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<mods:title id="EEA335C0F43508B7FEC56CC122979C30">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<taxonomicName id="9BEF3AD6FFDDFFF58C92FB2CFDE8049B" ID-CoL="7SXBJ" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[151,609,1153,1180]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="paraleucozona" status="comb. nov.">
<emphasis id="6E9B9D47FFDDFFF58C92FB2CFE1C049C" bold="true" box="[151,405,1153,1180]" italics="true" pageId="40" pageNumber="41">Pusia paraleucozona</emphasis>
(
<bibRefCitation id="387E3CA4FFDDFFF58DA1FB2FFDD0049B" author="Boettger, O." box="[420,601,1153,1179]" pageId="40" pageNumber="41" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
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<taxonomicNameLabel id="75A8203CFFDDFFF58E6CFB2CFD66049C" box="[617,751,1154,1180]" pageId="40" pageNumber="41" rank="species">nov. comb.</taxonomicNameLabel>
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<subSubSection id="14F512DEFFDDFFF58C92FB08FE0A04C6" box="[151,387,1190,1222]" pageId="40" pageNumber="41" type="description">
<paragraph id="5C504155FFDDFFF58C92FB08FE0A04C6" blockId="40.[151,751,1153,1222]" box="[151,387,1190,1222]" pageId="40" pageNumber="41">
<figureCitation id="C4D45DD0FFDDFFF58C92FB08FF7704C0" box="[151,254,1190,1216]" captionStart="FIGURE 14" captionStartId="41.[151,250,1546,1570]" captionTargetBox="[151,1436,561,1522]" captionTargetId="figure-182@41.[151,1436,561,1522]" captionTargetPageId="41" captionText="FIGURE 14 A1A2. Pusia paraleucozona (Boettger, 1906), Drnovice u Vyškova (Czech Republic), NHMW 1865/0036/0010. B1B2. Pusia paraleucozona (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360345. C1C2. Pusia szobbiensis (Halaváts, 1884), lectotype, Inv. nr. M107, Szob (Hungary). D. Vexillum szobbiensis (Halaváts, 1884), paralectotype, Inv. nr. M107, Szob (Hungary). E1E2. Vexillum pseudoschafferi Biskupič, 2020, holotype, Z 40033, RohožníkKonopiská (Slovakia). E1E2. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40034, RohožníkKonopiská (Slovakia). G. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40035, RohožníkKonopiská (Slovakia)." figureDoi="http://doi.org/10.5281/zenodo.4921870" httpUri="https://zenodo.org/record/4921870/files/figure.png" pageId="40" pageNumber="41">Figs 14A</figureCitation>
<figureCitation id="C4D45DD0FFDDFFF58CFAFB16FEA004C0" box="[255,297,1190,1222]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="40" pageNumber="41">
<subScript id="C06B4310FFDDFFF58CFAFB16FE8E04C6" attach="right" box="[255,263,1208,1222]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
A
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<figureCitation id="C4D45DD0FFDDFFF58D2CFB16FEB804C6" box="[297,305,1208,1222]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="40" pageNumber="41">
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, B
<figureCitation id="C4D45DD0FFDDFFF58D57FB16FEF304C0" box="[338,378,1190,1222]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="40" pageNumber="41">
<subScript id="C06B4310FFDDFFF58D57FB16FED304C6" attach="left" box="[338,346,1208,1222]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
B
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<figureCitation id="C4D45DD0FFDDFFF58D7EFB16FE0A04C6" box="[379,387,1208,1222]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="40" pageNumber="41">
<subScript id="C06B4310FFDDFFF58D7EFB16FE0A04C6" attach="right" box="[379,387,1208,1222]" fontSize="6" pageId="40" pageNumber="41">2</subScript>
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<emphasis id="6E9B9D47FFDDFFF58C92FB45FF2D0502" box="[151,164,1259,1282]" italics="true" pageId="40" pageNumber="41">*</emphasis>
<taxonomicName id="9BEF3AD6FFDDFFF58CAFFB45FD250502" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[170,684,1258,1282]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="variety" species="ebenus" status="var. nov." subGenus="Uromitra" variety="paraleucozona">
<emphasis id="6E9B9D47FFDDFFF58CAFFB45FF6D0502" box="[170,228,1259,1282]" italics="true" pageId="40" pageNumber="41">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFDDFFF58CF6FB45FEDB0502" box="[243,338,1259,1282]" italics="true" pageId="40" pageNumber="41">Uromitra</emphasis>
)
<emphasis id="6E9B9D47FFDDFFF58D65FB44FE2F0502" box="[352,422,1258,1282]" italics="true" pageId="40" pageNumber="41">ebenus</emphasis>
Lmk. var.
<emphasis id="6E9B9D47FFDDFFF58E10FB44FD250502" box="[533,684,1258,1282]" italics="true" pageId="40" pageNumber="41">paraleucozona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFDDFFF58EB7FB45FD790503" box="[690,752,1259,1283]" pageId="40" pageNumber="41" rank="variety">n. var.</taxonomicNameLabel>
<treatmentCitation id="DD4E6744FFDDFFF58F0CFB45FC3E0502" author="Boettger, O." box="[777,951,1258,1283]" page="8" pageId="40" pageNumber="41" year="1906">
<bibRefCitation id="387E3CA4FFDDFFF58F0CFB45FC3E0502" author="Boettger, O." box="[777,951,1258,1283]" pageId="40" pageNumber="41" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger 1906: 8</bibRefCitation>
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.
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</paragraph>
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<taxonomicName id="9BEF3AD6FFDDFFF58C92FAA7FD320521" authority="(Boettger)" baseAuthorityName="Boettger" box="[151,699,1289,1313]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="subSpecies" species="ebenus" subGenus="Uromitra" subSpecies="paraleucozona">
<emphasis id="6E9B9D47FFDDFFF58C92FAA7FF670521" box="[151,238,1289,1313]" italics="true" pageId="40" pageNumber="41">Vexillum</emphasis>
(
<emphasis id="6E9B9D47FFDDFFF58CF8FAA4FED50521" box="[253,348,1290,1313]" italics="true" pageId="40" pageNumber="41">Uromitra</emphasis>
)
<emphasis id="6E9B9D47FFDDFFF58D6EFAA7FDC70521" box="[363,590,1289,1313]" italics="true" pageId="40" pageNumber="41">ebenus paraleucozona</emphasis>
(Boettger)
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<treatmentCitation id="DD4E6744FFDDFFF58ED0FAA7FCF30521" author="Zilch, A." box="[725,890,1289,1313]" page="259" pageId="40" pageNumber="41" year="1934">
<bibRefCitation id="387E3CA4FFDDFFF58ED0FAA7FCF30521" author="Zilch, A." box="[725,890,1289,1313]" pageId="40" pageNumber="41" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch 1934: 259</bibRefCitation>
</treatmentCitation>
, pl. 17, fig. 18.
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</paragraph>
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<taxonomicName id="9BEF3AD6FFDDFFF58C92FA86FD2D053F" authority="(Boettger, 1906)" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[151,676,1319,1344]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="paraleucozona" subGenus="Vexillum">
<emphasis id="6E9B9D47FFDDFFF58C92FA86FF670540" box="[151,238,1320,1344]" italics="true" pageId="40" pageNumber="41">Vexillum</emphasis>
(
<emphasis id="6E9B9D47FFDDFFF58CF8FA86FEDD0540" box="[253,340,1320,1344]" italics="true" pageId="40" pageNumber="41">Vexillum</emphasis>
)
<emphasis id="6E9B9D47FFDDFFF58D66FA86FE730540" box="[355,506,1320,1344]" italics="true" pageId="40" pageNumber="41">paraleucozona</emphasis>
(
<bibRefCitation id="387E3CA4FFDDFFF58E02FA86FD15053F" author="Boettger, O." box="[519,668,1319,1344]" pageId="40" pageNumber="41" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
<treatmentCitation id="DD4E6744FFDDFFF58EBBFA89FCD7053F" author="Baluk, W." box="[702,862,1319,1344]" page="35" pageId="40" pageNumber="41" year="1997">
<bibRefCitation id="387E3CA4FFDDFFF58EBBFA89FCD7053F" author="Baluk, W." box="[702,862,1319,1344]" pageId="40" pageNumber="41" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997: 35</bibRefCitation>
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, pl. 9, fig. 6.
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</paragraph>
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<emphasis id="6E9B9D47FFDDFFF58C92FAC0FECC0587" bold="true" box="[151,325,1389,1416]" pageId="40" pageNumber="41">Type material.</emphasis>
<materialsCitation id="EC874B08FFDDFFF58D4EFAC0FADB05B2" ID-GBIF-Occurrence="3311814350" collectionCode="SMF" collectorName="de Sus" country="Germany" location="Costeiu de Sus" pageId="40" pageNumber="41" specimenCode="SMF 360345, SMF 12.2208" specimenCount="1" typeStatus="lectotype">
<typeStatus id="8354FFF7FFDDFFF58D4EFAC0FE360588" box="[331,447,1390,1416]" pageId="40" pageNumber="41" type="lectotype">Lectotype</typeStatus>
:
<specimenCode id="0C49E92EFFDDFFF58DCCFAC0FDD50589" box="[457,604,1390,1417]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="40" pageNumber="41">SMF 360345</specimenCode>
(=
<specimenCode id="0C49E92EFFDDFFF58E87FAC0FCA90588" box="[642,800,1390,1416]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="40" pageNumber="41">SMF 12.2208</specimenCode>
a), Senckenberg Museum, Frankfurt/Main,
<collectingCountry id="24F801C5FFDDFFF58905FAC0FAE00588" box="[1280,1385,1390,1416]" name="Germany" pageId="40" pageNumber="41">Germany</collectingCountry>
, SL:
<quantity id="9B17ECB0FFDDFFF58C92FA3CFF7405AC" box="[151,253,1426,1453]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="40" pageNumber="41" unit="mm" value="11.5">11.5 mm</quantity>
, MD: 5.0 mm,
<location id="5930178EFFDDFFF58DA8FA3CFDDC05AC" LSID="urn:lsid:plazi:treatment:D446F043FFDDFFF48C92FB2CFE46021A:5930178EFFDDFFF58DA8FA3CFDDC05AC" box="[429,597,1426,1452]" country="Germany" name="Costeiu de Sus" pageId="40" pageNumber="41">
Coşteiu
<collectorName id="F11A2483FFDDFFF58E0FFA3CFDDC05AC" box="[522,597,1426,1452]" pageId="40" pageNumber="41">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFDDFFF58E60FA3CFD5905AD" box="[613,720,1426,1453]" name="Romania" pageId="40" pageNumber="41">Romania</collectingCountry>
), illustrated in
<bibRefCitation id="387E3CA4FFDDFFF58F7FFA3CFB8B05AC" author="Zilch, A." box="[890,1026,1426,1452]" pageId="40" pageNumber="41" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch (1934</bibRefCitation>
, pl. 17, fig. 18), figs 14B
<subScript id="C06B4310FFDDFFF58924FA0AFAA005B2" attach="left" box="[1313,1321,1444,1458]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
B
<subScript id="C06B4310FFDDFFF5894FFA0AFADB05B2" attach="right" box="[1354,1362,1444,1458]" fontSize="6" pageId="40" pageNumber="41">2</subScript>
</materialsCitation>
.
<materialsCitation id="EC874B08FFDDFFF58964FA3CFBAD05D0" ID-GBIF-Occurrence="3311814383" collectionCode="SMF" country="Germany" county="Main" location="Senckenberg Museum" municipality="Frankfurt" pageId="40" pageNumber="41" specimenCode="SMF 360.2208" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="8354FFF7FFDDFFF58964FA3CFE8405D0" pageId="40" pageNumber="41" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCode id="0C49E92EFFDDFFF58D1DFA18FE4D05D0" box="[280,452,1462,1488]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="40" pageNumber="41">SMF 360.2208</specimenCode>
bc,
<location id="5930178EFFDDFFF58DFDFA18FD7D05D0" LSID="urn:lsid:plazi:treatment:D446F043FFDDFFF48C92FB2CFE46021A:5930178EFFDDFFF58DFDFA18FD7D05D0" box="[504,756,1462,1488]" country="Germany" county="Main" municipality="Frankfurt" name="Senckenberg Museum" pageId="40" pageNumber="41">Senckenberg Museum</location>
,
<collectingMunicipality id="BC34DB2FFFDDFFF58F05FA18FCE405D0" box="[768,877,1462,1488]" pageId="40" pageNumber="41">Frankfurt</collectingMunicipality>
/
<collectingCounty id="B53139D9FFDDFFF58F70FA18FC3905D0" box="[885,944,1462,1488]" pageId="40" pageNumber="41">Main</collectingCounty>
,
<collectingCountry id="24F801C5FFDDFFF58FBEFA18FBAD05D0" box="[955,1060,1462,1488]" name="Germany" pageId="40" pageNumber="41">Germany</collectingCountry>
</materialsCitation>
,
<materialsCitation id="EC874B08FFDDFFF58835FA18FE9A05F5" ID-GBIF-Occurrence="3311814446" collectionCode="SMF" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="40" pageNumber="41" specimenCount="7" typeStatus="paralectotype">
17+
<specimenCount id="4AE98ADCFFDDFFF58859FA18FB6005D0" box="[1116,1257,1462,1488]" pageId="40" pageNumber="41" type="generic">7 specimens</specimenCount>
,
<location id="5930178EFFDDFFF588F0FA18FA1505D0" LSID="urn:lsid:plazi:treatment:D446F043FFDDFFF48C92FB2CFE46021A:5930178EFFDDFFF588F0FA18FA1505D0" box="[1269,1436,1462,1488]" country="Romania" name="Costeiu de Sus" pageId="40" pageNumber="41">
Coşteiu
<collectorName id="F11A2483FFDDFFF58957FA18FA1505D0" box="[1362,1436,1462,1488]" pageId="40" pageNumber="41">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFDDFFF58C9AFA74FE8205F5" box="[159,267,1498,1525]" name="Romania" pageId="40" pageNumber="41">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFDDFFF58CC2FA53FE06063C" blockId="40.[151,1437,1389,2028]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2FA53FE350617" bold="true" box="[199,444,1533,1559]" pageId="40" pageNumber="41">Additional material.</emphasis>
<collectionCode id="3AFED990FFDDFFF58DC1FA50FD960618" box="[452,543,1534,1560]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="41" type="Museum">NHMW</collectionCode>
1865/0036/0010, SL: 9.0 mm, MD:
<quantity id="9B17ECB0FFDDFFF58FC0FA50FB970618" box="[965,1054,1534,1560]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="40" pageNumber="41" unit="mm" value="3.9">3.9 mm</quantity>
, Drnovice u Vyškova (
<collectingCountry id="24F801C5FFDDFFF58926FA50FF68063D" name="Czech Republic" pageId="40" pageNumber="41">Czech Republic</collectingCountry>
), figs 14A
<subScript id="C06B4310FFDDFFF58D53F99AFED70642" attach="right" box="[342,350,1588,1602]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
A
<subScript id="C06B4310FFDDFFF58D85F99AFE010642" attach="left" box="[384,392,1588,1602]" fontSize="6" pageId="40" pageNumber="41">2</subScript>
.
</paragraph>
<paragraph id="5C504155FFDDFFF58CC2F9EBFA0F075C" blockId="40.[151,1437,1389,2028]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2F9EBFE30065F" bold="true" box="[199,441,1605,1631]" pageId="40" pageNumber="41">Revised description.</emphasis>
Shell moderately large, broad fusiform, with distinctly incised suture, and broad conical to weakly cyrtoconoid spire. Protoconch unknown. Teleoconch of seven whorls. Spire whorls weakly convex, with periphery at abapical suture. Axial sculpture of orthocline, convex ribs, separated by interspaces of about same width (about 2025 axial ribs on last whorl). Adapical tips of axial ribs may be slightly swollen resulting in slightly gradate spire. Last whorl strongly convex, rapidly contracting. Aperture wide, ovoid with indistinct anal sinus. Columellar callus indistinct. Columella with four prominent folds, weakening abapically. Outer lip thin, abapically contracting, about 1012 lirae within. No spiral sculpture, except four to six spiral cords on weak fasciole, partly extending from columellar folds. Siphonal canal straight, narrow, moderately short, with shallow siphonal notch.
</paragraph>
<paragraph id="5C504155FFDDFFF58CC2F8CBFEF807A4" blockId="40.[151,1437,1389,2028]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2F8CBFDB2077F" bold="true" box="[199,571,1893,1920]" pageId="40" pageNumber="41">Shell measurements and ratios.</emphasis>
SL: 9.012.0 mm, MD:
<quantity id="9B17ECB0FFDDFFF58F4AF8C8FC510780" box="[847,984,1894,1921]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="5.5" metricValueMin="3.9" pageId="40" pageNumber="41" unit="mm" value="4.7" valueMax="5.5" valueMin="3.9">3.95.5 mm</quantity>
; AA: 4555°, SL/MD: 2.32.4, AL/AW: 3.7, AH/S: 2.83.0.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDDFFF48CC2F803FCEC01AD" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="discussion">
<paragraph id="5C504155FFDDFFF48CC2F803FA9B0141" blockId="40.[151,1437,1389,2028]" lastBlockId="41.[151,1437,151,538]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2F803FEC507C7" bold="true" box="[199,332,1965,1991]" pageId="40" pageNumber="41">Discussion.</emphasis>
<bibRefCitation id="387E3CA4FFDDFFF58D56F800FD8D07C9" author="Boettger, O." box="[339,516,1966,1993]" pageId="40" pageNumber="41" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger (1906)</bibRefCitation>
described this species as variety of “
<taxonomicName id="9BEF3AD6FFDDFFF58F9BF801FBBA07C8" authorityName="Hoernes &amp; Auinger" authorityYear="1880" box="[926,1075,1966,1992]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="ebenus">
<emphasis id="6E9B9D47FFDDFFF58F9BF801FBBA07C8" box="[926,1075,1966,1992]" italics="true" pageId="40" pageNumber="41">Mitra ebenus</emphasis>
</taxonomicName>
” [=
<taxonomicName id="9BEF3AD6FFDDFFF58860F800FED707EC" authority="(Andrzejowski 1830)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Andrzejowski" baseAuthorityYear="1830" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="leucozona">
<emphasis id="6E9B9D47FFDDFFF58860F800FAE807C8" box="[1125,1377,1966,1992]" italics="true" pageId="40" pageNumber="41">Ebenomitra leucozona</emphasis>
(
<bibRefCitation id="387E3CA4FFDDFFF5896AF800FEDF07EC" author="Andrzejowski, A." pageId="40" pageNumber="41" pagination="90 - 104" refId="ref53874" refString="Andrzejowski, A. (1830) Notice sur quelques coquilles fossiles de Volhynie, Podolie etc. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 2, 90 - 104. [https: // www. biodiversitylibrary. org / item / 124281 # page / 94 / mode / 1 up]" type="journal article" year="1830">Andrzejowski 1830</bibRefCitation>
)
</taxonomicName>
], which is clearly incorrect.
<taxonomicName id="9BEF3AD6FFDDFFF58E93F87DFC0B07EC" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[662,898,2002,2028]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="paraleucozona">
<emphasis id="6E9B9D47FFDDFFF58E93F87DFC0B07EC" box="[662,898,2002,2028]" italics="true" pageId="40" pageNumber="41">Pusia paraleucozona</emphasis>
</taxonomicName>
has four prominent columellar folds and numerous delicate axial ribs, whereas
<taxonomicName id="9BEF3AD6FFDCFFF48DF0FF39FD0D00B1" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Andrzejowski" baseAuthorityYear="1830" box="[501,644,151,177]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="leucozona">
<emphasis id="6E9B9D47FFDCFFF48DF0FF39FD0D00B1" box="[501,644,151,177]" italics="true" pageId="41" pageNumber="42">E. leucozona</emphasis>
</taxonomicName>
has three folds and few blunt axial swellings. At first glance it is similar to
<taxonomicName id="9BEF3AD6FFDCFFF48CB1FF15FD2900D5" authority="(Hoernes &amp; Auinger, 1880)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[180,672,187,213]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="moravica">
<emphasis id="6E9B9D47FFDCFFF48CB1FF15FEEC00D4" box="[180,357,187,212]" italics="true" pageId="41" pageNumber="42">Pusia moravica</emphasis>
(
<bibRefCitation id="387E3CA4FFDCFFF48D71FF15FD1E00D5" author="Hoernes, R. &amp; Auinger, M." box="[372,663,187,213]" pageId="41" pageNumber="42" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
in being broadly fusiform with narrow ribs persisting on all whorls, but that species is larger [reconstructed height about
<quantity id="9B17ECB0FFDCFFF48EE6FF71FCD500F9" box="[739,860,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="41" pageNumber="42" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">1316 mm</quantity>
vs 9.012.0 mm in
<taxonomicName id="9BEF3AD6FFDCFFF4883CFF71FF54011D" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="paraleucozona">
<emphasis id="6E9B9D47FFDCFFF4883CFF71FAAC00F9" box="[1081,1317,223,249]" italics="true" pageId="41" pageNumber="42">Pusia paraleucozona</emphasis>
(
<bibRefCitation id="387E3CA4FFDCFFF48931FF71FF5C011D" author="Boettger, O." pageId="41" pageNumber="42" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
], at the same size the axial ribs are wider spaced in
<taxonomicName id="9BEF3AD6FFDCFFF48F13FEADFC13011C" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[790,922,259,284]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="moravica">
<emphasis id="6E9B9D47FFDCFFF48F13FEADFC13011C" box="[790,922,259,284]" italics="true" pageId="41" pageNumber="42">P. moravica</emphasis>
</taxonomicName>
, and the spiral sculpture in the subsutural area is weak, but present, whereas in
<taxonomicName id="9BEF3AD6FFDCFFF48E06FE89FD4C0141" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[515,709,295,321]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="paraleucozona">
<emphasis id="6E9B9D47FFDCFFF48E06FE89FD4C0141" box="[515,709,295,321]" italics="true" pageId="41" pageNumber="42">P. paraleucozona</emphasis>
</taxonomicName>
spiral sculpture is restricted to the siphonal fasciole.
</paragraph>
<paragraph id="5C504155FFDCFFF48CC2FEE5FBB00189" blockId="41.[151,1437,151,538]" pageId="41" pageNumber="42">
<taxonomicName id="9BEF3AD6FFDCFFF48CC2FEE5FC630165" authority="Landau, Ceulemans &amp; Van Dingenen, 2019" authorityName="Landau, Ceulemans &amp; Van Dingenen" authorityYear="2019" box="[199,1002,331,357]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="renauleauensis">
<emphasis id="6E9B9D47FFDCFFF48CC2FEE5FE720165" box="[199,507,331,357]" italics="true" pageId="41" pageNumber="42">Ebenomitra renauleauensis</emphasis>
<bibRefCitation id="387E3CA4FFDCFFF48E04FEE5FC630165" author="Landau, B. M. &amp; Ceulemans, L. &amp; Van Dingenen, F." box="[513,1002,331,357]" pageId="41" pageNumber="42" pagination="135 - 215" refId="ref58790" refString="Landau, B. M., Ceulemans, L. &amp; Van Dingenen, F. (2019) The upper Miocene gastropods of northwestern France, 4. Neogastropoda, Cainozoic Research, 19 / 2, 135 - 215." type="journal article" year="2019">Landau, Ceulemans &amp; Van Dingenen, 2019</bibRefCitation>
</taxonomicName>
, from the Tortonian of
<collectingCountry id="24F801C5FFDCFFF488E9FEE5FAB20165" box="[1260,1339,331,357]" name="France" pageId="41" pageNumber="42">France</collectingCountry>
, is reminiscent of
<taxonomicName id="9BEF3AD6FFDCFFF48D09FEC1FE440189" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[268,461,367,393]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="paraleucozona">
<emphasis id="6E9B9D47FFDCFFF48D09FEC1FE440189" box="[268,461,367,393]" italics="true" pageId="41" pageNumber="42">P. paraleucozona</emphasis>
</taxonomicName>
but differs in its smaller size and the coarser axial ribs.
</paragraph>
<paragraph id="5C504155FFDCFFF48CC2FE3CFCEC01AD" blockId="41.[151,1437,151,538]" box="[199,869,402,429]" pageId="41" pageNumber="42">
<emphasis id="6E9B9D47FFDCFFF48CC2FE3CFE3F01AC" bold="true" box="[199,438,402,428]" pageId="41" pageNumber="42">Palaeoenvironment.</emphasis>
Unknown; probably offshore settings.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDCFFF48CC2FE18FE46021A" pageId="41" pageNumber="42" type="distribution">
<paragraph id="5C504155FFDCFFF48CC2FE18FE46021A" blockId="41.[151,1437,151,538]" pageId="41" pageNumber="42">
<emphasis id="6E9B9D47FFDCFFF48CC2FE18FDEE01D1" bold="true" box="[199,615,438,465]" pageId="41" pageNumber="42">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFDCFFF48FB6FE19FAB401D1" box="[947,1341,439,465]" italics="true" pageId="41" pageNumber="42">North Alpine-Carpathian Foredeep</emphasis>
: Drnovice u Vyškova (
<collectingCountry id="24F801C5FFDCFFF48D41FE75FE7701F6" box="[324,510,475,502]" name="Czech Republic" pageId="41" pageNumber="42">Czech Republic</collectingCountry>
);
<emphasis id="6E9B9D47FFDCFFF48E14FE75FD4001F4" box="[529,713,475,500]" italics="true" pageId="41" pageNumber="42">Korytnica Basin</emphasis>
: Korytnica (
<collectingCountry id="24F801C5FFDCFFF48F52FE75FC2101F6" box="[855,936,475,502]" name="Poland" pageId="41" pageNumber="42">Poland</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFDCFFF48FBAFE75FBC101F5" author="Baluk, W." box="[959,1096,475,501]" pageId="41" pageNumber="42" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997</bibRefCitation>
);
<emphasis id="6E9B9D47FFDCFFF48859FE75FB6C01F4" box="[1116,1253,475,501]" italics="true" pageId="41" pageNumber="42">Făget Basin</emphasis>
: Coşteiu de Sus (
<collectingCountry id="24F801C5FFDCFFF48C9AFE51FE81021A" box="[159,264,511,538]" name="Romania" pageId="41" pageNumber="42">Romania</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFDCFFF48D18FE51FE4A021A" author="Boettger, O." box="[285,451,511,538]" pageId="41" pageNumber="42" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger 1906</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>