453 lines
53 KiB
XML
453 lines
53 KiB
XML
<document id="CDDC77BD6AFC71620EC1B11A5AE417CC" ID-DOI="10.5281/zenodo.5354097" ID-ISSN="1942-1354" ID-Zenodo-Dep="5354097" ID-ZooBank="44CB9DF0-0165-4365-AACF-BF4DE1842503" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212997409" checkinUser="felipe" docAuthor="Schnepp, Kyle E. & Ashman, Krystal L." docDate="2020" docId="A02CFF4AFFB8FFE5FF59F9E2FA001D10" docLanguage="en" docName="0841.pdf" docOrigin="Insecta Mundi 2020 (841)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Ataenius thomasi Schnepp and Ashman 2020, new species" docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="5C158732FFBBFFEDFF99FFB6FF9D1E09" masterDocTitle="A new species of Ataenius Harold (Coleoptera: Scarabaeidae: Aphodiinae) from the southeastern United States, with a lectotype designation" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1699188182883" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="8C421CA2D3A2AB53AB27470A0B49EF66">A new species of Ataenius Harold (Coleoptera: Scarabaeidae: Aphodiinae) from the southeastern United States, with a lectotype designation</mods:title>
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<mods:name id="57258C8D28482981D786FACED718A08B" type="personal">
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<mods:roleTerm id="76A8035854FA9E5F12D0C8A1A0179A5F">Author</mods:roleTerm>
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<mods:namePart id="FD1ECAADBBB3EEEFD241044BF194C766">Schnepp, Kyle E.</mods:namePart>
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<mods:affiliation id="DEC0A5727EDB1755FBA7F221F7095A00">Florida State Collection of Arthropods Division of Plant Industry, Florida Department of Agriculture and Consumer Services 1911 SW 34 th Street Gainesville, FL 32608, USA</mods:affiliation>
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<mods:roleTerm id="87657AFFB70CF1EA16EF00DA0C2E7F7A">Author</mods:roleTerm>
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<mods:namePart id="5A1CF55C7FAD32443072A439A37163C7">Ashman, Krystal L.</mods:namePart>
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<mods:affiliation id="21246F94093CEBB9A141A8549720326B">Florida State Collection of Arthropods Division of Plant Industry, Florida Department of Agriculture and Consumer Services 1911 SW 34 th Street Gainesville, FL 32608, USA</mods:affiliation>
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<mods:typeOfResource id="7CA28DCB60F275EE9B30F3158947A59D">text</mods:typeOfResource>
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<mods:title id="4CC3B5255ADE1128EF36142F6C91DE5E">Insecta Mundi</mods:title>
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<mods:part id="53FC77756BC2ABCFC0E102A33CD246A0">
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<mods:date id="A8F1A57CECD0D14386C149A19678DC3C">2020</mods:date>
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<mods:number id="3F5516114636AA5A75EDA7B704CFC891">2020-12-25</mods:number>
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<mods:number id="69046771FDEC03195F1C40DD2AFE1AAC">2020</mods:number>
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<mods:number id="F6D457683850A5A2B423E33DA064E05B">841</mods:number>
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<mods:identifier id="6268E205C02FB15F26BE3EB1E6543C2B" type="DOI">10.5281/zenodo.5354097</mods:identifier>
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<mods:identifier id="E137312BE30DD5A0700660D75A40F071" type="ISSN">1942-1354</mods:identifier>
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<treatment id="A02CFF4AFFB8FFE5FF59F9E2FA001D10" LSID="urn:lsid:plazi:treatment:A02CFF4AFFB8FFE5FF59F9E2FA001D10" httpUri="http://treatment.plazi.org/id/A02CFF4AFFB8FFE5FF59F9E2FA001D10" lastPageId="8" lastPageNumber="7" pageId="3" pageNumber="2">
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<subSubSection id="609F1DD7FFB8FFEEFF59F9E2FCC11879" box="[192,860,1619,1648]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="283A4E5CFFB8FFEEFF59F9E2FCC11879" blockId="3.[192,860,1619,1681]" box="[192,860,1619,1648]" pageId="3" pageNumber="2">
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<heading id="7372F930FFB8FFEEFF59F9E2FCC11879" box="[192,860,1619,1648]" fontSize="12" level="2" pageId="3" pageNumber="2" reason="2">
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<emphasis id="1AF1924EFFB8FFEEFF59F9E2FCC11879" bold="true" box="[192,860,1619,1648]" pageId="3" pageNumber="2">
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<taxonomicName id="EF8535DFFFB8FFEEFF59F9E2FD271879" authority="Schnepp and Ashman" authorityName="Schnepp and Ashman" authorityYear="2020" box="[192,698,1619,1648]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
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<emphasis id="1AF1924EFFB8FFEEFF59F9E2FE061879" bold="true" box="[192,411,1619,1648]" italics="true" pageId="3" pageNumber="2">Ataenius thomasi</emphasis>
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Schnepp and Ashman
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</taxonomicName>
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,
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<taxonomicNameLabel id="01C22F35FFB8FFEEFD5EF9E2FCC11879" box="[711,860,1619,1648]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="609F1DD7FFB8FFEEFF59F9C1FEE31898" box="[192,382,1655,1681]" pageId="3" pageNumber="2" type="description">
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<paragraph id="283A4E5CFFB8FFEEFF59F9C1FEE31898" blockId="3.[192,860,1619,1681]" box="[192,382,1655,1681]" pageId="3" pageNumber="2">
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<figureCitation id="B0BE52D9FFB8FFEEFF59F9C1FECC1898" box="[192,337,1655,1681]" captionStart="Figures 7–12" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[222,1405,223,1802]" captionTargetId="figure-57@5.[219,1414,213,1804]" captionTargetPageId="5" captionText="Figures 7–12. Ataenius thomasi Schnepp and Ashman. 7) Dorsal view of holotype. 8) Ventral view of holotype. 9) Frontal view of holotype. 10) Holotype labels. 11) Ventral view of metatibia of holotype, arrow indicates reduced accessory spine. 12) Male genitalia of paratype." figureDoi="http://doi.org/10.5281/zenodo.5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="3" pageNumber="2">Figures 7–12</figureCitation>
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,
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<figureCitation id="B0BE52D9FFB8FFEEFEC5F9C1FEE41898" box="[348,377,1655,1681]" captionStart="Figures 13–14" captionStartId="6.[192,279,1864,1890]" captionTargetBox="[231,1403,217,1830]" captionTargetId="figure-40@6.[216,1416,210,1836]" captionTargetPageId="6" captionText="Figures 13–14. Distribution of Ataenius brevis and Ataenius thomasi in the southeastern United States. 13) Ataenius brevis Fall. 14) Ataenius thomasi, new species." figureDoi="http://doi.org/10.5281/zenodo.5354103" httpUri="https://zenodo.org/record/5354103/files/figure.png" pageId="3" pageNumber="2">14</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="609F1DD7FFB8FFEAFF59F917FDFA1F18" lastPageId="7" lastPageNumber="6" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="283A4E5CFFB8FFEEFF59F917FB9D18D4" blockId="3.[191,1441,1697,1927]" pageId="3" pageNumber="2">
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<emphasis id="1AF1924EFFB8FFEEFF59F917FEF418B2" bold="true" box="[192,361,1697,1723]" pageId="3" pageNumber="2">Type material.</emphasis>
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<materialsCitation id="98ED4401FFB8FFEEFEF7F917FB9D18D4" collectingDate="1977-03-28" collectionCode="R, FSCA" county="Fla. Alachua Co." location="Fla. Alachua Co." pageId="3" pageNumber="2" specimenCount="2" typeStatus="holotype">
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<emphasis id="1AF1924EFFB8FFEEFEF7F917FE4618B2" bold="true" box="[366,475,1697,1723]" pageId="3" pageNumber="2">
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<typeStatus id="F73EF0FEFFB8FFEEFEF7F917FE4618B2" box="[366,475,1697,1723]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
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</emphasis>
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(
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<figureCitation id="B0BE52D9FFB8FFEEFE70F917FDF318B2" box="[489,622,1697,1723]" captionStart="Figures 7–12" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[222,1405,223,1802]" captionTargetId="figure-57@5.[219,1414,213,1804]" captionTargetPageId="5" captionText="Figures 7–12. Ataenius thomasi Schnepp and Ashman. 7) Dorsal view of holotype. 8) Ventral view of holotype. 9) Frontal view of holotype. 10) Holotype labels. 11) Ventral view of metatibia of holotype, arrow indicates reduced accessory spine. 12) Male genitalia of paratype." figureDoi="http://doi.org/10.5281/zenodo.5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="3" pageNumber="2">Fig. 7–9, 11</figureCitation>
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): “
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<collectingCounty id="C15B36D0FFB8FFEEFD15F917FCDB18B2" box="[652,838,1697,1723]" pageId="3" pageNumber="2">Fla. Alachua Co.</collectingCounty>
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/ NE Gainesville /
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<date id="5C3B689CFFB8FFEEFB8EF917FB3518B2" box="[1047,1192,1697,1724]" pageId="3" pageNumber="2" value="1977-03-28">
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<collectingDate id="4C7F9174FFB8FFEEFB8EF917FB3518B2" box="[1047,1192,1697,1724]" pageId="3" pageNumber="2" value="1977-03-28">28 Mar. 1977</collectingDate>
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</date>
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/ L.
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<collectionCode id="4E94D699FFB8FFEEFB40F914FB7318B5" box="[1241,1262,1698,1724]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="2">R</collectionCode>
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. Davis, Jr. // [on red]
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<typeStatus id="F73EF0FEFFB8FFEEFF6CF975FE1B18D4" box="[245,390,1731,1757]" pageId="3" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="EF8535DFFFB8FFEEFE05F975FDC618D4" box="[412,603,1731,1757]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thomasi">
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<emphasis id="1AF1924EFFB8FFEEFE05F975FDC618D4" box="[412,603,1731,1757]" italics="true" pageId="3" pageNumber="2">Ataenius thomasi</emphasis>
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</taxonomicName>
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/ Schnepp & Ashman, 2020” (
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<collectionCode id="4E94D699FFB8FFEEFC32F975FC6918D4" box="[939,1012,1731,1757]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="283A4E5CFFB8FFEEFE99F953FAE1198F" blockId="3.[191,1441,1697,1927]" pageId="3" pageNumber="2">
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<materialsCitation id="98ED4401FFB8FFEEFE99F953FAA318F6" box="[256,1342,1765,1791]" collectingDate="1977-03-28" collectionCode="R, FSCA" county="Fla. Alachua Co." location="Gainesville" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
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<emphasis id="1AF1924EFFB8FFEEFE99F953FEE718F6" bold="true" box="[256,378,1765,1791]" pageId="3" pageNumber="2">
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<typeStatus id="F73EF0FEFFB8FFEEFE99F953FEE818F6" box="[256,373,1765,1791]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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“
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<collectingCounty id="C15B36D0FFB8FFEEFE13F953FDD918F6" box="[394,580,1765,1791]" pageId="3" pageNumber="2">Fla. Alachua Co.</collectingCounty>
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/ NE
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<location id="2D5A1887FFB8FFEEFD1FF953FC9F18F6" LSID="urn:lsid:plazi:treatment:A02CFF4AFFB8FFE5FF59F9E2FA001D10:2D5A1887FFB8FFEEFD1FF953FC9F18F6" box="[646,770,1765,1791]" county="Fla. Alachua Co." name="Gainesville" pageId="3" pageNumber="2">Gainesville</location>
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/
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<date id="5C3B689CFFB8FFEEFC81F953FC3718F6" box="[792,938,1765,1791]" pageId="3" pageNumber="2" value="1977-03-28">
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<collectingDate id="4C7F9174FFB8FFEEFC81F953FC3718F6" box="[792,938,1765,1791]" pageId="3" pageNumber="2" value="1977-03-28">28 Mar. 1977</collectingDate>
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</date>
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||
/ L.
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||
<collectionCode id="4E94D699FFB8FFEEFC45F953FC6C18F6" box="[988,1009,1765,1791]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="2">R</collectionCode>
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. Davis, Jr.” (2
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<collectionCode id="4E94D699FFB8FFEEFB13F953FB5218F6" box="[1162,1231,1765,1791]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
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) (
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<figureCitation id="B0BE52D9FFB8FFEEFB7CF953FAA918F6" box="[1253,1332,1765,1791]" captionStart="Figures 7–12" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[222,1405,223,1802]" captionTargetId="figure-57@5.[219,1414,213,1804]" captionTargetPageId="5" captionText="Figures 7–12. Ataenius thomasi Schnepp and Ashman. 7) Dorsal view of holotype. 8) Ventral view of holotype. 9) Frontal view of holotype. 10) Holotype labels. 11) Ventral view of metatibia of holotype, arrow indicates reduced accessory spine. 12) Male genitalia of paratype." figureDoi="http://doi.org/10.5281/zenodo.5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="3" pageNumber="2">Fig. 12</figureCitation>
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)
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</materialsCitation>
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;
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<materialsCitation id="98ED4401FFB8FFEEFAD0F953FB1C1928" collectingDate="1977-03-25" collectionCode="R, FSCA" county="Alachua Co." location="Gainesville" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
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||
“FLOR- IDA:
|
||
<collectingCounty id="C15B36D0FFB8FFEEFF64F8B1FE151928" box="[253,392,1799,1825]" pageId="3" pageNumber="2">Alachua Co.</collectingCounty>
|
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, /
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<location id="2D5A1887FFB8FFEEFE3BF8B1FD831928" LSID="urn:lsid:plazi:treatment:A02CFF4AFFB8FFE5FF59F9E2FA001D10:2D5A1887FFB8FFEEFE3BF8B1FD831928" box="[418,542,1799,1825]" county="Alachua Co." name="Gainesville" pageId="3" pageNumber="2">Gainesville</location>
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/ [handwritten]
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<date id="5C3B689CFFB8FFEEFD4FF8B0FCCA1928" box="[726,855,1798,1825]" pageId="3" pageNumber="2" value="1977-03-25">
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<collectingDate id="4C7F9174FFB8FFEEFD4FF8B0FCCA1928" box="[726,855,1798,1825]" pageId="3" pageNumber="2" value="1977-03-25">25-III-1977</collectingDate>
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</date>
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/ L.
|
||
<collectionCode id="4E94D699FFB8FFEEFC1EF8B1FC011928" box="[903,924,1799,1825]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="2">R</collectionCode>
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. Davis, Jr.” (1
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<collectionCode id="4E94D699FFB8FFEEFBAAF8B1FBE71928" box="[1075,1146,1799,1825]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
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)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="98ED4401FFB8FFEEFB12F8B1FCE5194A" collectionCode="R, FSCA" county="Baker County" location="St. Mary's" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
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“FLA.
|
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<collectingCounty id="C15B36D0FFB8FFEEFB4DF8B1FAF21928" box="[1236,1391,1799,1825]" pageId="3" pageNumber="2">Baker County</collectingCounty>
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/
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<location id="2D5A1887FFB8FFEEFA1DF8B1FE97194A" LSID="urn:lsid:plazi:treatment:A02CFF4AFFB8FFE5FF59F9E2FA001D10:2D5A1887FFB8FFEEFA1DF8B1FE97194A" county="Baker County" name="St. Mary's" pageId="3" pageNumber="2">St. Mary’s</location>
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<collectionCode id="4E94D699FFB8FFEEFE89F89FFEB8194A" box="[272,293,1833,1859]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="2">R</collectionCode>
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. at / Fla. Hgw. 23 #484 / xii.30.47 FNYoung” (1
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<collectionCode id="4E94D699FFB8FFEEFCB3F89FFCEC194A" box="[810,881,1833,1859]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
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)
|
||
</materialsCitation>
|
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;
|
||
<materialsCitation id="98ED4401FFB8FFEEFC1BF89FFE8E196D" collectionCode="R, SEMC" collectorName="H. Beamer" location="Orange Grove" municipality="Miss" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
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“
|
||
<location id="2D5A1887FFB8FFEEFC12F89FFBB5194A" LSID="urn:lsid:plazi:treatment:A02CFF4AFFB8FFE5FF59F9E2FA001D10:2D5A1887FFB8FFEEFC12F89FFBB5194A" box="[907,1064,1833,1859]" municipality="Miss" name="Orange Grove" pageId="3" pageNumber="2">Orange Grove</location>
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/
|
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<collectingMunicipality id="C85ED426FFB8FFEEFBA4F89EFBEF194B" box="[1085,1138,1832,1858]" pageId="3" pageNumber="2">Miss</collectingMunicipality>
|
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7-10-34 /
|
||
<collectionCode id="4E94D699FFB8FFEEFB7AF89FFB65194A" box="[1251,1272,1833,1859]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="2">R</collectionCode>
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.
|
||
<collectorName id="85702B8AFFB8FFEEFA99F89FFAE7194A" box="[1280,1402,1833,1859]" pageId="3" pageNumber="2">H. Beamer</collectorName>
|
||
” (4
|
||
<collectionCode id="4E94D699FFB8FFEEFF26F8FCFE91196D" box="[191,268,1866,1892]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="3" pageNumber="2" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="98ED4401FFB8FFEEFE87F8FCFAE5198F" collectingDate="1912-06" collectionCode="USNM" location="Billy's Island" municipality="Okefenokee Swamp" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
||
“
|
||
<location id="2D5A1887FFB8FFEEFEBEF8FCFE29196D" LSID="urn:lsid:plazi:treatment:A02CFF4AFFB8FFE5FF59F9E2FA001D10:2D5A1887FFB8FFEEFEBEF8FCFE29196D" box="[295,436,1866,1892]" municipality="Okefenokee Swamp" name="Billy's Island" pageId="3" pageNumber="2">Billy’s Island</location>
|
||
/
|
||
<collectingMunicipality id="C85ED426FFB8FFEEFE53F8FCFD3A196D" box="[458,679,1866,1892]" pageId="3" pageNumber="2">Okefenokee Swamp</collectingMunicipality>
|
||
/
|
||
<date id="5C3B689CFFB8FFEEFD24F8FDFCB2196D" box="[701,815,1866,1893]" pageId="3" pageNumber="2" value="1912-06">
|
||
<collectingDate id="4C7F9174FFB8FFEEFD24F8FDFCB2196D" box="[701,815,1866,1893]" pageId="3" pageNumber="2" value="1912-06">June 1912</collectingDate>
|
||
</date>
|
||
. Ga. // Cornell U. / Lot. [handwritten] 440 / Sub. [handwritten] 37 // Cornell U. / Lot. 539 / Sub. [handwritten] 459 // OLCartwright / Collection / 1961” (1
|
||
<collectionCode id="4E94D699FFB8FFEEFA85F8DAFAED198F" box="[1308,1392,1900,1926]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="2" type="Museum">USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="7CFA1ED4FFBFFFE9FF59F89FFC9D198E" ID-DOI="http://doi.org/10.5281/zenodo.5354099" ID-Zenodo-Dep="5354099" httpUri="https://zenodo.org/record/5354099/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,279,1833,1859]" targetBox="[216,1417,215,1802]" targetPageId="4">
|
||
<paragraph id="283A4E5CFFBFFFE9FF59F89FFC9D198E" blockId="4.[192,1441,1832,1927]" pageId="4" pageNumber="3">
|
||
<emphasis id="1AF1924EFFBFFFE9FF59F89FFECC194A" bold="true" box="[192,337,1833,1859]" pageId="4" pageNumber="3">Figures 1–6.</emphasis>
|
||
<taxonomicName id="EF8535DFFFBFFFE9FEC0F89FFDA1194A" authority="Fall." authorityName="Fall." box="[345,572,1833,1859]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="1AF1924EFFBFFFE9FEC0F89FFD9E194A" box="[345,515,1833,1859]" italics="true" pageId="4" pageNumber="3">Ataenius brevis</emphasis>
|
||
Fall.
|
||
</taxonomicName>
|
||
<emphasis id="1AF1924EFFBFFFE9FDDFF89EFDC3194B" bold="true" box="[582,606,1832,1858]" pageId="4" pageNumber="3">1)</emphasis>
|
||
Dorsal view of lectotype.
|
||
<emphasis id="1AF1924EFFBFFFE9FC13F89EFC3F194B" bold="true" box="[906,930,1832,1858]" pageId="4" pageNumber="3">2)</emphasis>
|
||
Ventral view of lectotype.
|
||
<emphasis id="1AF1924EFFBFFFE9FB4CF89EFB70194B" bold="true" box="[1237,1261,1832,1858]" pageId="4" pageNumber="3">3)</emphasis>
|
||
Frontal view of lectotype.
|
||
<emphasis id="1AF1924EFFBFFFE9FEABF8FCFED7196D" bold="true" box="[306,330,1866,1892]" pageId="4" pageNumber="3">4)</emphasis>
|
||
Lectotype labels.
|
||
<emphasis id="1AF1924EFFBFFFE9FD96F8FCFDBA196D" bold="true" box="[527,551,1866,1892]" pageId="4" pageNumber="3">5)</emphasis>
|
||
Ventral view of metatibia of lectotype, arrow indicates outlined accessory spine.
|
||
<emphasis id="1AF1924EFFBFFFE9FF59F8DAFF45198F" bold="true" box="[192,216,1900,1926]" pageId="4" pageNumber="3">6)</emphasis>
|
||
Male genitalia, specimen from Citico, Tennessee.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7CFA1ED4FFBEFFE8FF59F89FFC85198E" ID-DOI="http://doi.org/10.5281/zenodo.5354101" ID-Zenodo-Dep="5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,279,1833,1859]" targetBox="[222,1405,223,1802]" targetPageId="5">
|
||
<paragraph id="283A4E5CFFBEFFE8FF59F89FFC85198E" blockId="5.[192,1441,1832,1927]" pageId="5" pageNumber="4">
|
||
<emphasis id="1AF1924EFFBEFFE8FF59F89FFEC0194A" bold="true" box="[192,349,1833,1859]" pageId="5" pageNumber="4">Figures 7–12.</emphasis>
|
||
<taxonomicName id="EF8535DFFFBEFFE8FEFDF89FFCB4194A" authority="Schnepp and Ashman." authorityName="Schnepp and Ashman." box="[356,809,1833,1859]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFBEFFE8FEFDF89FFDB9194A" box="[356,548,1833,1859]" italics="true" pageId="5" pageNumber="4">Ataenius thomasi</emphasis>
|
||
Schnepp and Ashman.
|
||
</taxonomicName>
|
||
<emphasis id="1AF1924EFFBEFFE8FCA9F89EFCD5194B" bold="true" box="[816,840,1832,1858]" pageId="5" pageNumber="4">7)</emphasis>
|
||
Dorsal view of holotype.
|
||
<emphasis id="1AF1924EFFBEFFE8FBF1F89EFB1D194B" bold="true" box="[1128,1152,1832,1858]" pageId="5" pageNumber="4">8)</emphasis>
|
||
Ventral view of holotype.
|
||
<emphasis id="1AF1924EFFBEFFE8FF59F8FCFF45196D" bold="true" box="[192,216,1866,1892]" pageId="5" pageNumber="4">9)</emphasis>
|
||
Frontal view of holotype.
|
||
<emphasis id="1AF1924EFFBEFFE8FD9AF8FCFDB4196D" bold="true" box="[515,553,1866,1892]" pageId="5" pageNumber="4">10)</emphasis>
|
||
Holotype labels.
|
||
<emphasis id="1AF1924EFFBEFFE8FD74F8FCFC89196D" bold="true" box="[749,788,1866,1892]" pageId="5" pageNumber="4">11)</emphasis>
|
||
Ventral view of metatibia of holotype, arrow indicates reduced accessory spine.
|
||
<emphasis id="1AF1924EFFBEFFE8FE5AF8DAFE75198F" bold="true" box="[451,488,1900,1926]" pageId="5" pageNumber="4">12)</emphasis>
|
||
Male genitalia of paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7CFA1ED4FFBDFFEBFF59F8FEFD6E198D" ID-DOI="http://doi.org/10.5281/zenodo.5354103" ID-Zenodo-Dep="5354103" httpUri="https://zenodo.org/record/5354103/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,279,1864,1890]" targetBox="[231,1403,217,1830]" targetPageId="6">
|
||
<paragraph id="283A4E5CFFBDFFEBFF59F8FEFD6E198D" blockId="6.[192,1440,1864,1925]" pageId="6" pageNumber="5">
|
||
<emphasis id="1AF1924EFFBDFFEBFF59F8FEFEF7196A" bold="true" box="[192,362,1864,1891]" pageId="6" pageNumber="5">Figures 13–14.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="EF8535DFFFBDFFEBFD85F8FFFD5E196A" box="[540,707,1865,1891]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="1AF1924EFFBDFFEBFD85F8FFFD5E196A" box="[540,707,1865,1891]" italics="true" pageId="6" pageNumber="5">Ataenius brevis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EF8535DFFFBDFFEBFD60F8FFFC25196A" box="[761,952,1865,1891]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFBDFFEBFD60F8FFFC25196A" box="[761,952,1865,1891]" italics="true" pageId="6" pageNumber="5">Ataenius thomasi</emphasis>
|
||
</taxonomicName>
|
||
in the southeastern United States.
|
||
<emphasis id="1AF1924EFFBDFFEBFAA1F8FEFAC0196B" bold="true" box="[1336,1373,1864,1890]" pageId="6" pageNumber="5">13)</emphasis>
|
||
<taxonomicName id="EF8535DFFFBDFFEBFAFDF8FFFEF6198D" authority="Fall." authorityName="Fall." class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="1AF1924EFFBDFFEBFAFDF8FFFEA8198D" italics="true" pageId="6" pageNumber="5">Ataenius brevis</emphasis>
|
||
Fall.
|
||
</taxonomicName>
|
||
<emphasis id="1AF1924EFFBDFFEBFEE8F8DCFE0B198D" bold="true" box="[369,406,1898,1924]" pageId="6" pageNumber="5">14)</emphasis>
|
||
<taxonomicName id="EF8535DFFFBDFFEBFE04F8DDFDC1198D" authority="Schnepp & Ashman, 2020" authorityName="Schnepp & Ashman" authorityYear="2020" box="[413,604,1898,1925]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
|
||
<emphasis id="1AF1924EFFBDFFEBFE04F8DDFDC1198D" box="[413,604,1898,1925]" italics="true" pageId="6" pageNumber="5">Ataenius thomasi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="01C22F35FFBDFFEBFDF0F8DDFD6D198D" box="[617,752,1898,1925]" pageId="6" pageNumber="5" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="283A4E5CFFBCFFEAFE99FF60FDFA1F18" blockId="7.[192,1442,213,273]" pageId="7" pageNumber="6">
|
||
In addition, all
|
||
<typeStatus id="F73EF0FEFFBCFFEAFE29FF60FD8C1EF9" box="[432,529,214,240]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
specimens have the following bottom label: “[on yellow]
|
||
<typeStatus id="F73EF0FEFFBCFFEAFB06FF60FABB1EF9" box="[1183,1318,214,240]" pageId="7" pageNumber="6" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="EF8535DFFFBCFFEAFAD8FF60FE851F18" authorityName="Schnepp & Ashman" authorityYear="2020" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFBCFFEAFAD8FF60FE851F18" italics="true" pageId="7" pageNumber="6">Ataenius thomasi</emphasis>
|
||
</taxonomicName>
|
||
/ Schnepp & / Ashman 2020”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="609F1DD7FFBCFFEAFF59FE97FEFE1C0D" pageId="7" pageNumber="6" type="diagnosis">
|
||
<paragraph id="283A4E5CFFBCFFEAFF59FE97FEFE1C0D" blockId="7.[192,1441,289,516]" pageId="7" pageNumber="6">
|
||
<emphasis id="1AF1924EFFBCFFEAFF59FE97FEA11F32" bold="true" box="[192,316,289,315]" pageId="7" pageNumber="6">Diagnosis.</emphasis>
|
||
<taxonomicName id="EF8535DFFFBCFFEAFEDFFE97FD951F32" authorityName="Schnepp & Ashman" authorityYear="2020" box="[326,520,289,315]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFBCFFEAFEDFFE97FD951F32" box="[326,520,289,315]" italics="true" pageId="7" pageNumber="6">Ataenius thomasi</emphasis>
|
||
</taxonomicName>
|
||
is distinguishable from other North American
|
||
<taxonomicName id="EF8535DFFFBCFFEAFBB6FE97FB0D1F32" authorityName="Harold" authorityYear="1867" box="[1071,1168,289,315]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1AF1924EFFBCFFEAFBB6FE97FB0D1F32" box="[1071,1168,289,315]" italics="true" pageId="7" pageNumber="6">Ataenius</emphasis>
|
||
</taxonomicName>
|
||
by the combination of: shortened body form; clypeus and frons lacking fine, transverse wrinkles; meso- and metatibiae with apical accessory spine strongly reduced or absent; clypeus lacking teeth; base of head with band of punctures coarser than those on clypeus; scutellar shield lacking punctures or depressions; pronotum apparently lacking marginal setae (possibly abraded in all specimens examined); elytra with striae impressed and distinctly punctate, intervals weakly convex and glossy, with minute punctures; anterolateral margins of pronotum deflexed, scarcely visible in dorsal view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="609F1DD7FFBCFFEAFF59FDA2FD451B55" pageId="7" pageNumber="6" type="description">
|
||
<paragraph id="283A4E5CFFBCFFEAFF59FDA2FE2A1B18" blockId="7.[191,1442,532,1298]" pageId="7" pageNumber="6">
|
||
<emphasis id="1AF1924EFFBCFFEAFF59FDA2FE501C27" bold="true" box="[192,461,532,558]" pageId="7" pageNumber="6">
|
||
Description.
|
||
<typeStatus id="F73EF0FEFFBCFFEAFEC0FDA2FE551C27" box="[345,456,532,558]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Sex unknown. Body short, elongate oval,
|
||
<quantity id="EF7DE3B9FFBCFFEAFC38FDA2FC6A1C26" box="[929,1015,532,559]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="6" unit="mm" value="4.2">4.2 mm</quantity>
|
||
long, width at elytral humeri
|
||
<quantity id="EF7DE3B9FFBCFFEAFADDFDA2FA011C26" box="[1348,1436,532,559]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="6" unit="mm" value="1.7">1.7 mm</quantity>
|
||
, width at middle of elytra
|
||
<quantity id="EF7DE3B9FFBCFFEAFE73FD80FDD81C59" box="[490,581,566,592]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="6" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; dorsum glossy black; legs and venter dark rufous.
|
||
<emphasis id="1AF1924EFFBCFFEAFB01FD80FB401C59" bold="true" box="[1176,1245,566,592]" pageId="7" pageNumber="6">Head:</emphasis>
|
||
Clypeus broadly rounded on each side of shallow median emargination, sides straight to sharply rounded, right-angled gena; surface finely, evenly punctate, punctures separated by about their diameter, surface between punctures smooth; occipital area with crossband of coarser punctures than frons, separated by 1–2 times their diameter.
|
||
<emphasis id="1AF1924EFFBCFFEAFAB9FD2DFA3D1CBC" bold="true" box="[1312,1440,667,693]" pageId="7" pageNumber="6">Pronotum:</emphasis>
|
||
About one-fourth wider than long,
|
||
<quantity id="EF7DE3B9FFBCFFEAFDD2FD0AFD3D1CDE" box="[587,672,700,727]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="6" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long at middle,
|
||
<quantity id="EF7DE3B9FFBCFFEAFCCCFD0AFC361CDE" box="[853,939,700,727]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="6" unit="mm" value="1.5">1.5 mm</quantity>
|
||
wide at base and apex; sides broadly arcuate, anterior angles obtuse, posterior angles broadly rounded; anterolateral margins deflexed, scarcely visible in dorsal view; sides and base margined, finely crenate around posterior angles; surface with mix of evenly spaced fine punctures and irregularly placed, moderately coarse punctures separated by 1–2 times their diameter, becoming denser laterally and finer apically on disk. Scutellar shield short, triangular, impunctate.
|
||
<emphasis id="1AF1924EFFBCFFEAFB0AFCF5FB6E1D54" bold="true" box="[1171,1267,835,861]" pageId="7" pageNumber="6">Elytron:</emphasis>
|
||
Short and convex,
|
||
<quantity id="EF7DE3B9FFBCFFEAFF6BFCD2FEDA1D76" box="[242,327,868,895]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="6" unit="mm" value="2.4">2.4 mm</quantity>
|
||
long, together almost one-half longer than wide, sides arcuate, humeral angle strongly dentate; striae strong, distinctly punctate; intervals weakly convex, smooth and glossy but with scattered minute punctures.
|
||
<emphasis id="1AF1924EFFBCFFEAFF59FC11FE8B1DC8" bold="true" box="[192,278,935,961]" pageId="7" pageNumber="6">Venter:</emphasis>
|
||
Mesoventrite weakly carinate posteriorly between coxae. Metaventrite glossy, middle line strong and deep, approximately the length of the first two abdominal ventrites combined; discal area finely punctate, lateral one-sixth becoming scabrous at sides; metaventrite with triangle anterior to metacoxae moderate, not sharply defined. Abdominal ventrites glossy, finely punctate throughout, punctures becoming coarser laterally where they are separated by 2–4 times their diameter; ventrites finely fluted along anterior margins, the fluting increasingly longer on each ventrite posteriorly.
|
||
<emphasis id="1AF1924EFFBCFFEAFD07FBF9FD471A60" bold="true" box="[670,730,1103,1129]" pageId="7" pageNumber="6">Legs:</emphasis>
|
||
Profemur with posterior face smooth, glossy, finely punctate, punctures separated by 3–4 times their diameter. Meso- and metafemur glossy, finely punctate, punctures separated by 3–5 times their diameter; posterior marginal line strong, extending from tibia halfway to the trochanter. Meso- and metatibiae with minute accessory spine, appearing as a slight projection at apex of tibia; lower spurs fine and slender, approximately as long as first tarsomere; first metatarsomere slightly longer than following three tarsomeres combined.
|
||
</paragraph>
|
||
<paragraph id="283A4E5CFFBCFFEAFF59FA97FD451B55" blockId="7.[192,1440,1313,1373]" pageId="7" pageNumber="6">
|
||
<emphasis id="1AF1924EFFBCFFEAFF59FA97FEB31B32" bold="true" box="[192,302,1313,1339]" pageId="7" pageNumber="6">Variation</emphasis>
|
||
(n = 9). Body length of
|
||
<typeStatus id="F73EF0FEFFBCFFEAFDA6FA97FD361B32" box="[575,683,1313,1339]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
|
||
<quantity id="EF7DE3B9FFBCFFEAFD2BFA97FCA71B32" box="[690,826,1313,1339]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="7" pageNumber="6" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8–4.2 mm</quantity>
|
||
; width at elytral humeri
|
||
<quantity id="EF7DE3B9FFBCFFEAFBD6FA97FB4A1B32" box="[1103,1239,1313,1340]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="7" pageNumber="6" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
|
||
; coarse punctures on the pronotal disc sparser in some specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="609F1DD7FFBCFFEAFF59FADAFB741B8F" box="[192,1257,1388,1415]" pageId="7" pageNumber="6" type="distribution">
|
||
<paragraph id="283A4E5CFFBCFFEAFF59FADAFB741B8F" blockId="7.[192,1257,1388,1415]" box="[192,1257,1388,1415]" pageId="7" pageNumber="6">
|
||
<emphasis id="1AF1924EFFBCFFEAFF59FADAFEC71B8F" bold="true" box="[192,346,1388,1414]" pageId="7" pageNumber="6">Distribution.</emphasis>
|
||
<taxonomicName id="EF8535DFFFBCFFEAFEF8FADBFDBD1B8F" authorityName="Schnepp & Ashman" authorityYear="2020" box="[353,544,1388,1415]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFBCFFEAFEF8FADBFDBD1B8F" box="[353,544,1388,1415]" italics="true" pageId="7" pageNumber="6">Ataenius thomasi</emphasis>
|
||
</taxonomicName>
|
||
appears to be restricted to the Southeast Coastal Plain (
|
||
<figureCitation id="B0BE52D9FFBCFFEAFB15FADAFB411B8F" box="[1164,1244,1388,1414]" captionStart="Figures 13–14" captionStartId="6.[192,279,1864,1890]" captionTargetBox="[231,1403,217,1830]" captionTargetId="figure-40@6.[216,1416,210,1836]" captionTargetPageId="6" captionText="Figures 13–14. Distribution of Ataenius brevis and Ataenius thomasi in the southeastern United States. 13) Ataenius brevis Fall. 14) Ataenius thomasi, new species." figureDoi="http://doi.org/10.5281/zenodo.5354103" httpUri="https://zenodo.org/record/5354103/files/figure.png" pageId="7" pageNumber="6">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="609F1DD7FFBCFFEAFF59FA20FB511BB9" box="[192,1228,1430,1456]" pageId="7" pageNumber="6" type="etymology">
|
||
<paragraph id="283A4E5CFFBCFFEAFF59FA20FB511BB9" blockId="7.[192,1228,1430,1456]" box="[192,1228,1430,1456]" pageId="7" pageNumber="6">
|
||
<emphasis id="1AF1924EFFBCFFEAFF59FA20FED91BB9" bold="true" box="[192,324,1430,1456]" pageId="7" pageNumber="6">Etymology.</emphasis>
|
||
Named in honor of the late Michael C. Thomas, coleopterist, mentor, and friend.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="609F1DD7FFBCFFE5FF59FA09FA001D10" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6" type="discussion">
|
||
<paragraph id="283A4E5CFFBCFFE5FF59FA09FDAD1F1B" blockId="7.[191,1443,1471,1868]" lastBlockId="8.[192,1440,214,375]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
|
||
<emphasis id="1AF1924EFFBCFFEAFF59FA09FEB21BD0" bold="true" box="[192,303,1471,1497]" pageId="7" pageNumber="6">Remarks.</emphasis>
|
||
Intrageneric groupings in
|
||
<taxonomicName id="EF8535DFFFBCFFEAFDCDFA76FD281BD3" authorityName="Harold" authorityYear="1867" box="[596,693,1472,1498]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1AF1924EFFBCFFEAFDCDFA76FD281BD3" box="[596,693,1472,1498]" italics="true" pageId="7" pageNumber="6">Ataenius</emphasis>
|
||
</taxonomicName>
|
||
are not based on comprehensive phylogenetic analyses of the genus and are poorly defined. These assemblages often have members that do not fit easily into any particular species group and could be placed in monotypic “groups”, as well as species that share similar combinations of external character states but may belong to different species groups (
|
||
<bibRefCitation id="4C1433ADFFBCFFEAFCE1F992FB3D1836" author="Stebnicka ZT & Lago PK" box="[888,1184,1572,1599]" pageId="7" pageNumber="6" pagination="55 - 83" refId="ref4064" refString="Stebnicka ZT, Lago PK. 2005. The New World species of Ataenius Harold, 1867. V. Revision of the A. strigatus group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi 19 (1 - 2): 55 - 83." type="journal article" year="2005">Stebnicka and Lago 2005</bibRefCitation>
|
||
). These issues make it extremely difficult to place new species into established groups. Based on general body form,
|
||
<taxonomicName id="EF8535DFFFBCFFEAFB26F9F0FAAB1869" authorityName="Schnepp & Ashman" authorityYear="2020" box="[1215,1334,1606,1632]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFBCFFEAFB26F9F0FB4F1869" box="[1215,1234,1606,1632]" italics="true" pageId="7" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFBCFFEAFB47F9F0FAAB1869" box="[1246,1334,1606,1632]" italics="true" pageId="7" pageNumber="6">thomasi</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="EF8535DFFFBCFFEAFF45F9DEFEA61888" box="[220,315,1639,1666]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="1AF1924EFFBCFFEAFF45F9DEFF72188B" box="[220,239,1640,1666]" italics="true" pageId="7" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFBCFFEAFF62F9D1FEA61888" box="[251,315,1639,1665]" italics="true" pageId="7" pageNumber="6">brevis</emphasis>
|
||
</taxonomicName>
|
||
, sharing with it many of the characters that would place it in the strigatus group, but is distinguishable from that species by its smooth, punctate head, reduced or absent meso- and metatibial apical accessory spine, deflexed anterolateral margins of the pronotum, and male genitalia. Shared character states that would suggest placement in the aequalis-platensis group (Stebnicka 2005) are the meso- and metatibiae with accessory spine reduced or lacking, the surface of the head lacking wrinkles, and to a lesser degree the form of the male genitalia. The parameres of
|
||
<taxonomicName id="EF8535DFFFBCFFEAFE16F8A6FE731920" box="[399,494,1807,1834]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="1AF1924EFFBCFFEAFE16F8A6FE3D1923" box="[399,416,1808,1834]" italics="true" pageId="7" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFBCFFEAFE37F8B9FE731920" box="[430,494,1807,1833]" italics="true" pageId="7" pageNumber="6">brevis</emphasis>
|
||
</taxonomicName>
|
||
are evenly narrowed (in lateral view) from the base to near the apex with the apex slightly deflexed while
|
||
<taxonomicName id="EF8535DFFFBCFFEAFE23F887FDAC1942" authorityName="Schnepp & Ashman" authorityYear="2020" box="[442,561,1841,1867]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFBCFFEAFE23F887FE501942" box="[442,461,1841,1867]" italics="true" pageId="7" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFBCFFEAFE40F887FDAC1942" box="[473,561,1841,1867]" italics="true" pageId="7" pageNumber="6">thomasi</emphasis>
|
||
</taxonomicName>
|
||
are abruptly narrowed basally then parallel sided (in lateral view) to the broadly rounded apex. Future phylogenetic analyses are required to confidently place
|
||
<taxonomicName id="EF8535DFFFB3FFE5FBB6FF60FB3B1EF9" authorityName="Schnepp & Ashman" authorityYear="2020" box="[1071,1190,214,240]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFB3FFE5FBB6FF60FBDD1EF9" box="[1071,1088,214,240]" italics="true" pageId="8" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFB3FFE5FBD7FF60FB3B1EF9" box="[1102,1190,214,240]" italics="true" pageId="8" pageNumber="7">thomasi</emphasis>
|
||
</taxonomicName>
|
||
, as well as confirm or reclassify current species groups.
|
||
</paragraph>
|
||
<paragraph id="283A4E5CFFB3FFE5FE99FEAFFA851F3A" blockId="8.[192,1440,214,375]" box="[256,1304,281,307]" pageId="8" pageNumber="7">
|
||
From the available label data, it appears neither
|
||
<taxonomicName id="EF8535DFFFB3FFE5FC8DFEAFFCE91F3A" box="[788,884,281,307]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="1AF1924EFFB3FFE5FC8DFEAFFCBA1F3A" box="[788,807,281,307]" italics="true" pageId="8" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFB3FFE5FCADFEAFFCE91F3A" box="[820,884,281,307]" italics="true" pageId="8" pageNumber="7">brevis</emphasis>
|
||
</taxonomicName>
|
||
nor
|
||
<taxonomicName id="EF8535DFFFB3FFE5FC31FEAFFBBD1F3A" authorityName="Schnepp & Ashman" authorityYear="2020" box="[936,1056,281,307]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFB3FFE5FC31FEAFFC261F3A" box="[936,955,281,307]" italics="true" pageId="8" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFB3FFE5FC51FEAFFBBD1F3A" box="[968,1056,281,307]" italics="true" pageId="8" pageNumber="7">thomasi</emphasis>
|
||
</taxonomicName>
|
||
are attracted to lights.
|
||
</paragraph>
|
||
<paragraph id="283A4E5CFFB3FFE5FE99FE8CFC701F7E" blockId="8.[192,1440,214,375]" pageId="8" pageNumber="7">
|
||
Though the new species is not placed into a group, the following amended key for the
|
||
<taxonomicName id="EF8535DFFFB3FFE5FB43FE8DFA3D1F5D" baseAuthorityName="Say" baseAuthorityYear="1823" box="[1242,1440,314,341]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="strigatus">
|
||
<emphasis id="1AF1924EFFB3FFE5FB43FE8DFAA61F5C" box="[1242,1339,315,341]" italics="true" pageId="8" pageNumber="7">Ataenius</emphasis>
|
||
strigatus
|
||
</taxonomicName>
|
||
group is modified from
|
||
<bibRefCitation id="4C1433ADFFB3FFE5FE53FEEAFD6E1F7E" author="Stebnicka ZT & Lago PK" box="[458,755,348,375]" pageId="8" pageNumber="7" pagination="55 - 83" refId="ref4064" refString="Stebnicka ZT, Lago PK. 2005. The New World species of Ataenius Harold, 1867. V. Revision of the A. strigatus group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi 19 (1 - 2): 55 - 83." type="journal article" year="2005">Stebnicka and Lago (2005)</bibRefCitation>
|
||
to include
|
||
<taxonomicName id="EF8535DFFFB3FFE5FCE9FEEBFC7A1F7F" authorityName="Schnepp & Ashman" authorityYear="2020" box="[880,999,348,375]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="thomasi">
|
||
<emphasis id="1AF1924EFFB3FFE5FCE9FEEBFC1E1F7E" box="[880,899,349,375]" italics="true" pageId="8" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFB3FFE5FC16FEEAFC7A1F7F" box="[911,999,348,374]" italics="true" pageId="8" pageNumber="7">thomasi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="283A4E5CFFB3FFE5FF59FE30FA031FC8" blockId="8.[192,1441,390,517]" pageId="8" pageNumber="7">
|
||
4(1). Elytra short, oval with convex margins, length less than 2 times as long as pronotum; humeral denticles strong, acutely pointed.
|
||
<collectingCountry id="50920ECCFFB3FFE5FDC5FE11FD0C1FC8" box="[604,657,423,449]" name="United States of America" pageId="8" pageNumber="7">USA</collectingCountry>
|
||
..........................................................
|
||
<emphasis id="1AF1924EFFB3FFE5FA1BFE11FA031FC8" bold="true" box="[1410,1438,423,449]" pageId="8" pageNumber="7">4a</emphasis>
|
||
</paragraph>
|
||
<paragraph id="283A4E5CFFB3FFE5FF59FE7FFA001C0C" blockId="8.[192,1441,390,517]" pageId="8" pageNumber="7">
|
||
— Elytra elongate, nearly parallel-sided, length 2.1 times as long as pronotum or longer, humeral denticles fine to moderate, usually obtuse........................................................
|
||
<emphasis id="1AF1924EFFB3FFE5FA16FE5DFA001C0C" bold="true" box="[1423,1437,491,517]" pageId="8" pageNumber="7">5</emphasis>
|
||
</paragraph>
|
||
<paragraph id="283A4E5CFFB3FFE5FF59FDA2FA001C9A" blockId="8.[192,1442,532,794]" pageId="8" pageNumber="7">
|
||
4a(4). Meso- and metatibiae with apical accessory spine conspicuous (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FC44FDA2FB801C26" box="[989,1053,532,559]" captionStart="Figures 1–6" captionStartId="4.[192,279,1833,1859]" captionTargetBox="[216,1417,215,1802]" captionTargetId="figure-13@4.[214,1419,213,1802]" captionTargetPageId="4" captionText="Figures 1–6. Ataenius brevis Fall. 1) Dorsal view of lectotype. 2) Ventral view of lectotype. 3) Frontal view of lectotype.4) Lectotype labels. 5) Ventral view of metatibia of lectotype, arrow indicates outlined accessory spine. 6) Male genitalia, specimen from Citico, Tennessee." figureDoi="http://doi.org/10.5281/zenodo.5354099" httpUri="https://zenodo.org/record/5354099/files/figure.png" pageId="8" pageNumber="7">Fig. 5</figureCitation>
|
||
); punctures of abdominal ventrites coarse (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FE3EFD80FE761C59" box="[423,491,566,592]" captionStart="Figures 1–6" captionStartId="4.[192,279,1833,1859]" captionTargetBox="[216,1417,215,1802]" captionTargetId="figure-13@4.[214,1419,213,1802]" captionTargetPageId="4" captionText="Figures 1–6. Ataenius brevis Fall. 1) Dorsal view of lectotype. 2) Ventral view of lectotype. 3) Frontal view of lectotype.4) Lectotype labels. 5) Ventral view of metatibia of lectotype, arrow indicates outlined accessory spine. 6) Male genitalia, specimen from Citico, Tennessee." figureDoi="http://doi.org/10.5281/zenodo.5354099" httpUri="https://zenodo.org/record/5354099/files/figure.png" pageId="8" pageNumber="7">Fig. 2</figureCitation>
|
||
); clypeus and frons with transverse sculpturing, appearing finely wrinkled (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FAD4FD80FA0C1C59" box="[1357,1425,566,592]" captionStart="Figures 1–6" captionStartId="4.[192,279,1833,1859]" captionTargetBox="[216,1417,215,1802]" captionTargetId="figure-13@4.[214,1419,213,1802]" captionTargetPageId="4" captionText="Figures 1–6. Ataenius brevis Fall. 1) Dorsal view of lectotype. 2) Ventral view of lectotype. 3) Frontal view of lectotype.4) Lectotype labels. 5) Ventral view of metatibia of lectotype, arrow indicates outlined accessory spine. 6) Male genitalia, specimen from Citico, Tennessee." figureDoi="http://doi.org/10.5281/zenodo.5354099" httpUri="https://zenodo.org/record/5354099/files/figure.png" pageId="8" pageNumber="7">Fig. 3</figureCitation>
|
||
); anterolateral margins of pronotum slightly explanate, clearly visible in dorsal view (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FB77FDE1FAB11C78" box="[1262,1324,599,625]" captionStart="Figures 1–6" captionStartId="4.[192,279,1833,1859]" captionTargetBox="[216,1417,215,1802]" captionTargetId="figure-13@4.[214,1419,213,1802]" captionTargetPageId="4" captionText="Figures 1–6. Ataenius brevis Fall. 1) Dorsal view of lectotype. 2) Ventral view of lectotype. 3) Frontal view of lectotype.4) Lectotype labels. 5) Ventral view of metatibia of lectotype, arrow indicates outlined accessory spine. 6) Male genitalia, specimen from Citico, Tennessee." figureDoi="http://doi.org/10.5281/zenodo.5354099" httpUri="https://zenodo.org/record/5354099/files/figure.png" pageId="8" pageNumber="7">Fig. 1</figureCitation>
|
||
)................................................................................
|
||
<taxonomicName id="EF8535DFFFB3FFE5FA9FFDCFFA001C9A" authority="Fall" authorityName="Fall" box="[1286,1437,632,659]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="brevis">
|
||
<emphasis id="1AF1924EFFB3FFE5FA9FFDCFFA001C9A" bold="true" box="[1286,1437,632,659]" pageId="8" pageNumber="7">
|
||
<emphasis id="1AF1924EFFB3FFE5FA9FFDCFFA841C9A" bold="true" box="[1286,1305,633,659]" italics="true" pageId="8" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFB3FFE5FABFFDCEFAF61C9A" bold="true" box="[1318,1387,632,659]" italics="true" pageId="8" pageNumber="7">brevis</emphasis>
|
||
Fall
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="283A4E5CFFB3FFE5FF59FD2DFA001D10" blockId="8.[192,1442,532,794]" pageId="8" pageNumber="7">
|
||
— Meso- and metatibiae with apical accessory spine absent or reduced (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FBB4FD2CFBE01CBC" box="[1069,1149,666,693]" captionStart="Figures 7–12" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[222,1405,223,1802]" captionTargetId="figure-57@5.[219,1414,213,1804]" captionTargetPageId="5" captionText="Figures 7–12. Ataenius thomasi Schnepp and Ashman. 7) Dorsal view of holotype. 8) Ventral view of holotype. 9) Frontal view of holotype. 10) Holotype labels. 11) Ventral view of metatibia of holotype, arrow indicates reduced accessory spine. 12) Male genitalia of paratype." figureDoi="http://doi.org/10.5281/zenodo.5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="8" pageNumber="7">Fig. 11</figureCitation>
|
||
); punctures of abdominal ventrites fine (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FE6BFD0AFDA91CDF" box="[498,564,700,726]" captionStart="Figures 7–12" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[222,1405,223,1802]" captionTargetId="figure-57@5.[219,1414,213,1804]" captionTargetPageId="5" captionText="Figures 7–12. Ataenius thomasi Schnepp and Ashman. 7) Dorsal view of holotype. 8) Ventral view of holotype. 9) Frontal view of holotype. 10) Holotype labels. 11) Ventral view of metatibia of holotype, arrow indicates reduced accessory spine. 12) Male genitalia of paratype." figureDoi="http://doi.org/10.5281/zenodo.5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="8" pageNumber="7">Fig. 8</figureCitation>
|
||
); clypeus and frons lacking transverse sculpturing, smooth and punctate (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FAEFFD0AFEFD1CF1" captionStart="Figures 7–12" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[222,1405,223,1802]" captionTargetId="figure-57@5.[219,1414,213,1804]" captionTargetPageId="5" captionText="Figures 7–12. Ataenius thomasi Schnepp and Ashman. 7) Dorsal view of holotype. 8) Ventral view of holotype. 9) Frontal view of holotype. 10) Holotype labels. 11) Ventral view of metatibia of holotype, arrow indicates reduced accessory spine. 12) Male genitalia of paratype." figureDoi="http://doi.org/10.5281/zenodo.5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="8" pageNumber="7">Fig. 9</figureCitation>
|
||
); anterolateral margins of pronotum deflexed, scarcely visible in dorsal view (
|
||
<figureCitation id="B0BE52D9FFB3FFE5FB27FD68FB631CF1" box="[1214,1278,734,760]" captionStart="Figures 7–12" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[222,1405,223,1802]" captionTargetId="figure-57@5.[219,1414,213,1804]" captionTargetPageId="5" captionText="Figures 7–12. Ataenius thomasi Schnepp and Ashman. 7) Dorsal view of holotype. 8) Ventral view of holotype. 9) Frontal view of holotype. 10) Holotype labels. 11) Ventral view of metatibia of holotype, arrow indicates reduced accessory spine. 12) Male genitalia of paratype." figureDoi="http://doi.org/10.5281/zenodo.5354101" httpUri="https://zenodo.org/record/5354101/files/figure.png" pageId="8" pageNumber="7">Fig. 7</figureCitation>
|
||
)............................................................
|
||
<emphasis id="1AF1924EFFB3FFE5FC4BFD49FA001D10" bold="true" box="[978,1437,767,794]" pageId="8" pageNumber="7">
|
||
<taxonomicName id="EF8535DFFFB3FFE5FC4BFD49FAC81D10" authority="Schnepp and Ashman" authorityName="Schnepp and Ashman" authorityYear="2020" box="[978,1365,767,794]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
|
||
<emphasis id="1AF1924EFFB3FFE5FC4BFD49FC781D10" bold="true" box="[978,997,767,793]" italics="true" pageId="8" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="1AF1924EFFB3FFE5FC6BFD49FBD31D13" bold="true" box="[1010,1102,767,794]" italics="true" pageId="8" pageNumber="7">thomasi</emphasis>
|
||
Schnepp and Ashman
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="01C22F35FFB3FFE5FAF9FD49FA001D10" box="[1376,1437,767,793]" pageId="8" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |