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<mods:title id="01C3687FF22CBAEB1DABD56F5E70B5C6">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<emphasis id="6E9B9D47FFC4FFEC8C92FE50FEE30219" bold="true" box="[151,362,510,537]" italics="true" pageId="49" pageNumber="50">Vexillum harmati</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC8D7CFE50FD5F0218" author="Csepreghy-Meznerics, I." box="[377,726,510,536]" pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics, 1954</bibRefCitation>
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<figureCitation id="C4D45DD0FFC4FFEC8C92FD8DFF76023D" box="[151,255,547,573]" captionStart="FIGURE 17" captionStartId="49.[151,250,1862,1886]" captionTargetBox="[151,1436,1378,1838]" captionTargetId="figure-447@49.[151,1436,1378,1838]" captionTargetPageId="49" captionText="FIGURE 17 A1A2. Vexillum harmati (Csepreghy-Meznerics, 1954), holotype, M.61.4361, Sámsonháza (Hungary). B1B2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary). C1C2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary). D1D2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary). E1E2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.4921876" httpUri="https://zenodo.org/record/4921876/files/figure.png" pageId="49" pageNumber="50">Figs 17A</figureCitation>
<figureCitation id="C4D45DD0FFC4FFEC8CFAFD9BFEA0023D" box="[255,297,547,579]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="49" pageNumber="50">
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A
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, B
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<subScript id="C06B4310FFC4FFEC8D57FD9BFED30243" attach="left" box="[338,346,565,579]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
B
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<figureCitation id="C4D45DD0FFC4FFEC8D7EFD9BFE0A0243" box="[379,387,565,579]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="49" pageNumber="50">
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, C
<figureCitation id="C4D45DD0FFC4FFEC8DA1FD9BFE45023D" box="[420,460,547,579]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC8DA1FD9BFE250243" attach="left" box="[420,428,565,579]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
C
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<figureCitation id="C4D45DD0FFC4FFEC8DC9FD9BFE5D0243" box="[460,468,565,579]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="49" pageNumber="50">
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, D
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D
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, E
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E
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<figureCitation id="C4D45DD0FFC4FFEC8E75FD9BFDF10243" box="[624,632,565,579]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="49" pageNumber="50">
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<emphasis id="6E9B9D47FFC4FFEC8C92FDC6FF2D027F" box="[151,164,616,639]" italics="true" pageId="49" pageNumber="50">*</emphasis>
<taxonomicName id="9BEF3AD6FFC4FFEC8CAFFDC9FE68027F" ID-CoL="8KRVT" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[170,481,615,639]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="harmati" status="sp. nov." subGenus="Costellaria">
<emphasis id="6E9B9D47FFC4FFEC8CAFFDC9FE88027F" box="[170,257,615,639]" italics="true" pageId="49" pageNumber="50">Vexillum</emphasis>
(
<emphasis id="6E9B9D47FFC4FFEC8D15FDC9FE0B027F" box="[272,386,615,639]" italics="true" pageId="49" pageNumber="50">Costellaria</emphasis>
)
<emphasis id="6E9B9D47FFC4FFEC8D94FDC9FE68027F" box="[401,481,615,639]" italics="true" pageId="49" pageNumber="50">harmati</emphasis>
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<taxonomicNameLabel id="75A8203CFFC4FFEC8DEDFDC6FD93027F" box="[488,538,615,639]" pageId="49" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
<treatmentCitation id="DD4E6744FFC4FFEC8E36FDC9FCF1027F" author="Csepreghy-Meznerics, I." box="[563,888,615,639]" page="45" pageId="49" pageNumber="50" year="1954">
<bibRefCitation id="387E3CA4FFC4FFEC8E36FDC9FCF1027F" author="Csepreghy-Meznerics, I." box="[563,888,615,639]" pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics 1954: 45</bibRefCitation>
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, pl. 5, figs 29, 3133.
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<treatmentCitationGroup id="7CFF667BFFC4FFEC8C92FD28FB90029E" box="[151,1049,646,670]" pageId="49" pageNumber="50">
<taxonomicName id="9BEF3AD6FFC4FFEC8C92FD28FD31029E" authority="Csepreghy-Meznerics, 1954" authorityName="Csepreghy-Meznerics" authorityYear="1954" box="[151,696,646,670]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="harmati" subGenus="Vexillum">
<emphasis id="6E9B9D47FFC4FFEC8C92FD28FF58029D" box="[151,209,646,669]" italics="true" pageId="49" pageNumber="50">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFC4FFEC8CE5FD28FEBE029E" box="[224,311,646,670]" italics="true" pageId="49" pageNumber="50">Vexillum</emphasis>
)
<emphasis id="6E9B9D47FFC4FFEC8D43FD28FE1F029E" box="[326,406,646,670]" italics="true" pageId="49" pageNumber="50">harmati</emphasis>
<bibRefCitation id="387E3CA4FFC4FFEC8D99FD28FD31029E" author="Csepreghy-Meznerics, I." box="[412,696,646,670]" pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics, 1954</bibRefCitation>
</taxonomicName>
<treatmentCitation id="DD4E6744FFC4FFEC8ED6FD28FC05029E" author="Strausz, L." box="[723,908,646,670]" page="368" pageId="49" pageNumber="50" year="1966">
<bibRefCitation id="387E3CA4FFC4FFEC8ED6FD28FC05029E" author="Strausz, L." box="[723,908,646,670]" pageId="49" pageNumber="50" refId="ref62393" refString="Strausz, L. (1966) Die Miozan-Mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp." type="book" year="1966">Strausz 1966: 368</bibRefCitation>
</treatmentCitation>
, pl. 12, fig. 9.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFC4FFEC8C92FD62FA970496" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="5C504155FFC4FFEC8C92FD62FF560353" blockId="49.[151,1437,716,1355]" pageId="49" pageNumber="50">
<emphasis id="6E9B9D47FFC4FFEC8C92FD62FECF02E6" bold="true" box="[151,326,716,742]" pageId="49" pageNumber="50">Type material.</emphasis>
<materialsCitation id="EC874B08FFC4FFEC8D48FD62FD490310" ID-GBIF-Occurrence="3311814490" collectorName="Hungarian Natural History Museum, SL" country="Hungary" location="Samsonhaza" pageId="49" pageNumber="50" specimenCount="1" typeStatus="holotype">
<typeStatus id="8354FFF7FFC4FFEC8D48FD62FE3302E6" box="[333,442,716,742]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
: M.61.4361,
<collectorName id="F11A2483FFC4FFEC8E4AFD62FB9302E6" box="[591,1050,716,743]" pageId="49" pageNumber="50">Hungarian Natural History Museum, SL</collectorName>
: 24.0 mm, MD:
<quantity id="9B17ECB0FFC4FFEC88D0FD62FAA702E7" box="[1237,1326,716,743]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="49" pageNumber="50" unit="mm" value="6.2">6.2 mm</quantity>
,
<location id="5930178EFFC4FFEC893FFD62FF43030A" LSID="urn:lsid:plazi:treatment:D446F043FFC4FFEF8C92FE50FECA011E:5930178EFFC4FFEC893FFD62FF43030A" country="Hungary" name="Samsonhaza" pageId="49" pageNumber="50">Sámsonháza</location>
(
<collectingCountry id="24F801C5FFC4FFEC8CDFFD5EFEB6030A" box="[218,319,752,778]" name="Hungary" pageId="49" pageNumber="50">Hungary</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFC4FFEC8D53FD5EFD91030A" author="Palfy, J. &amp; Dulai, A. &amp; Gasparik, M. &amp; Oszvart, P. &amp; Pazonyi, P. &amp; Szives O." box="[342,536,752,778]" pageId="49" pageNumber="50" refId="ref60193" refString="Palfy, J., Dulai, A., Gasparik, M., Oszvart, P., Pazonyi, P. &amp; Szives O. (2008) Catalogue of Invertebrate and Vertebrate Paleontological Type Specimens of the Hungarian Natural History Museum. Hungarian Natural History Museum, Budapest, 209 pp. [http: // www. nhmus. hu / modules / Tar-Konyv / pdf / paleontological _ type _ spec. pdf]" type="book" year="2008">
Pálfy
<emphasis id="6E9B9D47FFC4FFEC8D9EFD5FFE5D030A" box="[411,468,752,778]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
2008
</bibRefCitation>
), figs 17A
<subScript id="C06B4310FFC4FFEC8E8BFCACFD1F0310" attach="left" box="[654,662,770,784]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
A
<subScript id="C06B4310FFC4FFEC8EBDFCACFD490310" attach="left" box="[696,704,770,784]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
</materialsCitation>
.
<materialsCitation id="EC874B08FFC4FFEC8ECAFD5FFF510358" ID-GBIF-Occurrence="3311814431" collectorName="Hungarian Natural History Museum, SL" country="Hungary" location="Samsonhaza" pageId="49" pageNumber="50" specimenCount="3" typeStatus="paratype">
<typeStatus id="8354FFF7FFC4FFEC8ECAFD5FFCC9030B" box="[719,832,753,779]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
: M.61.4362,
<specimenCount id="4AE98ADCFFC4FFEC8FD0FD5EFBEA030A" box="[981,1123,752,778]" pageId="49" pageNumber="50" type="generic">3 specimens</specimenCount>
,
<collectorName id="F11A2483FFC4FFEC886BFD5EFEA6032E" pageId="49" pageNumber="50">Hungarian Natural History Museum, SL</collectorName>
: 15.0 mm, MD:
<quantity id="9B17ECB0FFC4FFEC8DF4FCBAFDC2032F" box="[497,587,788,815]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="49" pageNumber="50" unit="mm" value="3.9">3.9 mm</quantity>
,
<location id="5930178EFFC4FFEC8E5CFCBAFD6C032E" LSID="urn:lsid:plazi:treatment:D446F043FFC4FFEF8C92FE50FECA011E:5930178EFFC4FFEC8E5CFCBAFD6C032E" box="[601,741,788,814]" country="Hungary" name="Samsonhaza" pageId="49" pageNumber="50">Sámsonháza</location>
(
<collectingCountry id="24F801C5FFC4FFEC8EF3FCBAFCD2032E" box="[758,859,788,814]" name="Hungary" pageId="49" pageNumber="50">Hungary</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFC4FFEC8F71FCBAFBB2032E" author="Palfy, J. &amp; Dulai, A. &amp; Gasparik, M. &amp; Oszvart, P. &amp; Pazonyi, P. &amp; Szives O." box="[884,1083,788,814]" pageId="49" pageNumber="50" refId="ref60193" refString="Palfy, J., Dulai, A., Gasparik, M., Oszvart, P., Pazonyi, P. &amp; Szives O. (2008) Catalogue of Invertebrate and Vertebrate Paleontological Type Specimens of the Hungarian Natural History Museum. Hungarian Natural History Museum, Budapest, 209 pp. [http: // www. nhmus. hu / modules / Tar-Konyv / pdf / paleontological _ type _ spec. pdf]" type="book" year="2008">
Pálfy
<emphasis id="6E9B9D47FFC4FFEC8FBEFCBBFC7C032E" box="[955,1013,788,814]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
2008
</bibRefCitation>
),
<figureCitation id="C4D45DD0FFC4FFEC8855FCBAFB30032E" box="[1104,1209,788,814]" captionStart="FIGURE 17" captionStartId="49.[151,250,1862,1886]" captionTargetBox="[151,1436,1378,1838]" captionTargetId="figure-447@49.[151,1436,1378,1838]" captionTargetPageId="49" captionText="FIGURE 17 A1A2. Vexillum harmati (Csepreghy-Meznerics, 1954), holotype, M.61.4361, Sámsonháza (Hungary). B1B2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary). C1C2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary). D1D2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary). E1E2. Vexillum harmati (Csepreghy-Meznerics, 1954), paratype, M.61.4362, Sámsonháza (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.4921876" httpUri="https://zenodo.org/record/4921876/files/figure.png" pageId="49" pageNumber="50">Figs 17B</figureCitation>
<figureCitation id="C4D45DD0FFC4FFEC88BCFC88FB68032F" box="[1209,1249,789,820]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC88BCFC88FB480334" attach="left" box="[1209,1217,806,820]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
B
</figureCitation>
<figureCitation id="C4D45DD0FFC4FFEC88E7FC88FB630334" box="[1250,1258,806,820]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC88E7FC88FB630334" attach="right" box="[1250,1258,806,820]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
</figureCitation>
, C
<figureCitation id="C4D45DD0FFC4FFEC8908FC88FABF032E" box="[1293,1334,788,820]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC8908FC88FA9C0334" attach="left" box="[1293,1301,806,820]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
C
</figureCitation>
<figureCitation id="C4D45DD0FFC4FFEC8933FC88FAB70334" box="[1334,1342,806,820]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC8933FC88FAB70334" attach="left" box="[1334,1342,806,820]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
</figureCitation>
, D
<figureCitation id="C4D45DD0FFC4FFEC8966FC88FA04032F" box="[1379,1421,789,820]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC8966FC88FAE20334" attach="right" box="[1379,1387,806,820]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
D
</figureCitation>
<figureCitation id="C4D45DD0FFC4FFEC8988FC88FA1C0334" box="[1421,1429,806,820]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC8988FC88FA1C0334" attach="left" box="[1421,1429,806,820]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
</figureCitation>
, E
<figureCitation id="C4D45DD0FFC4FFEC8CADFCE4FF590353" box="[168,208,825,856]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC8CADFCE4FF390358" attach="left" box="[168,176,842,856]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
E
</figureCitation>
<figureCitation id="C4D45DD0FFC4FFEC8CD5FCE4FF510358" box="[208,216,842,856]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="49" pageNumber="50">
<subScript id="C06B4310FFC4FFEC8CD5FCE4FF510358" attach="left" box="[208,216,842,856]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
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</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFC4FFEC8CC2FCF2FD2F0472" blockId="49.[151,1437,716,1355]" pageId="49" pageNumber="50">
<emphasis id="6E9B9D47FFC4FFEC8CC2FCF2FE350376" bold="true" box="[199,444,860,886]" pageId="49" pageNumber="50">Revised description.</emphasis>
Shell moderately large, slender fusiform, with very high spire. Protoconch unknown. Teleoconch of nine high, subcylindrical to weakly convex whorls, with periphery mid-whorl, separated by incised suture. Sculpture of widely spaced, sharp-crested, orthocline to weakly opisthocline axial ribs, most prominent along periphery, weakening towards adapical and abapical suture. Axial sculpture fading out on seventh spire whorl; penultimate and last whorls smooth. Last whorl high, subcylindrical, weakly shouldered, moderately constricted at base. No spiral sculpture except for faint spiral cords on weak fasciole. Aperture narrow elongate. Columellar callus indistinct, columella with four oblique folds. Outer lip not preserved. Siphonal canal moderately long, wide, slightly bent to left, with wide, shallow siphonal notch.
</paragraph>
<paragraph id="5C504155FFC4FFEC8CC2FBD2FA970496" blockId="49.[151,1437,716,1355]" box="[199,1310,1148,1175]" pageId="49" pageNumber="50">
<emphasis id="6E9B9D47FFC4FFEC8CC2FBD2FDB50496" bold="true" box="[199,572,1148,1174]" pageId="49" pageNumber="50">Shell measurements and ratios.</emphasis>
SL: 15.024.0 mm, MD:
<quantity id="9B17ECB0FFC4FFEC8F5AFBD2FC600497" box="[863,1001,1148,1175]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="6.2" metricValueMin="3.9" pageId="49" pageNumber="50" unit="mm" value="5.05" valueMax="6.2" valueMin="3.9">3.96.2 mm</quantity>
; AA: 2430°, A/S: 0.60.7.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFC4FFEF8CC2FB0EFEDA00D5" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="discussion">
<paragraph id="5C504155FFC4FFEC8CC2FB0EFB5D054A" blockId="49.[151,1437,716,1355]" pageId="49" pageNumber="50">
<emphasis id="6E9B9D47FFC4FFEC8CC2FB0EFEC504BA" bold="true" box="[199,332,1184,1210]" pageId="49" pageNumber="50">Discussion.</emphasis>
This species is reminiscent of
<taxonomicName id="9BEF3AD6FFC4FFEC8EB7FB0EFB3304BA" authority="(Hoernes &amp; Auinger, 1880)" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[690,1210,1184,1210]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="badense">
<emphasis id="6E9B9D47FFC4FFEC8EB7FB0EFCF104BA" box="[690,888,1184,1210]" italics="true" pageId="49" pageNumber="50">Vexillum badense</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC8F8DFB0EFB3B04BA" author="Hoernes, R. &amp; Auinger, M." box="[904,1202,1184,1210]" pageId="49" pageNumber="50" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="9BEF3AD6FFC4FFEC88F0FB0EFDAE04DE" authority="(Hoernes &amp; Auinger, 1880)" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="intermittens">
<emphasis id="6E9B9D47FFC4FFEC88F0FB0EFF6104DE" italics="true" pageId="49" pageNumber="50">Vexillum intermittens</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC8CF2FB6AFDA904DE" author="Hoernes, R. &amp; Auinger, M." box="[247,544,1220,1246]" pageId="49" pageNumber="50" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
, but is distinguished from both species by its distinctly more slender shell and higher spire whorls. Its axial ribs are most prominent mid-whorl, whereas in the other two species they strengthen towards the abapical suture. Finally, the occurrence in silty-sandy nearshore deposits of Sámsonháza indicates an ecological separation from
<taxonomicName id="9BEF3AD6FFC4FFEC8DCCFA9FFDC9054A" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[457,576,1328,1354]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="badense">
<emphasis id="6E9B9D47FFC4FFEC8DCCFA9FFDC9054A" box="[457,576,1328,1354]" italics="true" pageId="49" pageNumber="50">V. badense</emphasis>
</taxonomicName>
and
<emphasis id="6E9B9D47FFC4FFEC8E73FA9FFC97054A" box="[630,798,1329,1354]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="9BEF3AD6FFC4FFEC8E73FA9FFC93054A" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[630,794,1329,1354]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="intermittens">V. intermittens</taxonomicName>
,
</emphasis>
which are confined to basinal settings.
</paragraph>
<caption id="089011DDFFC4FFEC8C92F8E8FD8007EE" ID-DOI="http://doi.org/10.5281/zenodo.4921876" ID-Zenodo-Dep="4921876" httpUri="https://zenodo.org/record/4921876/files/figure.png" pageId="49" pageNumber="50" startId="49.[151,250,1862,1886]" targetBox="[151,1436,1378,1838]" targetPageId="49">
<paragraph id="5C504155FFC4FFEC8C92F8E8FD8007EE" blockId="49.[151,1437,1862,2030]" pageId="49" pageNumber="50">
<emphasis id="6E9B9D47FFC4FFEC8C92F8E8FEE5075E" bold="true" box="[151,364,1862,1892]" pageId="49" pageNumber="50">
FIGURE 17 A
<subScript id="C06B4310FFC4FFEC8D3DF8F8FEB60764" attach="left" box="[312,319,1878,1892]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
A
<subScript id="C06B4310FFC4FFEC8D5BF8F8FEEC0764" attach="left" box="[350,357,1878,1892]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFC4FFEC8D70F8E8FCEA075E" authority="(Csepreghy-Meznerics, 1954)" baseAuthorityName="Csepreghy-Meznerics" baseAuthorityYear="1954" box="[373,867,1862,1886]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="harmati">
<emphasis id="6E9B9D47FFC4FFEC8D70F8E8FDAF075E" box="[373,550,1862,1886]" italics="true" pageId="49" pageNumber="50">Vexillum harmati</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC8E33F8E8FCD5075E" author="Csepreghy-Meznerics, I." box="[566,860,1862,1886]" pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics, 1954</bibRefCitation>
)
</taxonomicName>
, holotype, M.61.4361, Sámsonháza (Hungary).
<emphasis id="6E9B9D47FFC4FFEC895CF8E8FA15075E" bold="true" box="[1369,1436,1862,1892]" pageId="49" pageNumber="50">
B
<subScript id="C06B4310FFC4FFEC896FF8F8FAF80764" attach="left" box="[1386,1393,1878,1892]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
B
<subScript id="C06B4310FFC4FFEC898AF8F8FA1F0764" attach="right" box="[1423,1430,1878,1892]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFC4FFEC8C92F8C4FD060782" authority="(Csepreghy-Meznerics, 1954)" baseAuthorityName="Csepreghy-Meznerics" baseAuthorityYear="1954" box="[151,655,1898,1922]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="harmati">
<emphasis id="6E9B9D47FFC4FFEC8C92F8C4FEC20782" box="[151,331,1898,1922]" italics="true" pageId="49" pageNumber="50">Vexillum harmati</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC8D5AF8C4FD010782" author="Csepreghy-Meznerics, I." box="[351,648,1898,1922]" pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics, 1954</bibRefCitation>
)
</taxonomicName>
, paratype, M.61.4362, Sámsonháza (Hungary).
<emphasis id="6E9B9D47FFC4FFEC8890F8C4FB520782" bold="true" box="[1173,1243,1898,1928]" pageId="49" pageNumber="50">
C
<subScript id="C06B4310FFC4FFEC88A2F8D4FB270788" attach="left" box="[1191,1198,1914,1928]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
C
<subScript id="C06B4310FFC4FFEC88C8F8D4FB5D0788" attach="left" box="[1229,1236,1914,1928]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFC4FFEC88EDF8C4FE4207A6" authority="(Csepreghy-Meznerics, 1954)" baseAuthorityName="Csepreghy-Meznerics" baseAuthorityYear="1954" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="harmati">
<emphasis id="6E9B9D47FFC4FFEC88EDF8C4FA150782" box="[1256,1436,1898,1922]" italics="true" pageId="49" pageNumber="50">Vexillum harmati</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC8C9BF820FE4D07A6" author="Csepreghy-Meznerics, I." box="[158,452,1934,1958]" pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics, 1954</bibRefCitation>
)
</taxonomicName>
, paratype, M.61.4362, Sámsonháza (Hungary).
<emphasis id="6E9B9D47FFC4FFEC8FC5F820FB8F07A6" bold="true" box="[960,1030,1934,1964]" pageId="49" pageNumber="50">
D
<subScript id="C06B4310FFC4FFEC8FD7F830FC5007AC" attach="left" box="[978,985,1950,1964]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
D
<subScript id="C06B4310FFC4FFEC8FFDF830FC7607AC" attach="left" box="[1016,1023,1950,1964]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFC4FFEC880AF820FE8907CA" authority="(Csepreghy-Meznerics, 1954)" baseAuthorityName="Csepreghy-Meznerics" baseAuthorityYear="1954" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="harmati">
<emphasis id="6E9B9D47FFC4FFEC880AF820FB4907A6" box="[1039,1216,1934,1958]" italics="true" pageId="49" pageNumber="50">Vexillum harmati</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC88D5F820FF7007CA" author="Csepreghy-Meznerics, I." pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics, 1954</bibRefCitation>
)
</taxonomicName>
, paratype, M.61.4362, Sámsonháza (Hungary).
<emphasis id="6E9B9D47FFC4FFEC8EF3F81CFCB007CA" bold="true" box="[758,825,1970,2000]" pageId="49" pageNumber="50">
E
<subScript id="C06B4310FFC4FFEC8F02F86CFC8707D0" attach="left" box="[775,782,1986,2000]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
E
<subScript id="C06B4310FFC4FFEC8F2EF86CFCBB07D0" attach="left" box="[811,818,1986,2000]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFC4FFEC8F46F81CFAB807CA" authority="(Csepreghy-Meznerics, 1954)" baseAuthorityName="Csepreghy-Meznerics" baseAuthorityYear="1954" box="[835,1329,1970,1994]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="harmati">
<emphasis id="6E9B9D47FFC4FFEC8F46F81CFC7A07CA" box="[835,1011,1970,1994]" italics="true" pageId="49" pageNumber="50">Vexillum harmati</emphasis>
(
<bibRefCitation id="387E3CA4FFC4FFEC8801F81CFAA307CA" author="Csepreghy-Meznerics, I." box="[1028,1322,1970,1994]" pageId="49" pageNumber="50" pagination="1 - 185" refId="ref55088" refString="Csepreghy-Meznerics, I. (1954). Helvetische und tortonische Fauna aus dem ostlichen Cserhatgebirge. Annales Instituti Regii Hungarici Geologici, 41, 1 - 185. [http: // olivirv. myspecies. info / sites / olivirv. myspecies. info / files / EPA 03274 _ mafi _ evkonyv _ 1954 _ 04. pdf]" type="journal article" year="1954">Csepreghy-Meznerics, 1954</bibRefCitation>
)
</taxonomicName>
, paratype, M.61.4362, Sámsonháza (Hungary).
</paragraph>
</caption>
<paragraph id="5C504155FFC7FFEF8CC2FF38FEDA00D5" blockId="50.[151,1437,150,286]" pageId="50" pageNumber="51">
<emphasis id="6E9B9D47FFC7FFEF8CC2FF38FE3F00B0" bold="true" box="[199,438,150,176]" pageId="50" pageNumber="51">Palaeoenvironment.</emphasis>
The Sámsonháza Formation represents inner neritic shallow water environments (
<bibRefCitation id="387E3CA4FFC7FFEF8942FF39FECE00D5" author="Kovacs, Z. &amp; Vician, Z." pageId="50" pageNumber="51" pagination="75 - 104" refId="ref58494" refString="Kovacs, Z. &amp; Vician, Z. (2017) Middle Miocene Tonnoidea and Ficoidea (Caenogastropoda) assemblages from Letkes (Hungary). Fragmenta Palaeontologica Hungarica, 34, 75 - 104. [https: // core. ac. uk / reader / 159127466] https: // doi. org / 10.17111 / FragmPalHung. 2017.34.75" type="journal article" year="2017">Kovács &amp; Vicián, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFC7FFEF8CC2FF70FECA011E" pageId="50" pageNumber="51" type="distribution">
<paragraph id="5C504155FFC7FFEF8CC2FF70FECA011E" blockId="50.[151,1437,150,286]" pageId="50" pageNumber="51">
<emphasis id="6E9B9D47FFC7FFEF8CC2FF70FDE400F9" bold="true" box="[199,621,222,249]" pageId="50" pageNumber="51">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFC7FFEF8FC4FF71FB0D00F8" box="[961,1156,223,248]" italics="true" pageId="50" pageNumber="51">Pannonian Basin</emphasis>
: Sámsonháza (
<collectingCountry id="24F801C5FFC7FFEF892AFF71FA1D00F9" box="[1327,1428,223,249]" name="Hungary" pageId="50" pageNumber="51">Hungary</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFC7FFEF8C9AFEADFEBF011E" author="Strausz, L." box="[159,310,259,286]" pageId="50" pageNumber="51" refId="ref62393" refString="Strausz, L. (1966) Die Miozan-Mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp." type="book" year="1966">Strausz 1966</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>