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<document id="8E889B5EE1D333BBA041355914B610C8" ID-DOI="10.5281/zenodo.208726" ID-GBIF-Dataset="1d584fe5-b319-4d05-a0b4-affacba792b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="208726" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460512631462" checkinUser="plazi" docAuthor="Perger, Robert &amp; Guerra, Fernando" docDate="2012" docId="E50287DEDB0E986AFF68D4F256770DC4" docLanguage="en" docName="zt03434p058.pdf" docOrigin="Zootaxa 3434" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cylindera (Plectographa) yaguaree Perger &amp; Guerra, n.sp." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="193BFFA6DB0D986FFFFFD46552090F40" masterDocTitle="Two new tiger beetle (Coleoptera, Carabidae, Cicindelitae) species from the Tucuman-Bolivian forest in the National Tariquia Reserve, Bolivia" masterLastPageNumber="58" masterPageNumber="49" pageNumber="52" updateTime="1698313507748" updateUser="plazi">
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<mods:title id="A724ACCFE8CB90BC85ECD4277344AE7C">Two new tiger beetle (Coleoptera, Carabidae, Cicindelitae) species from the Tucuman-Bolivian forest in the National Tariquia Reserve, Bolivia</mods:title>
</mods:titleInfo>
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<mods:namePart id="7DF50EEBFB593E30C40F9A07341C26E7">Perger, Robert</mods:namePart>
</mods:name>
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<mods:namePart id="19E8B17CD3CFED8FABA519DD83F5225F">Guerra, Fernando</mods:namePart>
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<mods:date id="CC73F00B9401D3741C08DCC326C9CE71">2012</mods:date>
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<mods:number id="DB226AA8B6CF01A843C1DE3B2E2C1D2E">3434</mods:number>
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<mods:classification id="FEEB7037EDFA0097F36E74144F481524">journal article</mods:classification>
<mods:identifier id="81EC1269769EF3C5245E02C31E5AE6B8" type="DOI">10.5281/zenodo.208726</mods:identifier>
<mods:identifier id="0D0818DA3AE7DC9F2401BE8353AC536A" type="GBIF-Dataset">1d584fe5-b319-4d05-a0b4-affacba792b7</mods:identifier>
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<treatment id="E50287DEDB0E986AFF68D4F256770DC4" ID-DOI="http://doi.org/10.5281/zenodo.6491791" ID-GBIF-Taxon="119523421" ID-Zenodo-Dep="6491791" LSID="urn:lsid:plazi:treatment:E50287DEDB0E986AFF68D4F256770DC4" httpUri="http://treatment.plazi.org/id/E50287DEDB0E986AFF68D4F256770DC4" lastPageId="5" lastPageNumber="54" pageId="3" pageNumber="52">
<subSubSection id="25B16543DB0E986CFF68D4F253700F92" pageId="3" pageNumber="52" type="nomenclature">
<paragraph id="6D1436C8DB0E986CFF68D4F251680FF1" blockId="3.[151,865,151,210]" box="[151,865,151,177]" pageId="3" pageNumber="52">
<heading id="365C81A4DB0E986CFF68D4F251680FF1" bold="true" box="[151,865,151,177]" fontSize="11" level="1" pageId="3" pageNumber="52" reason="1">
<emphasis id="5FDFEADADB0E986CFF68D4F251680FF1" bold="true" box="[151,865,151,177]" pageId="3" pageNumber="52">
<taxonomicName id="AAAB4D4BDB0E986CFF68D4F251160FF1" ID-CoL="3329Z" authority="Perger &amp; Guerra" authorityName="Perger &amp; Guerra" box="[151,799,151,177]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB0E986CFF68D4F253070FF1" bold="true" box="[151,270,151,177]" italics="true" pageId="3" pageNumber="52">Cylindera</emphasis>
(
<emphasis id="5FDFEADADB0E986CFEE0D4F253C90FF1" bold="true" box="[287,448,151,177]" italics="true" pageId="3" pageNumber="52">Plectographa</emphasis>
)
<emphasis id="5FDFEADADB0E986CFE2ED4F250490FF1" bold="true" box="[465,576,151,177]" italics="true" pageId="3" pageNumber="52">yaguaree</emphasis>
Perger &amp; Guerra
</taxonomicName>
<taxonomicNameLabel id="44EC57A1DB0E986CFCD9D4F251680FF1" box="[806,865,151,177]" pageId="3" pageNumber="52" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6D1436C8DB0E986CFF68D4DF53700F92" blockId="3.[151,865,151,210]" box="[151,377,186,210]" pageId="3" pageNumber="52">
(
<figureCitation id="F5902A4DDB0E986CFF60D4DF52E00F92" box="[159,233,186,210]" captionStart="FIGURE 1. A" captionStartId="2.[151,248,1801,1824]" captionTargetBox="[171,1417,379,1770]" captionTargetId="figure@2.[169,1417,379,1780]" captionTargetPageId="2" captionText="FIGURE 1. A, Relief map Bolivia, subhumid Tucuman-Bolivian forest highlighted green, according to Navarro &amp; Ferreira (2007); (+) most southern records for Pseudoxycheila andina Cassola 1997, (*) Pseudoxycheila tucumana n. sp. (coll. MHN- NKM); B, Subandean region of Tarija department, study areas indicated by solid white line, limit of Tariquia by white line; C, study areas, view from the Andes; D, E, surveyed transects (white lines), collection location of Cylindera (Plectographa) yaguaree n. sp. (red points) and P. tucumana n. sp (red lines), (a) S 21 ° 45 ' 18, W 64 ° 12 ' 32, (b) S 21 ° 49 ' 23, W 64 ° 14 ' 35; F, habitat of C. yaguaree n. sp. and P. tucumana n. sp.; Life habitus and preferred substrate, G, C. (P.) yaguaree n. sp.; H, P. t u c u m a n a n. sp." httpUri="https://zenodo.org/record/208727/files/figure.png" pageId="3" pageNumber="52">Figs. 1</figureCitation>
G; 2; 3A, B)
</paragraph>
</subSubSection>
<subSubSection id="25B16543DB0E986CFF68D56653720D56" pageId="3" pageNumber="52" type="materials_examined">
<paragraph id="6D1436C8DB0E986CFF68D56656BC0EEA" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">
<emphasis id="5FDFEADADB0E986CFF68D566534B0E5B" bold="true" box="[151,322,258,283]" pageId="3" pageNumber="52">
<typeStatus id="B210886ADB0E986CFF68D56652D90E5B" box="[151,208,259,283]" pageId="3" pageNumber="52">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="15BC7658DB0E986CFEB4D56753A90E5A" box="[331,416,258,282]" name="Bolivia" pageId="3" pageNumber="52">Bolivia</collectingCountry>
, Tarija, O`Connor province, Tariquia:
<specimenCount id="7BADFD41DB0E986CFCB3D56751930E5A" box="[844,922,258,282]" pageId="3" pageNumber="52" type="male">1 male</specimenCount>
,
<typeStatus id="B210886ADB0E986CFC5AD56756030E5A" box="[933,1034,258,282]" pageId="3" pageNumber="52" type="holotype">holotype</typeStatus>
,
<collectionCode id="0BBAAE0DDB0E986CFBEBD56756430E5A" box="[1044,1098,258,282]" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="3" pageNumber="52">CBF</collectionCode>
,
<quantity id="AA539B2DDB0E986CFBAAD56756AB0E5A" box="[1109,1186,258,282]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="52" unit="km" value="1.4">1.4 km</quantity>
south west of Salinas,
<quantity id="AA539B2DDB0E986CFF68D54352FD0E7E" box="[151,244,294,318]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="52" unit="km" value="0.66">0.66 km</quantity>
east of Salinas River,
<geoCoordinate id="089F500FDB0E986CFE0BD54350450E7E" box="[500,588,294,318]" direction="south" orientation="latitude" pageId="3" pageNumber="52" precision="925" value="-21.816668">S21°49'</geoCoordinate>
20,
<geoCoordinate id="089F500FDB0E986CFD88D54350D50E7E" box="[631,732,294,318]" direction="west" orientation="longitude" pageId="3" pageNumber="52" precision="925" value="-64.23333">W64°14'</geoCoordinate>
45, 1093 m a.s.l., Tucuman-Bolivian subhumid forest, bank of small mountain river,
<date id="19151008DB0E986CFE71D52F504F0E22" box="[398,582,330,354]" pageId="3" pageNumber="52" value="2011-11">November 2011</date>
, F. Guerra and R. Perger;
<specimenCount id="7BADFD41DB0E986CFC94D52F51D90E22" box="[875,976,330,354]" pageId="3" pageNumber="52" type="female">1 female</specimenCount>
, allotype, same data;
<specimenCount id="7BADFD41DB0E986CFB3CD52F57140E22" box="[1219,1309,330,354]" pageId="3" pageNumber="52" type="male">2 males</specimenCount>
,
<specimenCount id="7BADFD41DB0E986CFAD7D52F57910E22" box="[1320,1432,330,354]" pageId="3" pageNumber="52" type="female">3 females</specimenCount>
,
<typeStatus id="B210886ADB0E986CFF68D50A530C0EC7" box="[151,261,367,391]" pageId="3" pageNumber="52" type="paratype">paratypes</typeStatus>
,
<collectionCode id="0BBAAE0DDB0E986CFEEFD50B534F0EC6" box="[272,326,366,390]" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="3" pageNumber="52">CBF</collectionCode>
,
<quantity id="AA539B2DDB0E986CFEAED50B53960EC6" box="[337,415,366,390]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="52" unit="km" value="3.2">3.2 km</quantity>
east of Salinas River,
<quantity id="AA539B2DDB0E986CFD64D50B50E00EC6" box="[667,745,366,390]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="52" unit="km" value="2.5">2.5 km</quantity>
east of road to Salinas,
<geoCoordinate id="089F500FDB0E986CFC09D50B56590EC6" box="[1014,1104,366,390]" direction="south" orientation="latitude" pageId="3" pageNumber="52" precision="925" value="-21.75">S21°45'</geoCoordinate>
18,
<geoCoordinate id="089F500FDB0E986CFB87D50B56D50EC6" box="[1144,1244,366,390]" direction="west" orientation="longitude" pageId="3" pageNumber="52" precision="925" value="-64.2">W64°12'</geoCoordinate>
32,
<quantity id="AA539B2DDB0E986CFAFAD50B57530EC6" box="[1285,1370,366,390]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="52" unit="m" value="1200.0">1200 m</quantity>
a.s.l., Tucuman-Bolivian subhumid forest, bank of small mountain river,
<date id="19151008DB0E986CFC7AD5F756300EEA" box="[901,1081,402,426]" pageId="3" pageNumber="52" value="2011-12">December 2011</date>
, F. Guerra.
</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D5D353720D56" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">
<emphasis id="5FDFEADADB0E986CFF38D5D350240E8F" bold="true" box="[199,557,438,463]" pageId="3" pageNumber="52">Derivation of specific epithet.</emphasis>
The epithet “
<taxonomicName id="AAAB4D4BDB0E986CFD33D5D3513C0E8E" box="[716,821,438,462]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="yaguaree">yaguaree</taxonomicName>
” (pronounced a-wa-ree) is derived from “yaguarete”, which is the original name of the jaguar and means “true beast” in Tupi-Guaraní, a language spoken by the local indigenous Guaraní.
</paragraph>
</subSubSection>
<subSubSection id="25B16543DB0E986CFF38D64750E20C3E" pageId="3" pageNumber="52" type="diagnosis">
<paragraph id="6D1436C8DB0E986CFF38D64750450DE6" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">
<emphasis id="5FDFEADADB0E986CFF38D64753480D7B" bold="true" box="[199,321,546,571]" pageId="3" pageNumber="52">Diagnosis.</emphasis>
<taxonomicName id="AAAB4D4BDB0E986CFEB7D64650D10D7B" box="[328,728,546,571]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB0E986CFEB7D64653B10D7B" box="[328,440,547,571]" italics="true" pageId="3" pageNumber="52">Cylindera</emphasis>
(
<emphasis id="5FDFEADADB0E986CFE37D64650690D7B" box="[456,608,547,571]" italics="true" pageId="3" pageNumber="52">Plectographa</emphasis>
)
<emphasis id="5FDFEADADB0E986CFD8ED64150D10D7B" box="[625,728,548,571]" italics="true" pageId="3" pageNumber="52">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB0E986CFD1FD646511E0D7B" bold="true" box="[736,791,547,571]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="44EC57A1DB0E986CFD1FD646511E0D7B" box="[736,791,547,571]" pageId="3" pageNumber="52" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
is separated from other Neotropical congeners by a combination of the following characters: 1) setose genae and head; 2) posteriorly tapering pronotum, pronotal disc laterally convex; 3) elytral surface completely covered with stiff, appressed setae; 4) three complete narrow elytral maculations; 5) sutural spines reduced.
</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D6D750E20C3E" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">
Three complete narrow elytral maculations and setose gena are also typical of
<taxonomicName id="AAAB4D4BDB0E986CFBCCD6D6575E0D8A" authority="Horn 1915" authorityName="Horn" authorityYear="1915" box="[1075,1367,690,715]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="mixtula" subGenus="Plectographa">
<emphasis id="5FDFEADADB0E986CFBCCD6D656450D8A" box="[1075,1100,691,714]" italics="true" pageId="3" pageNumber="52">C.</emphasis>
(
<emphasis id="5FDFEADADB0E986CFBA3D6D156670D8B" box="[1116,1134,692,715]" italics="true" pageId="3" pageNumber="52">P.</emphasis>
)
<emphasis id="5FDFEADADB0E986CFB7ED6D656DC0D8B" box="[1153,1237,691,715]" italics="true" pageId="3" pageNumber="52">mixtula</emphasis>
<bibRefCitation id="093A4B39DB0E986CFB23D6D6575E0D8A" author="Horn" box="[1244,1367,690,714]" pageId="3" pageNumber="52" refString="Horn, W. (1915) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. Genera Insectorum, 82, 209 - 486, pls. 16 - 23." type="journal article" year="1915">Horn 1915</bibRefCitation>
</taxonomicName>
, but it lacks the elytral setae and the pronotum tapers in the opposite direction. In
<taxonomicName id="AAAB4D4BDB0E986CFC1BD6B256830DAF" box="[996,1162,726,751]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="sinuosa" subGenus="Plectographa">
<emphasis id="5FDFEADADB0E986CFC1BD6B251F70DAE" box="[996,1022,727,750]" italics="true" pageId="3" pageNumber="52">C.</emphasis>
(
<emphasis id="5FDFEADADB0E986CFBF0D6BD56280DAF" box="[1039,1057,728,751]" italics="true" pageId="3" pageNumber="52">P.</emphasis>
)
<emphasis id="5FDFEADADB0E986CFBCBD6BD56830DAF" box="[1076,1162,728,751]" italics="true" pageId="3" pageNumber="52">sinuosa</emphasis>
</taxonomicName>
and
<taxonomicName id="AAAB4D4BDB0E986CFB3ED6B2577B0DAE" box="[1217,1394,726,751]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="suturalis" subGenus="Plectographa">
<emphasis id="5FDFEADADB0E986CFB3ED6B256D20DAE" box="[1217,1243,727,750]" italics="true" pageId="3" pageNumber="52">C.</emphasis>
(
<emphasis id="5FDFEADADB0E986CFB14D6BD56F40DAF" box="[1259,1277,728,751]" italics="true" pageId="3" pageNumber="52">P.</emphasis>
)
<emphasis id="5FDFEADADB0E986CFAEFD6B3577B0DAE" box="[1296,1394,726,750]" italics="true" pageId="3" pageNumber="52">suturalis</emphasis>
</taxonomicName>
the elytra have three complete maculations, but these are wider and the genae are bare.
<taxonomicName id="AAAB4D4BDB0E986CFB89D69E52F70C77" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB0E986CFB89D69E56EF0C53" box="[1142,1254,763,787]" italics="true" pageId="3" pageNumber="52">Cylindera</emphasis>
(
<emphasis id="5FDFEADADB0E986CFB05D69E579B0C53" box="[1274,1426,763,787]" italics="true" pageId="3" pageNumber="52">Plectographa</emphasis>
)
<emphasis id="5FDFEADADB0E986CFF68D74552F70C77" box="[151,254,800,823]" italics="true" pageId="3" pageNumber="52">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB0E986CFEF6D77A53490C77" bold="true" box="[265,320,799,823]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="44EC57A1DB0E986CFEF6D77A53490C77" box="[265,320,799,823]" pageId="3" pageNumber="52" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
shares the posteriorly tapering pronotum and setose elytra with the Argentinean species
<taxonomicName id="AAAB4D4BDB0E986CFAAFD77A53910C1A" authority="Castelnau 1835" authorityName="Castelnau" authorityYear="1835" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="eugeni" subGenus="Plectographa">
<emphasis id="5FDFEADADB0E986CFAAFD77A57630C76" box="[1360,1386,799,822]" italics="true" pageId="3" pageNumber="52">C.</emphasis>
(
<emphasis id="5FDFEADADB0E986CFA81D74557990C77" box="[1406,1424,800,823]" italics="true" pageId="3" pageNumber="52">P.</emphasis>
)
<emphasis id="5FDFEADADB0E986CFF68D72152EB0C1B" box="[151,226,836,859]" italics="true" pageId="3" pageNumber="52">eugeni</emphasis>
Castelnau 1835
</taxonomicName>
; however, this latter species is distinguished by glabrous head, large subsutural foveae, wide elytral maculations and elytra glabrous in apical half.
</paragraph>
</subSubSection>
<subSubSection id="25B16543DB0E986AFF38D7EF56770DC4" lastPageId="5" lastPageNumber="54" pageId="3" pageNumber="52" type="description">
<paragraph id="6D1436C8DB0E986CFF38D7EF53550CAA" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">
<emphasis id="5FDFEADADB0E986CFF38D7EF53500CE3" bold="true" box="[199,345,906,931]" pageId="3" pageNumber="52">Description.</emphasis>
General. Relatively large (
<quantity id="AA539B2DDB0E986CFD69D7EF51560CE2" box="[662,863,906,930]" metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.095" metricValueMin="1.015" pageId="3" pageNumber="52" unit="mm" value="10.55" valueMax="10.95" valueMin="10.15">10.15–10.95 mm</quantity>
), dorsum matte to slightly shining cupreous-dark olive green, elytra with distinct, separate appressed setae covering most of the surface and three complete, narrow pale maculations.
</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D79350E60B82" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Setation. Dense appressed or suberect setae on body laterally (except area behind genae and on mesepisternum), on pronotal dorsum, on femora and laterally on meso- and metacoxa. Labial palpi and proepisternum with suberect to erect setae. Frons, vertex and elytra with separate, distinct setae. Distal 7 antennomeres, tibia and tarsi longitudinally lined with very fine and short setae. Labrum with 9 to 12 submarginal setae. Antennal scape with 1 erect subapical seta distal, antennomere 4 with three or more setae, apex of front trochanters with a single sensory seta. Apical setae on eighth abdominal sternum short.</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D0AB50BD0A6E" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Head slightly shining cupreous, vertex rugose, frons and gena striate. Labrum narrower or as wide as clypeus, subrectangular, straight, short, uni- or obtuse tridentate, ferrugineo-testaceous to dark brown. Eyes prominent, not bulging laterally. Mentum tooth well developed.</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D15F505B0ADA" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Mandibular base ferrugineo-testaceous, third terebral tooth shiny green with black tip, incisor tooth black. Antennal socket and antennomeres 1-4 shiny cupreous, with green reflections or completely green, distal 7 antennomeres testaceous, densely pubescent.</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D1C253D7096A" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Pronotum slightly shining cupreous, broad, widest in the middle, anterior margin noticably wider than posterior, posterior constricted, both margins with deep transverse sulci that are shiny cupreous; pronotal disc laterally convex, rugose, median groove distinct; proepisternum wrinkled, shiny cupreous, coupling sulcus with distinct groove. Scutellum triangular.</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D25351E5099E" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Elytra gradually widened to apical 1/4, then narrow until just before apex, sutural spine reduced; color matt brown cupreous with small shallow, weakly reflective green punctures, giving an overall olive impression; subsutural row of large foveae absent, three narrow, pale maculations, laterally connected, anterior G-shaped, median sharply elbowed, bend without additional bracket, not connected to posthumeral spot, sometimes slightly lacerated, mostly continuous, apical maculation U-shaped, apical-sutural arm reduced.</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D28F53120866" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Abdominal sternites dark brown with cupreous-green reflections. Ventral sclerite with two elongate posterior projections.</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D3575073082E" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Coxae shiny cupreous with green reflections; trochanters testaceous, femora cupreous, tibia and tarsomeres cupreous with or without green reflections.</paragraph>
<paragraph id="6D1436C8DB0E986CFF38D31F5154089A" blockId="3.[151,1437,258,2010]" pageId="3" pageNumber="52">Aedeagus strongly bent in basal quarter, in distal moderately bent; slightly widening distally in basal third, then considerably thickened median to shortly to the base of the apex and finally strongly narrowing on one side to a pointing apex. Laterally longitudinal concavity in distal third.</paragraph>
<caption id="39D46640DB09986BFF68D28953760862" httpUri="https://zenodo.org/record/208728/files/figure.png" pageId="4" pageNumber="53" targetBox="[302,1296,206,1706]" targetPageId="4">
<paragraph id="6D1436C8DB09986BFF68D28953760862" blockId="4.[151,1436,1772,1826]" pageId="4" pageNumber="53">
<emphasis id="5FDFEADADB09986BFF68D289531B0842" bold="true" box="[151,274,1772,1795]" pageId="4" pageNumber="53">FIGURE 2.</emphasis>
<taxonomicName id="AAAB4D4BDB09986BFEE7D28850890843" box="[280,640,1773,1795]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB09986BFEE7D28853740843" box="[280,381,1773,1795]" italics="true" pageId="4" pageNumber="53">Cylindera</emphasis>
(
<emphasis id="5FDFEADADB09986BFE74D288501C0843" box="[395,533,1773,1795]" italics="true" pageId="4" pageNumber="53">Plectographa</emphasis>
)
<emphasis id="5FDFEADADB09986BFDDCD28B50890843" box="[547,640,1774,1795]" italics="true" pageId="4" pageNumber="53">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB09986BFD79D28850B60842" bold="true" box="[646,703,1773,1794]" pageId="4" pageNumber="53">
<taxonomicNameLabel id="44EC57A1DB09986BFD79D28850B60842" box="[646,703,1773,1794]" pageId="4" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, habitus dorsal, male, holotype; B, idem, female, allotype; C, habitus lateral, male, paratype.
</paragraph>
</caption>
<paragraph id="6D1436C8DB09986BFF38D30250D708AB" blockId="4.[151,1436,1895,2027]" pageId="4" pageNumber="53">
<emphasis id="5FDFEADADB09986BFF38D30253E608C0" bold="true" box="[199,495,1895,1920]" pageId="4" pageNumber="53">Geographic distribution.</emphasis>
This species is currently known only from the two locations where
<typeStatus id="B210886ADB09986BFB10D30D572308C0" box="[1263,1322,1896,1920]" pageId="4" pageNumber="53">types</typeStatus>
and
<typeStatus id="B210886ADB09986BFA9DD30D52DB08E4" pageId="4" pageNumber="53" type="paratype">paratypes</typeStatus>
were collected, in Tariquia, close to its North-western limit. Several collections (see
<bibRefCitation id="093A4B39DB09986BFB65D3E9577C08E3" author="Pearson" box="[1178,1397,1931,1956]" pageId="4" pageNumber="53" refString="Pearson, D. L., Guerra, J. F. &amp; Brzoska, D. W. (1999) The tiger beetles of Bolivia: Their identification, distribution and natural history. Contributions on Entomology, International, 3, 379 - 523." type="journal article" year="1999">
Pearson
<emphasis id="5FDFEADADB09986BFB04D3E8573C08E4" box="[1275,1333,1932,1956]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
1999
</bibRefCitation>
) in the adjacent Chaco Serrano, the elbow of the Andes and the Bolivian Yungas failed to find this species; therefore we consider it endemic to Tucuman-Bolivian forest.
</paragraph>
<paragraph id="6D1436C8DB08986AFF38D4FD57270FB7" blockId="5.[151,1437,151,644]" pageId="5" pageNumber="54">
<emphasis id="5FDFEADADB08986AFF38D4FD53220FF0" bold="true" box="[199,299,152,176]" pageId="5" pageNumber="54">Ecology.</emphasis>
<taxonomicName id="AAAB4D4BDB08986AFECDD4FD50CB0FF0" box="[306,706,151,176]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB08986AFECDD4FD53AB0FF0" box="[306,418,152,176]" italics="true" pageId="5" pageNumber="54">Cylindera</emphasis>
(
<emphasis id="5FDFEADADB08986AFE4DD4FD50430FF0" box="[434,586,152,176]" italics="true" pageId="5" pageNumber="54">Plectographa</emphasis>
)
<emphasis id="5FDFEADADB08986AFDA4D4FC50CB0FF0" box="[603,706,153,176]" italics="true" pageId="5" pageNumber="54">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB08986AFD36D4FD51090FF0" bold="true" box="[713,768,152,176]" pageId="5" pageNumber="54">
<taxonomicNameLabel id="44EC57A1DB08986AFD36D4FD51090FF0" box="[713,768,152,176]" pageId="5" pageNumber="54" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
was observed during November and December on two narrow sand stone banks of a medium forest stream. The activity of adults was apparently triggered by initial rainfalls of the rainy season, as we observed no individuals at the same sites earlier in the drier season of October.
</paragraph>
<paragraph id="6D1436C8DB08986AFF38D561571A0D7C" blockId="5.[151,1437,151,644]" pageId="5" pageNumber="54">
At midday on sunny days, individuals perched on stones close to a moist, relatively steep sloping wall of sandstone. This microhabitat was temporarily insolated, and shaded, with conspicuous and highly localized patches of liverworts and mosses growing on the sand stonewall. This tiger beetle was absent from shady banks of smaller forest rivers and on sunny banks of larger rivers. We assume that humid sand stone microhabitats lined with trees and with sunny patches are the preferred microhabitat. Such habitats are apparently the result of a specific tectonical setting and were only observed twice in the study area. We collected the specimens of
<taxonomicName id="AAAB4D4BDB08986AFB1BD5DD57950E90" box="[1252,1436,439,464]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB08986AFB1BD5DD56F70E8F" box="[1252,1278,440,463]" italics="true" pageId="5" pageNumber="54">C.</emphasis>
(
<emphasis id="5FDFEADADB08986AFAF0D5DC57280E90" box="[1295,1313,441,464]" italics="true" pageId="5" pageNumber="54">P.</emphasis>
)
<emphasis id="5FDFEADADB08986AFACAD5DC57950E90" box="[1333,1436,441,464]" italics="true" pageId="5" pageNumber="54">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB08986AFF68D5B952C70EB4" bold="true" box="[151,206,476,500]" pageId="5" pageNumber="54">
<taxonomicNameLabel id="44EC57A1DB08986AFF68D5B952C70EB4" box="[151,206,476,500]" pageId="5" pageNumber="54" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
together with
<taxonomicName id="AAAB4D4BDB08986AFE8BD5B851110EB4" box="[372,792,476,500]" class="Insecta" family="Carabidae" genus="Pentacomia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="cribrata" subGenus="Mesacanthina">
<emphasis id="5FDFEADADB08986AFE8BD5B853F30EB4" box="[372,506,477,500]" italics="true" pageId="5" pageNumber="54">Pentacomia</emphasis>
(
<emphasis id="5FDFEADADB08986AFDF3D5B950A20EB4" box="[524,683,476,500]" italics="true" pageId="5" pageNumber="54">Mesacanthina</emphasis>
)
<emphasis id="5FDFEADADB08986AFD43D5B951110EB4" box="[700,792,476,500]" italics="true" pageId="5" pageNumber="54">cribrata</emphasis>
</taxonomicName>
in the same microhabitat. As we approached individuals of
<taxonomicName id="AAAB4D4BDB08986AFF44D66553720D58" box="[187,379,511,536]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB08986AFF44D66552DD0D57" box="[187,212,512,535]" italics="true" pageId="5" pageNumber="54">C.</emphasis>
(
<emphasis id="5FDFEADADB08986AFF15D66452F50D58" box="[234,252,513,536]" italics="true" pageId="5" pageNumber="54">P.</emphasis>
)
<emphasis id="5FDFEADADB08986AFEEBD66453720D58" box="[276,379,513,536]" italics="true" pageId="5" pageNumber="54">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB08986AFE77D66553B60D58" bold="true" box="[392,447,512,536]" pageId="5" pageNumber="54">
<taxonomicNameLabel id="44EC57A1DB08986AFE77D66553B60D58" box="[392,447,512,536]" pageId="5" pageNumber="54" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="AAAB4D4BDB08986AFDFED66451390D58" box="[513,816,511,536]" class="Insecta" family="Carabidae" genus="Pentacomia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="cribrata" subGenus="Mesacanthina">
<emphasis id="5FDFEADADB08986AFDFED664508E0D58" box="[513,647,513,536]" italics="true" pageId="5" pageNumber="54">Pentacomia</emphasis>
(
<emphasis id="5FDFEADADB08986AFD63D66450BA0D58" box="[668,691,513,536]" italics="true" pageId="5" pageNumber="54">M</emphasis>
.)
<emphasis id="5FDFEADADB08986AFD2ED66551390D58" box="[721,816,512,536]" italics="true" pageId="5" pageNumber="54">cribrata</emphasis>
</taxonomicName>
, they quickly flew away from us. Several times we observed
<taxonomicName id="AAAB4D4BDB08986AFEFCD64053B00D7C" box="[259,441,548,572]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB08986AFEFCD64053140D7C" box="[259,285,549,572]" italics="true" pageId="5" pageNumber="54">C.</emphasis>
(
<emphasis id="5FDFEADADB08986AFED2D64053360D7C" box="[301,319,549,572]" italics="true" pageId="5" pageNumber="54">P.</emphasis>
)
<emphasis id="5FDFEADADB08986AFEADD64053B00D7C" box="[338,441,549,572]" italics="true" pageId="5" pageNumber="54">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB08986AFE3FD64153F10D7C" bold="true" box="[448,504,548,572]" pageId="5" pageNumber="54">
<taxonomicNameLabel id="44EC57A1DB08986AFE3FD64153F10D7C" box="[448,504,548,572]" pageId="5" pageNumber="54" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
flying to land on stones lying in the river or to the opposite river bank.
</paragraph>
<paragraph id="6D1436C8DB08986AFF38D62256770DC4" blockId="5.[151,1437,151,644]" pageId="5" pageNumber="54">
Also observed in this habitat of
<taxonomicName id="AAAB4D4BDB08986AFDD4D62D50E90D20" box="[555,736,583,608]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="yaguaree" status="sp. nov." subGenus="Plectographa">
<emphasis id="5FDFEADADB08986AFDD4D62D504C0D1F" box="[555,581,584,607]" italics="true" pageId="5" pageNumber="54">C.</emphasis>
(
<emphasis id="5FDFEADADB08986AFDAAD62C506E0D20" box="[597,615,585,608]" italics="true" pageId="5" pageNumber="54">P.</emphasis>
)
<emphasis id="5FDFEADADB08986AFD86D62C50E90D20" box="[633,736,585,608]" italics="true" pageId="5" pageNumber="54">yaguaree</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB08986AFD18D62D51170D20" bold="true" box="[743,798,584,608]" pageId="5" pageNumber="54">
<taxonomicNameLabel id="44EC57A1DB08986AFD18D62D51170D20" box="[743,798,584,608]" pageId="5" pageNumber="54" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
was
<taxonomicName id="AAAB4D4BDB08986AFCA7D62D56740D20" box="[856,1149,584,608]" class="Insecta" family="Carabidae" genus="Pseudoxycheila" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="tucumana" status="sp. nov.">
<emphasis id="5FDFEADADB08986AFCA7D62D56740D20" box="[856,1149,584,608]" italics="true" pageId="5" pageNumber="54">Pseudoxycheila tucumana</emphasis>
</taxonomicName>
<emphasis id="5FDFEADADB08986AFB7BD62D56B20D20" bold="true" box="[1156,1211,584,608]" pageId="5" pageNumber="54">
<taxonomicNameLabel id="44EC57A1DB08986AFB7BD62D56B20D20" box="[1156,1211,584,608]" pageId="5" pageNumber="54" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
between stones and
<taxonomicName id="AAAB4D4BDB08986AFF68D60953EF0DC4" authority="Fleutiaux" authorityName="Fleutiaux" box="[151,486,620,644]" class="Insecta" family="Carabidae" genus="Oxycheila" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="germaini">
<emphasis id="5FDFEADADB08986AFF68D609537D0DC4" box="[151,372,620,644]" italics="true" pageId="5" pageNumber="54">Oxycheila germaini</emphasis>
Fleutiaux
</taxonomicName>
hiding between and under stones close to the water’s edge.
</paragraph>
</subSubSection>
</treatment>
</document>
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