B17FFDBE31FF864FB5DF87B" box="[1072,1174,1993,2020]" captionStart="FIGURE 45" captionStartId="83.[151,250,1527,1551]" captionTargetBox="[170,1409,197,1483]" captionTargetId="figure-15@83.[151,1436,181,1503]" captionTargetPageId="83" captionText="FIGURE 45. Alpheus nipa Banner & Banner, 1985: paratype, male (cl indet., range: 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526); A—rostro-orbital region of carapace, dorsal view; B—frontal region, lateral view; C— ventromesial carina of first article of antennular peduncle, lateral view; D—antennal scaphocerite, dorsal view; E—third maxilliped, lateral view [arthrobranch not drawn]; F—same, detail of branchial structures on exopod, mesial view; G—major (right) cheliped, chela, mesial view; H—same, distal part of chela, fingers open, lateral view; I—second pleopod, appendices masculina and interna; J—right uropod, detail of distolateral angle, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912433" httpUri="https://zenodo.org/record/7912433/files/figure.png" pageId="83" pageNumber="84">Fig. 45B</figureCitation>
|
||
) and the presence of a faint mesial subdistal ridge on the major cheliped pollex (
|
||
<figureCitation id="3AB4AFFE4B10FFDCE403FF3AFC55FF2E" box="[812,926,151,177]" captionStart="FIGURE 44" captionStartId="82.[151,250,1710,1734]" captionTargetBox="[151,1435,987,1687]" captionTargetId="figure-336@82.[151,1435,987,1687]" captionTargetPageId="82" captionText="FIGURE 44. Alpheus nipa Banner & Banner, 1985: paratypes, males (cl 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526), detached chelipeds; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (right) cheliped, lateral view; D—same, dorsolateral view; E—same, mesial view." figureDoi="http://doi.org/10.5281/zenodo.7912431" httpUri="https://zenodo.org/record/7912431/files/figure.png" pageId="84" pageNumber="85">Figs. 44B</figureCitation>
|
||
,
|
||
<figureCitation id="3AB4AFFE4B10FFDCE483FF3BFC2BFF2E" box="[940,992,150,177]" captionStart="FIGURE 45" captionStartId="83.[151,250,1527,1551]" captionTargetBox="[170,1409,197,1483]" captionTargetId="figure-15@83.[151,1436,181,1503]" captionTargetPageId="83" captionText="FIGURE 45. Alpheus nipa Banner & Banner, 1985: paratype, male (cl indet., range: 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526); A—rostro-orbital region of carapace, dorsal view; B—frontal region, lateral view; C— ventromesial carina of first article of antennular peduncle, lateral view; D—antennal scaphocerite, dorsal view; E—third maxilliped, lateral view [arthrobranch not drawn]; F—same, detail of branchial structures on exopod, mesial view; G—major (right) cheliped, chela, mesial view; H—same, distal part of chela, fingers open, lateral view; I—second pleopod, appendices masculina and interna; J—right uropod, detail of distolateral angle, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912433" httpUri="https://zenodo.org/record/7912433/files/figure.png" pageId="84" pageNumber="85">45G</figureCitation>
|
||
), which may link this species to the
|
||
<emphasis id="90FB6F694B10FFDCE2ABFF3AFE31FF4A" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="658FC8F84B10FFDCE2ABFF3AFEDCFF4A" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
|
||
—
|
||
<taxonomicName id="658FC8F84B10FFDCE61CFF16FE31FF4A" authorityName="sensu De Man" authorityYear="1897" box="[307,506,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">A. microrhynchus</taxonomicName>
|
||
</emphasis>
|
||
complex. Another previously unknown feature of
|
||
<taxonomicName id="658FC8F84B10FFDCE315FF16FB47FF4B" authorityName="Banner & Banner" authorityYear="1985" box="[1082,1164,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE315FF16FB47FF4B" box="[1082,1164,187,213]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
is the presence of three small, gill-like leaflets at the base of the third maxilliped exopod; this feature was observed on two dissected third maxillipeds and one maxilliped still
|
||
<emphasis id="90FB6F694B10FFDCE5ADFEAEFD00FE83" box="[642,715,259,284]" italics="true" pageId="84" pageNumber="85">in situ</emphasis>
|
||
. This gill-like structure does not appear to be homologous to the “rudimentary pleurobranch at the base of the third maxilliped” in
|
||
<taxonomicName id="658FC8F84B10FFDCE4ECFE8AFAA1FEDE" authority="(Christoffersen 1984)" baseAuthorityName="Christoffersen" baseAuthorityYear="1984" box="[963,1386,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="pontederiae">
|
||
<emphasis id="90FB6F694B10FFDCE4ECFE8AFBA0FEDE" box="[963,1131,295,321]" italics="true" pageId="84" pageNumber="85">A. pontederiae</emphasis>
|
||
(
|
||
<bibRefCitation id="C61ECE8A4B10FFDCE353FE8AFAAAFEDE" author="Christoffersen, M. L." box="[1148,1377,295,321]" pageId="84" pageNumber="85" pagination="189 - 208" refId="ref70049" refString="Christoffersen, M. L. (1984) The western Atlantic snapping shrimps related to Alpheus heterochaelis Say (Crustacea, Caridea) with the description of a new species. Papeis Avulsos de Zoologia, 35, 189 - 208." type="journal article" year="1984">Christoffersen 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and “une petite pleurobranchie de forme très spéciale sur le 3
|
||
<superScript id="55FA1E334B10FFDCE431FEE6FCEEFEC6" attach="left" box="[798,805,331,345]" fontSize="6" pageId="84" pageNumber="85">e</superScript>
|
||
mxp.” in several other species of
|
||
<taxonomicName id="658FC8F84B10FFDCE386FEE6FAC8FEFA" authorityName="Fabricius" authorityYear="1798" box="[1193,1283,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="90FB6F694B10FFDCE386FEE6FAC8FEFA" box="[1193,1283,331,357]" italics="true" pageId="84" pageNumber="85">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="658FC8F84B10FFDCE2ABFEE6FEDCFE16" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">
|
||
<emphasis id="90FB6F694B10FFDCE2ABFEE6FEDCFE16" italics="true" pageId="84" pageNumber="85">A. euphrosyne</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE60AFEC2FE21FE16" authorityName="sensu De Man" authorityYear="1897" box="[293,490,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="90FB6F694B10FFDCE60AFEC2FE21FE16" box="[293,490,367,393]" italics="true" pageId="84" pageNumber="85">A. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="658FC8F84B10FFDCE50EFEC2FD2FFE16" authorityName="Ortmann" authorityYear="1890" box="[545,740,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="macrodactylus">
|
||
<emphasis id="90FB6F694B10FFDCE50EFEC2FD2FFE16" box="[545,740,367,393]" italics="true" pageId="84" pageNumber="85">A. macrodactylus</emphasis>
|
||
</taxonomicName>
|
||
(see discussion in
|
||
<bibRefCitation id="C61ECE8A4B10FFDCE497FEC2FB06FE16" author="Coutiere, H." box="[952,1229,367,393]" pageId="84" pageNumber="85" pagination="1 - 559" refId="ref70196" refString="Coutiere, H. (1899) Les " Alpheidae ". Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, Serie 8, 9, 1 - 559, pls. 1 - 6. Masson, Paris. https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière 1899: 279–281</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A230B37B4B10FFDCE7E8FE3EFC41FCC2" blockId="84.[151,1437,150,1257]" pageId="84" pageNumber="85">
|
||
As noted by
|
||
<bibRefCitation id="C61ECE8A4B10FFDCE64AFE3EFD45FE32" author="Banner, D. M. & Banner, A. H." box="[357,654,402,429]" pageId="84" pageNumber="85" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner (1985)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE5B0FE3EFD3EFE33" authorityName="Banner & Banner" authorityYear="1985" box="[671,757,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE5B0FE3EFD3EFE33" box="[671,757,403,429]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="658FC8F84B10FFDCE33CFE3EFB73FE32" authorityName="sensu De Man" authorityYear="1897" box="[1043,1208,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">
|
||
<emphasis id="90FB6F694B10FFDCE33CFE3EFB73FE32" box="[1043,1208,403,429]" italics="true" pageId="84" pageNumber="85">A. euphrosyne</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE3E4FE3EFABAFE32" authorityName="Yaldwyn" authorityYear="1971" box="[1227,1393,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="richardsoni">
|
||
<emphasis id="90FB6F694B10FFDCE3E4FE3EFABAFE32" box="[1227,1393,403,429]" italics="true" pageId="84" pageNumber="85">A. richardsoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE2ABFE3EFEC4FE4E" authorityName="Kemp" authorityYear="1915" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="paludicola">
|
||
<emphasis id="90FB6F694B10FFDCE2ABFE3EFEC4FE4E" italics="true" pageId="84" pageNumber="85">A. paludicola</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="658FC8F84B10FFDCE666FE1AFDDBFE4E" authorityName="sensu De Man" authorityYear="1897" box="[329,528,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="90FB6F694B10FFDCE666FE1AFDDBFE4E" box="[329,528,439,465]" italics="true" pageId="84" pageNumber="85">A. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
by the dactyli of the third and fourth pereiopods being short and conical rather than spatulate (in the first three species) or trigonal-subspatulate (in the latter species). This distinguishing character is also valid for
|
||
<taxonomicName id="658FC8F84B10FFDCE662FE52FE30FD86" authorityName="De Man" authorityYear="1920" box="[333,507,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="eurydactylus">
|
||
<emphasis id="90FB6F694B10FFDCE662FE52FE30FD86" box="[333,507,511,537]" italics="true" pageId="84" pageNumber="85">A. eurydactylus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE525FE52FD43FD87" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[522,648,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
|
||
<emphasis id="90FB6F694B10FFDCE525FE52FD43FD87" box="[522,648,511,537]" italics="true" pageId="84" pageNumber="85">A. nomurai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90FB6F694B10FFDCE5A0FE52FD22FD86" bold="true" box="[655,745,511,537]" pageId="84" pageNumber="85">
|
||
<taxonomicNameLabel id="8BC8D2124B10FFDCE5A0FE52FD22FD86" box="[655,745,511,537]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE5D7FE52FCB5FD86" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[760,894,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
|
||
<emphasis id="90FB6F694B10FFDCE5D7FE52FCB5FD86" box="[760,894,511,537]" italics="true" pageId="84" pageNumber="85">A. mangalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90FB6F694B10FFDCE4AAFE52FC14FD86" bold="true" box="[901,991,511,537]" pageId="84" pageNumber="85">
|
||
<taxonomicNameLabel id="8BC8D2124B10FFDCE4AAFE52FC14FD86" box="[901,991,511,537]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE4C2FE52FB8EFD86" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1005,1093,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
|
||
<emphasis id="90FB6F694B10FFDCE4C2FE52FB8EFD86" box="[1005,1093,511,537]" italics="true" pageId="84" pageNumber="85">A. takla</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="90FB6F694B10FFDCE363FE52FB6DFD86" bold="true" box="[1100,1190,511,537]" pageId="84" pageNumber="85">
|
||
<taxonomicNameLabel id="8BC8D2124B10FFDCE363FE52FB6DFD86" box="[1100,1190,511,537]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="658FC8F84B10FFDCE3F2FE52FA98FD86" authorityName="sensu Banner & Banner" authorityYear="1966" baseAuthorityName="Banner & Banner" baseAuthorityYear="1966" box="[1245,1363,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="songkla">
|
||
<emphasis id="90FB6F694B10FFDCE3F2FE52FA98FD86" box="[1245,1363,511,537]" italics="true" pageId="84" pageNumber="85">A. songkla</emphasis>
|
||
</taxonomicName>
|
||
, all of which with spatulate dactyli on the ambulatory pereiopods. In addition,
|
||
<taxonomicName id="658FC8F84B10FFDCE4E7FD8EFBD1FDA3" authorityName="Banner & Banner" authorityYear="1985" box="[968,1050,547,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE4E7FD8EFBD1FDA3" box="[968,1050,547,573]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="658FC8F84B10FFDCE39EFD8EFAB3FDA2" authorityName="sensu De Man" authorityYear="1897" box="[1201,1400,547,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="90FB6F694B10FFDCE39EFD8EFAB3FDA2" box="[1201,1400,547,573]" italics="true" pageId="84" pageNumber="85">A. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
by the markedly different ratio of the first two carpal subarticles of the second pereiopod (1:
|
||
<quantity id="65771E9E4B10FFDCE3A0FDEAFB04FDFE" box="[1167,1231,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" pageId="84" pageNumber="85" unit="in" value="0.8">0.8 in</quantity>
|
||
<emphasis id="90FB6F694B10FFDCE3F9FDEAFA8DFDFF" box="[1238,1350,583,609]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="658FC8F84B10FFDCE3F9FDEAFAE3FDFF" authorityName="Banner & Banner" authorityYear="1985" box="[1238,1320,583,609]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">A. nipa</taxonomicName>
|
||
vs
|
||
</emphasis>
|
||
. 1:
|
||
<quantity id="65771E9E4B10FFDCE256FDEAFF66FD1A" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="84" pageNumber="85" unit="in" value="0.3">0.3 in</quantity>
|
||
<taxonomicName id="658FC8F84B10FFDCE799FDC6FEB7FD1A" authorityName="sensu De Man" authorityYear="1897" box="[182,380,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="90FB6F694B10FFDCE799FDC6FEB7FD1A" box="[182,380,619,645]" italics="true" pageId="84" pageNumber="85">A. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
); from both
|
||
<taxonomicName id="658FC8F84B10FFDCE53FFDC6FD1CFD1A" authorityName="sensu De Man" authorityYear="1897" box="[528,727,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="90FB6F694B10FFDCE53FFDC6FD1CFD1A" box="[528,727,619,645]" italics="true" pageId="84" pageNumber="85">A. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="658FC8F84B10FFDCE43EFDC6FC63FD1A" authorityName="Yeo & Ng" authorityYear="1996" box="[785,936,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="cyanoteles">
|
||
<emphasis id="90FB6F694B10FFDCE43EFDC6FC63FD1A" box="[785,936,619,645]" italics="true" pageId="84" pageNumber="85">A. cyanoteles</emphasis>
|
||
</taxonomicName>
|
||
by the much stronger sculpture of the male minor chela, with both dorsal and ventral transverse grooves well pronounced and bordered by shoulders (
|
||
<emphasis id="90FB6F694B10FFDCE21DFD22FA82FD37" box="[1330,1353,655,680]" italics="true" pageId="84" pageNumber="85">vs</emphasis>
|
||
. feebly delineated, if it all, and without shoulders, in
|
||
<taxonomicName id="658FC8F84B10FFDCE5A2FD1EFC9AFD52" authorityName="sensu De Man" authorityYear="1897" box="[653,849,691,717]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="90FB6F694B10FFDCE5A2FD1EFC9AFD52" box="[653,849,691,717]" italics="true" pageId="84" pageNumber="85">A. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="658FC8F84B10FFDCE4A9FD1EFBD0FD52" authorityName="Yeo & Ng" authorityYear="1996" box="[902,1051,691,717]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="cyanoteles">
|
||
<emphasis id="90FB6F694B10FFDCE4A9FD1EFBD0FD52" box="[902,1051,691,717]" italics="true" pageId="84" pageNumber="85">A. cyanoteles</emphasis>
|
||
</taxonomicName>
|
||
); and the scaphocerite blade much shorter than the distolateral tooth (
|
||
<emphasis id="90FB6F694B10FFDCE536FD7AFDFBFD6F" box="[537,560,727,752]" italics="true" pageId="84" pageNumber="85">vs</emphasis>
|
||
. reaching slightly or far beyond the distolateral tooth in
|
||
<taxonomicName id="658FC8F84B10FFDCE388FD7AFAA7FD6E" authorityName="sensu De Man" authorityYear="1897" box="[1191,1388,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="90FB6F694B10FFDCE388FD7AFAA7FD6E" box="[1191,1388,727,753]" italics="true" pageId="84" pageNumber="85">A. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="658FC8F84B10FFDCE7B8FD56FEE5FC8A" authorityName="Yeo & Ng" authorityYear="1996" box="[151,302,763,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="cyanoteles">
|
||
<emphasis id="90FB6F694B10FFDCE7B8FD56FEE5FC8A" box="[151,302,763,789]" italics="true" pageId="84" pageNumber="85">A. cyanoteles</emphasis>
|
||
</taxonomicName>
|
||
). Another diagnostic character of
|
||
<taxonomicName id="658FC8F84B10FFDCE599FD56FCC2FC8B" authorityName="Banner & Banner" authorityYear="1985" box="[694,777,763,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE599FD56FCC2FC8B" box="[694,777,763,789]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
is the presence of several long erect setae on the rostrum (
|
||
<figureCitation id="3AB4AFFE4B10FFDCE7B0FCB2FEEDFCA6" box="[159,294,798,825]" captionStart="FIGURE 45" captionStartId="83.[151,250,1527,1551]" captionTargetBox="[170,1409,197,1483]" captionTargetId="figure-15@83.[151,1436,181,1503]" captionTargetPageId="83" captionText="FIGURE 45. Alpheus nipa Banner & Banner, 1985: paratype, male (cl indet., range: 6.2–7.5 mm) from off Medan, Sumatra, Indonesia (ZMUC CRU-7526); A—rostro-orbital region of carapace, dorsal view; B—frontal region, lateral view; C— ventromesial carina of first article of antennular peduncle, lateral view; D—antennal scaphocerite, dorsal view; E—third maxilliped, lateral view [arthrobranch not drawn]; F—same, detail of branchial structures on exopod, mesial view; G—major (right) cheliped, chela, mesial view; H—same, distal part of chela, fingers open, lateral view; I—second pleopod, appendices masculina and interna; J—right uropod, detail of distolateral angle, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7912433" httpUri="https://zenodo.org/record/7912433/files/figure.png" pageId="84" pageNumber="85">Fig. 45A, B</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="C61ECE8A4B10FFDCE6A0FCB2FD5EFCA6" author="Banner, D. M. & Banner, A. H." box="[399,661,798,825]" pageId="84" pageNumber="85" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner 1985</bibRefCitation>
|
||
: fig. 2a, b); these setae are absent in the other species being dealt with in this study (or if present, not nearly as conspicuous as in
|
||
<taxonomicName id="658FC8F84B10FFDCE407FCEEFCB2FCC3" authorityName="Banner & Banner" authorityYear="1985" box="[808,889,835,861]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE407FCEEFCB2FCC3" box="[808,889,835,861]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A230B37B4B10FFDCE7E8FCCAFE00FB76" blockId="84.[151,1437,150,1257]" pageId="84" pageNumber="85">
|
||
<bibRefCitation id="C61ECE8A4B10FFDCE7E8FCCAFE17FC1E" author="Banner, D. M. & Banner, A. H." box="[199,476,870,897]" pageId="84" pageNumber="85" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. & Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's " The Decapoda of the Siboga Expedition, Part II. Family Alpheidae " (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner & Banner (1985)</bibRefCitation>
|
||
suggested that
|
||
<taxonomicName id="658FC8F84B10FFDCE5A7FCCAFD12FC1F" authorityName="Banner & Banner" authorityYear="1985" box="[648,729,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE5A7FCCAFD12FC1F" box="[648,729,871,897]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
“is derived from one of these euryhaline mud-or soft sedimentdwelling species, possibly from the most common and widespread
|
||
<emphasis id="90FB6F694B10FFDCE4B5FC26FB81FC3A" box="[922,1098,907,933]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="658FC8F84B10FFDCE4B5FC26FBF4FC3A" authorityName="sensu De Man" authorityYear="1897" box="[922,1087,907,933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
|
||
*
|
||
</emphasis>
|
||
and has left the mud and silt where the other three species [
|
||
<emphasis id="90FB6F694B10FFDCE52FFC02FD7AFC56" box="[512,689,943,969]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="658FC8F84B10FFDCE52FFC02FD6DFC56" authorityName="sensu De Man" authorityYear="1897" box="[512,678,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
|
||
*
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE5ECFC02FCA3FC56" authorityName="Yaldwyn" authorityYear="1971" box="[707,872,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="richardsoni">
|
||
<emphasis id="90FB6F694B10FFDCE5ECFC02FCA3FC56" box="[707,872,943,969]" italics="true" pageId="84" pageNumber="85">A. richardsoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="658FC8F84B10FFDCE456FC02FBDFFC56" authorityName="Kemp" authorityYear="1915" box="[889,1044,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="paludicola">
|
||
<emphasis id="90FB6F694B10FFDCE456FC02FBDFFC56" box="[889,1044,943,969]" italics="true" pageId="84" pageNumber="85">A. paludicola</emphasis>
|
||
</taxonomicName>
|
||
] dwell and has invaded a unique habitat, the nipa palms. In so doing, it has lost the spatulate dactyli on the walking legs, so necessary for digging in soft sediments” [*note:
|
||
<taxonomicName id="658FC8F84B10FFDCE6E6FC5AFDA0FB8E" authorityName="sensu De Man" authorityYear="1897" box="[457,619,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">
|
||
<emphasis id="90FB6F694B10FFDCE6E6FC5AFDA0FB8E" box="[457,619,1015,1041]" italics="true" pageId="84" pageNumber="85">A. euphrosyne</emphasis>
|
||
</taxonomicName>
|
||
not
|
||
<emphasis id="90FB6F694B10FFDCE58DFC5AFD2BFB8F" box="[674,736,1015,1040]" italics="true" pageId="84" pageNumber="85">sensu</emphasis>
|
||
<bibRefCitation id="C61ECE8A4B10FFDCE5C5FC5AFC6AFB8E" author="De Man, J. G." box="[746,929,1015,1041]" pageId="84" pageNumber="85" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man (1897)</bibRefCitation>
|
||
]. This is indeed an interesting hypothesis to be tested in the future, and to be contrasted to an alternative hypothesis of
|
||
<taxonomicName id="658FC8F84B10FFDCE4CAFBB6FBFCFBAB" authorityName="Banner & Banner" authorityYear="1985" box="[997,1079,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE4CAFBB6FBFCFBAB" box="[997,1079,1051,1077]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
not being closely related to the
|
||
<emphasis id="90FB6F694B10FFDCE7B8FB92FDD2FBC6" box="[151,537,1087,1113]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="658FC8F84B10FFDCE7B8FB92FEF3FBC6" authorityName="sensu De Man" authorityYear="1897" box="[151,312,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
|
||
—
|
||
<taxonomicName id="658FC8F84B10FFDCE67AFB92FDD2FBC6" authorityName="sensu De Man" authorityYear="1897" box="[341,537,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">A. microrhynchus</taxonomicName>
|
||
</emphasis>
|
||
complex and being derived, for instance, from the morphologically similar
|
||
<taxonomicName id="658FC8F84B10FFDCE2ABFB92FF3DFBE2" authorityName="De Haan" authorityYear="1849" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="90FB6F694B10FFDCE2ABFB92FF3DFBE2" italics="true" pageId="84" pageNumber="85">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex, in which the walking leg dactyli are also conical. Whatever the case may be, the origin and phylogenetic affinites of
|
||
<taxonomicName id="658FC8F84B10FFDCE697FB2AFDC7FB3F" authorityName="Banner & Banner" authorityYear="1985" box="[440,524,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="nipa">
|
||
<emphasis id="90FB6F694B10FFDCE697FB2AFDC7FB3F" box="[440,524,1159,1185]" italics="true" pageId="84" pageNumber="85">A. nipa</emphasis>
|
||
</taxonomicName>
|
||
are difficult to ascertain without a comprehensive phylogenetic analysis of the entire
|
||
<taxonomicName id="658FC8F84B10FFDCE7F1FB06FEA1FB5A" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[222,362,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="edwardsii">
|
||
<emphasis id="90FB6F694B10FFDCE7F1FB06FEA1FB5A" box="[222,362,1195,1221]" italics="true" pageId="84" pageNumber="85">A. edwardsii</emphasis>
|
||
</taxonomicName>
|
||
group, with inclusion of as many members as possible of the
|
||
<emphasis id="90FB6F694B10FFDCE330FB06FA57FB5A" box="[1055,1436,1195,1221]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="658FC8F84B10FFDCE330FB06FB76FB5A" authorityName="sensu De Man" authorityYear="1897" box="[1055,1213,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
|
||
—
|
||
<taxonomicName id="658FC8F84B10FFDCE3F5FB06FA57FB5A" authorityName="sensu De Man" authorityYear="1897" box="[1242,1436,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microrhynchus">A. microrhynchus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="658FC8F84B10FFDCE7E8FB62FE8EFB76" authorityName="De Haan" authorityYear="1849" box="[199,325,1231,1257]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="90FB6F694B10FFDCE7E8FB62FE8EFB76" box="[199,325,1231,1257]" italics="true" pageId="84" pageNumber="85">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complexes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |