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<document id="1BA127F015C4052232F69B077313A774" ID-DOI="10.5852/ejt.2023.914.2377" ID-ISSN="2118-9773" ID-Zenodo-Dep="10407071" ID-ZooBank="2ECE4DA6-97CA-4917-97F1-FE506998A2D0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703002881060" checkinUser="plazi" docAuthor="Basílio, Daniel Silva, Vaz-De-Mello, Fernando Zagury, Cherman, Mariana Alejandra &amp; Almeida, Lúcia Massutti de" docDate="2023" docId="EA056F037426FFF5BBA2FDE64481FB43" docLanguage="en" docName="EJT.2023.914.1-53.pdf" docOrigin="European Journal of Taxonomy 914 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2377/10437" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Coilodes mayae Basilio" docType="treatment" docVersion="2" lastPageNumber="31" masterDocId="163C177B743AFFEBBA58FFE4400FFFA7" masterDocTitle="A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species" masterLastPageNumber="53" masterPageNumber="1" pageNumber="29" updateTime="1703003437132" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="716FD4F29AAAFA26E55A554DB0FAA4B4">A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species</mods:title>
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<mods:namePart id="06E3DDD987000025C012BE7C854DD384">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="93E86D57AC391425357C664920E85602">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
<mods:nameIdentifier id="BD729289E4C5362FC791B9447D86421C" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="0A1363AFFD4C4C5E1617689A08F44205">Author</mods:roleTerm>
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<mods:namePart id="BE0B299988E018B597D41C35B434F28F">Vaz-De-Mello, Fernando Zagury</mods:namePart>
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<mods:affiliation id="45A822D8A5190E2ADC101423420A07E9">Laboratório de Scarabaeoidologia, Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="6BD76C57E16A7FD68A94791EE0C7F8E9" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="F8B1DB1D45E655590296CBC4DF5D453F">Author</mods:roleTerm>
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<mods:namePart id="ED5434FD7890FA8620D7055A14E8F611">Cherman, Mariana Alejandra</mods:namePart>
<mods:nameIdentifier id="9625324D9D4D9C485D6C93AC068882E3" type="LSID">EAF28053-7ABC-4F2D-9C46-CA7B1AEC2C5A</mods:nameIdentifier>
<mods:affiliation id="2F1F36460A67044359FB0016537AEBDA">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
<mods:nameIdentifier id="CF3A4DE57F082F3F1BE9781A2DD0454D" type="email">marianabioar@gmail.com</mods:nameIdentifier>
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<mods:namePart id="644D6DF8D9FDCA152F106F49D2C8DCA7">Almeida, Lúcia Massutti de</mods:namePart>
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<mods:affiliation id="8A1C4A35C524856E010802BACEB6B9EB">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
<mods:nameIdentifier id="93F85BB39436EE5ECE5923F33A981044" type="email">lalmeida51@gmail.com</mods:nameIdentifier>
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<mods:title id="268CA6B83A464A65C16326BD37AB1711">European Journal of Taxonomy</mods:title>
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<mods:part id="3558088184DD6A8911182883B697E5A9">
<mods:date id="703761FFBBF40F58CE74E64B0623B5C1">2023</mods:date>
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<mods:number id="25F2702C65AE2B08E56F6CF126B35E13">2023-12-19</mods:number>
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<mods:number id="8DA4130EC3F263D7660AE086FEBA0530">914</mods:number>
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<mods:classification id="3007B2FF2730731460E484CBBA919771">journal article</mods:classification>
<mods:identifier id="02DC41223D523EF7B2118D5DB92BBA32" type="DOI">10.5852/ejt.2023.914.2377</mods:identifier>
<mods:identifier id="1F4C37AC00A5DBE389A45C02EDC9E170" type="ISSN">2118-9773</mods:identifier>
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<treatment id="EA056F037426FFF5BBA2FDE64481FB43" LSID="urn:lsid:plazi:treatment:EA056F037426FFF5BBA2FDE64481FB43" httpUri="http://treatment.plazi.org/id/EA056F037426FFF5BBA2FDE64481FB43" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="2AB68D9E7426FFF7BBA2FDE644CBFDE6" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="6213DE157426FFF7BBA2FDE64435FDBA" blockId="28.[368,1220,514,613]" box="[506,1082,514,541]" pageId="28" pageNumber="29">
<heading id="395B69797426FFF7BBA2FDE64435FDBA" box="[506,1082,514,541]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="A5ACA5967426FFF7BBA2FDE6431DFDBA" authority="Basilio" authorityName="Basilio" box="[506,786,514,541]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mayae">
<emphasis id="50D802077426FFF7BBA2FDE642B8FDBA" bold="true" box="[506,695,514,541]" italics="true" pageId="28" pageNumber="29">Coilodes mayae</emphasis>
Basílio
</taxonomicName>
&amp; Vaz-de-Mello sp. nov.
</heading>
</paragraph>
<paragraph id="6213DE157426FFF7BB28FDC244CBFDE6" blockId="28.[368,1220,514,613]" box="[368,1220,550,577]" pageId="28" pageNumber="29">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="2AB68D9E7426FFF7B88EFDAF4351FDC2" box="[726,862,587,613]" pageId="28" pageNumber="29" type="description">
<paragraph id="6213DE157426FFF7B88EFDAF4351FDC2" blockId="28.[368,1220,514,613]" box="[726,862,587,613]" pageId="28" pageNumber="29">
<figureCitation id="FA97C2907426FFF7B88EFDAF432EFDC2" box="[726,801,587,613]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="28" pageNumber="29">Figs 7</figureCitation>
,
<figureCitation id="FA97C2907426FFF7B975FDAF4351FDC2" box="[813,862,587,613]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1739]" captionTargetId="figure-12@47.[272,1315,265,1739]" captionTargetPageId="47" captionText="Fig. 14. Distribution map of species of Coilodes Westwood, 1846. A. Coilodes bezerrai Basílio &amp; Vazde-Mello sp. nov.; Coilodes castaneus Westwood, 1846; and Coilodes edeiltae Basílio &amp; Vaz-de-Mello sp. nov. B. Coilodes fumipennis Arrow, 1909; Coilodes humeralis (Mannerheim, 1829); and Coilodes lunae Basílio &amp; Vaz-de-Mello sp. nov. C. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov.;Coilodes niger (Mannerheim, 1829) stat. rev.; Coilodes ovalis Robinson, 1948; and Coilodes parvulus Westwood 1846. D. Coilodes punctipennis Arrow, 1909; Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov.; and Coilodes skelleyi Basílio &amp; Vaz-de-Mello sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10407111" httpUri="https://zenodo.org/record/10407111/files/figure.png" pageId="28" pageNumber="29">14C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2AB68D9E7426FFF7BAE5FD69417DFCC0" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="6213DE157426FFF7BAE5FD694139FD00" blockId="28.[189,310,653,679]" box="[189,310,653,679]" pageId="28" pageNumber="29">
<heading id="395B69797426FFF7BAE5FD694139FD00" bold="true" box="[189,310,653,679]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="50D802077426FFF7BAE5FD694139FD00" bold="true" box="[189,310,653,679]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157426FFF7BAE5FD58417DFCC0" blockId="28.[189,1399,700,871]" pageId="28" pageNumber="29">
Similar to
<taxonomicName id="A5ACA5967426FFF7BB67FD584228FD71" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[319,551,700,726]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="50D802077426FFF7BB67FD584228FD71" box="[319,551,700,726]" italics="true" pageId="28" pageNumber="29">Coilodes humeralis</emphasis>
</taxonomicName>
and to
<taxonomicName id="A5ACA5967426FFF7B8DCFD5942E5FD71" authority="Basílio &amp; Vaz-De-Mello &amp; Cherman &amp; Almeida, 2023" authorityName="Basílio &amp; Vaz-De-Mello &amp; Cherman &amp; Almeida" authorityYear="2023" box="[644,746,700,727]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="lunae" status="sp. nov.">
<emphasis id="50D802077426FFF7B8DCFD5942E5FD71" box="[644,746,700,727]" italics="true" pageId="28" pageNumber="29">C. lunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BEBBF7C7426FFF7B8A9FD594341FD70" box="[753,846,701,727]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
Distinguished from both by the presence of a perpendicular laminar structure on the internal region of the tegmen. It is distinguished from
<taxonomicName id="A5ACA5967426FFF7BEB8FD05457BFD5C" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1248,1396,737,763]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="50D802077426FFF7BEB8FD05457BFD5C" box="[1248,1396,737,763]" italics="true" pageId="28" pageNumber="29">C. humeralis</emphasis>
</taxonomicName>
by the absence of a dorsal lobe on the tegmen, and from
<taxonomicName id="A5ACA5967426FFF7B909FCE143BBFCB8" authority="Basílio &amp; Vaz-De-Mello &amp; Cherman &amp; Almeida, 2023" authorityName="Basílio &amp; Vaz-De-Mello &amp; Cherman &amp; Almeida" authorityYear="2023" box="[849,948,773,799]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="lunae" status="sp. nov.">
<emphasis id="50D802077426FFF7B909FCE143BBFCB8" box="[849,948,773,799]" italics="true" pageId="28" pageNumber="29">C. lunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BEBBF7C7426FFF7B9E4FCE14419FCB8" box="[956,1046,773,799]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
by the sharp constriction next to the base of the right paramere and by the absence of a transparent region on the superior half of the right paramere.
</paragraph>
</subSubSection>
<subSubSection id="2AB68D9E7426FFF7BAE5FC6B42F8FC5A" pageId="28" pageNumber="29" type="etymology">
<paragraph id="6213DE157426FFF7BAE5FC6B414BFC0E" blockId="28.[189,324,911,937]" box="[189,324,911,937]" pageId="28" pageNumber="29">
<heading id="395B69797426FFF7BAE5FC6B414BFC0E" bold="true" box="[189,324,911,937]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="50D802077426FFF7BAE5FC6B414BFC0E" bold="true" box="[189,324,911,937]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157426FFF7BAE5FC5B42F8FC5A" blockId="28.[189,1398,958,1021]" pageId="28" pageNumber="29">
Noun in the genitive singular.
<taxonomicName id="A5ACA5967426FFF7B874FC5A4348FC7F" authority="Basilio" authorityName="Basilio" box="[556,839,958,984]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mayae">
<emphasis id="50D802077426FFF7B874FC5A42E4FC7F" box="[556,747,958,984]" italics="true" pageId="28" pageNumber="29">Coilodes mayae</emphasis>
Basílio
</taxonomicName>
&amp; Vaz-de-Mello sp. nov. is named after Maya Basílio Dantas, younger niece of the first author.
</paragraph>
</subSubSection>
<subSubSection id="2AB68D9E7426FFF7BAE5FBC142D1FA76" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="6213DE157426FFF7BAE5FBC14163FB98" blockId="28.[189,364,1061,1087]" box="[189,364,1061,1087]" pageId="28" pageNumber="29">
<heading id="395B69797426FFF7BAE5FBC14163FB98" bold="true" box="[189,364,1061,1087]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="50D802077426FFF7BAE5FBC14163FB98" bold="true" box="[189,364,1061,1087]" pageId="28" pageNumber="29">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157426FFF7BAB2FBB04153FBC9" blockId="28.[189,1091,1108,1171]" box="[234,348,1108,1134]" pageId="28" pageNumber="29">
<emphasis id="50D802077426FFF7BAB2FBB04153FBC9" bold="true" box="[234,348,1108,1134]" pageId="28" pageNumber="29">
<typeStatus id="BD1760B77426FFF7BAB2FBB04153FBC9" box="[234,348,1108,1134]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6213DE157426FFF7BAE5FB9D444CFB34" blockId="28.[189,1091,1108,1171]" box="[189,1091,1144,1171]" pageId="28" pageNumber="29">
<materialsCitation id="D2C4D4487426FFF7BAE5FB9D444CFB34" box="[189,1091,1144,1171]" collectingDate="1996-12" collectionCode="CEMT" collectorName="Louzada" country="Brazil" location="Castelo" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo" typeStatus="holotype">
<collectingCountry id="1ABB9E857426FFF7BAE5FB9D4129FB34" box="[189,294,1145,1171]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
•
<specimenCount id="74AA159C7426FFF7BB67FB9D4157FB34" box="[319,344,1145,1171]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
;
<collectingRegion id="A06810F77426FFF7BB3CFB9C4201FB34" box="[356,526,1144,1171]" country="Brazil" name="Espirito Santo" pageId="28" pageNumber="29">Espírito Santo</collectingRegion>
,
<location id="677388CE7426FFF7B842FB9C4279FB35" LSID="urn:lsid:plazi:treatment:EA056F037426FFF5BBA2FDE64481FB43:677388CE7426FFF7B842FB9C4279FB35" box="[538,630,1144,1170]" country="Brazil" name="Castelo" pageId="28" pageNumber="29" stateProvince="Espirito Santo">Castelo</location>
;
<date id="1612F8D57426FFF7B8D9FB9D42F2FB35" box="[641,765,1144,1171]" pageId="28" pageNumber="29" value="1996-12">
<collectingDate id="0656013D7426FFF7B8D9FB9D42F2FB35" box="[641,765,1144,1171]" pageId="28" pageNumber="29" value="1996-12">Dec. 1996</collectingDate>
</date>
; JNC
<collectorName id="CF59BBC37426FFF7B91DFB9C43A5FB35" box="[837,938,1144,1170]" pageId="28" pageNumber="29">Louzada</collectorName>
leg.;
<collectionCode id="04BD46D07426FFF7B9B4FB9D4430FB34" box="[1004,1087,1145,1171]" pageId="28" pageNumber="29">CEMT</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="6213DE157426FFF7BAB2FB5F4241FB72" blockId="28.[189,1399,1211,1489]" box="[234,590,1211,1237]" pageId="28" pageNumber="29">
<emphasis id="50D802077426FFF7BAB2FB5F4168FB72" bold="true" box="[234,359,1211,1237]" pageId="28" pageNumber="29">
<typeStatus id="BD1760B77426FFF7BAB2FB5F4168FB72" box="[234,359,1211,1237]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="74AA159C7426FFF7BB20FB5F41C7FB72" box="[376,456,1211,1237]" count="11" pageId="28" pageNumber="29" type="male">11 ♂♂</specimenCount>
and
<specimenCount id="74AA159C7426FFF7B859FB5F424EFB72" box="[513,577,1211,1237]" count="6" pageId="28" pageNumber="29" type="female">6 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="6213DE157426FFF7BAE5FB3B42D1FA76" blockId="28.[189,1399,1211,1489]" pageId="28" pageNumber="29">
<materialsCitation id="D2C4D4487426FFF7BAE5FB3B440EFB5E" box="[189,1025,1246,1274]" collectingDate="1996-12" collectionCode="CEMT" collectorName="Louzada" country="Brazil" location="Castelo" pageId="28" pageNumber="29" specimenCount="5" specimenCount-male="5" stateProvince="Espirito Santo">
<collectingCountry id="1ABB9E857426FFF7BAE5FB3B4129FB5E" box="[189,294,1247,1273]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
–
<emphasis id="50D802077426FFF7BB19FB3A41FAFB5E" bold="true" box="[321,501,1246,1273]" pageId="28" pageNumber="29">
<collectingRegion id="A06810F77426FFF7BB19FB3A41FAFB5E" box="[321,501,1246,1273]" country="Brazil" name="Espirito Santo" pageId="28" pageNumber="29">Espírito Santo</collectingRegion>
</emphasis>
•
<specimenCount id="74AA159C7426FFF7B855FB3B425DFB5E" box="[525,594,1247,1274]" count="5" pageId="28" pageNumber="29" type="male">5 ♂♂</specimenCount>
; same collection data as for holotype
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7BE41FB3B4146FABA" collectingDate="1996-12" collectionCode="CEMT" collectorName="Louzada" country="Brazil" location="Castelo" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7BE41FB3B4454FB5E" box="[1049,1115,1247,1273]" count="1" pageId="28" pageNumber="29" type="female">1 ♀♀</specimenCount>
; same collection data as for holotype
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7BB06FAE74389FABA" box="[350,902,1283,1309]" collectingDate="1996-12" collectionCode="DZUP" collectorName="Louzada" country="Brazil" location="Castelo" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7BB06FAE74184FABA" box="[350,395,1283,1309]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="04BD46D07426FFF7B963FAE74389FABA" box="[827,902,1283,1309]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="28" pageNumber="29" type="University or college">DZUP</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7B9C3FAE74578FABA" box="[923,1399,1283,1309]" collectingDate="1996-12" collectionCode="DZUP" collectorName="Louzada" country="Brazil" location="Castelo" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7B9C3FAE743C9FABA" box="[923,966,1283,1309]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; same collection data as for preceding
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7BA97FAC34311FAE6" box="[207,798,1319,1346]" collectingDate="1996-12" collectionCode="CERPE" collectorName="Louzada" country="Brazil" location="Castelo" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7BA97FAC340F1FAE5" box="[207,254,1319,1346]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="04BD46D07426FFF7B89BFAC34311FAE6" box="[707,798,1319,1345]" pageId="28" pageNumber="29">CERPE</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7B96FFAC34526FAE6" box="[823,1321,1319,1346]" collectingDate="1996-12" collectionCode="CERPE" collectorName="Louzada" country="Brazil" location="Castelo" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7B96FFAC3436AFAE5" box="[823,869,1319,1346]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; same collection data as for preceding
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7BF1AFAC343EBFAC2" collectingDate="2000-12" collectionCode="CEMT" collectorName="F. Z. Vaz-de-Mello" country="Brazil" location="Venda Nova do Imigrante" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7BF1AFAC3457EFAE5" box="[1346,1393,1319,1346]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
;
<location id="677388CE7426FFF7BAE5FAAF41E3FAC2" LSID="urn:lsid:plazi:treatment:EA056F037426FFF5BBA2FDE64481FB43:677388CE7426FFF7BAE5FAAF41E3FAC2" box="[189,492,1355,1382]" country="Brazil" name="Venda Nova do Imigrante" pageId="28" pageNumber="29" stateProvince="Espirito Santo">Venda Nova do Imigrante</location>
;
<date id="1612F8D57426FFF7BBAEFAA8427EFAC2" box="[502,625,1355,1382]" pageId="28" pageNumber="29" value="2000-12">
<collectingDate id="0656013D7426FFF7BBAEFAA8427EFAC2" box="[502,625,1355,1382]" pageId="28" pageNumber="29" value="2000-12">Dec. 2000</collectingDate>
</date>
;
<collectorName id="CF59BBC37426FFF7B824FAA8435EFAC2" box="[636,849,1355,1382]" pageId="28" pageNumber="29">F.Z. Vaz-de-Mello</collectorName>
leg.;
<collectionCode id="04BD46D07426FFF7B9C9FAAF43EBFAC2" box="[913,996,1355,1381]" pageId="28" pageNumber="29">CEMT</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7B9ABFAAF413FFA2E" collectingDate="2000-12" collectionCode="CEMT" collectorName="F. Z. Vaz-de-Mello" country="Brazil" location="Venda Nova do Imigrante" pageId="28" pageNumber="29" specimenCount="2" specimenCount-female="2" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7B9ABFAAF443AFAC1" box="[1011,1077,1355,1382]" count="2" pageId="28" pageNumber="29" type="female">2 ♀♀</specimenCount>
; same collection data as for preceding
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7BB1DFA8B4370FA2D" box="[325,895,1391,1418]" collectingDate="2000-12" collectionCode="DZUP" collectorName="F. Z. Vaz-de-Mello" country="Brazil" location="Venda Nova do Imigrante" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7BB1DFA8B417DFA2D" box="[325,370,1391,1418]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="04BD46D07426FFF7B96CFA944370FA2D" box="[820,895,1392,1418]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="28" pageNumber="29" type="University or college">DZUP</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7B9CDFA8B4579FA2E" box="[917,1398,1391,1418]" collectingDate="2000-12" collectionCode="DZUP" collectorName="F. Z. Vaz-de-Mello" country="Brazil" location="Venda Nova do Imigrante" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7B9CDFA8B43CFFA2D" box="[917,960,1391,1418]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; same collection data as for preceding
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7BA97FA774325FA0A" box="[207,810,1427,1454]" collectingDate="2000-12" collectionCode="CERPE" collectorName="F. Z. Vaz-de-Mello" country="Brazil" location="Venda Nova do Imigrante" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7BA97FA7740F1FA09" box="[207,254,1427,1454]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="04BD46D07426FFF7B897FA774325FA0A" box="[719,810,1427,1453]" pageId="28" pageNumber="29">CERPE</collectionCode>
</materialsCitation>
•
<materialsCitation id="D2C4D4487426FFF7B91CFA7742D5FA76" collectingDate="1998-10" collectionCode="CEMT" collectorName="Falqueto &amp; Vaz-de Mello" country="Brazil" location="Venda Nova do Imigrante" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo">
<specimenCount id="74AA159C7426FFF7B91CFA77437CFA09" box="[836,883,1427,1454]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<date id="1612F8D57426FFF7BF1DFA7740F4FA76" pageId="28" pageNumber="29" value="1998-10">
<collectingDate id="0656013D7426FFF7BF1DFA7740F4FA76" pageId="28" pageNumber="29" value="1998-10">Oct. 1998</collectingDate>
</date>
;
<collectorName id="CF59BBC37426FFF7BB5FFA534161FA76" box="[263,366,1463,1489]" pageId="28" pageNumber="29">Falqueto</collectorName>
and
<collectorName id="CF59BBC37426FFF7BBFEFA53424AFA76" box="[422,581,1463,1489]" pageId="28" pageNumber="29">Vaz-de Mello</collectorName>
leg.;
<collectionCode id="04BD46D07426FFF7B8DFFA5342D5FA76" box="[647,730,1463,1489]" pageId="28" pageNumber="29">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2AB68D9E7426FFF5BAE5FA1D4481FB43" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="6213DE157426FFF7BAE5FA1D4141F9B4" blockId="28.[189,334,1529,1555]" box="[189,334,1529,1555]" pageId="28" pageNumber="29">
<heading id="395B69797426FFF7BAE5FA1D4141F9B4" bold="true" box="[189,334,1529,1555]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="50D802077426FFF7BAE5FA1D4141F9B4" bold="true" box="[189,334,1529,1555]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157426FFF7BAB2F9CD41A4F9E4" blockId="28.[189,1263,1577,1640]" box="[234,427,1577,1603]" pageId="28" pageNumber="29">
<emphasis id="50D802077426FFF7BAB2F9CD4125F9E4" bold="true" box="[234,298,1577,1603]" pageId="28" pageNumber="29">Male</emphasis>
(
<typeStatus id="BD1760B77426FFF7BB61F9CD41ACF9E4" box="[313,419,1577,1603]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="6213DE157426FFF7BAE5F9A944E0F9C0" blockId="28.[189,1263,1577,1640]" box="[189,1263,1613,1640]" pageId="28" pageNumber="29">
MEASUREMENTS. Length
<quantity id="A55473F07426FFF7BB86F9A94234F9CF" box="[478,571,1613,1640]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="28" pageNumber="29" unit="mm" value="5.6">5.6 mm</quantity>
. Width
<quantity id="A55473F07426FFF7B8CDF9A942FDF9CF" box="[661,754,1613,1640]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="28" pageNumber="29" unit="mm" value="3.6">3.6 mm</quantity>
. Body (
<figureCitation id="FA97C2907426FFF7B915F9A943C3F9C0" box="[845,972,1613,1639]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="28" pageNumber="29">Fig. 7A–B</figureCitation>
) convex, oval and shiny.
</paragraph>
<paragraph id="6213DE157426FFF7BAE5F974428AF96A" blockId="28.[189,1398,1679,1741]" pageId="28" pageNumber="29">COLOUR. Head and pronotum dark brown; scutellar shield, venter and legs yellowish brown; elytra yellowish brown, margins dark brown.</paragraph>
<paragraph id="6213DE157426FFF6BAE5F9124546FECE" blockId="28.[187,1400,1782,2025]" lastBlockId="29.[188,1398,265,362]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
HEAD (
<figureCitation id="FA97C2907426FFF7BB49F912419BF8B7" box="[273,404,1782,1808]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="28" pageNumber="29">Fig. 7C–D</figureCitation>
). Surface strigulate. Frons, in dorsal view, subrectangular; one small tubercle in the middle; proximal border rounded; posterior margin as wide as the anterior one; lateral margin slightly arched; sparse punctures, restricted to region next to clypeus. Eye barely visible dorsally. Canthus strong; distinct area expanded downwards. Clypeus subquadrate; lateral margins weakly rounded; anterior borders rounded; apex straight; setae thin and sparse, throughout the anterior margin dorsally; sparse punctures. Labrum semicircular; slight medial projection; setae sparse, spreading along the entire dorsal border. Mandibles dorsally excavated, protruding beyond the apex of labrum. Labium with mentum subquadrate; inferior margin arched; lateral margins slightly rounded and superior margin curved in the middle; disc strigulate, long setae on the margins. Maxilla subtriangular; long setae throughout surface. Antenna with 10 antennomeres; antennal club with three antennomeres; club with basal antennomere cupuliform.
</paragraph>
<paragraph id="6213DE157427FFF6BAE5FE7544E4FE56" blockId="29.[188,1399,401,498]" pageId="29" pageNumber="30">
PRONOTUM (
<figureCitation id="FA97C2907427FFF6BB17FE7541A6FE0C" box="[335,425,401,427]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="29" pageNumber="30">Fig. 7E</figureCitation>
). Convex; strongly excavated medially; subtrapezoidal, posterior margin wider than anterior margin; anterior margin straight, weakly produced medially, angles subacute; posterior margin rounded and weakly produced medially; lateral margin convex, setae and punctures absent.
</paragraph>
<paragraph id="6213DE157427FFF6BAE5FDFD4446FD94" blockId="29.[189,1097,537,563]" box="[189,1097,537,563]" pageId="29" pageNumber="30">SCUTELLAR SHIELD. Subtriangular; setae and punctures absent; apex punctate.</paragraph>
<paragraph id="6213DE157427FFF6BAE5FDBE44FDFDD2" blockId="29.[189,1266,602,629]" box="[189,1266,602,629]" pageId="29" pageNumber="30">ELYTRA. Double row of longitudinal punctures, sutural stria complete, formed by punctures.</paragraph>
<paragraph id="6213DE157427FFF6BAE5FD784578FD7E" blockId="29.[188,1399,667,729]" pageId="29" pageNumber="30">
VENTER (
<figureCitation id="FA97C2907427FFF6BB72FD7F4189FD11" box="[298,390,667,694]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="29" pageNumber="30">Fig. 7B</figureCitation>
). Hypomeral surface strigulate, setose; prosternum (
<figureCitation id="FA97C2907427FFF6B9BFFD7F446BFD11" box="[999,1124,667,694]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="29" pageNumber="30">Fig. 7B, D</figureCitation>
) elevated at midlength, transversally grooved; anterior margin rounded and slightly jagged, mesoventrite and metaventrite smooth.
</paragraph>
<caption id="36D38E9D7427FFF6BAE5F8804516F84E" ID-DOI="http://doi.org/10.5281/zenodo.10407095" ID-Zenodo-Dep="10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,231,1892,1918]" targetBox="[198,1398,774,1811]" targetPageId="29">
<paragraph id="6213DE157427FFF6BAE5F8804516F84E" blockId="29.[188,1399,1892,2025]" pageId="29" pageNumber="30">
<emphasis id="50D802077427FFF6BAE5F8804108F8D8" bold="true" box="[189,263,1892,1919]" pageId="29" pageNumber="30">Fig. 7.</emphasis>
<taxonomicName id="A5ACA5967427FFF6BB54F8814217F8D8" authority="Basilio" authorityName="Basilio" box="[268,536,1893,1919]" class="Insecta" family="Hybosoridae" genus="Coilodes" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mayae">
<emphasis id="50D802077427FFF6BB54F88141CEF8D8" box="[268,449,1893,1919]" italics="true" pageId="29" pageNumber="30">Coilodes mayae</emphasis>
Basílio
</taxonomicName>
&amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I).
<emphasis id="50D802077427FFF6BB57F86C412BF805" bold="true" box="[271,292,1928,1954]" pageId="29" pageNumber="30">A</emphasis>
. Body, dorsal view.
<emphasis id="50D802077427FFF6B847F86C423EF805" bold="true" box="[543,561,1928,1954]" pageId="29" pageNumber="30">B</emphasis>
. Body, ventral view.
<emphasis id="50D802077427FFF6B96EF86C4345F805" bold="true" box="[822,842,1928,1954]" pageId="29" pageNumber="30">C</emphasis>
. Head, dorsal view.
<emphasis id="50D802077427FFF6BE1CF86C4457F805" bold="true" box="[1092,1112,1928,1954]" pageId="29" pageNumber="30">D</emphasis>
. Head, ventral view.
<emphasis id="50D802077427FFF6BF05F86C4560F805" bold="true" box="[1373,1391,1928,1954]" pageId="29" pageNumber="30">E</emphasis>
. Pronotum, dorsal view.
<emphasis id="50D802077427FFF6BB88F84F4207F862" bold="true" box="[464,520,1963,1990]" pageId="29" pageNumber="30">F–K</emphasis>
. Male genitalia.
<emphasis id="50D802077427FFF6B891F84F42D4F862" bold="true" box="[713,731,1963,1989]" pageId="29" pageNumber="30">F</emphasis>
. Dorsal view.
<emphasis id="50D802077427FFF6B9DAF84F4396F862" bold="true" box="[898,921,1963,1989]" pageId="29" pageNumber="30">G</emphasis>
. Ventral view.
<emphasis id="50D802077427FFF6BE1DF84F4453F862" bold="true" box="[1093,1116,1963,1989]" pageId="29" pageNumber="30">H</emphasis>
. Right paramere.
<emphasis id="50D802077427FFF6BF73F84F4539F862" bold="true" box="[1323,1334,1963,1989]" pageId="29" pageNumber="30">I</emphasis>
. Left paramere.
<emphasis id="50D802077427FFF6BB61F82A4147F84F" bold="true" box="[313,328,1998,2024]" pageId="29" pageNumber="30">J</emphasis>
. Right side.
<emphasis id="50D802077427FFF6BB85F82A41FBF84F" bold="true" box="[477,500,1998,2024]" pageId="29" pageNumber="30">K</emphasis>
. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm.
</paragraph>
</caption>
<paragraph id="6213DE157424FFF5BAE5FEED44B6FE5C" blockId="30.[187,1400,265,508]" pageId="30" pageNumber="31">LEGS. Procoxa conical; surface strigulate. Protrochanter joint with the procoxa rounded, distally angulate. Profemur with posterior margin carinate. Protibia with inner margin convex; carinate; single spur, slightly curved at the apex; external margin with three larger teeth and series of smaller denticles along the entire margin. Protarsi with tarsal insertion beneath the protibial second tooth. Mesothoracic and metathoracic legs smooth. Mesotrochanter and metatrochanter subtriangular with bifurcated apex. Mesofemur and metafemur carinate on posterior margin. Mesotibia and metatibia with erect setae; apex weakly expanded; pair of spurs with sharp apex. Tarsi with five tarsomeres; tarsal claws toothed medially.</paragraph>
<paragraph id="6213DE157424FFF5BAE5FDC742D2FD99" blockId="30.[189,733,547,574]" box="[189,733,547,574]" pageId="30" pageNumber="31">ABDOMEN. Six ventrites with recumbent setae.</paragraph>
<paragraph id="6213DE157424FFF5BAE5FD8242C1FCB7" blockId="30.[187,1400,614,784]" pageId="30" pageNumber="31">
GENITALIA (
<figureCitation id="FA97C2907424FFF5BB1FFD8241CAFD27" box="[327,453,614,640]" captionStart="Fig" captionStartId="29.[189,231,1892,1918]" captionTargetBox="[198,1398,774,1811]" captionTargetId="figure-12@29.[189,1398,766,1854]" captionTargetPageId="29" captionText="Fig. 7. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov., holotype ♂ (A–G, J–K) and graphic representation (H–I). A. Body, dorsal view. B. Body, ventral view. C. Head, dorsal view. D. Head, ventral view. E. Pronotum, dorsal view. F–K. Male genitalia. F. Dorsal view. G. Ventral view. H. Right paramere. I. Left paramere. J. Right side. K. Left side. Scale bars: A–B = 1 mm; C–E = 0.5 mm; F–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10407095" httpUri="https://zenodo.org/record/10407095/files/figure.png" pageId="30" pageNumber="31">Fig. 7F–K</figureCitation>
). Lobe of tegmen absent; region with strongly sclerotized projection next to right paramere base, sclerotized laminar structure on inferior ventral region of tegmen, parallel to the sides of tegmen; parameres asymmetrical; right paramere longer than it is wide, base as wide as apex, strong constriction next to paramere base, gradually increasing towards the apex, rounded vertices; left paramere, wide base and apex strongly curved outward.
</paragraph>
<paragraph id="6213DE157424FFF5BAB2FCDC414AFCF5" blockId="30.[189,663,824,887]" box="[234,325,824,850]" pageId="30" pageNumber="31">
<emphasis id="50D802077424FFF5BAB2FCDC414AFCF5" bold="true" box="[234,325,824,850]" pageId="30" pageNumber="31">Female</emphasis>
</paragraph>
<paragraph id="6213DE157424FFF5BAE5FCB84298FCD1" blockId="30.[189,663,824,887]" box="[189,663,860,887]" pageId="30" pageNumber="31">
Length 5.5–6.0 mm. Width
<quantity id="A55473F07424FFF5B85AFCB8429DFCD1" box="[514,658,860,887]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="30" pageNumber="31" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.1–3.5 mm</quantity>
.
</paragraph>
<paragraph id="6213DE157424FFF5BAB2FC7A416FFC1F" blockId="30.[189,1398,926,1025]" box="[234,352,926,952]" pageId="30" pageNumber="31">
<emphasis id="50D802077424FFF5BAB2FC7A416FFC1F" bold="true" box="[234,352,926,952]" pageId="30" pageNumber="31">Variation</emphasis>
</paragraph>
<paragraph id="6213DE157424FFF5BAE5FC2643E4FBA7" blockId="30.[189,1398,926,1025]" pageId="30" pageNumber="31">
Length ranging from
<quantity id="A55473F07424FFF5BBE2FC26426DFC7A" box="[442,610,962,989]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="6.2" metricValueMin="5.3" pageId="30" pageNumber="31" unit="mm" value="5.75" valueMax="6.2" valueMin="5.3">5.3 to 6.2 mm</quantity>
. Width ranging from
<quantity id="A55473F07424FFF5B906FC264409FC7A" box="[862,1030,962,989]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.6" metricValueMin="3.1" pageId="30" pageNumber="31" unit="mm" value="3.35" valueMax="3.6" valueMin="3.1">3.1 to 3.6 mm</quantity>
. Elytra with dark brown region restricted to the margins or extended to the entire distal half of elytra.
</paragraph>
<paragraph id="6213DE157424FFF5BAE5FBCC420DFBE5" blockId="30.[189,514,1064,1090]" box="[189,514,1064,1090]" pageId="30" pageNumber="31">
<heading id="395B69797424FFF5BAE5FBCC420DFBE5" bold="true" box="[189,514,1064,1090]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="50D802077424FFF5BAE5FBCC420DFBE5" bold="true" box="[189,514,1064,1090]" pageId="30" pageNumber="31">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157424FFF5BAE5FBBC4240FBD5" blockId="30.[189,591,1112,1138]" box="[189,591,1112,1138]" pageId="30" pageNumber="31">
<collectingCountry id="1ABB9E857424FFF5BAE5FBBC410AFBD5" box="[189,261,1112,1138]" name="Brazil" pageId="30" pageNumber="31">Brazil</collectingCountry>
(
<collectingRegion id="A06810F77424FFF5BB4CFBBC41B0FBD5" box="[276,447,1112,1138]" country="Brazil" name="Espirito Santo" pageId="30" pageNumber="31">Espírito Santo</collectingRegion>
) (
<figureCitation id="FA97C2907424FFF5BB8FFBBC424DFBD5" box="[471,578,1112,1138]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1739]" captionTargetId="figure-12@47.[272,1315,265,1739]" captionTargetPageId="47" captionText="Fig. 14. Distribution map of species of Coilodes Westwood, 1846. A. Coilodes bezerrai Basílio &amp; Vazde-Mello sp. nov.; Coilodes castaneus Westwood, 1846; and Coilodes edeiltae Basílio &amp; Vaz-de-Mello sp. nov. B. Coilodes fumipennis Arrow, 1909; Coilodes humeralis (Mannerheim, 1829); and Coilodes lunae Basílio &amp; Vaz-de-Mello sp. nov. C. Coilodes mayae Basílio &amp; Vaz-de-Mello sp. nov.;Coilodes niger (Mannerheim, 1829) stat. rev.; Coilodes ovalis Robinson, 1948; and Coilodes parvulus Westwood 1846. D. Coilodes punctipennis Arrow, 1909; Coilodes ravii Basílio &amp; Vaz-de-Mello sp. nov.; and Coilodes skelleyi Basílio &amp; Vaz-de-Mello sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10407111" httpUri="https://zenodo.org/record/10407111/files/figure.png" pageId="30" pageNumber="31">Fig. 14C</figureCitation>
).
</paragraph>
<paragraph id="6213DE157424FFF5BAE5FB7E4178FB13" blockId="30.[189,375,1178,1204]" box="[189,375,1178,1204]" pageId="30" pageNumber="31">
<heading id="395B69797424FFF5BAE5FB7E4178FB13" bold="true" box="[189,375,1178,1204]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="50D802077424FFF5BAE5FB7E4178FB13" bold="true" box="[189,375,1178,1204]" pageId="30" pageNumber="31">Biological data</emphasis>
</heading>
</paragraph>
<paragraph id="6213DE157424FFF5BAE5FB2D4481FB43" blockId="30.[189,1166,1225,1252]" box="[189,1166,1225,1252]" pageId="30" pageNumber="31">Species collected in: pitfall trap with human faeces bait and flight interception trap.</paragraph>
</subSubSection>
</treatment>
</document>
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