153 lines
19 KiB
XML
153 lines
19 KiB
XML
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<mods:title id="BA289772681D83370A805411AEE36793">Cave millipeds of the United States. V. The genus Idagona Buckett & Gardner (Chordeumatida, Conotylidae, Idagoninae)</mods:title>
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<mods:namePart id="16614FB715DA3D45B3BCBCDE93CFC7EF">Shear, William A.</mods:namePart>
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<mods:date id="D51131E28FD71344CCF341339776A4C6">2007</mods:date>
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<treatment id="0393879DFFFD962BFF7C559CFE71C385" ID-DOI="http://doi.org/10.5281/zenodo.6236699" ID-GBIF-Taxon="119358473" ID-Zenodo-Dep="6236699" LSID="urn:lsid:plazi:treatment:0393879DFFFD962BFF7C559CFE71C385" httpUri="http://treatment.plazi.org/id/0393879DFFFD962BFF7C559CFE71C385" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<subSubSection id="C3206500FFFD9628FF7C559CFD9EC2C1" box="[151,594,1232,1258]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8B85368BFFFD9628FF7C559CFD9EC2C1" blockId="1.[151,594,1232,1258]" box="[151,594,1232,1258]" pageId="1" pageNumber="2">
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<heading id="D0CD81E7FFFD9628FF7C559CFD9EC2C1" bold="true" box="[151,594,1232,1258]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<taxonomicName id="4C3A4D08FFFD9628FF7C559CFD9EC2C1" ID-CoL="7P6YB" authority="Buckett and Gardner 1967" authorityName="Buckett and Gardner" authorityYear="1967" box="[151,594,1232,1258]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B94EEA99FFFD9628FF7C559CFD9EC2C1" bold="true" box="[151,594,1232,1258]" pageId="1" pageNumber="2">
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<emphasis id="B94EEA99FFFD9628FF7C559CFF30C2C1" bold="true" box="[151,252,1232,1258]" italics="true" pageId="1" pageNumber="2">Idagona</emphasis>
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<bibRefCitation id="EFAB4B7AFFFD9628FEE8559CFD9EC2C1" author="Buckett" box="[259,594,1232,1258]" pageId="1" pageNumber="12" refString="Buckett, J. S. & Garner, M. R. (1967) A new family of cavernicolous millipedes with description of a new genus and species from Idaho (Diplopoda: Chordeumida: Chordeumidea). The Michigan Entomologist, 1, 117 - 126." type="journal article" year="1967">Buckett and Gardner 1967</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3206500FFFD9628FF7C545CFC86C30C" box="[151,842,1296,1319]" pageId="1" pageNumber="2" type="reference_group">
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<paragraph id="8B85368BFFFD9628FF7C545CFC86C30C" blockId="1.[151,842,1296,1319]" box="[151,842,1296,1319]" pageId="1" pageNumber="2">
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<taxonomicName id="4C3A4D08FFFD9628FF7C545CFDDFC30C" ID-CoL="7P6YB" authority="Buckett and Gardner 1967" authorityName="Buckett and Gardner" authorityYear="1967" box="[151,531,1296,1319]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B94EEA99FFFD9628FF7C545CFF3CC30C" box="[151,240,1296,1319]" italics="true" pageId="1" pageNumber="2">Idagona</emphasis>
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<bibRefCitation id="EFAB4B7AFFFD9628FF1D545CFDDFC30C" author="Buckett" box="[246,531,1296,1319]" pageId="1" pageNumber="12" refString="Buckett, J. S. & Garner, M. R. (1967) A new family of cavernicolous millipedes with description of a new genus and species from Idaho (Diplopoda: Chordeumida: Chordeumidea). The Michigan Entomologist, 1, 117 - 126." type="journal article" year="1967">Buckett and Gardner 1967</bibRefCitation>
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</taxonomicName>
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, p. 120.
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<bibRefCitation id="EFAB4B7AFFFD9628FD9F545CFD39C30C" author="Shear" box="[628,757,1296,1319]" pageId="1" pageNumber="12" refString="Shear, W. A. (1972) Studies in the milliped order Chordeumida (Diplopoda): a revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology 144, 151 - 352." type="journal article" year="1972">Shear, 1972</bibRefCitation>
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, p. 270.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3206500FFFD962BFF7C5414FE71C385" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="diagnosis">
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<paragraph id="8B85368BFFFD9628FF7C5414FE1AC039" blockId="1.[151,1437,1368,1874]" pageId="1" pageNumber="2">
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<emphasis id="B94EEA99FFFD9628FF7C5414FEC2C35A" box="[151,270,1368,1393]" italics="true" pageId="1" pageNumber="2">Diagnosis</emphasis>
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: Distinct from all other North American conotylid millipeds in the complete loss of the telopodite articles of the posterior gonopods and the reduction of those appendages to a small pair of coxites which pass anteriorly between the anterior gonopods (
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<figureCitation id="13012A0EFFFD9628FD6554E4FD11C3E9" box="[654,733,1448,1474]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
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-8, 19, 20). Additionally, the third legpair femora in males are inflated and bear an adenostyle (
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<figureCitation id="13012A0EFFFD9628FDD5549CFD28C3C1" box="[574,740,1488,1514]" captionStart="FIGURES 9 – 14" captionStartId="6.[151,269,1722,1746]" captionTargetBox="[151,1436,357,1706]" captionTargetId="figure@6.[151,1436,357,1707]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Pregonopodal leg modifications of Idagona species. 9 – 12, third legs, anterior views. 9. Idagona westcotti, Giant Arch Cave. 10.. I. westcotti, Ice Capades Cave. 11. I. jasperi n. sp. 12. I. lehmanensis n. sp. 13. Leg 4 of I. lehmanensis n. sp., anterior view. 14. Denticles near adenostyle of leg 3 femur of I. lehmanensis n. sp." httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="1" pageNumber="2">Figs. 9-12, 14</figureCitation>
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); the tenth coxae in males are enlarged and lobed; the eleventh coxae are unmodified.
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</paragraph>
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<paragraph id="8B85368BFFFD9628FF2D576CFDC4C1DF" blockId="1.[151,1437,1368,1874]" lastBlockId="1.[151,1415,1951,2036]" pageId="1" pageNumber="2">
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<emphasis id="B94EEA99FFFD9628FF2D576CFEDFC012" box="[198,275,1568,1593]" italics="true" pageId="1" pageNumber="2">Notes:</emphasis>
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The species of
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<taxonomicName id="4C3A4D08FFFD9628FE3E576CFDFBC012" box="[469,567,1568,1593]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B94EEA99FFFD9628FE3E576CFDFBC012" box="[469,567,1568,1593]" italics="true" pageId="1" pageNumber="2">Idagona</emphasis>
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</taxonomicName>
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have been recorded from five discrete areas (Map 1). The new species
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<taxonomicName id="4C3A4D08FFFD9628FA60576CFEE4C04A" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="lehmanensis">
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<emphasis id="B94EEA99FFFD9628FA60576CFEE4C04A" italics="true" pageId="1" pageNumber="2">I. lehmanensis</emphasis>
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</taxonomicName>
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occurs in southeastern Nevada, far separated from the other localities; this species is highly distinctive: the fourth as well as the third prefemora are modified in males, and the gonopods differ in their much greater complexity and the proportions of the several elements.
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<taxonomicName id="4C3A4D08FFFD9628FC7157D4FBD0C09A" box="[922,1052,1688,1713]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="westcotti">
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<emphasis id="B94EEA99FFFD9628FC7157D4FBD0C09A" box="[922,1052,1688,1713]" italics="true" pageId="1" pageNumber="2">I. westcotti</emphasis>
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</taxonomicName>
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is known from three clusters of lava tubes in southern Idaho, and
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<taxonomicName id="4C3A4D08FFFD9628FDDD578CFD68C0F2" box="[566,676,1728,1753]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="jasperi">
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<emphasis id="B94EEA99FFFD9628FDDD578CFD68C0F2" box="[566,676,1728,1753]" italics="true" pageId="1" pageNumber="2">I. jasperi</emphasis>
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</taxonomicName>
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is from two high-altitude caves in Logan Canyon, Cache Co., Utah, not far over the border between Idaho and Utah. Accordingly,
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<taxonomicName id="4C3A4D08FFFD9628FC2B57A4FBDDC12A" box="[960,1041,1768,1793]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="jasperi">
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<emphasis id="B94EEA99FFFD9628FC2B57A4FBDDC12A" box="[960,1041,1768,1793]" italics="true" pageId="1" pageNumber="2">jasperi</emphasis>
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</taxonomicName>
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more closely resembles
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<taxonomicName id="4C3A4D08FFFD9628FADF57A4FA50C12A" box="[1332,1436,1768,1793]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="westcotti">
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<emphasis id="B94EEA99FFFD9628FADF57A4FA50C12A" box="[1332,1436,1768,1793]" italics="true" pageId="1" pageNumber="2">westcotti</emphasis>
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</taxonomicName>
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and can be separated from that species only by the details of gonopod shape (
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<figureCitation id="13012A0EFFFD9628FBCC565CFBB6C101" box="[1063,1146,1808,1834]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="1" pageNumber="2">Figs. 3</figureCitation>
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,
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<date id="FF84104BFFFD9628FB63565CFB3FC101" box="[1160,1267,1808,1834]" pageId="1" pageNumber="2" value="1920-07-19">7. 19. 20</date>
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). The distinct habitat and locality reinforces the hypothesis that
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<taxonomicName id="4C3A4D08FFFD9628FD185674FC89C17A" box="[755,837,1848,1873]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="jasperi">
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<emphasis id="B94EEA99FFFD9628FD185674FC89C17A" box="[755,837,1848,1873]" italics="true" pageId="1" pageNumber="2">jasperi</emphasis>
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</taxonomicName>
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is a reproductively isolated population, while the 1. In point of fact, it is the reduced coxites of the posterior gonopods that are extended
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<emphasis id="B94EEA99FFFD9628FC1F56D3FB98C19F" box="[1012,1108,1951,1972]" italics="true" pageId="1" pageNumber="2">anteriorly</emphasis>
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between the anterior gonopods, but this is an homologous arrangement necessitated by the extreme reduction of the posterior gonopods, which in
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<taxonomicName id="4C3A4D08FFFD9628FADD56F3FA4AC1FF" box="[1334,1414,1983,2004]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B94EEA99FFFD9628FADD56F3FA4AC1FF" box="[1334,1414,1983,2004]" italics="true" pageId="1" pageNumber="2">Idagona</emphasis>
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</taxonomicName>
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consist only of the small coxites.
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</paragraph>
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<paragraph id="8B85368BFFFE962BFF7C51D4FE36C6F1" blockId="2.[151,1436,152,418]" pageId="2" pageNumber="3">
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three clusters of
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<taxonomicName id="4C3A4D08FFFE962BFE8A51D4FE05C69A" box="[353,457,152,177]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="westcotti">
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<emphasis id="B94EEA99FFFE962BFE8A51D4FE05C69A" box="[353,457,152,177]" italics="true" pageId="2" pageNumber="3">westcotti</emphasis>
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</taxonomicName>
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localities harbor populations not distinguishable by gonopod or other differences (
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<figureCitation id="13012A0EFFFE962BFF74518CFEDCC6F1" box="[159,272,192,218]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 2</figureCitation>
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, 5, 6,
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<date id="FF84104BFFFE962BFEBC518CFE08C6F1" box="[343,452,192,218]" pageId="2" pageNumber="3" value="1910-09-15">9. 10, 15.</date>
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16).
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</paragraph>
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<paragraph id="8B85368BFFFE962BFF2D51A4FD58C779" blockId="2.[151,1436,152,418]" pageId="2" pageNumber="3">For conservation reasons, and at the request of the collectors, the exact locations of caves mentioned in this paper are not given. Map 1 represents geometrically average locations for clusters of caves, or the approximate location of an individual cave.</paragraph>
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<paragraph id="8B85368BFFFE962BFF2D502CFDC7C789" blockId="2.[151,1436,152,418]" pageId="2" pageNumber="3">
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The
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<typeStatus id="54818829FFFE962BFF10502CFEF5C751" box="[251,313,352,378]" pageId="2" pageNumber="3">types</typeStatus>
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of the new species described below are deposited in the collections of the Field Museum of Natural History (FMNH), Chicago.
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</paragraph>
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<paragraph id="8B85368BFFFE962BFF7C5440FE71C385" blockId="2.[151,1437,1292,1454]" pageId="2" pageNumber="3">
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<emphasis id="B94EEA99FFFE962BFF7C5440FF22C30F" bold="true" box="[151,238,1292,1316]" pageId="2" pageNumber="3">MAP 1.</emphasis>
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Central Rocky Mountain and Great Basin region of the
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<collectingCountry id="F32D761BFFFE962BFCA25440FC16C308" box="[841,986,1292,1315]" name="United States of America" pageId="2" pageNumber="3">United States</collectingCountry>
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, showing distribution of
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<taxonomicName id="4C3A4D08FFFE962BFB025440FA8EC308" box="[1257,1346,1292,1315]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B94EEA99FFFE962BFB025440FA8EC308" box="[1257,1346,1292,1315]" italics="true" pageId="2" pageNumber="3">Idagona</emphasis>
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</taxonomicName>
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species. Shaded areas are karst or pseudokarst (lava beds). Triangles,
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<emphasis id="B94EEA99FFFE962BFCF45463FC50C36D" box="[799,924,1327,1350]" italics="true" pageId="2" pageNumber="3">
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<taxonomicName id="4C3A4D08FFFE962BFCF45463FC5AC36D" box="[799,918,1327,1350]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="westcotti">I. westcotti</taxonomicName>
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;
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</emphasis>
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square,
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<emphasis id="B94EEA99FFFE962BFC1D5463FB93C36D" box="[1014,1119,1327,1350]" italics="true" pageId="2" pageNumber="3">
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<taxonomicName id="4C3A4D08FFFE962BFC1D5463FB94C36D" box="[1014,1112,1327,1350]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="jasperi">I. jasperi</taxonomicName>
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;
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</emphasis>
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circle,
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<emphasis id="B94EEA99FFFE962BFB465463FA82C36D" box="[1197,1358,1327,1350]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C3A4D08FFFE962BFB465463FA86C36D" box="[1197,1354,1327,1350]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lehmanensis">I. lehmanensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Triangle 1, general location of Blowhole-Sand Lake, Ice Capades, and London Tunnel Caves; triangle 2, general location of Government Cave; triangle 3, general location of Giant Arch and Pot O’ Gold Caves (may be two names for the same cave). Map by S. J. Taylor.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |