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<document ID-DOI="http://doi.org/10.5281/zenodo.3792794" ID-GBIF-Dataset="142a25ed-47c3-4361-bc24-6f1b3e8dba5c" ID-GBIF-Taxon="163755050" ID-ISSN="13132970" ID-Zenodo-Dep="3792794" ID-ZooBank="664C49F1-5384-43C4-8BF1-CE76AC11D32E" approvalRequired="10" approvalRequired_for_treatments="10" checkinTime="1587998787238" checkinUser="plazi" docAuthor="Klimaszewski, Jan, Savard, Karine, Pelletier, Georges &amp; Webster, Reginald" docDate="2008" docId="D43E034BFF8E4179FF17FE5A6628FDCD" docLanguage="en" docName="ZK_article_1943.pdf" docOrigin="ZooKeys 2 (2)" docStyle="DocumentStyle:BEF2356135E79670CA200BCAD40C4E3A:zookeys.2008.journal_article" docStyleId="BEF2356135E79670CA200BCAD40C4E3A" docStyleName="zookeys.2008.journal_article" docTitle="Gnypeta brincki Palm 1966" docType="treatment" docVersion="11" lastPageId="21" lastPageNumber="32" masterDocId="28077B33FF9D416CFF9BFFC76718FFEC" masterDocTitle="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution" masterLastPageNumber="84" masterPageNumber="11" pageId="19" pageNumber="30" updateTime="1643574247635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimaszewski, Jan</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Savard, Karine</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pelletier, Georges</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Webster, Reginald</mods:namePart>
<mods:affiliation>24 Millstream Drive, Fredericton, NB, Canada &amp; F-</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>84</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.2.4</mods:identifier>
<mods:identifier type="GBIF-Dataset">142a25ed-47c3-4361-bc24-6f1b3e8dba5c</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576410</mods:identifier>
<mods:identifier type="ZooBank">664C49F1-5384-43C4-8BF1-CE76AC11D32E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792794" ID-GBIF-Taxon="163755050" ID-Zenodo-Dep="3792794" LSID="urn:lsid:plazi:treatment:D43E034BFF8E4179FF17FE5A6628FDCD" httpUri="http://treatment.plazi.org/id/D43E034BFF8E4179FF17FE5A6628FDCD" lastPageId="21" lastPageNumber="32" pageId="19" pageNumber="30">
<subSubSection box="[140,432,412,439]" pageId="19" pageNumber="30" type="nomenclature">
<paragraph blockId="19.[140,459,412,475]" box="[140,432,412,439]" pageId="19" pageNumber="30">
<heading box="[140,432,412,439]" fontSize="11" level="2" pageId="19" pageNumber="30" reason="3">
<emphasis bold="true" box="[140,432,412,439]" pageId="19" pageNumber="30">
2.
<taxonomicName authority="Palm" authorityName="Palm" authorityYear="1966" box="[170,432,412,439]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="30" phylum="Arthropoda" rank="species" species="brincki">
<emphasis bold="true" box="[170,364,412,439]" italics="true" pageId="19" pageNumber="30">Gnypeta brincki</emphasis>
Palm
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,459,448,475]" pageId="19" pageNumber="30" type="description">
<paragraph blockId="19.[140,459,412,475]" box="[140,459,448,475]" pageId="19" pageNumber="30">
(
<figureCitation box="[147,218,448,474]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="19" pageNumber="30">Figs 4</figureCitation>
,
<figureCitation box="[231,311,448,475]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="19" pageNumber="30">22 a, b</figureCitation>
,
<figureCitation box="[323,393,448,475]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">48-56</figureCitation>
,
<figureCitation box="[406,451,448,474]" captionStart="Fig" captionStartId="60.[140,174,1526,1548]" captionTargetBox="[140,1103,451,785]" captionTargetPageId="60" captionText="Fig. 191. Collection localities in Canada and Alaska of G. brincki." figureDoi="http://doi.org/10.5281/zenodo.3774134" httpUri="https://zenodo.org/record/3774134/files/figure.png" pageId="19" pageNumber="30">191</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="30" type="materials_examined">
<paragraph blockId="19.[140,1108,518,651]" pageId="19" pageNumber="30">
<emphasis bold="true" box="[140,359,518,545]" pageId="19" pageNumber="30">
<collectionCode box="[140,211,519,545]" pageId="19" pageNumber="30">TYPE</collectionCode>
<collectionCode box="[217,359,518,544]" pageId="19" pageNumber="30">LOCALITY</collectionCode>
</emphasis>
:
<collectingCountry box="[373,499,519,545]" name="Sweden" pageId="19" pageNumber="30">SWEDEN</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:D43E034BFF8E4179FF17FE5A6628FDCD:5948E486FF8E417FFE60FDC065A5FDCC" box="[507,701,518,545]" country="Sweden" municipality="Abisco" name="Torne Lappmark" pageId="19" pageNumber="30">Torne Lappmark</location>
,
<collectingMunicipality box="[711,790,518,544]" pageId="19" pageNumber="30">Abisco</collectingMunicipality>
, 1947 (
<collectionCode box="[873,937,519,545]" pageId="19" pageNumber="30">LUC</collectionCode>
).
<typeStatus box="[955,1062,518,544]" pageId="19" pageNumber="30">Holotype</typeStatus>
not examined.
</paragraph>
<paragraph blockId="19.[140,1108,518,651]" box="[140,363,589,615]" pageId="19" pageNumber="30">
<emphasis bold="true" box="[140,363,589,615]" pageId="19" pageNumber="30">
<emphasis box="[140,241,589,615]" inLineHeading="true" pageId="19" pageNumber="30" reason="1">Material</emphasis>
examined
</emphasis>
</paragraph>
<paragraph blockId="19.[140,1108,518,651]" box="[185,592,624,651]" pageId="19" pageNumber="30">Specimens are listed in Appendix A.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="30" type="diagnosis">
<paragraph blockId="19.[140,1107,694,968]" box="[140,259,694,720]" pageId="19" pageNumber="30">
<emphasis bold="true" box="[140,259,694,720]" inLineHeading="true" pageId="19" pageNumber="30" reason="1">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="19.[140,1107,694,968]" pageId="19" pageNumber="30">
This species can be recognized by the following combination of characters: body length 3.2-3.6 mm (
<figureCitation box="[295,365,765,791]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="19" pageNumber="30">Fig. 4</figureCitation>
); elytra (at suture) slightly longer than pronotum and 1/3 wider than maximum width of pronotum (
<figureCitation box="[553,621,800,826]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="19" pageNumber="30">Fig. 4</figureCitation>
); abdomen at base distinctly narrower than elytra (
<figureCitation box="[221,293,835,861]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="19" pageNumber="30">Fig. 4</figureCitation>
); antennal articles 4-6 elongate in males and subquadrate in females, articles 7-9 transverse (
<figureCitation box="[397,532,870,897]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="19" pageNumber="30">Figs 22 a, b</figureCitation>
); median lobe of aedeagus with apical part broadly triangular in lateral view (
<figureCitation box="[447,534,906,932]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 48</figureCitation>
); spermatheca with capsule wider than long and funnel-shaped (
<figureCitation box="[316,399,941,968]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 54</figureCitation>
); stem sinuate and slightly swollen basally (
<figureCitation box="[889,973,941,968]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 54</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="30" type="description">
<paragraph blockId="19.[140,1108,1011,1672]" box="[140,282,1011,1037]" pageId="19" pageNumber="30">
<emphasis bold="true" box="[140,282,1011,1037]" inLineHeading="true" pageId="19" pageNumber="30" reason="1">Description</emphasis>
</paragraph>
<paragraph blockId="19.[140,1108,1011,1672]" pageId="19" pageNumber="30">
Body length 3.2-3.6 mm; black, sometimes central part of elytra and tarsi reddish brown (
<figureCitation box="[229,297,1082,1108]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="19" pageNumber="30">Fig. 4</figureCitation>
); integument strongly glossy; pubescence yellowish grey and moderately long and dense; antennal article 4 slightly shorter than preceding one, 4-6 elongate in males and subquadrate in females, articles 7-9 transverse (
<figureCitation box="[839,979,1152,1179]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="19" pageNumber="30">Figs 22 a, b</figureCitation>
); head and pronotum of about the same width, abdomen wider than either head or pronotum; elytra large and distinctly broader than remainder of body (
<figureCitation box="[848,920,1222,1249]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="19" pageNumber="30">Fig. 4</figureCitation>
); head rounded posteriorly; pronotum broadest in apical third, pubescence directed anterad along midline and laterad elsewhere; elytra (at suture) slightly longer than pronotum and 1/3 broader than maximum width of pronotum (
<figureCitation box="[727,800,1328,1354]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="19" pageNumber="30">Fig. 4</figureCitation>
), pubescence directed obliquely laterad; abdomen subparallel; metatarsus with two basal articles of about the same length and the third one slightly shorter.
<emphasis bold="true" box="[666,726,1399,1425]" pageId="19" pageNumber="30">Male</emphasis>
. Tergite 8 transverse and truncate apically (
<figureCitation box="[241,326,1433,1460]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 52</figureCitation>
). Sternite 8 slightly transverse and rounded posteriorly (
<figureCitation box="[956,1040,1433,1460]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 53</figureCitation>
). Median lobe of aedeagus with broadly triangular apical part of tubus in lateral view (
<figureCitation captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 48</figureCitation>
), bulbus enlarged and bearing two small angular anterior projections in dorsal view (
<figureCitation box="[207,291,1539,1565]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 49</figureCitation>
), internal sac with structures as illustrated (
<figureCitation box="[782,909,1539,1566]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Figs 48-50</figureCitation>
).
<emphasis bold="true" box="[930,1015,1539,1565]" pageId="19" pageNumber="30">Female</emphasis>
. Tergite 8 similar to that of male (
<figureCitation box="[439,525,1574,1601]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 55</figureCitation>
). Sternite 8 broadly rounded posteriorly (
<figureCitation box="[1007,1094,1574,1601]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 56</figureCitation>
). Spermatheca with capsule wider than long and funnel-shaped (
<figureCitation box="[859,944,1609,1636]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 54</figureCitation>
); stem sinuate and slightly swollen basally (
<figureCitation box="[465,551,1645,1672]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="19" pageNumber="30">Fig. 54</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774098" ID-Zenodo-Dep="3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="20" pageNumber="31" startId="20.[140,184,1590,1612]" targetBox="[152,1097,238,1512]" targetPageId="20">
<paragraph blockId="20.[140,1108,1590,1676]" pageId="20" pageNumber="31">
<emphasis bold="true" box="[140,260,1590,1612]" pageId="20" pageNumber="31">Figs 48-56.</emphasis>
Genital structures of
<taxonomicName authorityName="Palm" authorityYear="1966" box="[466,611,1590,1612]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="31" phylum="Arthropoda" rank="species" species="brincki">
<emphasis box="[466,611,1590,1612]" italics="true" pageId="20" pageNumber="31">Gnypeta brincki</emphasis>
</taxonomicName>
: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8.
</paragraph>
</caption>
<subSubSection pageId="21" pageNumber="32" type="distribution">
<paragraph blockId="21.[140,1108,166,333]" box="[140,411,166,193]" pageId="21" pageNumber="32">
<emphasis bold="true" box="[140,291,166,192]" inLineHeading="true" pageId="21" pageNumber="32" reason="1">Distribution</emphasis>
(Fig. 191)
</paragraph>
<paragraph blockId="21.[140,1108,166,333]" pageId="21" pageNumber="32">
<taxonomicName authorityName="Palm" authorityYear="1966" box="[140,317,202,228]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="32" phylum="Arthropoda" rank="species" species="brincki">
<emphasis box="[140,317,202,228]" italics="true" pageId="21" pageNumber="32">Gnypeta brincki</emphasis>
</taxonomicName>
is a Holarctic species with probably circumpolar distribution. In the Palaearctic region it was recorded from
<collectingCountry box="[608,699,237,263]" name="Norway" pageId="21" pageNumber="32">Norway</collectingCountry>
,
<collectingCountry box="[715,801,237,263]" name="Sweden" pageId="21" pageNumber="32">Sweden</collectingCountry>
and
<collectingCountry box="[865,952,237,263]" name="Finland" pageId="21" pageNumber="32">Finland</collectingCountry>
(
<bibRefCitation author="Palm" box="[971,1102,237,263]" firstAuthor="Palm" pageId="21" pageNumber="32" pagination="136 - 141" refId="ref23890" refString="Palm T (1966) De svenska Gnypeta - arterna (Col. Staphylinidae, Aleocharinae). Entomologisk Tidskrift 87: 136 - 141." type="journal article" year="1966">Palm 1966</bibRefCitation>
;
<bibRefCitation author="Smetana" box="[140,306,272,298]" firstAuthor="Smetana" pageId="21" pageNumber="32" pagination="353 - 494" refId="ref24122" refString="Smetana A (2004) Subfamily Aleocharinae Fleming, 1821 In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 2. Apollo Books, Stenstrup. 353 - 494." type="book chapter" year="2004">Smetana 2004</bibRefCitation>
). Here, it is recorded for the first time in North America from Quebec, Yukon Territory, Northwest Territories and Alaska.
</paragraph>
<paragraph blockId="21.[140,1108,377,545]" box="[140,467,377,404]" pageId="21" pageNumber="32">
<emphasis bold="true" box="[140,467,377,404]" pageId="21" pageNumber="32">
<emphasis box="[140,265,377,403]" inLineHeading="true" pageId="21" pageNumber="32" reason="1">Collection</emphasis>
and habitat data
</emphasis>
</paragraph>
<paragraph blockId="21.[140,1108,377,545]" pageId="21" pageNumber="32">
Adults were collected in June, July, and August. The
<collectingRegion box="[724,812,413,439]" country="Canada" name="Quebec" pageId="21" pageNumber="32">Quebec</collectingRegion>
specimen was found in an estuary of a river,
<quantity box="[345,415,448,475]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="21" pageNumber="32" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1-2 m</quantity>
above high tide line in organic mat on sand and gravel. The
<collectingRegion box="[140,213,483,509]" country="Canada" name="Yukon" pageId="21" pageNumber="32">Yukon</collectingRegion>
and
<collectingRegion box="[274,518,483,509]" country="Canada" name="Northwest Territories" pageId="21" pageNumber="32">Northwest Territories</collectingRegion>
specimens were captured at altitudes ranging from
<quantity box="[140,301,518,545]" metricMagnitude="3" metricUnit="m" metricValue="1.1425" metricValueMax="1.28" metricValueMin="1.005" pageId="21" pageNumber="32" unit="m" value="1142.5" valueMax="1280.0" valueMin="1005.0">1005-1280 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>