345 lines
36 KiB
XML
345 lines
36 KiB
XML
<document id="2B1473C48B839B5659732B8D65690706" ID-DOI="10.5281/zenodo.3726987" ID-GBIF-Dataset="80783542-5446-42b3-a9bd-177b0e4ec73a" ID-ISSN="1942-1354" ID-Zenodo-Dep="3726987" ID-ZooBank="CF6C1B9C-2EC0-4220-BED7-6402A6C9D175" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1585149460741" checkinUser="felipe" docAuthor="Zidek, Jiri & Pokorný, Svatopluk" docDate="2018" docId="1E76DE47FFEE312152FAF919EF54FE27" docLanguage="en" docName="InsectaMundi.0611.1-35.pdf.imf" docOrigin="Insecta Mundi 611" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Scarabaeus (Scarabaeolus) lizleri Zidek and Pokorny 2018, sp. n." docType="treatment" docVersion="5" lastPageNumber="5" masterDocId="E24FA63FFFEA3127523AFFD7EC13FFD4" masterDocTitle="New species of the Scarabaeus subgenus Scarabaeolus Balthasar, with a review of the subgenus (Scarabaeidae: Scarabaeinae: Scarabaeini)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="3" updateTime="1698731770912" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="C2BD7D064B220C481877143B0F123BE6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="67CD5ACFF4B8E1235D4544B804CA1469">
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<mods:title id="90C47B3BB663691892D91AB1251AEB56">New species of the Scarabaeus subgenus Scarabaeolus Balthasar, with a review of the subgenus (Scarabaeidae: Scarabaeinae: Scarabaeini)</mods:title>
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</mods:titleInfo>
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<mods:name id="D4DDC14B17E2D5C480B3727DE4E5E849" type="personal">
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<mods:roleTerm id="CDEA63247A1AEC09E8A33E508E884493">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="52C3DA8E80C1B5DB2666C445FDF17A76">Zidek, Jiri</mods:namePart>
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</mods:name>
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<mods:name id="5F9B2043D2D5A311237BD8749CE1DB1F" type="personal">
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<mods:role id="DEFB87770C5B548589994AC36D5BD9A1">
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<mods:roleTerm id="6260A28097D3E20DCF04BD6A3E5D0307">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0F7F8F759E65ADB4A45452B5AAA5E2C8">Pokorný, Svatopluk</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2D822C7C1922199C3BAAA3D70847D7FE">text</mods:typeOfResource>
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<mods:relatedItem id="CED9042CC9276CAFC687356070787892" type="host">
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<mods:titleInfo id="3F9BACA01990C69426D7EC06D81DB809">
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<mods:title id="2A7CC07799A775C8FBFE2200FCC2CE6A">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="23CF908C9616B3612BB77BC6EBE205EF">
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<mods:date id="87898974A24291DDB1598351447A84E2">2018</mods:date>
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<mods:detail id="1B939F5AF68F35738034B76E4983BBBB" type="pubDate">
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<mods:number id="F985740F1BB6841B9560B96DCCC2F424">2018-03-30</mods:number>
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</mods:detail>
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<mods:detail id="BB56AFBAEDE7C5EBBF514EB6E6ADB1D5" type="issue">
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<mods:number id="80DA5256FE89F145A1F339BDB0F6ACA0">611</mods:number>
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</mods:detail>
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<mods:extent id="034F5CCBC6D461921516156D1F7C1B78" unit="page">
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<mods:start id="A74D6AD66ACA1B742A32FC47F4D2707D">1</mods:start>
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<mods:end id="60CA1917813F6DA6FDCCEAA598A9D66A">35</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="482A5CEC72EEB1F9F80E7A0547BFB7EB">journal article</mods:classification>
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<mods:identifier id="F26945F887D32E1E474CC939AE2359C1" type="DOI">10.5281/zenodo.3726987</mods:identifier>
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<mods:identifier id="A86029DC249D8F6D397B4177FDFD2C75" type="GBIF-Dataset">80783542-5446-42b3-a9bd-177b0e4ec73a</mods:identifier>
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<mods:identifier id="5E2B7F05AF1C6CA985D7C32FDC86E9D7" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="DD6BB0287C35D898B5AF75279509BD0D" type="Zenodo-Dep">3726987</mods:identifier>
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<mods:identifier id="8707CFC0435ABC088715AD433FBFB0EE" type="ZooBank">CF6C1B9C-2EC0-4220-BED7-6402A6C9D175</mods:identifier>
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</mods:mods>
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<treatment id="1E76DE47FFEE312152FAF919EF54FE27" ID-GBIF-Taxon="163288763" LSID="urn:lsid:plazi:treatment:1E76DE47FFEE312152FAF919EF54FE27" httpUri="http://treatment.plazi.org/id/1E76DE47FFEE312152FAF919EF54FE27" lastPageId="6" lastPageNumber="5" pageId="4" pageNumber="3">
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<subSubSection id="DEC53CDAFFEE312352FAF919EFE5F933" box="[192,1014,1742,1767]" pageId="4" pageNumber="3" type="nomenclature">
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<paragraph id="96606F51FFEE312352FAF919EFE5F933" blockId="4.[192,1014,1742,1799]" box="[192,1014,1742,1767]" pageId="4" pageNumber="3">
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<heading id="CD28D83DFFEE312352FAF919EFE5F933" bold="true" box="[192,1014,1742,1767]" fontSize="10" level="2" pageId="4" pageNumber="3" reason="3">
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<emphasis id="A4ABB343FFEE312352FAF919EFE5F933" bold="true" box="[192,1014,1742,1767]" pageId="4" pageNumber="3">
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||
<taxonomicName id="51DF14D2FFEE312352FAF919EFB2F932" ID-CoL="6Y869" authority="Zidek and Pokorny" authorityName="Zidek and Pokorny" authorityYear="2018" box="[192,929,1742,1767]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov." subGenus="Scarabaeolus">
|
||
<emphasis id="A4ABB343FFEE312352FAF919ED73F932" bold="true" box="[192,352,1742,1766]" italics="true" pageId="4" pageNumber="3">Scarabaeus</emphasis>
|
||
(
|
||
<emphasis id="A4ABB343FFEE31235348F919EE3FF932" bold="true" box="[370,556,1742,1766]" italics="true" pageId="4" pageNumber="3">Scarabaeolus</emphasis>
|
||
)
|
||
<emphasis id="A4ABB343FFEE31235004F919EE83F932" bold="true" box="[574,656,1742,1766]" italics="true" pageId="4" pageNumber="3">lizleri</emphasis>
|
||
Zidek and Pokorný
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="BF980E38FFEE31235195F918EFE5F933" box="[943,1014,1743,1767]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFEE312352FAF938ED21F8D3" box="[192,306,1775,1799]" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="96606F51FFEE312352FAF938ED21F8D3" blockId="4.[192,1014,1742,1799]" box="[192,306,1775,1799]" pageId="4" pageNumber="3">
|
||
<figureCitation id="0EE473D4FFEE312352FAF938ED21F8D3" box="[192,306,1775,1799]" captionStart="Figures 9–16" captionStartId="26.[192,285,1819,1841]" captionTargetBox="[229,1402,308,1746]" captionTargetId="figure@26.[806,1402,1046,1747]" captionTargetPageId="26" captionText="Figures 9–16. Scarabaeus (Scarabaeolus) spp. 9–12) Scarabaeus (Scarabaeolus) lizleri sp. n., HTm, 15 mm. 9) Dorsal habitus. 10) Ventral habitus. 11) Aedeagus, dorsal. 12) Aedeagus, left lateral. 13–16) Scarabaeus (Scarabaeolus) rubripennis (Boheman), 14 mm. 13) Dorsal habitus. 14) Ventral habitus. 15) Aedeagus, dorsal. 16) Aedeagus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3726991" httpUri="https://zenodo.org/record/3726991/files/figure.png" pageId="4" pageNumber="3">Fig. 9–12</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFEE312352FAF8F7E849F8EC" box="[192,1114,1823,1848]" pageId="4" pageNumber="3" type="distribution">
|
||
<paragraph id="96606F51FFEE312352FAF8F7E849F8EC" blockId="4.[192,1114,1823,1848]" box="[192,1114,1823,1848]" pageId="4" pageNumber="3">
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||
<emphasis id="A4ABB343FFEE312352FAF8F7ED68F8E3" bold="true" box="[192,379,1823,1848]" pageId="4" pageNumber="3">
|
||
<typeStatus id="4964D1F3FFEE312352FAF8F7ED17F8EC" box="[192,260,1824,1848]" pageId="4" pageNumber="3">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="EEC82FC1FFEE312353B8F8F7EDAFF8EC" box="[386,444,1824,1848]" name="South Africa" pageId="4" pageNumber="3">RSA</collectingCountry>
|
||
,
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||
<collectingRegion id="541BA1B3FFEE312353F2F8F7EEE7F8EC" box="[456,756,1824,1848]" country="South Africa" name="Northern Cape" pageId="4" pageNumber="3">Northern Cape Province</collectingRegion>
|
||
,
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||
<quantity id="5127C2B4FFEE3123513AF8F7EF5FF8EC" box="[768,844,1824,1848]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="3" unit="km" value="30.0">30 km</quantity>
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||
SE of Alexander Bay.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFEE312252FAF887ED2FFEA6" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3" type="materials_examined">
|
||
<paragraph id="96606F51FFEE312352FAF887E9B3F85C" blockId="4.[192,1440,1872,1929]" pageId="4" pageNumber="3">
|
||
<materialsCitation id="26B7650CFFEE312352FAF887EFDDF8BC" ID-GBIF-Occurrence="2580374386" box="[192,974,1872,1897]" collectingDate="2000-10-09" collectionCode="R" country="South Africa" location="Lizler" pageId="4" pageNumber="3" specimenCount="2" typeStatus="type">
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||
<emphasis id="A4ABB343FFEE312352FAF887ED9AF8BC" bold="true" box="[192,393,1872,1896]" pageId="4" pageNumber="3">
|
||
<typeStatus id="4964D1F3FFEE312352FAF887ED17F8BC" box="[192,260,1872,1896]" pageId="4" pageNumber="3">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
HTm from
|
||
<typeStatus id="4964D1F3FFEE31235022F886EE5EF8BD" box="[536,589,1873,1897]" pageId="4" pageNumber="3">type</typeStatus>
|
||
locality, leg.
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||
<collectionCode id="F0CEF794FFEE312350D5F886EF15F8BC" box="[751,774,1873,1896]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
|
||
.
|
||
<location id="9300398AFFEE3123512AF887EF4BF8BC" LSID="urn:lsid:plazi:treatment:1E76DE47FFEE312152FAF919EF54FE27:9300398AFFEE3123512AF887EF4BF8BC" box="[784,856,1872,1896]" country="South Africa" name="Lizler" pageId="4" pageNumber="3">Lízler</location>
|
||
<date id="E2614991FFEE3123515AF887EFDDF8BC" box="[864,974,1872,1896]" pageId="4" pageNumber="3" value="2000-10-09">
|
||
<collectingDate id="F225B079FFEE3123515AF887EFDDF8BC" box="[864,974,1872,1896]" pageId="4" pageNumber="3" value="2000-10-09">9.X.2000</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="26B7650CFFEE312352D2F8A6EF9DF85D" ID-GBIF-Occurrence="2580374351" box="[232,910,1904,1929]" collectingDate="2000-10-09" collectionCode="R" country="South Africa" location="Lizler" pageId="4" pageNumber="3" specimenCount="1" typeStatus="type">
|
||
PT (179): 172 from
|
||
<typeStatus id="4964D1F3FFEE312353E9F8A6EE1BF85D" box="[467,520,1905,1929]" pageId="4" pageNumber="3">type</typeStatus>
|
||
locality, leg.
|
||
<collectionCode id="F0CEF794FFEE31235090F8A6EED2F85C" box="[682,705,1905,1928]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="3">R</collectionCode>
|
||
.
|
||
<location id="9300398AFFEE312350F6F8A7EF07F85C" LSID="urn:lsid:plazi:treatment:1E76DE47FFEE312152FAF919EF54FE27:9300398AFFEE312350F6F8A7EF07F85C" box="[716,788,1904,1928]" country="South Africa" name="Lizler" pageId="4" pageNumber="3">Lízler</location>
|
||
<date id="E2614991FFEE31235121F8A6EF9AF85D" box="[795,905,1905,1929]" pageId="4" pageNumber="3" value="2000-10-09">
|
||
<collectingDate id="F225B079FFEE31235121F8A6EF9AF85D" box="[795,905,1905,1929]" pageId="4" pageNumber="3" value="2000-10-09">9.X.2000</collectingDate>
|
||
</date>
|
||
;
|
||
</materialsCitation>
|
||
\\
|
||
<materialsCitation id="26B7650CFFEE31235186F8A6E9B3F85C" ID-GBIF-Occurrence="2580374316" box="[956,1440,1904,1929]" collectingDate="1976-10-04" collectorName="Endrodi-Younga" country="South Africa" location="Port Nolloth" municipality="Richtersveld" pageId="4" pageNumber="3" specimenCount="1">
|
||
2 from
|
||
<collectingCountry id="EEC82FC1FFEE31235629F8A6E85EF85D" box="[1043,1101,1905,1929]" name="South Africa" pageId="4" pageNumber="3">RSA</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="7604F52BFFEE31235662F8A7E8E6F85C" box="[1112,1269,1904,1928]" pageId="4" pageNumber="3">Richtersveld</collectingMunicipality>
|
||
,
|
||
<location id="9300398AFFEE3123573BF8A6E988F85C" LSID="urn:lsid:plazi:treatment:1E76DE47FFEE312152FAF919EF54FE27:9300398AFFEE3123573BF8A6E988F85C" box="[1281,1435,1904,1928]" country="South Africa" municipality="Richtersveld" name="Port Nolloth" pageId="4" pageNumber="3">Port Nolloth</location>
|
||
,
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252FAFF0EE854FEE6" blockId="5.[192,1440,217,370]" pageId="5" pageNumber="4">
|
||
leg.
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||
<collectorName id="3B2A0A87FFEF312252CAFF0EEDA4FF25" box="[240,439,217,241]" pageId="5" pageNumber="4">Endrödi-Younga</collectorName>
|
||
<date id="E2614991FFEF31225386FF0EEE3BFF25" box="[444,552,217,241]" pageId="5" pageNumber="4" value="1976-10-04">
|
||
<collectingDate id="F225B079FFEF31225386FF0EEE3BFF25" box="[444,552,217,241]" pageId="5" pageNumber="4" value="1976-10-04">4.X.1976</collectingDate>
|
||
</date>
|
||
;
|
||
<emphasis id="A4ABB343FFEF3122500BFF0EEE42FF25" bold="true" box="[561,593,217,241]" pageId="5" pageNumber="4">\\</emphasis>
|
||
<materialsCitation id="26B7650CFFEF3122506CFF0DE98FFF25" ID-GBIF-Occurrence="2580374313" box="[598,1436,217,242]" collectingDate="1976-10-10" collectorName="Endrodi-Younga" location="Manganese" municipality="Richtersveld" pageId="5" pageNumber="4" specimenCount="1">
|
||
1 from
|
||
<collectingMunicipality id="7604F52BFFEF31225092FF0EEF50FF25" box="[680,835,217,241]" pageId="5" pageNumber="4">Richtersveld</collectingMunicipality>
|
||
,
|
||
<location id="9300398AFFEF31225176FF0DEFC5FF26" LSID="urn:lsid:plazi:treatment:1E76DE47FFEE312152FAF919EF54FE27:9300398AFFEF31225176FF0DEFC5FF26" box="[844,982,218,242]" municipality="Richtersveld" name="Manganese" pageId="5" pageNumber="4">Manganese</location>
|
||
mine, leg.
|
||
<collectorName id="3B2A0A87FFEF3122566FFF0EE90FFF25" box="[1109,1308,217,241]" pageId="5" pageNumber="4">Endrödi-Younga</collectorName>
|
||
<date id="E2614991FFEF3122571AFF0EE98FFF25" box="[1312,1436,217,241]" pageId="5" pageNumber="4" value="1976-10-10">
|
||
<collectingDate id="F225B079FFEF3122571AFF0EE98FFF25" box="[1312,1436,217,241]" pageId="5" pageNumber="4" value="1976-10-10">10.X.1976</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; \\
|
||
<materialsCitation id="26B7650CFFEF312252D3FF2DE8C5FEC6" ID-GBIF-Occurrence="2580374333" box="[233,1238,250,274]" collectingDate="1973-09-20" collectorName="Endrodi-Younga" country="Namibia" location="Obib" municipality="Namib" pageId="5" pageNumber="4" specimenCount="1">
|
||
2 from S.W. Afr. [
|
||
<collectingCountry id="EEC82FC1FFEF312253FEFF2DEE26FEC6" box="[452,565,250,274]" name="Namibia" pageId="5" pageNumber="4">Namibia</collectingCountry>
|
||
],
|
||
<collectingMunicipality id="7604F52BFFEF31225073FF2DEE8CFEC6" box="[585,671,250,274]" pageId="5" pageNumber="4">Namib</collectingMunicipality>
|
||
,
|
||
<location id="9300398AFFEF31225096FF2DEEF5FEC6" LSID="urn:lsid:plazi:treatment:1E76DE47FFEE312152FAF919EF54FE27:9300398AFFEF31225096FF2DEEF5FEC6" box="[684,742,250,274]" country="Namibia" municipality="Namib" name="Obib" pageId="5" pageNumber="4">Obib</location>
|
||
dunes, leg.
|
||
<collectorName id="3B2A0A87FFEF31225147FF2DE855FEC6" box="[893,1094,250,274]" pageId="5" pageNumber="4">Endrödi-Younga</collectorName>
|
||
<date id="E2614991FFEF31225675FF2DE8C5FEC6" box="[1103,1238,250,274]" pageId="5" pageNumber="4" value="1973-09-20">
|
||
<collectingDate id="F225B079FFEF31225675FF2DE8C5FEC6" box="[1103,1238,250,274]" pageId="5" pageNumber="4" value="1973-09-20">20.IX.1973</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; \\
|
||
<materialsCitation id="26B7650CFFEF31225737FF2DE850FEE6" ID-GBIF-Occurrence="2580374346" collectingDate="1981-07-28" collectorName="S. Namib & Endrodi-Younga" country="Namibia" location="Oranjemund" pageId="5" pageNumber="4" specimenCount="1">
|
||
2 from S.W. Afr. [
|
||
<collectingCountry id="EEC82FC1FFEF312252C7FECDED7DFEE6" box="[253,366,282,306]" name="Namibia" pageId="5" pageNumber="4">Namibia</collectingCountry>
|
||
],
|
||
<collectorName id="3B2A0A87FFEF312253BBFECDEDE5FEE6" box="[385,502,282,306]" pageId="5" pageNumber="4">S. Namib</collectorName>
|
||
,
|
||
<location id="9300398AFFEF31225038FECDEEB2FEE6" LSID="urn:lsid:plazi:treatment:1E76DE47FFEE312152FAF919EF54FE27:9300398AFFEF31225038FECDEEB2FEE6" box="[514,673,282,306]" country="Namibia" name="Oranjemund" pageId="5" pageNumber="4">Oranjemund</location>
|
||
, leg.
|
||
<collectorName id="3B2A0A87FFEF312250E5FECDEFBBFEE6" box="[735,936,282,306]" pageId="5" pageNumber="4">Endrödi-Younga</collectorName>
|
||
<date id="E2614991FFEF3122518AFECDE850FEE6" box="[944,1091,282,306]" pageId="5" pageNumber="4" value="1981-07-28">
|
||
<collectingDate id="F225B079FFEF3122518AFECDE850FEE6" box="[944,1091,282,306]" pageId="5" pageNumber="4" value="1981-07-28">28.VII.1981</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FEEDED2FFEA6" blockId="5.[192,1440,217,370]" pageId="5" pageNumber="4">
|
||
<materialsCitation id="26B7650CFFEF312252D2FEEDED2FFEA6" ID-GBIF-Occurrence="2580374390" collectionCode="TMSA, BMNH, OXUM, MNHB, NMPC, GWPC" pageId="5" pageNumber="4" specimenCount="1">
|
||
HT+10 PT at
|
||
<collectionCode id="F0CEF794FFEF312253A9FEEDEDF6FE86" box="[403,485,314,338]" collectionName="South Africa, Gauteng, Pretoria, Transvaal Museum" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="4">TMSA</collectionCode>
|
||
, 10 PT at
|
||
<collectionCode id="F0CEF794FFEF3122505EFEEDEEACFE85" box="[612,703,314,337]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="5" pageNumber="4">BMNH</collectionCode>
|
||
, 10 PT at
|
||
<collectionCode id="F0CEF794FFEF31225105FEEDEF8BFE86" box="[831,920,314,338]" collectionName="United Kingdom, Oxford, University Museum of Natural History" pageId="5" pageNumber="4">OXUM</collectionCode>
|
||
, 10 PT at
|
||
<collectionCode id="F0CEF794FFEF31225622FEEDE860FE85" box="[1048,1139,314,337]" collectionName="MNHB" pageId="5" pageNumber="4">MNHB</collectionCode>
|
||
, 8 PT at
|
||
<collectionCode id="F0CEF794FFEF312256DEFEEDE928FE86" box="[1252,1339,314,338]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="4">NMPC</collectionCode>
|
||
, 131 PT at
|
||
<collectionCode id="F0CEF794FFEF312252DAFE8DED2BFEA6" box="[224,312,346,370]" collectionName="GWPC" pageId="5" pageNumber="4">GWPC</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFEF312252FAFE5DED88FE16" pageId="5" pageNumber="4" type="etymology">
|
||
<paragraph id="96606F51FFEF312252FAFE5DED88FE16" blockId="5.[192,1440,394,451]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252FAFE5DED48FE76" bold="true" box="[192,347,394,418]" pageId="5" pageNumber="4">Etymology.</emphasis>
|
||
Named for Robert Lízler of
|
||
<collectingRegion id="541BA1B3FFEF312250FDFE5DEF9EFE76" box="[711,909,394,418]" country="Czech Republic" name="Kralovehradecky" pageId="5" pageNumber="4">Hradec Králové</collectingRegion>
|
||
,
|
||
<collectingCountry id="EEC82FC1FFEF312251A7FE5DE84CFE76" box="[925,1119,394,418]" name="Czech Republic" pageId="5" pageNumber="4">Czech Republic</collectingCountry>
|
||
, who collected nearly the entire
|
||
<typeStatus id="4964D1F3FFEF3122532AFE7CED56FE17" box="[272,325,427,451]" pageId="5" pageNumber="4">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFEF312252FAFE0DE9B3F943" pageId="5" pageNumber="4" type="description">
|
||
<paragraph id="96606F51FFEF312252FAFE0DE8D9FE26" blockId="5.[192,1441,474,1270]" box="[192,1226,474,498]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252FAFE0DEE1FFE26" bold="true" box="[192,524,474,498]" pageId="5" pageNumber="4">
|
||
Description of
|
||
<typeStatus id="4964D1F3FFEF312253B6FE0DEE15FE26" box="[396,518,474,498]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Length
|
||
<quantity id="5127C2B4FFEF31225048FE0DEED9FE26" box="[626,714,474,498]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="4" unit="mm" value="15.0">15 mm</quantity>
|
||
. Black with reddish-orange elytra, glossy.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FE2DEEF1FD07" blockId="5.[192,1441,474,1270]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FE2DED22FDC6" box="[232,305,506,530]" italics="true" pageId="5" pageNumber="4">Head.</emphasis>
|
||
Clypeus without ventral keel, teeth weekly upturned, spaces between them U-shaped, space between medial teeth wider than that between medial and lateral teeth; anterior part with sparse asperate and setose punctures that posteriorly change to setose granules. Genae separated from clypeus by narrow excisions, their sculpture somewhat finer than that of clypeus, front end of each gena sharply pointed. Clypeus/frons boundary with an inconspicuous broad and smooth tubercle. Smooth sagittal line present throughout length, narrow on clypeus, widest on frons and narrowing toward vertex. Pubescence rusty brown, antennal club brownish yellow.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FD0BEDDFFCA0" blockId="5.[192,1441,474,1270]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FD0BED7AFD27" box="[232,361,732,755]" italics="true" pageId="5" pageNumber="4">Pronotum.</emphasis>
|
||
Bordered all around, with a distinct smooth sagittal line, disc with sparse deep, setose punctures that become smaller and more closely spaced anteriorly and laterally. Lateral margins evenly rounded, coarsely serrate, with long hairs. Front angles obtuse, pointing outward, hind corners evenly rounded. Basal margin lined by a shallow groove with irregularly spaced setose punctures; medial part of base weakly lobate.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FCABEED3FC40" blockId="5.[192,1441,474,1270]" box="[232,704,892,916]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FCABED7FFC40" box="[232,364,892,916]" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="51DF14D2FFEF312252D2FCABED7BFC40" box="[232,360,892,916]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Exposed, small, triangular.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FC4BE9B3FC21" blockId="5.[192,1441,474,1270]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FC4BED2EFC60" box="[232,317,924,948]" italics="true" pageId="5" pageNumber="4">Elytra.</emphasis>
|
||
With humeri well developed, indistinctly striate, striae micropunctate. Intervals smooth and glossy, first interval flat on disc, convex and sparsely punctate posterolaterally. Second through fifth intervals nearly impunctate; sixth and seventh intervals weakly arched; suture and first interval black.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FC2BE999FBC1" blockId="5.[192,1441,474,1270]" box="[232,1418,1020,1045]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FC2BED75FBC0" box="[232,358,1020,1044]" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="51DF14D2FFEF312252D2FC2BED71FBC0" box="[232,354,1020,1044]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="4" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Obliquely triangular, bordered all around, smooth on disc and punctate near margins.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FBCAED8CFB81" blockId="5.[192,1441,474,1270]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FBCAED2CFBE0" box="[232,319,1053,1076]" italics="true" pageId="5" pageNumber="4">Venter.</emphasis>
|
||
Metasternum with posterior longitudinal furrow and keeled anterior process. Ventral pubescence dark brown.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FB89E8B3FB61" blockId="5.[192,1441,474,1270]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FB89ED34FBA1" box="[232,295,1118,1141]" italics="true" pageId="5" pageNumber="4">Legs.</emphasis>
|
||
Protibia curved inward at level of second tooth, lateral margin densely serrate, medial margin densely granulose. Profemoral anteroventral carina without proximal tooth. Mesotibia with two spurs. Lateral margin of metatibia without transverse carinae. Pubescence dark brown.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252D2FB6AEE83FB22" blockId="5.[192,1441,474,1270]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252D2FB6AED70FB01" box="[232,355,1213,1237]" italics="true" pageId="5" pageNumber="4">Aedeagus.</emphasis>
|
||
Parameres shorter than phallobase, in lateral view dorso-ventrally parallel- sided, without ventral tooth and with truncated tips.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252FAFAD9E8EFFA92" blockId="5.[192,1440,1294,1350]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252FAFAD9ED4DFAF2" bold="true" box="[192,350,1294,1318]" pageId="5" pageNumber="4">Variability.</emphasis>
|
||
Length
|
||
<quantity id="5127C2B4FFEF312253F8FAD9EE57FAF1" box="[450,580,1294,1318]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="4" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
|
||
, elytra prevalently bright to dark reddish orange, frequently with scattered small blackish spots, but in some specimens entirely dark brown with reddish hue.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEF312252FAFA89E9B3F943" blockId="5.[192,1440,1374,1687]" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252FAFA89ED7DFAA2" bold="true" box="[192,366,1374,1398]" pageId="5" pageNumber="4">Comparison.</emphasis>
|
||
The dorsal habitus of the new species is indistinguishable from
|
||
<taxonomicName id="51DF14D2FFEF31225652FA89E9B3FAA2" authority="(Boheman)" baseAuthorityName="Boheman" box="[1128,1440,1374,1398]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="rubripennis">
|
||
<emphasis id="A4ABB343FFEF31225652FA89E906FAA2" box="[1128,1301,1374,1398]" italics="true" pageId="5" pageNumber="4">S. rubripennis</emphasis>
|
||
(Boheman)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0EE473D4FFEF312252F3FAA9ED5FFA42" box="[201,332,1406,1430]" captionStart="Figures 9–16" captionStartId="26.[192,285,1819,1841]" captionTargetBox="[229,1402,308,1746]" captionTargetId="figure@26.[806,1402,1046,1747]" captionTargetPageId="26" captionText="Figures 9–16. Scarabaeus (Scarabaeolus) spp. 9–12) Scarabaeus (Scarabaeolus) lizleri sp. n., HTm, 15 mm. 9) Dorsal habitus. 10) Ventral habitus. 11) Aedeagus, dorsal. 12) Aedeagus, left lateral. 13–16) Scarabaeus (Scarabaeolus) rubripennis (Boheman), 14 mm. 13) Dorsal habitus. 14) Ventral habitus. 15) Aedeagus, dorsal. 16) Aedeagus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3726991" httpUri="https://zenodo.org/record/3726991/files/figure.png" pageId="5" pageNumber="4">Fig. 13–16</figureCitation>
|
||
). In ventral view
|
||
<taxonomicName id="51DF14D2FFEF3122501AFAA9EE94FA42" authority="Zidek & Pokorný, 2018" authorityName="Zidek & Pokorný" authorityYear="2018" box="[544,647,1406,1430]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">
|
||
<emphasis id="A4ABB343FFEF3122501AFAA9EE94FA42" box="[544,647,1406,1430]" italics="true" pageId="5" pageNumber="4">S. lizleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF980E38FFEF312250B4FAA8EEDDFA42" box="[654,718,1407,1430]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
has a larger and more keeled metasternal process, but this cannot serve as a reliable diagnostic character. However, the two species can be readily separated on the presence of a large, sharp anteroventral tooth at the proximal end of the profemur in
|
||
<taxonomicName id="51DF14D2FFEF31225736FA68ECF6FA22" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="rubripennis">
|
||
<emphasis id="A4ABB343FFEF31225736FA68E90DFA02" box="[1292,1310,1471,1494]" italics="true" pageId="5" pageNumber="4">S</emphasis>
|
||
.
|
||
<emphasis id="A4ABB343FFEF31225717FA69ECF6FA22" italics="true" pageId="5" pageNumber="4">rubripennis</emphasis>
|
||
</taxonomicName>
|
||
, which is absent in the new species. The aedeagi of both species do not have a ventral tooth (merely a minute, barely noticeable tubercle) and differ only in proportions, with the phallobase longer than the parameres in
|
||
<taxonomicName id="51DF14D2FFEF312253A4F9C8EE14F9E3" authorityName="Zidek & Pokorný" authorityYear="2018" box="[414,519,1567,1591]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="lizleri">
|
||
<emphasis id="A4ABB343FFEF312253A4F9C8EDA3F9E2" box="[414,432,1567,1590]" italics="true" pageId="5" pageNumber="4">S</emphasis>
|
||
.
|
||
<emphasis id="A4ABB343FFEF312253FAF9C8EE14F9E3" box="[448,519,1567,1591]" italics="true" pageId="5" pageNumber="4">lizleri</emphasis>
|
||
</taxonomicName>
|
||
and vice versa in
|
||
<taxonomicName id="51DF14D2FFEF312250D7F9C8EF8DF9E3" box="[749,926,1567,1591]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="rubripennis">
|
||
<emphasis id="A4ABB343FFEF312250D7F9C8EEECF9E2" box="[749,767,1567,1590]" italics="true" pageId="5" pageNumber="4">S</emphasis>
|
||
.
|
||
<emphasis id="A4ABB343FFEF31225135F9C8EF8DF9E3" box="[783,926,1567,1591]" italics="true" pageId="5" pageNumber="4">rubripennis</emphasis>
|
||
</taxonomicName>
|
||
. Another species with reddish-brown but much darker elytra is
|
||
<taxonomicName id="51DF14D2FFEF312253EBF9E8EEABF983" authority="Ferreira" authorityName="Ferreira" authorityYear="1958" box="[465,696,1599,1623]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="knobeli">
|
||
<emphasis id="A4ABB343FFEF312253EBF9E8EDF0F982" box="[465,483,1599,1622]" italics="true" pageId="5" pageNumber="4">S</emphasis>
|
||
.
|
||
<emphasis id="A4ABB343FFEF312253C8F9E8EE59F983" box="[498,586,1599,1623]" italics="true" pageId="5" pageNumber="4">knobeli</emphasis>
|
||
Ferreira
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="EEC82FC1FFEF3122514FF9E8EFD8F983" box="[885,971,1599,1623]" name="Angola" pageId="5" pageNumber="4">Angola</collectingCountry>
|
||
(HT) and northwestern
|
||
<collectingCountry id="EEC82FC1FFEF312256CCF9E8E973F983" box="[1270,1376,1599,1623]" name="Namibia" pageId="5" pageNumber="4">Namibia</collectingCountry>
|
||
(
|
||
<figureCitation id="0EE473D4FFEF3122574AF9E8ED1FF9A3" captionStart="Figures 17–24" captionStartId="27.[192,287,1819,1841]" captionTargetBox="[209,1426,270,1756]" captionTargetId="figure@27.[810,1426,270,1582]" captionTargetPageId="27" captionText="Figures 17–24. Scarabaeus (Scarabaeolus) spp. 17–20) Scarabaeus (Scarabaeolus) namibensis sp. n., HTm, 13.5 mm. 17) Dorsal habitus. 18) Ventral habitus. 19) Aedeagus, dorsal. 20) Aedeagus, left lateral. 21–24) Scarabaeus (Scarabaeolus) knobeli Ferreira, 12 mm. 21) Dorsal habitus. 22) Ventral habitus. 23) Aedeagus, dorsal. 24) Aedeagus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3726993" httpUri="https://zenodo.org/record/3726993/files/figure.png" pageId="5" pageNumber="4">Fig. 21–24</figureCitation>
|
||
), which can be separated from the other two species on having sparsely and shallowly punctate elytral intervals, only one mesotibial spur, and
|
||
<taxonomicName id="51DF14D2FFEF312250C7F9A8EF63F943" box="[765,880,1663,1687]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="11" phylum="Chordata" rank="genus">pygidium</taxonomicName>
|
||
terminating in a short, outward curved spine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFEF312152FAF978EF54FE27" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="biology_ecology">
|
||
<paragraph id="96606F51FFEF312152FAF978EED2FEC6" blockId="5.[192,1440,1711,1928]" lastBlockId="6.[192,1440,217,499]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
|
||
<emphasis id="A4ABB343FFEF312252FAF978ED23F913" bold="true" box="[192,304,1711,1735]" pageId="5" pageNumber="4">Biology.</emphasis>
|
||
According to R. Lízler (in litt.) the
|
||
<typeStatus id="4964D1F3FFEF312250D7F967EF31F91C" box="[749,802,1712,1736]" pageId="5" pageNumber="4">type</typeStatus>
|
||
locality is about
|
||
<quantity id="5127C2B4FFEF312251C0F967E85BF91C" box="[1018,1096,1712,1736]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="4" unit="km" value="10.0">10 km</quantity>
|
||
from the shore, only
|
||
<quantity id="5127C2B4FFEF3122576CF967ECCBF933" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="4" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">10–20 m</quantity>
|
||
above the sea level, and has the character of a semidesert with small dunes and interspersed rocky outcrops. Vegetation consists of sparse “tamarysk” brush and succulents, and herbivorous mammals include lagomorphs, small antelope, goat and sheep; large ruminants are absent. At the time of capture of
|
||
<taxonomicName id="51DF14D2FFEF312252DAF8E7ED59F89C" authorityName="Zidek & Pokorný" authorityYear="2018" box="[224,330,1840,1864]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="lizleri">
|
||
<emphasis id="A4ABB343FFEF312252DAF8E7ED59F89C" box="[224,330,1840,1864]" italics="true" pageId="5" pageNumber="4">S. lizleri</emphasis>
|
||
</taxonomicName>
|
||
the temperature fluctuated between close to freezing at night and 30–35° C at mid-day, and the area experienced strong winds. The beetles were taken at human dung between 8 and 11 a.m. and again in the late afternoon and at dusk. It thus is a diurnal–crepuscular species that avoids daily thermal maxima more than minima, because in the early morning beetles were observed flying although the temperature was still only about 5° C.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFEC312152D2FECDEF54FE27" blockId="6.[192,1440,217,499]" pageId="6" pageNumber="5">
|
||
The localities of the
|
||
<typeStatus id="4964D1F3FFEC312153E9FECDEE14FEE6" box="[467,519,282,306]" pageId="6" pageNumber="5">type</typeStatus>
|
||
series are in
|
||
<collectingRegion id="541BA1B3FFEC3121509BFECDEFD0FEE6" box="[673,963,282,306]" country="South Africa" name="Northern Cape" pageId="6" pageNumber="5">Northern Cape Province</collectingRegion>
|
||
close to the southwestern
|
||
<collectingCountry id="EEC82FC1FFEC312156C3FECDE922FEE6" box="[1273,1329,282,306]" name="South Africa" pageId="6" pageNumber="5">RSA</collectingCountry>
|
||
/
|
||
<collectingCountry id="EEC82FC1FFEC31215700FECDE9B3FEE6" box="[1338,1440,282,306]" name="Namibia" pageId="6" pageNumber="5">Namibia</collectingCountry>
|
||
border and just across the mouth of the Orange River in southernmost
|
||
<collectingCountry id="EEC82FC1FFEC31215611FEEDE886FE86" box="[1067,1173,314,338]" name="Namibia" pageId="6" pageNumber="5">Namibia</collectingCountry>
|
||
(Oranjemund, TMSA specimens), whereas localities of
|
||
<taxonomicName id="51DF14D2FFEC3121505AFE8DEF01FEA6" box="[608,786,346,370]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="rubripennis">
|
||
<emphasis id="A4ABB343FFEC3121505AFE8DEE61FEA5" box="[608,626,346,369]" italics="true" pageId="6" pageNumber="5">S</emphasis>
|
||
.
|
||
<emphasis id="A4ABB343FFEC312150B9FE8DEF01FEA6" box="[643,786,346,370]" italics="true" pageId="6" pageNumber="5">rubripennis</emphasis>
|
||
</taxonomicName>
|
||
are more to the north in the Namib desert—Kuisip (HTm), Namtib dunes (TMSA specimens) and Sessriem (BMNH and OXUM specimens). The two species thus are close neighbors and the question arises whether north of the Orange River they could not be sympatric, separated by diel activity or chemical stimuli. We have found no evidence of hybridization in the morphology of about
|
||
<specimenCount id="80D9A4D8FFEC3121502AFE0CEED2FE27" box="[528,705,475,499]" pageId="6" pageNumber="5" type="generic">200 specimens</specimenCount>
|
||
examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |