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<mods:title id="478EC510A1E3815FA16E0E981FA3A79B">A hypothetical evolutionary history of passalid beetles narrated by the comparative anatomy of the hindgut (Coleoptera: Passalidae)</mods:title>
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<mods:namePart id="B55E35F0EE3F0335141F5D9E3FABAEF5">Fonseca, Claudio Ruy Vasconcelos Da</mods:namePart>
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<mods:namePart id="1A212C4971DA07018BB1CEB93945E95B">Barbosa, Márcio Luís Leitão</mods:namePart>
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Subfamily
<taxonomicName id="A68B729CFFF4FF9F6743CF22FDADFAF2" ID-CoL="8RYRG" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" box="[287,575,1309,1335]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Passalinae">
Passalinae
<bibRefCitation id="051A74EEFFF4FF9F67F4CF22FDADFAF2" author="Leach" box="[424,575,1309,1335]" pageId="13" pageNumber="14" refString="Leach, W. E. (1815) Entomology. In: Brewster, D. (Ed). Brewster's Edinburgh Encyclopedia. Volume IX [part I]. W. Blackwood, J. Waugh, etc., Edinburgh, pp. 57 - 172." type="book chapter" year="1815">Leach, 1815</bibRefCitation>
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(subfamily here redefined to include a smaller group of taxa)
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<paragraph id="6134091FFFF4FF9F669BCF7FFD6EFA9D" blockId="13.[151,1437,1309,1512]" box="[199,764,1344,1368]" pageId="13" pageNumber="14">This branch was supported by one synapomorphy:</paragraph>
<paragraph id="6134091FFFF4FF9F6699CF5BFEE6FA65" blockId="13.[151,1437,1309,1512]" pageId="13" pageNumber="14">[Character 17] Ileum with symmetrical, lateral diverticula (0); asymmetrical (1). Unambiguous optimization (ci = 100; ri = 100).</paragraph>
<paragraph id="6134091FFFF4FF9F6699CF92FC69FA2D" blockId="13.[151,1437,1309,1512]" pageId="13" pageNumber="14">
At this level, lateral diverticula were asymmetrical in numbers and size on both surfaces of the ileum, giving it a more flattened appearance (
<figureCitation id="F9B0159AFFF4FF9F67BCCFEFFDBBFA2D" box="[480,553,1488,1512]" captionStart="FIGURE 6" captionStartId="9.[151,250,1347,1370]" captionTargetBox="[159,1432,198,1314]" captionTargetId="figure@9.[151,1436,193,1326]" captionTargetPageId="9" captionText="FIGURE 6. Passalinae hindgut. A— Ptichopus angulatus (Percheron), Costa Rica; B— Passalus interruptus (Linnaeus), Brazil; C— Spasalus crenatus (MacLeay), Puerto Rico; D— Paxillus leachi MacLeay, Brazil; E— Passipassalus buhrnheimi Fonseca &amp; Reyes-Castillo, Brazil (Fig. E from Fonseca &amp; Reyes Castillo 1993)." httpUri="https://zenodo.org/record/200840/files/figure.png" pageId="13" pageNumber="14">Figs 6</figureCitation>
,
<figureCitation id="F9B0159AFFF4FF9F6469CFEFFDD7FA2D" box="[565,581,1488,1512]" captionStart="FIGURE 7" captionStartId="10.[151,250,1899,1922]" captionTargetBox="[202,1411,197,1885]" captionTargetId="figure@10.[199,1420,193,1891]" captionTargetPageId="10" captionText="FIGURE 7. Macrolininae hindgut. A— Pleurarius pilipes Kaup, India; B— Ophrygonius inaequalis (Burmeister), India; C— Episphenus moorei Kaup, India; D— Aceraius grandis (Burmeister), Taiwan; E— Macrolinus latipennis (Percheron), Java; F— Pseudepisphenus perplexus Gravely, New Guinea; G— Protomocoelus australis (Boisduval), Solomon Islands; H— Labienus ptox (Kaup), New Guinea; I— Analaches puberilis Kuwert, New Guinea." httpUri="https://zenodo.org/record/200841/files/figure.png" pageId="13" pageNumber="14">7</figureCitation>
). This character state was apomorphic.
</paragraph>
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