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<document id="F53AA268C04117F01A3B793854D2705D" ID-DOI="10.5281/zenodo.5741537" ID-ISSN="1536-9307" ID-Zenodo-Dep="5741537" ID-ZooBank="6C4BC10A-F418-48E0-8BF8-D33DE4BA62A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638265185309" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2020" docId="9C4287CFFFB5FF87CFFAF98108A9F832" docLanguage="en" docName="Euscorpius.302.1-43.pdf" docOrigin="Euscorpius 302" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Scorpiops hofereki Kovařík 2020, sp. n." docType="treatment" docUuid="10A76C0E-2759-47A0-8063-7DE46ADA205C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="17" masterDocId="607BFFB7FFBEFF95CC19FFAD094CFFD1" masterDocTitle="Nine new species of Scorpiops Peters, 1861 (Scorpiones: Scorpiopidae) from China, India, Nepal, and Pakistan" masterLastPageNumber="43" masterPageNumber="1" pageNumber="10" updateTime="1699436699584" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="C011A1583B307A3787E9D8FFEFAFDC0B">Nine new species of Scorpiops Peters, 1861 (Scorpiones: Scorpiopidae) from China, India, Nepal, and Pakistan</mods:title>
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<mods:date id="148C32B5D7C3A76ED93671B2F91B9C89">2020</mods:date>
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<taxonomicName id="D3EB4D5AFFB5FF9ECFFAF9810D8AF996" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[995,1222,1580,1607]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="hofereki" status="sp. nov.">Scorpiops hofereki</taxonomicName>
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<emphasis id="269FEACBFFB5FF9EC8D7F9810DA6F997" bold="true" box="[1230,1258,1580,1606]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="269FEACBFFB5FF9EC8E1F9810C44F997" bold="true" box="[1272,1288,1580,1606]" pageId="11" pageNumber="10">n</emphasis>
.
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<paragraph id="145436D9FFB5FF9ECFC2F9E30CF0F957" blockId="11.[823,1468,1580,1702]" pageId="11" pageNumber="10">
(
<figureCitation id="8CD02A5CFFB5FF9ECFFBF9E30DCCF9B7" box="[994,1152,1614,1638]" captionStart-0="Figures 69–73" captionStart-1="Figures 74–82" captionStart-2="Figures 83–92" captionStartId-0="15.[117,194,1980,2002]" captionStartId-1="16.[114,191,1956,1978]" captionStartId-2="17.[113,190,1983,2005]" captionTargetBox-0="[130,1451,248,1967]" captionTargetBox-1="[124,1410,249,1926]" captionTargetBox-2="[113,1474,234,1954]" captionTargetId-0="figure-45@15.[113,1474,234,1973]" captionTargetId-1="figure-66@16.[114,1475,234,1945]" captionTargetId-2="figure-55@17.[113,1474,234,1955]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Figures 69–73. Scorpiops hofereki sp. n., female holotype. Dorsal (69) and ventral (70) views. Metasoma and telson lateral (71), ventral (72), and dorsal (73) views. Scale bars: 10 mm (69–70, 71–73)." captionText-1="Figures 74–82. Scorpiops hofereki sp. n., female holotype, pedipalp segments. Chela dorsal (74), external (75) and ventral (76) views. Patella dorsal (77), external (78) and ventral (79) views. Femur and trochanter dorsal (80) and ventral (81) views. Movable finger dentition (82). Trichobothrial pattern is indicated by white circles." captionText-2="Figures 83–92. Scorpiops hofereki sp. n., female holotype. Carapace and tergites I–III (83), posterior coxosternal area and sternites III–IV (84). Right chelicera in dorsal (85) and ventral (86) views. Telson lateral (87). Pectine (88). Left legs I–IV, retrolateral aspect (89–92 respectively)." figureDoi-0="http://doi.org/10.5281/zenodo.5741563" figureDoi-1="http://doi.org/10.5281/zenodo.5741565" figureDoi-2="http://doi.org/10.5281/zenodo.5741567" httpUri-0="https://zenodo.org/record/5741563/files/figure.png" httpUri-1="https://zenodo.org/record/5741565/files/figure.png" httpUri-2="https://zenodo.org/record/5741567/files/figure.png" pageId="11" pageNumber="10">Figures 69–92</figureCitation>
,
<figureCitation id="8CD02A5CFFB5FF9EC892F9E30DFAF9B7" box="[1163,1206,1614,1638]" captionStart="Figure 240" captionStartId="41.[113,181,1361,1383]" captionTargetBox="[113,1474,234,1350]" captionTargetId="figure-353@41.[113,1474,234,1350]" captionTargetPageId="41" captionText="Figure 240. Map of distribution of nine new species described here and Scorpiops vonwicki Birula, 1913." figureDoi="http://doi.org/10.5281/zenodo.5741606" httpUri="https://zenodo.org/record/5741606/files/figure.png" pageId="11" pageNumber="10">240</figureCitation>
,
<tableCitation id="59690362FFB5FF9EC8D9F9E30C41F9B7" box="[1216,1293,1614,1638]" captionStart="Table 1" captionStartId="3.[113,169,985,1006]" captionText="Table 1. Comparative measurements of adults of Scorpiops furai sp. n., S. grosseri sp. n., S. harmsi sp. n. and S. hofereki sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="11" pageNumber="10">Table 1</tableCitation>
)
<emphasis id="269FEACBFFB5FF9ECF2EF9C30CF0F957" bold="true" box="[823,1468,1646,1670]" pageId="11" pageNumber="10">http://zoobank.org/urn:lsid:zoobank.org:act:10A76C0E-</emphasis>
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.
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, Gilgit-
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,
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, ca
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(formerly North-West Frontier)
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<materialsCitation id="A4833C84FFB5FF99C965F8630991F97A" collectionCode="FKCP" country="Pakistan" lastPageId="12" lastPageNumber="11" location="David Hoferek" municipality="The" pageId="11" pageNumber="10" specimenCount="1" typeStatus="paratype">
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. ETYMOLOGY.
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specific epithet honors
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(Czech Republic) for his contribution to understanding scorpions
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.
</paragraph>
</subSubSection>
<caption id="40946651FFB2FF99CC6CF9B40DF0F9FE" ID-DOI="http://doi.org/10.5281/zenodo.5741557" ID-Zenodo-Dep="5741557" box="[117,1212,1561,1584]" httpUri="https://zenodo.org/record/5741557/files/figure.png" pageId="12" pageNumber="11" startId="12.[117,194,1561,1583]" targetBox="[120,1467,263,1541]" targetPageId="12">
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<emphasis id="269FEACBFFB2FF99CC6CF9B40848F9FF" bold="true" box="[117,260,1561,1583]" pageId="12" pageNumber="11">Figures 49–50</emphasis>
.
<taxonomicName id="D3EB4D5AFFB2FF99CD09F9B708F5F9FF" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[272,441,1561,1583]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="harmsi" status="sp. nov.">
<emphasis id="269FEACBFFB2FF99CD09F9B708F5F9FF" box="[272,441,1561,1583]" italics="true" pageId="12" pageNumber="11">Scorpiops harmsi</emphasis>
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<emphasis id="269FEACBFFB2FF99CDA7F9B70899F9FE" bold="true" box="[446,469,1562,1583]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="269FEACBFFB2FF99CDF8F9B708A2F9FE" bold="true" box="[481,494,1562,1583]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
, female holotype in dorsal (49) and ventral (50) views. Scale bar: 10 mm.
</paragraph>
</caption>
<subSubSection id="5CF16552FFB2FF99CC6AF97E0C3DF89A" pageId="12" pageNumber="11" type="diagnosis">
<paragraph id="145436D9FFB2FF99CC6AF97E0C3DF89A" blockId="12.[115,774,1747,2033]" lastBlockId="12.[818,1483,1619,1867]" pageId="12" pageNumber="11">
DIAGNOSIS (
<specimenCount id="02EDFD50FFB2FF99CD1FF97E085BF93A" box="[262,279,1747,1771]" pageId="12" pageNumber="11" type="female">♀</specimenCount>
). Total length
<quantity id="D3139B3CFFB2FF99CDFDF97E0B26F93A" box="[484,618,1747,1771]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" metricValueMax="6.0" metricValueMin="4.8" pageId="12" pageNumber="11" unit="mm" value="54.0" valueMax="60.0" valueMin="48.0">48–60 mm</quantity>
. Base color uniformly reddish brown to black, telson and tarsomere II of legs yellowish to reddish brown. Pectinal teeth number
<quantity id="D3139B3CFFB2FF99CEC4F8BE09C4F89A" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="12" pageNumber="11" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
females, fulcra absent; marginal lamella I (basal) present, the remaining pectinal surface forming one compact unit. Patella of pedipalp with 17 (5
<emphasis id="269FEACBFFB2FF99CDD7F8DE08ABF85A" box="[462,487,1907,1931]" italics="true" pageId="12" pageNumber="11">eb</emphasis>
, 2
<emphasis id="269FEACBFFB2FF99CE16F8DE0B7FF85A" box="[527,563,1907,1931]" italics="true" pageId="12" pageNumber="11">esb</emphasis>
, 2
<emphasis id="269FEACBFFB2FF99CE42F8D90B36F85A" box="[603,634,1908,1931]" italics="true" pageId="12" pageNumber="11">em</emphasis>
, 4
<emphasis id="269FEACBFFB2FF99CEBBF8D90B8CF85A" box="[674,704,1908,1931]" italics="true" pageId="12" pageNumber="11">est</emphasis>
, 4
<emphasis id="269FEACBFFB2FF99CEF1F8D90BB0F85A" box="[744,764,1908,1931]" italics="true" pageId="12" pageNumber="11">et</emphasis>
) external and 7 ventral trichobothria. Chela of pedipalp with 4 ventral trichobothria located on ventral surface. Chelal trichobothrium
<emphasis id="269FEACBFFB2FF99CD06F87E0809F820" box="[287,325,2003,2033]" italics="true" pageId="12" pageNumber="11">
Eb
<subScript id="886F349CFFB2FF99CD24F84E0809F820" attach="left" box="[317,325,2019,2033]" fontSize="6" pageId="12" pageNumber="11">3</subScript>
</emphasis>
located in lower half of external surface between trichobothria
<emphasis id="269FEACBFFB2FF99C823F9F90D19F9BA" box="[1082,1109,1620,1643]" italics="true" pageId="12" pageNumber="11">Dt</emphasis>
and
<emphasis id="269FEACBFFB2FF99C885F9FE0DF1F9BA" box="[1180,1213,1619,1643]" italics="true" pageId="12" pageNumber="11">Db</emphasis>
. Fingers of pedipalps undulate in females. Chela length to width ratio
<quantity id="D3139B3CFFB2FF99C94DF9DE0C88F95A" box="[1364,1476,1651,1675]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="7.874" metricValueMin="7.366" pageId="12" pageNumber="11" unit="in" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 in</quantity>
females. Pedipalp movable finger with ca 38 IAD, which form a second row, parallel to MD (
<quantity id="D3139B3CFFB2FF99C8A3F91E0C50F91A" box="[1210,1308,1715,1739]" metricMagnitude="0" metricUnit="m" metricValue="1.6509999999999998" metricValueMax="1.7271999999999998" metricValueMin="1.5748" pageId="12" pageNumber="11" unit="in" value="65.0" valueMax="68.0" valueMin="62.0">62–68 in</quantity>
number); there are also 5 ID and 9–10 OD present. Tarsomere II of legs with 6–9 stout median ventral spines in a row and two other parallel spines. Telson bulbous and finely granulate, length to depth ratio
<quantity id="D3139B3CFFB2FF99CFD2F89E0D69F89A" box="[971,1061,1843,1867]" metricMagnitude="-2" metricUnit="m" metricValue="7.111999999999999" metricValueMax="7.62" metricValueMin="6.604" pageId="12" pageNumber="11" unit="in" value="2.8" valueMax="3.0" valueMin="2.6">2.6–3 in</quantity>
females; annular ring present.
</paragraph>
</subSubSection>
<subSubSection id="5CF16552FFB2FF87CF2BF8DE08A9F832" lastPageId="18" lastPageNumber="17" pageId="12" pageNumber="11" type="description">
<paragraph id="145436D9FFB2FF99CF2BF8DE0C3AF83A" blockId="12.[818,1477,1907,2027]" pageId="12" pageNumber="11">
DESCRIPTION (
<specimenCount id="02EDFD50FFB2FF99CFC8F8DE0AAEF85A" box="[977,994,1907,1931]" pageId="12" pageNumber="11" type="female">♀</specimenCount>
). Total length
<quantity id="D3139B3CFFB2FF99C892F8DE0C4DF85A" box="[1163,1281,1907,1931]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" metricValueMax="6.0" metricValueMin="4.8" pageId="12" pageNumber="11" unit="mm" value="54.0" valueMax="60.0" valueMin="48.0">48–60 mm</quantity>
of females, male unknown. The habitus is shown in
<figureCitation id="8CD02A5CFFB2FF99C8ABF83E0C79F87A" box="[1202,1333,1939,1963]" captionStart="Figures 69–73" captionStartId="15.[117,194,1980,2002]" captionTargetBox="[130,1451,248,1967]" captionTargetId="figure-45@15.[113,1474,234,1973]" captionTargetPageId="15" captionText="Figures 69–73. Scorpiops hofereki sp. n., female holotype. Dorsal (69) and ventral (70) views. Metasoma and telson lateral (71), ventral (72), and dorsal (73) views. Scale bars: 10 mm (69–70, 71–73)." figureDoi="http://doi.org/10.5281/zenodo.5741563" httpUri="https://zenodo.org/record/5741563/files/figure.png" pageId="12" pageNumber="11">Figs. 69–70</figureCitation>
. For position and distribution of trichobothria of pedipalps see
<figureCitation id="8CD02A5CFFB2FF99C959F81E0C8DF81A" box="[1344,1473,1971,1995]" captionStart="Figures 74–82" captionStartId="16.[114,191,1956,1978]" captionTargetBox="[124,1410,249,1926]" captionTargetId="figure-66@16.[114,1475,234,1945]" captionTargetPageId="16" captionText="Figures 74–82. Scorpiops hofereki sp. n., female holotype, pedipalp segments. Chela dorsal (74), external (75) and ventral (76) views. Patella dorsal (77), external (78) and ventral (79) views. Femur and trochanter dorsal (80) and ventral (81) views. Movable finger dentition (82). Trichobothrial pattern is indicated by white circles." figureDoi="http://doi.org/10.5281/zenodo.5741565" httpUri="https://zenodo.org/record/5741565/files/figure.png" pageId="12" pageNumber="11">Figs. 74–80</figureCitation>
. Fingers of pedipalps are undulate in females (
<figureCitation id="8CD02A5CFFB2FF99C903F87E0C25F83A" box="[1306,1385,2003,2027]" captionStart="Figures 74–82" captionStartId="16.[114,191,1956,1978]" captionTargetBox="[124,1410,249,1926]" captionTargetId="figure-66@16.[114,1475,234,1945]" captionTargetPageId="16" captionText="Figures 74–82. Scorpiops hofereki sp. n., female holotype, pedipalp segments. Chela dorsal (74), external (75) and ventral (76) views. Patella dorsal (77), external (78) and ventral (79) views. Femur and trochanter dorsal (80) and ventral (81) views. Movable finger dentition (82). Trichobothrial pattern is indicated by white circles." figureDoi="http://doi.org/10.5281/zenodo.5741565" httpUri="https://zenodo.org/record/5741565/files/figure.png" pageId="12" pageNumber="11">Fig. 75</figureCitation>
).
</paragraph>
<caption id="40946651FFB3FF98CC6DF80B089CF827" ID-DOI="http://doi.org/10.5281/zenodo.5741559" ID-Zenodo-Dep="5741559" httpUri="https://zenodo.org/record/5741559/files/figure.png" pageId="13" pageNumber="12" startId="13.[116,193,1958,1980]" targetBox="[123,1474,262,1869]" targetPageId="13">
<paragraph id="145436D9FFB3FF98CC6DF80B089CF827" blockId="13.[116,1474,1958,2038]" pageId="13" pageNumber="12">
<emphasis id="269FEACBFFB3FF98CC6DF80B084FF86A" bold="true" box="[116,259,1958,1980]" pageId="13" pageNumber="12">Figures 51–58</emphasis>
.
<taxonomicName id="D3EB4D5AFFB3FF98CD16F80B08F5F86A" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[271,441,1958,1979]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="harmsi" status="sp. nov.">
<emphasis id="269FEACBFFB3FF98CD16F80B08F5F86A" box="[271,441,1958,1979]" italics="true" pageId="13" pageNumber="12">Scorpiops harmsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DAC57B0FFB3FF98CDA7F80B08B8F86D" box="[446,500,1958,1980]" pageId="13" pageNumber="12" rank="species">
<emphasis id="269FEACBFFB3FF98CDA7F80B0899F86A" bold="true" box="[446,469,1958,1979]" pageId="13" pageNumber="12">sp</emphasis>
.
<emphasis id="269FEACBFFB3FF98CDF8F80B08A2F86A" bold="true" box="[481,494,1958,1979]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
, female holotype, pedipalp segments. Chela dorsal (51), external (52) and ventral (53) views. Patella dorsal (54), external (55) and ventral (56) views. Movable finger dentition under white light (57) and UV fluorescence (58). Trichobothrial pattern is indicated by white circles.
</paragraph>
</caption>
<caption id="40946651FFB0FF9BCC6BF8130CF5F820" ID-DOI="http://doi.org/10.5281/zenodo.5741561" ID-Zenodo-Dep="5741561" httpUri="https://zenodo.org/record/5741561/files/figure.png" pageId="14" pageNumber="13" startId="14.[114,189,1982,2004]" targetBox="[113,1455,234,1964]" targetPageId="14">
<paragraph id="145436D9FFB0FF9BCC6BF8130CF5F820" blockId="14.[114,1473,1982,2033]" pageId="14" pageNumber="13">
<emphasis id="269FEACBFFB0FF9BCC6BF813084CF805" bold="true" box="[114,256,1982,2004]" pageId="14" pageNumber="13">Figures 59–68</emphasis>
.
<taxonomicName id="D3EB4D5AFFB0FF9BCD15F81208FFF805" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[268,435,1983,2004]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="harmsi" status="sp. nov.">
<emphasis id="269FEACBFFB0FF9BCD15F81208FFF805" box="[268,435,1983,2004]" italics="true" pageId="14" pageNumber="13">Scorpiops harmsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DAC57B0FFB0FF9BCDA3F81208BCF805" box="[442,496,1983,2004]" pageId="14" pageNumber="13" rank="species">
<emphasis id="269FEACBFFB0FF9BCDA3F812089CF805" bold="true" box="[442,464,1983,2004]" pageId="14" pageNumber="13">sp</emphasis>
.
<emphasis id="269FEACBFFB0FF9BCDC4F81208A6F805" bold="true" box="[477,490,1983,2004]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
, female holotype. Carapace and tergites I–III (59), posterior coxosternal area and sternites III–V (60). Left legs I–IV, retrolateral aspect (61–64 respectively). Telson lateral (65). Metasoma and telson lateral (66), ventral (67), and dorsal (68) views.
</paragraph>
</caption>
<caption id="40946651FFB1FF9ACC6CF8110BEFF83E" ID-DOI="http://doi.org/10.5281/zenodo.5741563" ID-Zenodo-Dep="5741563" httpUri="https://zenodo.org/record/5741563/files/figure.png" pageId="15" pageNumber="14" startId="15.[117,194,1980,2002]" targetBox="[130,1451,248,1967]" targetPageId="15">
<paragraph id="145436D9FFB1FF9ACC6CF8110BEFF83E" blockId="15.[117,1475,1980,2031]" pageId="15" pageNumber="14">
<emphasis id="269FEACBFFB1FF9ACC6CF811084FF800" bold="true" box="[117,259,1980,2002]" pageId="15" pageNumber="14">Figures 69–73</emphasis>
.
<taxonomicName id="D3EB4D5AFFB1FF9ACD16F810088FF800" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[271,451,1980,2002]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="hofereki" status="sp. nov.">
<emphasis id="269FEACBFFB1FF9ACD16F810088FF800" box="[271,451,1980,2002]" italics="true" pageId="15" pageNumber="14">Scorpiops hofereki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DAC57B0FFB1FF9ACDD1F81108B1F803" box="[456,509,1980,2002]" pageId="15" pageNumber="14" rank="species">
<emphasis id="269FEACBFFB1FF9ACDD1F8110893F800" bold="true" box="[456,479,1980,2001]" pageId="15" pageNumber="14">sp</emphasis>
.
<emphasis id="269FEACBFFB1FF9ACDF3F81108BBF800" bold="true" box="[490,503,1980,2001]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
, female holotype. Dorsal (69) and ventral (70) views. Metasoma and telson lateral (71), ventral (72), and dorsal (73) views. Scale bars: 10 mm (69–70, 71–73).
</paragraph>
</caption>
<caption id="40946651FFAEFF85CC6BF8090B12F825" ID-DOI="http://doi.org/10.5281/zenodo.5741565" ID-Zenodo-Dep="5741565" httpUri="https://zenodo.org/record/5741565/files/figure.png" pageId="16" pageNumber="15" startId="16.[114,191,1956,1978]" targetBox="[124,1410,249,1926]" targetPageId="16">
<paragraph id="145436D9FFAEFF85CC6BF8090B12F825" blockId="16.[114,1473,1956,2036]" pageId="16" pageNumber="15">
<emphasis id="269FEACBFFAEFF85CC6BF809084DF868" bold="true" box="[114,257,1956,1978]" pageId="16" pageNumber="15">Figures 74–82</emphasis>
.
<taxonomicName id="D3EB4D5AFFAEFF85CD15F808088CF868" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[268,448,1956,1978]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="hofereki" status="sp. nov.">
<emphasis id="269FEACBFFAEFF85CD15F808088CF868" box="[268,448,1956,1978]" italics="true" pageId="16" pageNumber="15">Scorpiops hofereki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DAC57B0FFAEFF85CDDCF80908B6F86B" box="[453,506,1956,1978]" pageId="16" pageNumber="15" rank="species">
<emphasis id="269FEACBFFAEFF85CDDCF8090890F868" bold="true" box="[453,476,1956,1977]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="269FEACBFFAEFF85CDF1F80908B9F868" bold="true" box="[488,501,1956,1977]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
, female holotype, pedipalp segments. Chela dorsal (74), external (75) and ventral (76) views. Patella dorsal (77), external (78) and ventral (79) views. Femur and trochanter dorsal (80) and ventral (81) views. Movable finger dentition (82). Trichobothrial pattern is indicated by white circles.
</paragraph>
</caption>
<caption id="40946651FFAFFF84CC68F8120CF3F823" ID-DOI="http://doi.org/10.5281/zenodo.5741567" ID-Zenodo-Dep="5741567" httpUri="https://zenodo.org/record/5741567/files/figure.png" pageId="17" pageNumber="16" startId="17.[113,190,1983,2005]" targetBox="[113,1474,234,1954]" targetPageId="17">
<paragraph id="145436D9FFAFFF84CC68F8120CF3F823" blockId="17.[113,1472,1983,2035]" pageId="17" pageNumber="16">
<emphasis id="269FEACBFFAFFF84CC68F81209B3F805" bold="true" box="[113,255,1983,2005]" pageId="17" pageNumber="16">Figures 83–92</emphasis>
.
<taxonomicName id="D3EB4D5AFFAFFF84CD13F86D08F1F804" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[266,445,1984,2005]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="hofereki" status="sp. nov.">
<emphasis id="269FEACBFFAFFF84CD13F86D08F1F804" box="[266,445,1984,2005]" italics="true" pageId="17" pageNumber="16">Scorpiops hofereki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DAC57B0FFAFFF84CDD8F86D08B9F804" box="[449,501,1984,2005]" pageId="17" pageNumber="16" rank="species">
<emphasis id="269FEACBFFAFFF84CDD8F86D0894F804" bold="true" box="[449,472,1984,2005]" pageId="17" pageNumber="16">sp</emphasis>
.
<emphasis id="269FEACBFFAFFF84CDFAF86D08BCF804" bold="true" box="[483,496,1984,2005]" pageId="17" pageNumber="16">n</emphasis>
.
</taxonomicNameLabel>
, female holotype. Carapace and tergites I–III (83), posterior coxosternal area and sternites III–IV (84). Right chelicera in dorsal (85) and ventral (86) views. Telson lateral (87). Pectine (88). Left legs I–IV, retrolateral aspect (89–92 respectively).
</paragraph>
</caption>
<paragraph id="145436D9FFACFF87CC68FF47088AFEB2" blockId="18.[113,772,234,1667]" pageId="18" pageNumber="17">
<emphasis id="269FEACBFFACFF87CC68FF4709A0FED3" bold="true" box="[113,236,234,258]" pageId="18" pageNumber="17">Coloration</emphasis>
(
<figureCitation id="8CD02A5CFFACFF87CD1FFF4608D9FED2" box="[262,405,235,259]" captionStart="Figures 69–73" captionStartId="15.[117,194,1980,2002]" captionTargetBox="[130,1451,248,1967]" captionTargetId="figure-45@15.[113,1474,234,1973]" captionTargetPageId="15" captionText="Figures 69–73. Scorpiops hofereki sp. n., female holotype. Dorsal (69) and ventral (70) views. Metasoma and telson lateral (71), ventral (72), and dorsal (73) views. Scale bars: 10 mm (69–70, 71–73)." figureDoi="http://doi.org/10.5281/zenodo.5741563" httpUri="https://zenodo.org/record/5741563/files/figure.png" pageId="18" pageNumber="17">Figs. 69–70</figureCitation>
). The base color is uniformly reddish brown (
<typeStatus id="CB50887BFFACFF87CD02FEA60830FEF2" box="[283,380,267,291]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
) to black (
<typeStatus id="CB50887BFFACFF87CDF6FEA60B1DFEF2" box="[495,593,267,291]" pageId="18" pageNumber="17" type="paratype">paratype</typeStatus>
). The telson and tarsomere II of legs yellowish to reddish brown. Chelicerae are reddish black and reticulate.
</paragraph>
<paragraph id="145436D9FFACFF87CC68FEC70B97FBD3" blockId="18.[113,772,234,1667]" pageId="18" pageNumber="17">
<emphasis id="269FEACBFFACFF87CC68FEC708D0FE52" bold="true" box="[113,412,362,387]" pageId="18" pageNumber="17">Carapace and mesosoma</emphasis>
(
<figureCitation id="8CD02A5CFFACFF87CDACFEC60B3CFE52" box="[437,624,363,387]" captionStart="Figures 83–92" captionStartId="17.[113,190,1983,2005]" captionTargetBox="[113,1474,234,1954]" captionTargetId="figure-55@17.[113,1474,234,1955]" captionTargetPageId="17" captionText="Figures 83–92. Scorpiops hofereki sp. n., female holotype. Carapace and tergites I–III (83), posterior coxosternal area and sternites III–IV (84). Right chelicera in dorsal (85) and ventral (86) views. Telson lateral (87). Pectine (88). Left legs I–IV, retrolateral aspect (89–92 respectively)." figureDoi="http://doi.org/10.5281/zenodo.5741567" httpUri="https://zenodo.org/record/5741567/files/figure.png" pageId="18" pageNumber="17">Figs. 83–84, 88</figureCitation>
). The entire carapace is covered with large and minute granules; carinae are absent but larger granules form two parallel rows in the anterior part of carapace. The anterior margin of the carapace is markedly depressed, convex in the middle. The carapace bears three lateral eyes, of which one can be reduced. The mesosoma is granulated, with one median carina. Tergite VII is pentacarinate. The sternites are finely granulated with two parallel furrows except sternite VII, which bears four sparsely granulate carinae. Pectinal teeth number
<quantity id="D3139B3CFFACFF87CE07FD260B2CFD72" box="[542,608,651,675]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="18" pageNumber="17" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
females, fulcra are absent. The pectens have marginal lamella I (basal) present, the remainder of the pectinal surface forming one compact unit.
<emphasis id="269FEACBFFACFF87CC68FD460827FCD2" bold="true" box="[113,363,747,771]" pageId="18" pageNumber="17">Metasoma and telson</emphasis>
(
<figureCitation id="8CD02A5CFFACFF87CD64FD460B4FFCD2" box="[381,515,747,771]" captionStart="Figures 69–73" captionStartId="15.[117,194,1980,2002]" captionTargetBox="[130,1451,248,1967]" captionTargetId="figure-45@15.[113,1474,234,1973]" captionTargetPageId="15" captionText="Figures 69–73. Scorpiops hofereki sp. n., female holotype. Dorsal (69) and ventral (70) views. Metasoma and telson lateral (71), ventral (72), and dorsal (73) views. Scale bars: 10 mm (69–70, 71–73)." figureDoi="http://doi.org/10.5281/zenodo.5741563" httpUri="https://zenodo.org/record/5741563/files/figure.png" pageId="18" pageNumber="17">Figs. 71–73</figureCitation>
,
<figureCitation id="8CD02A5CFFACFF87CE0BFD460B7CFCD2" box="[530,560,747,771]" captionStart="Figures 83–92" captionStartId="17.[113,190,1983,2005]" captionTargetBox="[113,1474,234,1954]" captionTargetId="figure-55@17.[113,1474,234,1955]" captionTargetPageId="17" captionText="Figures 83–92. Scorpiops hofereki sp. n., female holotype. Carapace and tergites I–III (83), posterior coxosternal area and sternites III–IV (84). Right chelicera in dorsal (85) and ventral (86) views. Telson lateral (87). Pectine (88). Left legs I–IV, retrolateral aspect (89–92 respectively)." figureDoi="http://doi.org/10.5281/zenodo.5741567" httpUri="https://zenodo.org/record/5741567/files/figure.png" pageId="18" pageNumber="17">87</figureCitation>
). The metasoma is very sparsely hirsute and granulated, with relatively large granules. Metasomal segment I with 10 carinae, II–IV with 8 carinae, and V with 5 carinae. The median lateral carinae of metasoma V are indicated by isolated granules that may coalesce into carinae. The dorsolateral carinae of segments III and IV are granulate with sharp granules, which are not posteriorly terminated with a pronounced tooth. The telson is bulbous and sparsely granulate, with annular ring present.
</paragraph>
<paragraph id="145436D9FFACFF87CC68FBA70889FA53" blockId="18.[113,772,234,1667]" pageId="18" pageNumber="17">
<emphasis id="269FEACBFFACFF87CC68FBA709ACFBF3" bold="true" box="[113,224,1034,1058]" pageId="18" pageNumber="17">Pedipalps</emphasis>
(
<figureCitation id="8CD02A5CFFACFF87CCEFFBA608CCFBF2" box="[246,384,1035,1059]" captionStart="Figures 74–82" captionStartId="16.[114,191,1956,1978]" captionTargetBox="[124,1410,249,1926]" captionTargetId="figure-66@16.[114,1475,234,1945]" captionTargetPageId="16" captionText="Figures 74–82. Scorpiops hofereki sp. n., female holotype, pedipalp segments. Chela dorsal (74), external (75) and ventral (76) views. Patella dorsal (77), external (78) and ventral (79) views. Femur and trochanter dorsal (80) and ventral (81) views. Movable finger dentition (82). Trichobothrial pattern is indicated by white circles." figureDoi="http://doi.org/10.5281/zenodo.5741565" httpUri="https://zenodo.org/record/5741565/files/figure.png" pageId="18" pageNumber="17">Figs. 74–82</figureCitation>
). The pedipalps are very sparsely hirsute. The patella bears 17 (5
<emphasis id="269FEACBFFACFF87CDCAFB8608A0FB92" box="[467,492,1067,1091]" italics="true" pageId="18" pageNumber="17">eb</emphasis>
, 2
<emphasis id="269FEACBFFACFF87CE0BFB860B7AFB92" box="[530,566,1067,1091]" italics="true" pageId="18" pageNumber="17">esb</emphasis>
, 2
<emphasis id="269FEACBFFACFF87CE44FB860B30FB93" box="[605,636,1067,1090]" italics="true" pageId="18" pageNumber="17">em</emphasis>
, 4
<emphasis id="269FEACBFFACFF87CEBAFB860B8DFB93" box="[675,705,1067,1090]" italics="true" pageId="18" pageNumber="17">est</emphasis>
, 4
<emphasis id="269FEACBFFACFF87CEFEFB860BB6FB93" box="[743,762,1067,1090]" italics="true" pageId="18" pageNumber="17">et</emphasis>
) external and 7 ventral trichobothria. The chela bears 4 ventral trichobothria located on ventral surface. The femur and patella are granulated. The femur has 5 granulose carinae and the patella has 5 complete carinae with dorsal and ventral patellar spurs present but reduced. The manus dorsally bears fine, rounded reticulated granules. The external surface of the chela is covered by minute and larger sparse granules, which form another almost complete carina. The movable fingers with 9–10 IAD, parallel with MD (
<quantity id="D3139B3CFFACFF87CDA8FAE60B5DFAB2" box="[433,529,1355,1379]" metricMagnitude="0" metricUnit="m" metricValue="1.6509999999999998" metricValueMax="1.7271999999999998" metricValueMin="1.5748" pageId="18" pageNumber="17" unit="in" value="65.0" valueMax="68.0" valueMin="62.0">62–68 in</quantity>
number) and there are also 5 ID and 9–10 OD present.
</paragraph>
<paragraph id="145436D9FFACFF87CC68FA260B59F9B2" blockId="18.[113,772,234,1667]" pageId="18" pageNumber="17">
<emphasis id="269FEACBFFACFF87CC68FA2609EAFA72" bold="true" box="[113,166,1419,1443]" pageId="18" pageNumber="17">Legs</emphasis>
(
<figureCitation id="8CD02A5CFFACFF87CCA3FA26080FFA72" box="[186,323,1419,1443]" captionStart="Figures 83–92" captionStartId="17.[113,190,1983,2005]" captionTargetBox="[113,1474,234,1954]" captionTargetId="figure-55@17.[113,1474,234,1955]" captionTargetPageId="17" captionText="Figures 83–92. Scorpiops hofereki sp. n., female holotype. Carapace and tergites I–III (83), posterior coxosternal area and sternites III–IV (84). Right chelicera in dorsal (85) and ventral (86) views. Telson lateral (87). Pectine (88). Left legs I–IV, retrolateral aspect (89–92 respectively)." figureDoi="http://doi.org/10.5281/zenodo.5741567" httpUri="https://zenodo.org/record/5741567/files/figure.png" pageId="18" pageNumber="17">Figs. 89–92</figureCitation>
). The tibia and tarsomeres of legs with several setae not arranged into bristlecombs on dorsal surfaces but with rows of spines on dorsolateral and ventrolateral surface. Tarsomere II of legs I–IV with 6–9 stout median ventral spines in a row and two other parallel spines. The femur bears 3–4 and patella, 4–5 incomplete carinae; both femur and patella are finely granulated.
</paragraph>
<paragraph id="145436D9FFACFF87CC68F9C608E7F952" blockId="18.[113,772,234,1667]" box="[113,427,1643,1667]" pageId="18" pageNumber="17">
<emphasis id="269FEACBFFACFF87CC68F9C60854F952" bold="true" box="[113,280,1643,1667]" pageId="18" pageNumber="17">Measurements</emphasis>
. See
<tableCitation id="59690362FFACFF87CD4AF9C608E9F952" box="[339,421,1643,1667]" captionStart="Table 1" captionStartId="3.[113,169,985,1006]" captionText="Table 1. Comparative measurements of adults of Scorpiops furai sp. n., S. grosseri sp. n., S. harmsi sp. n. and S. hofereki sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="18" pageNumber="17">Table 1</tableCitation>
.
</paragraph>
<paragraph id="145436D9FFACFF87CC68F9060864F873" blockId="18.[113,771,1707,1955]" pageId="18" pageNumber="17">
AFFINITIES. The described features distinguish
<taxonomicName id="D3EB4D5AFFACFF87CE71F9060B97F912" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[616,731,1707,1731]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="hofereki" status="sp. nov.">
<emphasis id="269FEACBFFACFF87CE71F9060B97F912" box="[616,731,1707,1731]" italics="true" pageId="18" pageNumber="17">S. hofereki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DAC57B0FFACFF87CEFAF90609CBF933" pageId="18" pageNumber="17" rank="species">
<emphasis id="269FEACBFFACFF87CEFAF9060BB0F912" bold="true" box="[739,764,1707,1731]" pageId="18" pageNumber="17">sp</emphasis>
.
<emphasis id="269FEACBFFACFF87CC68F96609CCF932" bold="true" box="[113,128,1739,1763]" pageId="18" pageNumber="17">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. The combination of four characters (chelal trichobothrium
<emphasis id="269FEACBFFACFF87CE13F9460B7CF8D8" box="[522,560,1771,1801]" italics="true" pageId="18" pageNumber="17">
Eb
<subScript id="886F349CFFACFF87CE31F9560B7CF8D8" attach="left" box="[552,560,1787,1801]" fontSize="6" pageId="18" pageNumber="17">3</subScript>
</emphasis>
located in proximal half of external surface between trichobothria
<emphasis id="269FEACBFFACFF87CE40F8A60B38F8F3" box="[601,628,1803,1826]" italics="true" pageId="18" pageNumber="17">Dt</emphasis>
and
<emphasis id="269FEACBFFACFF87CEBFF8A60B8BF8F2" box="[678,711,1803,1827]" italics="true" pageId="18" pageNumber="17">Db</emphasis>
; total length over
<quantity id="D3139B3CFFACFF87CCEAF887080EF893" box="[243,322,1834,1858]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="17" unit="mm" value="45.0">45 mm</quantity>
; fingers of pedipalps undulate in females; chela length to width ratio
<quantity id="D3139B3CFFACFF87CD8EF8E60B49F8B2" box="[407,517,1867,1891]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="7.874" metricValueMin="7.366" pageId="18" pageNumber="17" unit="in" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 in</quantity>
females; and patella of pedipalp with 7 ventral trichobothria) is unique in the entire genus
<taxonomicName id="D3EB4D5AFFACFF87CCAFF826086DF873" authorityName="Peters" authorityYear="1861" box="[182,289,1931,1954]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="269FEACBFFACFF87CCAFF826086DF873" box="[182,289,1931,1954]" italics="true" pageId="18" pageNumber="17">Scorpiops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="145436D9FFACFF87CC68F86608A9F832" blockId="18.[113,485,1995,2019]" box="[113,485,1995,2019]" pageId="18" pageNumber="17">
DISTRIBUTION.
<collectingCountry id="6CFC7649FFACFF87CD0BF8660820F832" box="[274,364,1995,2019]" name="Pakistan" pageId="18" pageNumber="17">Pakistan</collectingCountry>
(
<figureCitation id="8CD02A5CFFACFF87CD62F8660895F832" box="[379,473,1995,2019]" captionStart="Figure 240" captionStartId="41.[113,181,1361,1383]" captionTargetBox="[113,1474,234,1350]" captionTargetId="figure-353@41.[113,1474,234,1350]" captionTargetPageId="41" captionText="Figure 240. Map of distribution of nine new species described here and Scorpiops vonwicki Birula, 1913." figureDoi="http://doi.org/10.5281/zenodo.5741606" httpUri="https://zenodo.org/record/5741606/files/figure.png" pageId="18" pageNumber="17">Fig. 240</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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