198 lines
19 KiB
XML
198 lines
19 KiB
XML
<document id="CD3E00D3C8EA7CF8331B432EB20B2887" ID-CLB-Dataset="8002" ID-DOI="10.11646/zootaxa.4881.1.1" ID-GBIF-Dataset="d7c23423-af16-4c2d-bd68-3526fb8bb654" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425611" ID-ZooBank="DEEA669F-3A5F-4096-B005-B9CA3506F59E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610042036924" checkinUser="plazi" docAuthor="Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa & Gower, David J." docDate="2020" docId="144D8797FFBF752AFF0BFAC5FA04E08F" docLanguage="en" docName="zootaxa.4881.1.1.pdf" docOrigin="Zootaxa 4881 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhinophis" docType="key" docVersion="8" lastPageNumber="14" masterDocId="E874FFEFFFB27527FF9CFFBDFF98E776" masterDocTitle="A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="14" updateTime="1698923977174" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="86AA1DF13BEF0A647BF7F435E924AB18">
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<mods:title id="E9C04AFCB9D92B01FF7031989CC0F3C5">A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae)</mods:title>
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<mods:name id="97E9DCCDC59333EF412514B328CAB6FC" type="personal">
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<mods:roleTerm id="B545694EF6BE1A1FB0BF5544020EBCB1">Author</mods:roleTerm>
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<mods:namePart id="2C2FB9230A5E71A2E03B5C862AB10FC6">Sampaio, Filipa L.</mods:namePart>
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<mods:affiliation id="3EB2996F34E1549765D7C183EB1AD345">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK & Department of Genetics, Evolution and Environment, University College London, London WC 1 E 6 BT, UK</mods:affiliation>
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<mods:roleTerm id="16C448A3F885FFAB3F4594394B99F8EA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="07C389DE7CCDDBFFC0BB17C9F8C74886">Narayanan, Surya</mods:namePart>
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<mods:affiliation id="CE8A48034C1BC40F6B2BE98555195005">Wayanad Wild, Lakkidi, Vythiri, Wayanad, Kerala, India</mods:affiliation>
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<mods:name id="9675CDC0289CDD6A65A96EC98037069D" type="personal">
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<mods:roleTerm id="5798096457D6DF17F2497BC06C2B6E9B">Author</mods:roleTerm>
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<mods:namePart id="5FEEBFCBD02764E1DF4E353220FC85F6">Cyriac, Vivek Philip</mods:namePart>
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<mods:nameIdentifier id="9D16734CD83E38EE4078731EF0A7E982" type="ORCID">0000-0002-0465-0452</mods:nameIdentifier>
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<mods:affiliation id="098330FDAA2252A8FC265C471F800725">IISER-TVM Centre for Research and Education in Ecology and Evolution (ICREEE) and School of Biology, Indian Institute of Science Education and Research Thiruvananthapuram, Maruthamala P. O. Vithura, Thiruvananthapuram, Kerala 695551, India vivek. cyriac @ gmail. com; https: // orcid. org / 0000 - 0002 - 0465 - 0452</mods:affiliation>
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<mods:nameIdentifier id="C318D7FEEFDA2D7884F60588ACE6CA24" type="email">vivek.cyriac@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="5191A4AD9470A5FEF683CE8A28186D3C">Author</mods:roleTerm>
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<mods:namePart id="D0DAE2DC096FA2B9391D0FC5B6D4E0FA">Venu, Govindappa</mods:namePart>
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<mods:nameIdentifier id="EA8A569C5DEF99B85A2C34E7E9A3040A" type="ORCID">0000-0003-4873-7418</mods:nameIdentifier>
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<mods:affiliation id="231303D8AE3DA85CB4B9FD0A6B85C236">Centre for Applied Genetics, Department of Zoology, Bangalore University, Bengaluru, Karnataka, 560056, India venugcaecilian @ gmail. com; https: // orcid. org / 0000 - 0003 - 4873 - 7418</mods:affiliation>
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<mods:nameIdentifier id="22A5BCE9034850A886813326E74CA3DD" type="email">venugcaecilian@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4B33DD75F02F40B59BE3CB81F0A90729">Gower, David J.</mods:namePart>
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<mods:affiliation id="A8BF563250658B0D073A039F47D6BFBB">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
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<mods:typeOfResource id="9B9B310D0413C7927559B400C7985000">text</mods:typeOfResource>
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<mods:title id="FA28459C78C0BB1AB09B90D570AADF73">Zootaxa</mods:title>
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<mods:part id="880E148BBB829E126C22C61EC4B46F17">
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<mods:date id="788305C7F9AAB1D442CC1223B16BEB3E">2020</mods:date>
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<mods:number id="F9F46F35E24F25A15D2051F6D72279F8">2020-11-18</mods:number>
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<mods:number id="0665A262E01D2746CDC5FB9328D9321A">4881</mods:number>
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<mods:number id="A18E3CCA209278F22456D4C2B3FE90EC">1</mods:number>
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<mods:start id="A0FC3257EDCADF623E1849C936B02319">1</mods:start>
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<mods:classification id="D4CE156BBEDA3C449C2B2E4B671556D2">journal article</mods:classification>
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<mods:identifier id="5C3378D6B6A0E1B50BF33921F3A3B774" type="CLB-Dataset">8002</mods:identifier>
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<mods:identifier id="1825BE9559014AE3E8AFB40B04C6506D" type="DOI">10.11646/zootaxa.4881.1.1</mods:identifier>
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<mods:identifier id="99515D782A41C0ADF1B118D2F4068704" type="GBIF-Dataset">d7c23423-af16-4c2d-bd68-3526fb8bb654</mods:identifier>
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<mods:identifier id="9E4784E6DEAA180C4FDBD89461556A88" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="713CBD621ED9F1707E16143295C793BA" type="Zenodo-Dep">4425611</mods:identifier>
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</mods:mods>
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<treatment id="144D8797FFBF752AFF0BFAC5FA04E08F" ID-DOI="http://doi.org/10.5281/zenodo.4564927" ID-GBIF-Taxon="179917489" ID-Zenodo-Dep="4564927" LSID="urn:lsid:plazi:treatment:144D8797FFBF752AFF0BFAC5FA04E08F" httpUri="http://treatment.plazi.org/id/144D8797FFBF752AFF0BFAC5FA04E08F" lastPageNumber="14" pageId="13" pageNumber="14">
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<subSubSection id="D4FE650AFFBF752AFF0BFAC5FDEDE2E5" box="[151,629,1400,1427]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="9C5B3681FFBF752AFF0BFAC5FDEDE2E5" blockId="13.[151,629,1400,1427]" box="[151,629,1400,1427]" pageId="13" pageNumber="14">
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<heading id="C71381EDFFBF752AFF0BFAC5FDEDE2E5" bold="true" box="[151,629,1400,1427]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="AE90EA93FFBF752AFF0BFAC5FDEDE2E5" bold="true" box="[151,629,1400,1427]" pageId="13" pageNumber="14">
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Key to the Indian species of
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<taxonomicName id="5BE44D02FFBF752AFE6BFAC5FDEDE2E5" box="[503,629,1400,1427]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
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<emphasis id="AE90EA93FFBF752AFE6BFAC5FDEDE2E5" bold="true" box="[503,629,1400,1427]" italics="true" pageId="13" pageNumber="14">Rhinophis</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4FE650AFFBF752AFF0BFA06FCA1E137" pageId="13" pageNumber="14" type="discussion">
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<paragraph id="9C5B3681FFBF752AFF0BFA06FCA1E137" blockId="13.[151,1437,1467,1601]" pageId="13" pageNumber="14">
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This is an updated version of the key presented by
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<bibRefCitation id="F8754B70FFBF752AFD59FA06FC38E2A3" author="Cyriac, V. P. & Narayanan, S. & Sampaio, F. L. & Umesh, P. & Gower, D. J." box="[709,928,1467,1493]" pageId="13" pageNumber="14" pagination="329 - 342" refId="ref10962" refString="Cyriac, V. P., Narayanan, S., Sampaio, F. L., Umesh, P. & Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from the Wayanad region of peninsular India. Zootaxa, 4778 (2), 329 - 342. https: // doi. org / 10.11646 / zootaxa. 4778.2.5" type="journal article" year="2020">
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Cyriac
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<emphasis id="AE90EA93FFBF752AFC8AFA01FCD5E2A3" box="[790,845,1467,1493]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
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(2020)
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</bibRefCitation>
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, revised with the addition of
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<taxonomicName id="5BE44D02FFBF752AFB7CFA01FA05E2A3" authority="Sampaio & Narayanan & Cyriac & Venu & Gower, 2020" authorityName="Sampaio & Narayanan & Cyriac & Venu & Gower" authorityYear="2020" box="[1248,1437,1467,1493]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="karinthandani" status="sp. nov.">
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<emphasis id="AE90EA93FFBF752AFB7CFA01FA05E2A3" box="[1248,1437,1467,1493]" italics="true" pageId="13" pageNumber="14">R. karinthandani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B5A357E8FFBF752AFF0BFA62FF69E28F" box="[151,241,1503,1529]" pageId="13" pageNumber="14" rank="species">
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<emphasis id="AE90EA93FFBF752AFF0BFA62FF73E28F" bold="true" box="[151,235,1503,1529]" pageId="13" pageNumber="14">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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We have removed the relative rostral length character used in Cyriac
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<emphasis id="AE90EA93FFBF752AFC69FA5DFBBDE28F" box="[1013,1061,1503,1529]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
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.’s (2020) key because we are less convinced of its diagnostic value and because it can be difficult to measure. All six species are endemic to
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<collectingCountry id="E4F37611FFBF752AFAA8F9BEFAF6E16B" box="[1332,1390,1539,1565]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
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and are the only members of this genus known to occur in
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<collectingCountry id="E4F37611FFBF752AFD64F99AFCADE137" box="[760,821,1575,1601]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D4FE650AFFBF752AFF0BF9DAFA04E08F" pageId="13" pageNumber="14" type="key">
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<keyStep id="27102E24FFBF752AFF0BF9DAFA04E1EF" pageId="13" pageNumber="14">
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<paragraph id="9C5B3681FFBF752AFF0BF9DAFA04E10A" blockId="13.[151,1437,1638,2041]" box="[151,1436,1638,1660]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF9DAFA04E10A" box="[151,1436,1638,1660]" pageId="13" pageNumber="14">1 Number of dorsal scale rows just behind midbody 17......................................................... 2</keyLead>
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</paragraph>
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<paragraph id="9C5B3681FFBF752AFF0BF939FA04E1EF" blockId="13.[151,1437,1638,2041]" box="[151,1436,1668,1690]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF939FA04E1EF" box="[151,1436,1668,1690]" pageId="13" pageNumber="14">- Number of dorsal scale rows just behind midbody 15......................................................... 3</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="27102E24FFBF752AFF0BF91CFA05E1A2" pageId="13" pageNumber="14">
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<paragraph id="9C5B3681FFBF752AFF0BF91CFA04E1C1" blockId="13.[151,1437,1638,2041]" box="[151,1436,1697,1719]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF91CFA04E1C1" box="[151,1436,1697,1719]" pageId="13" pageNumber="14">
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2 Ventral shields fewer than 160...............................................................
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<taxonomicName id="5BE44D02FFBF752AFB60F91FFA04E1C1" box="[1276,1436,1698,1719]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="travancoricus">
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<emphasis id="AE90EA93FFBF752AFB60F91FFA04E1C1" box="[1276,1436,1698,1719]" italics="true" pageId="13" pageNumber="14">R. travancoricus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="9C5B3681FFBF752AFF0BF902FA05E1A2" blockId="13.[151,1437,1638,2041]" box="[151,1437,1726,1748]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF902FA05E1A2" box="[151,1437,1726,1748]" pageId="13" pageNumber="14">
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- Ventral shields more than 180....................................................................
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<taxonomicName id="5BE44D02FFBF752AFAA3F902FA05E1A2" authorityName="Aengals & Ganesh" authorityYear="2013" box="[1343,1437,1727,1748]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="goweri">
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<emphasis id="AE90EA93FFBF752AFAA3F902FA05E1A2" box="[1343,1437,1727,1748]" italics="true" pageId="13" pageNumber="14">R. goweri</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="27102E24FFBF752AFF0BF961FA04E079" pageId="13" pageNumber="14">
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<paragraph id="9C5B3681FFBF752AFF0BF961FA04E187" blockId="13.[151,1437,1638,2041]" box="[151,1436,1756,1778]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF961FA04E187" box="[151,1436,1756,1778]" pageId="13" pageNumber="14">3 Ventral surface with dark blotches in irregular, staggered, transverse half-bands.................................... 4</keyLead>
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</paragraph>
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<paragraph id="9C5B3681FFBF752AFF0BF944FA04E079" blockId="13.[151,1437,1638,2041]" box="[151,1436,1785,1807]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF944FA04E079" box="[151,1436,1785,1807]" pageId="13" pageNumber="14">- Ventral surface with irregular dark speckles or spots.......................................................... 5</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="27102E24FFBF752AFF0BF8ABFA04E0F5" pageId="13" pageNumber="14">
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<paragraph id="9C5B3681FFBF752AFF0BF8ABFA04E03F" blockId="13.[151,1437,1638,2041]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF8ABFA04E03F" pageId="13" pageNumber="14">
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4 Only known specimen a female with 195 ventral and 5,5 subcaudal shields (from Cardamom Hills of Western Ghats, South of Palghat Gap).............................................................................
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<taxonomicName id="5BE44D02FFBF752AFB66F889FA04E03F" authorityName="Boulenger" authorityYear="1896" box="[1274,1436,1844,1865]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="fergusonianus">
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<emphasis id="AE90EA93FFBF752AFB66F889FA04E03F" box="[1274,1436,1844,1865]" italics="true" pageId="13" pageNumber="14">R. fergusonianus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="9C5B3681FFBF752AFF0BF8ECFA04E0F5" blockId="13.[151,1437,1638,2041]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF8ECFA04E0F5" pageId="13" pageNumber="14">
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- Ventral shields more than 215; subcaudals 6 or more pairs (from Wayanad region of Western Ghats, North of Palghat Gap).........................................................................................
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<taxonomicName id="5BE44D02FFBF752AFB62F8D2FA04E0F5" box="[1278,1436,1902,1924]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="melanoleucus">
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<emphasis id="AE90EA93FFBF752AFB62F8D2FA04E0F5" box="[1278,1436,1902,1924]" italics="true" pageId="13" pageNumber="14">R. melanoleucus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="27102E24FFBF752AFF0BF831FA04E08F" pageId="13" pageNumber="14">
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<paragraph id="9C5B3681FFBF752AFF0BF831FA04E0C9" blockId="13.[151,1437,1638,2041]" pageId="13" pageNumber="14">
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<keyLead id="271595B4FFBF752AFF0BF831FA04E0C9" pageId="13" pageNumber="14">
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5 Subcaudals 5 (more typically 6)–11 (5–
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<specimenCount id="8AE2FD08FFBF752AFDC5F831FD21E0D4" box="[601,697,1932,1954]" pageId="13" pageNumber="14" type="female">7 females</specimenCount>
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; 8–
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<specimenCount id="8AE2FD08FFBF752AFD46F831FCA9E0D4" box="[730,817,1932,1954]" pageId="13" pageNumber="14" type="male">11 males</specimenCount>
|
||
); venter background colour bright red in life (paler, more orangeyellow in smaller specimens)..................................................................
|
||
<taxonomicName id="5BE44D02FFBF752AFA8AF817FA04E0C9" baseAuthorityName="Wall" baseAuthorityYear="1919" box="[1302,1436,1962,1983]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="sanguineus">
|
||
<emphasis id="AE90EA93FFBF752AFA8AF817FA04E0C9" box="[1302,1436,1962,1983]" italics="true" pageId="13" pageNumber="14">R. sanguineus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="9C5B3681FFBF752AFF0BF87AFA04E08F" blockId="13.[151,1437,1638,2041]" pageId="13" pageNumber="14">
|
||
<keyLead id="271595B4FFBF752AFF0BF87AFA04E08F" pageId="13" pageNumber="14">
|
||
- Subcaudals 4–8 (4–
|
||
<specimenCount id="8AE2FD08FFBF752AFE3CF87BFE67E0AA" box="[416,511,1990,2012]" pageId="13" pageNumber="14" type="female">5 females</specimenCount>
|
||
; 6–
|
||
<specimenCount id="8AE2FD08FFBF752AFDBEF87BFDF6E0AA" box="[546,622,1990,2012]" pageId="13" pageNumber="14" type="male">8 males</specimenCount>
|
||
); venter background colour whitish (to pale pink anteriorly) in life..............................................................................................
|
||
<taxonomicName id="5BE44D02FFBF752AFB3AF859FAD0E08F" authority="Sampaio & Narayanan & Cyriac & Venu & Gower, 2020" authorityName="Sampaio & Narayanan & Cyriac & Venu & Gower" authorityYear="2020" box="[1190,1352,2020,2041]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="karinthandani" status="sp. nov.">
|
||
<emphasis id="AE90EA93FFBF752AFB3AF859FAD0E08F" box="[1190,1352,2020,2041]" italics="true" pageId="13" pageNumber="14">R. karinthandani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B5A357E8FFBF752AFAD2F859FA04E08F" box="[1358,1436,2020,2041]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="AE90EA93FFBF752AFAD2F859FA0EE08F" bold="true" box="[1358,1430,2020,2041]" pageId="13" pageNumber="14">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |