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<mods:title id="73C9C6E7C2F8F1896AA9821E3C2C60AD">The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae)</mods:title>
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<mods:namePart id="00098759D23132689B445C760D3C6B35">Giroti, André Marsola</mods:namePart>
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<mods:namePart id="6F0BB57C8CEFBE96D14E6174E930F86C">Brescovit, Antonio Domingos</mods:namePart>
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<treatment id="039B87D1FFCFB131169BFC4FC402F94B" ID-DOI="http://doi.org/10.5281/zenodo.5987874" ID-GBIF-Taxon="142616433" ID-Zenodo-Dep="5987874" LSID="urn:lsid:plazi:treatment:039B87D1FFCFB131169BFC4FC402F94B" httpUri="http://treatment.plazi.org/id/039B87D1FFCFB131169BFC4FC402F94B" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection id="C328654CFFCFB131169BFC4FC545FC03" box="[151,540,908,934]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8B8D36C7FFCFB131169BFC4FC545FC03" blockId="43.[151,540,908,967]" box="[151,540,908,934]" pageId="43" pageNumber="44">
<heading id="D0C581ABFFCFB131169BFC4FC545FC03" bold="true" box="[151,540,908,934]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<taxonomicName id="4C324D44FFCFB131169BFC4FC545FC03" ID-CoL="GLNW" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[151,540,908,934]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tovarensis">
<emphasis id="B946EAD5FFCFB131169BFC4FC545FC03" bold="true" box="[151,540,908,934]" pageId="43" pageNumber="44">
<emphasis id="B946EAD5FFCFB131169BFC4FC622FC03" bold="true" box="[151,379,908,934]" italics="true" pageId="43" pageNumber="44">Ariadna tovarensis</emphasis>
<bibRefCitation id="EFA34B36FFCFB131178FFC4FC545FC03" author="Simon, E." bookContentInfo="26 pp." box="[387,540,908,934]" journalOrPublisher="Librarie Encyclopedique de Roret, Paris" pageId="43" pageNumber="44" pagination="257 - 488" refId="ref90788" refString="Simon, E. (1893 a) Histoire naturelle das araignees. Vol. 1. Librarie Encyclopedique de Roret, Paris, 26 pp. [pp. 257 - 488]" title="Histoire naturelle das araignees. Vol. 1" type="book chapter" year="1893">Simon, 1893</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C328654CFFCFB131169BFC6DC660FC62" box="[151,313,942,967]" pageId="43" pageNumber="44" type="description">
<paragraph id="8B8D36C7FFCFB131169BFC6DC660FC62" blockId="43.[151,540,908,967]" box="[151,313,942,967]" pageId="43" pageNumber="44">
<figureCitation id="13092A42FFCFB131169BFC6DC649FC62" box="[151,272,942,967]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Figures 26</figureCitation>
,
<figureCitation id="13092A42FFCFB1311710FC6DC660FC62" box="[284,313,942,967]" captionStart="FIGURE68" captionStartId="105.[151,250,1970,1992]" captionTargetBox="[151,1436,1244,1947]" captionTargetId="figure@105.[151,1436,1244,1948]" captionTargetPageId="105" captionText="FIGURE68. Locality of the specimens of A.caerulea, A.tovarensis, A. arthuri, A.multispinosa, A.isthmica and A. reginae n. sp. examined in the present study." httpUri="https://zenodo.org/record/1207783/files/figure.png" pageId="43" pageNumber="44">68</figureCitation>
</paragraph>
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<subSubSection id="C328654CFFCFB131169BFC35C2D1FBCD" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="8B8D36C7FFCFB131169BFC35C6A8FBEF" blockId="43.[151,1436,1014,1128]" pageId="43" pageNumber="44">
<taxonomicName id="4C324D44FFCFB131169BFC35C6BCFBA9" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[151,485,1014,1037]" class="Arachnida" family="Segestriidae" genus="Ariadne" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tovarensis">
<emphasis id="B946EAD5FFCFB131169BFC35C601FBA8" box="[151,344,1014,1037]" italics="true" pageId="43" pageNumber="44">Ariadne tovarensis</emphasis>
<bibRefCitation id="EFA34B36FFCFB131176DFC35C6BCFBA9" author="Simon, E." bookContentInfo="26 pp." box="[353,485,1014,1036]" journalOrPublisher="Librarie Encyclopedique de Roret, Paris" pageId="43" pageNumber="44" pagination="257 - 488" refId="ref90788" refString="Simon, E. (1893 a) Histoire naturelle das araignees. Vol. 1. Librarie Encyclopedique de Roret, Paris, 26 pp. [pp. 257 - 488]" title="Histoire naturelle das araignees. Vol. 1" type="book chapter" year="1893">Simon, 1893</bibRefCitation>
</taxonomicName>
b: 448 (♀ lectotype and 2♀ lectoparatypes from Colônia ToVar [10°24'30&quot;N; 67°17'22&quot;W], Aragua, Venezuela, Simon leg., deposited in MNHNP-AR 14388 and MNHNP-AR 14388; 14389, respectively, designated by
<bibRefCitation id="EFA34B36FFCFB13116EAFBF6C62CFBEF" author="Beatty, J. A." box="[230,373,1076,1098]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="43" pageNumber="44" pagination="433 - 518" part="139" refId="ref87290" refString="Beatty, J. A. (1970) The spider genus Ariadna in the Americas (Araneae, Dysderidae). Bulletin of the Museum of Comparative Zoology, 139 (8), 433 - 518." title="The spider genus Ariadna in the Americas (Araneae, Dysderidae)" type="journal article" year="1970">Beatty (1970)</bibRefCitation>
, examined).
</paragraph>
<paragraph id="8B8D36C7FFCFB131169BFB91C2D1FBCD" blockId="43.[151,1436,1014,1128]" box="[151,1416,1106,1128]" pageId="43" pageNumber="44">
<emphasis id="B946EAD5FFCFB131169BFB91C634FBCD" box="[151,365,1106,1128]" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="4C324D44FFCFB131169BFB91C7B3FBCD" ID-CoL="32TD" authorityName="Audouin" authorityYear="1826" box="[151,234,1106,1128]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">Ariadna</taxonomicName>
tovarensis—
</emphasis>
<bibRefCitation id="EFA34B36FFCFB1311762FB91C52AFBCD" author="Petrunkevitch, A." box="[366,627,1106,1128]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="43" pageNumber="44" pagination="1 - 791" part="29" publicationUrl="https://doi.org/10.5962/bhl.title.23819" refId="ref89889" refString="Petrunkevitch, A. (1911) A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History, 29, 1 - 791. https: // doi. org / 10.5962 / bhl. title. 23819" title="A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc" type="journal article" year="1911">Petrunkevitch (1911: 131)</bibRefCitation>
.—
<bibRefCitation id="EFA34B36FFCFB131149DFB91C405FBCD" author="Bonnet, P." bookContentInfo="919 pp." box="[657,860,1106,1128]" journalOrPublisher="Toulouse" pageId="43" pageNumber="44" refId="ref87444" refString="Bonnet, P. (1955) Bibliographia araneorum. 2 (1). Les Artisans de L'Imprimerie Douladoure, Toulouse, 919 pp." title="Bibliographia araneorum. 2 (1). Les Artisans de L'Imprimerie Douladoure" type="book" year="1955">Bonnet (1955: 738)</bibRefCitation>
.—
<bibRefCitation id="EFA34B36FFCFB1311576FB91C363FBCD" author="Beatty, J. A." box="[890,1082,1106,1128]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="43" pageNumber="44" pagination="433 - 518" part="139" refId="ref87290" refString="Beatty, J. A. (1970) The spider genus Ariadna in the Americas (Araneae, Dysderidae). Bulletin of the Museum of Comparative Zoology, 139 (8), 433 - 518." title="The spider genus Ariadna in the Americas (Araneae, Dysderidae)" type="journal article" year="1970">Beatty (1970: 472)</bibRefCitation>
.—
<bibRefCitation id="EFA34B36FFCFB131125BFB91C2DDFBCD" author="World Spider Catalog" box="[1111,1412,1106,1128]" journalOrPublisher="Avaliable from: http: // wsc. nmbe. ch / (accessed on" pageId="43" pageNumber="44" pagination="2018" part="16" refId="ref91516" refString="World Spider Catalog (2018) World Spider Catalog. Natural History Museum Bern, version 19.0. Avaliable from: http: // wsc. nmbe. ch / (accessed on 16 January 2018)" title="World Spider Catalog. Natural History Museum Bern, version 19.0" type="journal article" year="2018">World Spider Catalog (2018)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328654CFFCFB131169BFB56C239FB77" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8B8D36C7FFCFB131169BFB56C239FB77" blockId="43.[151,1437,1173,1774]" pageId="43" pageNumber="44">
<emphasis id="B946EAD5FFCFB131169BFB56C648FB0B" bold="true" box="[151,273,1173,1198]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
<taxonomicName id="4C324D44FFCFB131172CFB56C6A4FB08" authorityName="Simon" authorityYear="1893" box="[288,509,1173,1198]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tovarensis">
<emphasis id="B946EAD5FFCFB131172CFB56C6A4FB08" box="[288,509,1173,1198]" italics="true" pageId="43" pageNumber="44">Ariadna tovarensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C324D44FFCFB13114A3FB55C46DFB08" authority="Giroti &amp; Brescovit, 2018" authorityName="Giroti &amp; Brescovit" authorityYear="2018" box="[687,820,1174,1197]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="caparao" status="sp. nov.">
<emphasis id="B946EAD5FFCFB13114A3FB55C46DFB08" box="[687,820,1174,1197]" italics="true" pageId="43" pageNumber="44">A. caparao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557AEFFCFB131154FFB56C4D3FB0B" box="[835,906,1173,1198]" pageId="43" pageNumber="44" rank="species">
<emphasis id="B946EAD5FFCFB131154FFB56C4D3FB0B" bold="true" box="[835,906,1173,1198]" pageId="43" pageNumber="44">n. sp.</emphasis>
</taxonomicNameLabel>
due to the elongated lobes of the anterior receptaculum, but differs by presenting the ventral lobe almost straight and shorter than the DL (
<figureCitation id="13092A42FFCFB13112DCFB7AC20DFB77" box="[1232,1364,1209,1234]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26EF</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C328654CFFCFB13116CBFB1EC2CEF96F" pageId="43" pageNumber="44" type="description">
<paragraph id="8B8D36C7FFCFB13116CBFB1EC579FB53" blockId="43.[151,1437,1173,1774]" box="[199,544,1245,1270]" pageId="43" pageNumber="44">
<emphasis id="B946EAD5FFCFB13116CBFB1EC6FAFB53" bold="true" box="[199,419,1245,1270]" pageId="43" pageNumber="44">Description. Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B8D36C7FFCFB13116CBFAC2C34EF903" blockId="43.[151,1437,1173,1774]" pageId="43" pageNumber="44">
<emphasis id="B946EAD5FFCFB13116CBFAC2C6E0FABF" bold="true" box="[199,441,1281,1306]" pageId="43" pageNumber="44">
Female (
<typeStatus id="54898865FFCFB131173DFAC2C6F5FABF" box="[305,428,1281,1306]" pageId="43" pageNumber="44" type="lectotype">Lectotype</typeStatus>
).
</emphasis>
Carapace and chelicerae orange (
<figureCitation id="13092A42FFCFB131154BFAC2C4EDFABF" box="[839,948,1281,1306]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26A</figureCitation>
). Endites yellow, labium orange. Sternum yellow, with darker margins (
<figureCitation id="13092A42FFCFB13117E5FAE6C50AFA9B" box="[489,595,1317,1342]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26B</figureCitation>
). Palps yellow, darkening gradually in distal direction. Legs yellow, with pairs III darker (
<figureCitation id="13092A42FFCFB1311750FA8AC6BFFAC7" box="[348,486,1353,1378]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26CD</figureCitation>
). Abdomen light grey (
<figureCitation id="13092A42FFCFB13114E6FA8AC42DFAC7" box="[746,884,1353,1378]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26AB</figureCitation>
). Total length 7.4; carapace 3.04 long, 2.16 wide. Legs: robust and coated with setae; legs III with ventro-prolateral and ventro-retrolateral tibial macrosetae of similar size, and metatarsal macrosetae of different sizes (
<figureCitation id="13092A42FFCFB131158CFA52C34FFA0F" box="[896,1046,1425,1450]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26CD</figureCitation>
). Leg formula: I-II-IV-III. Leg measurements: I, femur 2.24, patella 0.96, tibia 1.6, metatarsus 1.52, tarsus 0.6, total 6.92; II, 2.0, 0.96, 1.6, 1.4, 0.52, 6.48; III, 1.6, 0.76, 1.0, 0.92, 0.44, 4.72; IV, 1.92, 0.96, 1.44, 1.12, 0.52, 5.96. Macrosetae: I, femur D0-0-0-0- 0-2-0, P0-0-0-0-2-1-0; tibia P0-1-0-1-1/0-1-0, vp1-0-1-0-1-0-1, vr1/2-0-1-0-1-0-1, R1-0-1-0-1-0-1; metatarsus vp1-0/1-1-1-1-1-1, vr1-0-1-1-0-1-1 (
<figureCitation id="13092A42FFCFB131143FF9E2C5E4F99F" box="[563,701,1569,1594]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26CD</figureCitation>
); II, femur D0-0-0-0-0-2-0, P0-0-0-0-0/1-1-0; tibia P0-0/1-0-1-1/ 0-1-0, vp0-0-0-1/0-0-1-1, vr1-0-1-0-1-0-1, R0-0-1-0-1-0-1/0; metatarsus vp1-0-1-1-0-1-1, vr1-0-1-1-0-1-1; IV, metatarsus vr metatarsal comb with 4 macrosetae. Internal genitalia: anterior receptaculum with glandular plate located in the median region, and expanded in its proximal region (
<figureCitation id="13092A42FFCFB131158BF94EC352F903" box="[903,1035,1677,1702]" captionStart="FIGURE 26" captionStartId="44.[151,250,1798,1820]" captionTargetBox="[151,1436,379,1760]" captionTargetId="figure@44.[151,1436,379,1776]" captionTargetPageId="44" captionText="FIGURE 26. Ariadna tovarensis Simon, 1893. Female (Lectoparatype) from Colônia Tovar (MNHNP-AR 14389) (AF): habitus dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207697/files/figure.png" pageId="43" pageNumber="44">Fig. 26EF</figureCitation>
).
</paragraph>
<paragraph id="8B8D36C7FFCFB13116CBF972C2CEF96F" blockId="43.[151,1437,1173,1774]" box="[199,1431,1713,1738]" pageId="43" pageNumber="44">
<emphasis id="B946EAD5FFCFB13116CBF972C666F96F" bold="true" box="[199,319,1713,1738]" pageId="43" pageNumber="44">Variation.</emphasis>
Female (n=3): Total length 6.047.52; carapace 2.63.04 long, 1.762.16 wide; femur I 1.962.24.
</paragraph>
</subSubSection>
<subSubSection id="C328654CFFCFB13116CBF916C402F94B" box="[199,859,1749,1774]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8B8D36C7FFCFB13116CBF916C402F94B" blockId="43.[151,1437,1173,1774]" box="[199,859,1749,1774]" pageId="43" pageNumber="44">
<emphasis id="B946EAD5FFCFB13116CBF916C639F94B" bold="true" box="[199,352,1749,1774]" pageId="43" pageNumber="44">Distribution.</emphasis>
Only known from the
<typeStatus id="54898865FFCFB131146CF915C5C9F94B" box="[608,656,1750,1774]" pageId="43" pageNumber="44">type</typeStatus>
locality (
<figureCitation id="13092A42FFCFB13114F6F916C416F94B" box="[762,847,1749,1774]" captionStart="FIGURE68" captionStartId="105.[151,250,1970,1992]" captionTargetBox="[151,1436,1244,1947]" captionTargetId="figure@105.[151,1436,1244,1948]" captionTargetPageId="105" captionText="FIGURE68. Locality of the specimens of A.caerulea, A.tovarensis, A. arthuri, A.multispinosa, A.isthmica and A. reginae n. sp. examined in the present study." httpUri="https://zenodo.org/record/1207783/files/figure.png" pageId="43" pageNumber="44">Fig. 68</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>