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<document ID-DOI="http://doi.org/10.5281/zenodo.3792786" ID-GBIF-Dataset="142a25ed-47c3-4361-bc24-6f1b3e8dba5c" ID-GBIF-Taxon="163755054" ID-ISSN="13132970" ID-Zenodo-Dep="3792786" ID-ZooBank="664C49F1-5384-43C4-8BF1-CE76AC11D32E" approvalRequired="10" approvalRequired_for_treatments="10" checkinTime="1587998787238" checkinUser="plazi" docAuthor="Klimaszewski, Jan, Savard, Karine, Pelletier, Georges &amp; Webster, Reginald" docDate="2008" docId="D43E034BFF88417BFF17FDB7655EFC48" docLanguage="en" docName="ZK_article_1943.pdf" docOrigin="ZooKeys 2 (2)" docStyle="DocumentStyle:BEF2356135E79670CA200BCAD40C4E3A:zookeys.2008.journal_article" docStyleId="BEF2356135E79670CA200BCAD40C4E3A" docStyleName="zookeys.2008.journal_article" docTitle="Gnypeta atrolucens Casey 1893" docType="treatment" docVersion="15" lastPageId="23" lastPageNumber="34" masterDocId="28077B33FF9D416CFF9BFFC76718FFEC" masterDocTitle="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution" masterLastPageNumber="84" masterPageNumber="11" pageId="21" pageNumber="32" updateTime="1643574247635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Klimaszewski, Jan</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Savard, Karine</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pelletier, Georges</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Webster, Reginald</mods:namePart>
<mods:affiliation>24 Millstream Drive, Fredericton, NB, Canada &amp; F-</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>11</mods:start>
<mods:end>84</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.2.4</mods:identifier>
<mods:identifier type="GBIF-Dataset">142a25ed-47c3-4361-bc24-6f1b3e8dba5c</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576410</mods:identifier>
<mods:identifier type="ZooBank">664C49F1-5384-43C4-8BF1-CE76AC11D32E</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792786" ID-GBIF-Taxon="163755054" ID-Zenodo-Dep="3792786" LSID="urn:lsid:plazi:treatment:D43E034BFF88417BFF17FDB7655EFC48" httpUri="http://treatment.plazi.org/id/D43E034BFF88417BFF17FDB7655EFC48" lastPageId="23" lastPageNumber="34" pageId="21" pageNumber="32">
<subSubSection box="[140,476,623,650]" pageId="21" pageNumber="32" type="nomenclature">
<paragraph blockId="21.[140,476,623,686]" box="[140,476,623,650]" pageId="21" pageNumber="32">
<heading box="[140,476,623,650]" fontSize="11" level="2" pageId="21" pageNumber="32" reason="3">
<emphasis bold="true" box="[140,476,623,650]" pageId="21" pageNumber="32">
3.
<taxonomicName authority="Casey" authorityName="Casey" authorityYear="1893" box="[170,476,623,650]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="32" phylum="Arthropoda" rank="species" species="atrolucens">
<emphasis bold="true" box="[170,400,623,650]" italics="true" pageId="21" pageNumber="32">Gnypeta atrolucens</emphasis>
Casey
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,458,659,686]" pageId="21" pageNumber="32" type="description">
<paragraph blockId="21.[140,476,623,686]" box="[140,458,659,686]" pageId="21" pageNumber="32">
(
<figureCitation box="[147,217,659,686]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="21" pageNumber="32">Figs 5</figureCitation>
,
<figureCitation box="[231,311,659,686]" captionStart="Figs 21-38" captionStart-0="Figs 21-38" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStartId="10.[140,184,1462,1484]" captionStartId-0="10.[140,184,1462,1484]" captionStartId-1="60.[140,174,1590,1612]" captionStartId-2="60.[140,174,1622,1644]" captionStartId-3="60.[140,174,1654,1676]" captionStartId-4="61.[140,174,883,905]" captionStartId-5="61.[140,174,915,937]" captionStartId-6="61.[140,174,947,969]" captionStartId-7="61.[140,174,979,1001]" captionTargetBox="[230,1007,164,1440]" captionTargetBox-0="[230,1007,164,1440]" captionTargetBox-1="[140,1106,794,1132]" captionTargetBox-2="[140,1101,1127,1467]" captionTargetBox-3="[140,1101,1127,1467]" captionTargetBox-4="[140,1106,165,502]" captionTargetBox-5="[140,1106,165,502]" captionTargetBox-6="[140,1106,515,855]" captionTargetBox-7="[140,1106,515,855]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetId-0="figure@10.[176,1072,134,1444]" captionTargetId-1="figure@60.[140,1108,447,1475]" captionTargetPageId="10" captionTargetPageId-0="10" captionTargetPageId-1="60" captionTargetPageId-2="60" captionTargetPageId-3="60" captionTargetPageId-4="61" captionTargetPageId-5="61" captionTargetPageId-6="61" captionTargetPageId-7="61" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" captionText-0="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" captionText-1="Fig. 193. Collection localities in Canada and Alaska of G. minuta and G. sellmani." captionText-2="Fig. 194. Collection localities in Canada of G. crebrepunctata, G. dentata, and G. helenae." captionText-3="Fig. 195. Collection localities in Canada and Alaska of G. groenlandica." captionText-4="Fig. 196. Collection localities in Canada of G. brevicornis and G. uteana." captionText-5="Fig. 197. Collection localities in Canada and Alaska of G. caerulea." captionText-6="Fig. 198. Collection localities in Canada and Alaska of G. lohsei and G. canadiensis." captionText-7="Fig. 199. Collection localities in Canada and Alaska of G. carbonaria and G. saccharina." figureDoi="http://doi.org/10.5281/zenodo.3774059" figureDoi-0="http://doi.org/10.5281/zenodo.3774059" figureDoi-1="http://doi.org/10.5281/zenodo.3774140" figureDoi-2="http://doi.org/10.5281/zenodo.3774142" figureDoi-3="http://doi.org/10.5281/zenodo.3774144" figureDoi-4="http://doi.org/10.5281/zenodo.3774146" figureDoi-5="http://doi.org/10.5281/zenodo.3774148" figureDoi-6="http://doi.org/10.5281/zenodo.3774150" figureDoi-7="http://doi.org/10.5281/zenodo.3774152" httpUri="https://zenodo.org/record/3774059/files/figure.png" httpUri-0="https://zenodo.org/record/3774059/files/figure.png" httpUri-1="https://zenodo.org/record/3774140/files/figure.png" httpUri-2="https://zenodo.org/record/3774142/files/figure.png" httpUri-3="https://zenodo.org/record/3774144/files/figure.png" httpUri-4="https://zenodo.org/record/3774146/files/figure.png" httpUri-5="https://zenodo.org/record/3774148/files/figure.png" httpUri-6="https://zenodo.org/record/3774150/files/figure.png" httpUri-7="https://zenodo.org/record/3774152/files/figure.png" pageId="21" pageNumber="32">23 a, b</figureCitation>
,
<figureCitation box="[323,394,659,686]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="21" pageNumber="32">57-65</figureCitation>
,
<figureCitation box="[406,451,659,685]" captionStart="Fig" captionStartId="60.[140,174,1558,1580]" captionTargetBox="[140,1106,794,1132]" captionTargetId="figure@60.[140,1108,447,1475]" captionTargetPageId="60" captionText="Fig. 192. Collection localities in Canada of G. atrolucens and G. nigrella." figureDoi="http://doi.org/10.5281/zenodo.3774138" httpUri="https://zenodo.org/record/3774138/files/figure.png" pageId="21" pageNumber="32">192</figureCitation>
)
</paragraph>
</subSubSection>
<paragraph blockId="21.[140,1108,729,967]" pageId="21" pageNumber="32">
<subSubSection box="[140,842,729,756]" pageId="21" pageNumber="32" type="reference_group">
<bibRefCitation author="Casey" box="[140,339,730,756]" firstAuthor="Casey" page="346" pageId="21" pageNumber="32" pagination="281 - 606" refId="ref23129" refString="Casey TL (1894) Coleopterological notices. V. Annals of the New York Academy of Sciences 7: 281 - 606." type="journal article" year="1894">Casey 1894: 346</bibRefCitation>
,
<bibRefCitation author="Casey" box="[353,475,730,756]" firstAuthor="Casey" page="197" pageId="21" pageNumber="32" pagination="125 - 434" refId="ref23153" refString="Casey TL (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis 16: 125 - 434." type="journal article" year="1906">1906: 197</bibRefCitation>
;
<bibRefCitation author="Moore and Legner" box="[490,839,729,756]" firstAuthor="Moore" page="421" pageId="21" pageNumber="32" pagination="1 - 514" refId="ref23790" refString="Moore I., Legner EF (1975) A catalogue of the Staphylinidae of America north of Mexico (Coleoptera). University of California Division of Agricultural Sciences. Special Publication No. 3015: 1 - 514." type="book chapter" year="1975">Moore and Legner 1975: 421</bibRefCitation>
.
</subSubSection>
<subSubSection pageId="21" pageNumber="32" type="materials_examined">
<materialsCitation ID-GBIF-Occurrence="2609050478" pageId="21" pageNumber="32">
<emphasis bold="true" box="[852,1018,729,755]" pageId="21" pageNumber="32">
<typeStatus box="[852,1018,729,755]" pageId="21" pageNumber="32">LECTOTYPE</typeStatus>
</emphasis>
(male):
<collectingCountry box="[185,410,765,791]" name="United States of America" pageId="21" pageNumber="32">UNITED STATES</collectingCountry>
:
<collectingRegion box="[421,534,765,791]" country="United States of America" name="New York" pageId="21" pageNumber="32">New York</collectingRegion>
[in original description],
<taxonomicName authority="Casey" authorityName="Casey" authorityYear="1893" box="[828,1107,765,791]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="32" phylum="Arthropoda" rank="species" species="atrolucens">
<emphasis box="[828,1033,765,791]" italics="true" pageId="21" pageNumber="32">Gnypeta atrolucens</emphasis>
Casey
</taxonomicName>
[Caseys handwriting]; Type USNM 38867; Casey bequest 1925;
<taxonomicName authorityName="Casey" authorityYear="1893" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="32" phylum="Arthropoda" rank="species" species="atrolucens">
<emphasis italics="true" pageId="21" pageNumber="32">Gnypeta atrolucens</emphasis>
</taxonomicName>
<typeStatus box="[239,363,836,862]" pageId="21" pageNumber="32">Lectotypus</typeStatus>
Lohse des. 1989 [designation not published]; present
<typeStatus box="[1005,1107,835,861]" pageId="21" pageNumber="32">lectotype</typeStatus>
designation label by Klimaszewski 2008 (USNM). Examined.
</materialsCitation>
</subSubSection>
</paragraph>
<subSubSection pageId="21" pageNumber="32" type="materials_examined">
<paragraph blockId="21.[140,1108,729,967]" box="[140,363,905,931]" pageId="21" pageNumber="32">
<emphasis bold="true" box="[140,363,905,931]" pageId="21" pageNumber="32">
<emphasis box="[140,241,905,931]" inLineHeading="true" pageId="21" pageNumber="32" reason="1">Material</emphasis>
examined
</emphasis>
</paragraph>
<paragraph blockId="21.[140,1108,729,967]" box="[185,592,941,967]" pageId="21" pageNumber="32">Specimens are listed in Appendix A.</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="32" type="diagnosis">
<paragraph blockId="21.[140,1108,1011,1285]" box="[140,259,1011,1037]" pageId="21" pageNumber="32">
<emphasis bold="true" box="[140,259,1011,1037]" inLineHeading="true" pageId="21" pageNumber="32" reason="1">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="21.[140,1108,1011,1285]" pageId="21" pageNumber="32">
This species can be recognized by the following combination of characters: body length 3.2-3.4 mm (
<figureCitation box="[291,358,1081,1108]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="21" pageNumber="32">Fig. 5</figureCitation>
); elytra (at suture) about 1/5 longer and 1/3 wider than maximum width of pronotum (
<figureCitation box="[376,444,1117,1144]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="21" pageNumber="32">Fig. 5</figureCitation>
); abdomen at base distinctly narrower than elytra (
<figureCitation box="[1024,1092,1117,1144]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="21" pageNumber="32">Fig. 5</figureCitation>
); antennal articles 4-7 strongly elongate (
<figureCitation box="[582,711,1152,1179]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="21" pageNumber="32">Fig. 23 a, b</figureCitation>
); median lobe of aedeagus with triangular and constricted subapically in lateral view (
<figureCitation box="[716,799,1187,1214]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="21" pageNumber="32">Fig. 57</figureCitation>
); spermatheca with capsule longer than wide and funnel-shaped (
<figureCitation box="[566,650,1222,1249]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="21" pageNumber="32">Fig. 63</figureCitation>
), stem approximately straight or slightly curved and slightly swollen basally (
<figureCitation box="[544,629,1258,1285]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="21" pageNumber="32">Fig. 63</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="34" pageId="21" pageNumber="32" type="description">
<paragraph blockId="21.[140,1108,1328,1672]" box="[140,282,1328,1354]" pageId="21" pageNumber="32">
<emphasis bold="true" box="[140,282,1328,1354]" inLineHeading="true" pageId="21" pageNumber="32" reason="1">Description</emphasis>
</paragraph>
<paragraph blockId="21.[140,1108,1328,1672]" lastBlockId="23.[140,1108,166,475]" lastPageId="23" lastPageNumber="34" pageId="21" pageNumber="32">
Body length 3.2-3.4 mm; dark brown to black, central part of elytra and tarsi or entire legs reddish brown (
<figureCitation box="[445,514,1398,1425]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="21" pageNumber="32">Fig. 5</figureCitation>
); integument strongly glossy; pubescence yellowish grey and moderately long and dense; antennal article 4 slightly shorter than preceding article, 5-7 elongate, 8-9 subquadrate or slightly elongate (
<figureCitation box="[878,1021,1469,1496]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="21" pageNumber="32">Figs 23 a, b</figureCitation>
); head, pronotum and abdomen of about the same width, elytra large and distinctly broader than remainder of body (
<figureCitation box="[470,542,1539,1566]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="21" pageNumber="32">Fig. 5</figureCitation>
); head rounded posteriorly; pronotum broadest in apical third, pubescence directed anterad along midline and laterad elsewhere; elytra (at suture) about 1/5 longer and 1/3 broader than maximum width of pronotum (
<figureCitation box="[236,307,1645,1672]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="21" pageNumber="32">Fig. 5</figureCitation>
), pubescence directed obliquely postero-laterad; abdomen subparallel; metatarsus with two basal articles of about the same length and the third one slightly shorter.
<emphasis bold="true" box="[330,392,202,228]" pageId="23" pageNumber="34">Male</emphasis>
. Tergite 8 transverse and truncate apically (
<figureCitation box="[906,995,201,228]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 61</figureCitation>
). Sternite 8 elongate and pointing medially (
<figureCitation box="[545,633,237,264]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 62</figureCitation>
). Median lobe of aedeagus with triangular and produced apical part of tubus in lateral view (
<figureCitation box="[762,849,272,299]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 57</figureCitation>
); bulbus enlarged and bearing two angular anterior projections in dorsal view (
<figureCitation box="[803,890,307,334]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 58</figureCitation>
); internal sac with structures as illustrated (
<figureCitation box="[422,557,342,369]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Figs 58, 59</figureCitation>
).
<emphasis bold="true" box="[578,665,343,369]" pageId="23" pageNumber="34">Female</emphasis>
. Tergite 8 similar to that of male (
<figureCitation captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 64</figureCitation>
). Sternite 8 broadly rounded apically (
<figureCitation box="[619,706,377,404]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 65</figureCitation>
). Spermatheca with capsule longer than wide and funnel-shaped (
<figureCitation box="[507,596,413,440]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 63</figureCitation>
), stem slightly curved and slightly swollen basally (
<figureCitation box="[234,322,448,475]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="23" pageNumber="34">Fig. 63</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774100" ID-Zenodo-Dep="3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="22" pageNumber="33" startId="22.[140,184,1590,1612]" targetBox="[152,1098,247,1512]" targetPageId="22">
<paragraph blockId="22.[140,1108,1590,1676]" pageId="22" pageNumber="33">
<emphasis bold="true" box="[140,261,1590,1612]" pageId="22" pageNumber="33">Figs 57-65.</emphasis>
Genital structures of
<taxonomicName authorityName="Casey" authorityYear="1893" box="[470,640,1590,1612]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="species" species="atrolucens">
<emphasis box="[470,640,1590,1612]" italics="true" pageId="22" pageNumber="33">Gnypeta atrolucens</emphasis>
</taxonomicName>
: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8.
</paragraph>
</caption>
<subSubSection box="[140,411,518,545]" pageId="23" pageNumber="34" type="distribution">
<paragraph blockId="23.[140,1108,518,650]" box="[140,411,518,545]" pageId="23" pageNumber="34">
<emphasis bold="true" box="[140,291,518,544]" inLineHeading="true" pageId="23" pageNumber="34" reason="1">Distribution</emphasis>
(Fig. 192)
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="34" type="reference_group">
<paragraph blockId="23.[140,1108,518,650]" pageId="23" pageNumber="34">
<taxonomicName authorityName="Casey" authorityYear="1893" box="[140,347,554,580]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="34" phylum="Arthropoda" rank="species" species="atrolucens">
<emphasis box="[140,347,554,580]" italics="true" pageId="23" pageNumber="34">Gnypeta atrolucens</emphasis>
</taxonomicName>
was described from New York (
<bibRefCitation author="Casey" box="[723,864,554,580]" firstAuthor="Casey" pageId="23" pageNumber="34" pagination="281 - 606" refId="ref23129" refString="Casey TL (1894) Coleopterological notices. V. Annals of the New York Academy of Sciences 7: 281 - 606." type="journal article" year="1894">Casey 1894</bibRefCitation>
), and it was later reported from the Catskill Mts., New York (
<bibRefCitation author="Casey" box="[611,806,589,615]" firstAuthor="Casey" page="197" pageId="23" pageNumber="34" pagination="125 - 434" refId="ref23153" refString="Casey TL (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis 16: 125 - 434." type="journal article" year="1906">Casey 1906: 197</bibRefCitation>
). We report this species for the first time from
<collectingCountry box="[355,441,624,650]" name="Canada" pageId="23" pageNumber="34">Canada</collectingCountry>
in
<collectingRegion box="[479,567,624,650]" country="Canada" name="Quebec" pageId="23" pageNumber="34">Quebec</collectingRegion>
and southeastern Labrador.
</paragraph>
<paragraph blockId="23.[140,1059,694,756]" box="[140,467,694,721]" pageId="23" pageNumber="34">
<emphasis bold="true" box="[140,467,694,721]" pageId="23" pageNumber="34">Collection and habitat data</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[140,1059,729,756]" pageId="23" pageNumber="34" type="description">
<paragraph blockId="23.[140,1059,694,756]" box="[140,1059,729,756]" pageId="23" pageNumber="34">
Adults were captured from July through August at altitudes from
<quantity box="[879,939,729,756]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="23" pageNumber="34" unit="m" value="61.0">61 m</quantity>
to
<specimenCount box="[977,1059,729,756]" pageId="23" pageNumber="34" type="generic">
<quantity box="[977,1055,729,756]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="23" pageNumber="34" unit="m" value="853.0">853 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="34" type="discussion">
<paragraph blockId="23.[140,1108,800,932]" box="[140,271,800,826]" pageId="23" pageNumber="34">
<emphasis bold="true" box="[140,271,800,826]" inLineHeading="true" pageId="23" pageNumber="34" reason="1">Comments</emphasis>
</paragraph>
<paragraph blockId="23.[140,1108,800,932]" pageId="23" pageNumber="34">
In the original description,
<bibRefCitation author="Casey" box="[451,602,835,861]" firstAuthor="Casey" pageId="23" pageNumber="34" pagination="281 - 606" refId="ref23129" refString="Casey TL (1894) Coleopterological notices. V. Annals of the New York Academy of Sciences 7: 281 - 606." type="journal article" year="1894">Casey (1894)</bibRefCitation>
did not specify the number of specimens on which the description was based and therefore we have designated the only male in Caseys type collection as the
<typeStatus box="[471,577,906,932]" pageId="23" pageNumber="34">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>