299 lines
43 KiB
XML
299 lines
43 KiB
XML
<document ID-CLB-Dataset="274274" ID-DOI="http://dx.doi.org/10.3897/jhr.96.105494" ID-GBIF-Dataset="ae19b649-bfd7-45e7-84d1-07980c685878" ID-Pensoft-Pub="1314-2607-96-879" ID-Pensoft-UUID="091C960727095BAD977C9A253E846630" ID-ZooBank="DB845B3662BA4DA183705B36E5916BB0" ModsDocID="1314-2607-96-879" checkinTime="1698104569869" checkinUser="pensoft" docAuthor="UImer, Jonah M., Jansta, Petr, Azar, Dany & Krogmann, Lars" docDate="2023" docId="573B9E1F7C215331BCAEFD011D4A9A03" docLanguage="en" docName="JourHymenoptRes 96: 879-924" docOrigin="Journal of Hymenoptera Research 96" docPubDate="2023-10-23" docSource="http://dx.doi.org/10.3897/jhr.96.105494" docTitle="Protoita bidentata Ulmer & Krogmann 2023, sp. nov." docType="treatment" docUuid="35C89B66-DAA3-4CE6-9DD5-9F9D1716E03B" docUuidSource="ZooBank" docVersion="4" id="091C960727095BAD977C9A253E846630" lastPageNumber="879" masterDocId="091C960727095BAD977C9A253E846630" masterDocTitle="At the dawn of megadiversity - Protoitidae, a new family of Chalcidoidea (Hymenoptera) from Lower Cretaceous Lebanese amber" masterLastPageNumber="924" masterPageNumber="879" pageNumber="879" updateTime="1732659028523" updateUser="ExternalLinkService">
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<mods:titleInfo id="B7320249F1A0ED211BD4590FD7E4C006">
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<mods:title id="7D3FEAA58C05FCDE4DE842367C40FA4B">At the dawn of megadiversity - Protoitidae, a new family of Chalcidoidea (Hymenoptera) from Lower Cretaceous Lebanese amber</mods:title>
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<mods:name id="79A28B9BB1E0D605848AE780DB73F1A2" type="personal">
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<mods:roleTerm id="A5C898466C033C6CB5C22276C57C6283">Author</mods:roleTerm>
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<mods:namePart id="7F34BCA29B4D6C32B50FF349CB802AD0">UImer, Jonah M.</mods:namePart>
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<mods:nameIdentifier id="10EAB1A2F7CAC54049E1967469411AFC" type="ORCID">https://orcid.org/0000-0002-9185-6378</mods:nameIdentifier>
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<mods:affiliation id="FF3CA27AD94516859773F95EA4FA7F32">Department of Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany & Institute of Biology, Biological Systematics (190 w), University of Hohenheim, Stuttgart, Germany</mods:affiliation>
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<mods:nameIdentifier id="80D75B944888D5D4C35D149AF86254E1" type="email">jonah.ulmer@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="B02252CDB12C41C70C6CBBEE3D397437" type="personal">
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<mods:roleTerm id="886760FF9A4F5F283991F520E94A0A4B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B17BECC9CD8076E92850825DEF480F66">Jansta, Petr</mods:namePart>
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<mods:nameIdentifier id="BF47ED10F5FD2D0E11C44D61A083E12C" type="ORCID">https://orcid.org/0000-0001-6409-3603</mods:nameIdentifier>
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<mods:affiliation id="01AAAC9BBD3F30EA877D4A776EE14133">Department of Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany & Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name id="37882C64BE6FDAFC10CB797CD17770C6" type="personal">
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<mods:roleTerm id="7052011CFBE149AE4A374B75AC6D7C7C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D5E69F0F453FE412BE0AE3F8B294E585">Azar, Dany</mods:namePart>
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<mods:affiliation id="A92848E9F4E0B84A48A8EF29283C21D2">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China & Lebanese University, Faculty of Sciences II, Department of Natural Sciences, Fanar, Matn, Lebanon</mods:affiliation>
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</mods:name>
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<mods:name id="3141AC4328E79F1C35D7028AE2604E4E" type="personal">
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</mods:role>
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<mods:namePart id="BFDAF7E593963D522817C2745D2651B3">Krogmann, Lars</mods:namePart>
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<mods:nameIdentifier id="889876C5FA2D191F6E421670A1B91CF1" type="ORCID">https://orcid.org/0000-0002-3724-1735</mods:nameIdentifier>
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<mods:affiliation id="3101C6FAD198CD22335CC5416726314A">Department of Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany & Institute of Biology, Biological Systematics (190 w), University of Hohenheim, Stuttgart, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="AD602D344F7F6117B39832CDF63A0B51">text</mods:typeOfResource>
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<mods:titleInfo id="EF62A60D722A1F9F6A6309912934537E">
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<mods:title id="6592779F67FC479CF50BF24418D7836E">Journal of Hymenoptera Research</mods:title>
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<mods:part id="8F88143D7E487BD04C908C19F7B7CF2F">
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<mods:date id="15D419AC65796AF984EB896881BD20AA">2023</mods:date>
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<mods:number id="73DC289F6DA7283F6502A1A1173F5700">2023-10-23</mods:number>
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<mods:number id="C342548A1705A0C30689B9B30D260592">96</mods:number>
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<mods:start id="86F8FDC67A448F971AB9FA3C892F7F27">879</mods:start>
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<mods:end id="76B86403FB9A3E0269FDEF4B04354FAD">924</mods:end>
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<mods:location id="CF05FDEC55A6A122705C6703C37A81BA">
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<mods:url id="99E83329CE975FD3D9116B5F4AD26EEE">http://dx.doi.org/10.3897/jhr.96.105494</mods:url>
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</mods:location>
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<mods:classification id="A850EB5AF28232F080A12EFC58667473">journal article</mods:classification>
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<mods:identifier id="BAE59270C6F4DA9C0D6CBD7481DD7204" type="DOI">http://dx.doi.org/10.3897/jhr.96.105494</mods:identifier>
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<treatment id="573B9E1F7C215331BCAEFD011D4A9A03" ID-GBIF-Taxon="214236618" LSID="urn:lsid:zoobank.org:act:35C89B66-DAA3-4CE6-9DD5-9F9D1716E03B" httpUri="http://treatment.plazi.org/id/573B9E1F7C215331BCAEFD011D4A9A03" lastPageNumber="879" pageId="0" pageNumber="879" scope_family="Protoitidae" scope_order="Hymenoptera" scope_superFamily="Chalcidoidea">
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<subSubSection id="F2CD99805547395EFDF2D4D0991E8018" pageId="0" pageNumber="879" type="nomenclature">
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<paragraph id="47DEC520D6EE1B40A5F4943F1100F432" pageId="0" pageNumber="879">
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||
<taxonomicName id="738A0BC1815DAE1218B840FAB9EF8D43" LSID="https://zoobank.org/35C89B66-DAA3-4CE6-9DD5-9F9D1716E03B" authority="Ulmer & Krogmann" authorityName="Ulmer & Krogmann" authorityYear="2023" class="Insecta" family="Protoitidae" genus="Protoita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protoita bidentata" order="Hymenoptera" pageId="0" pageNumber="879" phylum="Arthropoda" rank="species" species="bidentata" status="sp. nov.">Protoita bidentata Ulmer & Krogmann</taxonomicName>
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<taxonomicNameLabel id="092D286B91B117B90D23C0DCD23A96AF" pageId="0" pageNumber="879">sp. nov.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="25547A5A29345E499F33E7119DF1EA58" pageId="0" pageNumber="879" type="description">
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<paragraph id="22B8D0879C9F4C10B0ADC88EABDF8EBD" pageId="0" pageNumber="879">
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||
<figureCitation id="EB3161E20B6093D9A53CF6C205ED799E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">Fig. 5</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="57B7ED855ABC5F45A5D1841BFC6095E4" pageId="0" pageNumber="879" type="diagnosis">
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||
<paragraph id="6B0A52D4EF373BF57C5524BF425F9EDA" pageId="0" pageNumber="879">Diagnosis.</paragraph>
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<paragraph id="F670029B0EBBF4B43C25EE079CF830E6" pageId="0" pageNumber="879">
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<taxonomicName id="532B3D1DE22F5405FEA23FCE8EE9E546" authorityName="Ulmer & Krogmann" authorityYear="2023" class="Insecta" family="Protoitidae" genus="Protoita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protoita bidentata" order="Hymenoptera" pageId="0" pageNumber="879" phylum="Arthropoda" rank="species" species="bidentata">
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<emphasis id="01909EF753A055387E552D3897A54D12" italics="true" pageId="0" pageNumber="879">Protoita bidentata</emphasis>
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</taxonomicName>
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, the only species of the genus known from the male, differs from all other species in the genus by the following combination of characters: Body and antenna bicolored (Fig.
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<figureCitation id="8D57B47E82A45013486D9D924EF4A8D7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5A</figureCitation>
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||
). Mandibular formula 2:2 (Fig.
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||
<figureCitation id="E9DB308E6CBF053259B609F6143FA698" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5E</figureCitation>
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:md). Clava 3-segmented (Fig.
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<figureCitation id="1CB0F1889584621B5D21E8C050CC294A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5C</figureCitation>
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). Stigmal vein of fore wing extending at a right angle to wing margin (Fig.
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<figureCitation id="00973ABB434AB7A484BFC3A471B7FD20" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5A</figureCitation>
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). Basitarsal comb of fore leg lacking.
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</paragraph>
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<caption id="A3B83FF0666F4367887376EF9BD01F42" doi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879" start="Figure 5" startId="F5">
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<paragraph id="860CFE8AC8FA3C2610A288431A22DF15" pageId="0" pageNumber="879">
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<emphasis id="621155EF573A7D6DBDAAB8C201DA5C16" bold="true" pageId="0" pageNumber="879">Figure 5.</emphasis>
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<taxonomicName id="90726F6053BDF5FB6A7EB08FD0E3A229" authorityName="Ulmer & Krogmann" authorityYear="2023" class="Insecta" family="Protoitidae" genus="Protoita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protoita bidentata" order="Hymenoptera" pageId="0" pageNumber="879" phylum="Arthropoda" rank="species" species="bidentata">
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<emphasis id="A6DBE9513D76D0619BB25996BCD0BBE0" italics="true" pageId="0" pageNumber="879">Protoita bidentata</emphasis>
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</taxonomicName>
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, holotype
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<emphasis id="504A4182F66980D7EC00F4DC2CA7C5AD" bold="true" pageId="0" pageNumber="879">A</emphasis>
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dorsolateral habitus
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<emphasis id="9DB7105B38CE2F9597C6C98E6A33CE75" bold="true" pageId="0" pageNumber="879">B</emphasis>
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male genitalia, ventral
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<emphasis id="A3B14D70F85E07B39D48C82EDD0A109F" bold="true" pageId="0" pageNumber="879">C</emphasis>
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terminal funicular segments and clava
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<emphasis id="DCE571EF49BB2E64DEA494407183F38B" bold="true" pageId="0" pageNumber="879">D</emphasis>
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mesosoma, dorsolateral
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<emphasis id="18B23ECB06511EE9C13539D12CD5A549" bold="true" pageId="0" pageNumber="879">E</emphasis>
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head, posteroventral. Scale bars: 500
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<normalizedToken id="E918E4D290B971ABE6B06780EBBCF358" originalValue="µm">µm</normalizedToken>
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(
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<emphasis id="67999BD1080C622A5D3D8C302730852F" bold="true" pageId="0" pageNumber="879">A</emphasis>
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); 100
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<normalizedToken id="5C92944A4CEBD3CD1A7A53A08FC1F81A" originalValue="µm">µm</normalizedToken>
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||
(
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<emphasis id="48E920ACBB33ED282F1ABD5B3D8C9FB7" bold="true" pageId="0" pageNumber="879">C-E</emphasis>
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||
).
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<emphasis id="3221E8EA2DD4911E3F0F7F5B604C239A" bold="true" pageId="0" pageNumber="879">aed</emphasis>
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= aedeagus;
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<emphasis id="6B9F966F798C94958BFC27112D00E88F" bold="true" pageId="0" pageNumber="879">dig</emphasis>
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= digitus;
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<emphasis id="4992AFEF819080CF0475F63A32EE183A" bold="true" pageId="0" pageNumber="879">frn</emphasis>
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= frenum;
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<emphasis id="FB56DB5A844A6B4B811AE8AADB31CA18" bold="true" pageId="0" pageNumber="879">hyc</emphasis>
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= hypostomal carina;
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||
<emphasis id="86A3CE53ECE58744022C1EFA6E93EB80" bold="true" pageId="0" pageNumber="879">lbp</emphasis>
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= labial palp;
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||
<emphasis id="073D311F26A59766E9AD47381C751D0C" bold="true" pageId="0" pageNumber="879">md</emphasis>
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||
= mandible;
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||
<emphasis id="F7031AA0F0E7EC290A91324F375A46F1" bold="true" pageId="0" pageNumber="879">msp</emphasis>
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||
= mesothoracic spiracle;
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||
<emphasis id="159F3E4DD813F7882CD81B3215018629" bold="true" pageId="0" pageNumber="879">Mtn</emphasis>
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||
= metasomal tergum;
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||
<emphasis id="82205F3704FA22F8158602FD2110BC76" bold="true" pageId="0" pageNumber="879">mxp</emphasis>
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= maxillary palp;
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<emphasis id="BC471D99895080E838D3EBA7BABE7D8B" bold="true" pageId="0" pageNumber="879">no3</emphasis>
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||
= metanotum;
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<emphasis id="8E5FBE6DAD492727FC18A8EE670E4BFD" bold="true" pageId="0" pageNumber="879">par</emphasis>
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= paramere;
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||
<emphasis id="53CF1F1687827BE6D440758EBA7218C0" bold="true" pageId="0" pageNumber="879">psp</emphasis>
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= propodeal spiracle;
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||
<emphasis id="ABD0A11355C8835C9CCE79011DF5C04B" bold="true" pageId="0" pageNumber="879">tsa</emphasis>
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||
= transcutal articulation;
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||
<emphasis id="6AF07C49A261481C3ABBF0443D42D0AC" bold="true" pageId="0" pageNumber="879">vol</emphasis>
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||
= volsella.
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||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="8950905E3ED7CCAAFDB7BA717C2D3C26" pageId="0" pageNumber="879" type="description">
|
||
<paragraph id="DEBC8027828890F9399EA0224B82BC2A" pageId="0" pageNumber="879">Description.</paragraph>
|
||
<paragraph id="B91C9C34086A9AC24122CAE3964B90B5" pageId="0" pageNumber="879">
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<emphasis id="BCCEE3F05CC51BE341F7883FC9DBF871" bold="true" pageId="0" pageNumber="879">Male.</emphasis>
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Body length 855. Head, scape, pedicel and fu1-fu4 brown. Mesosoma, metasoma and fu5 to tip of clava dark brown. Legs pale. Sculpture on head, meso- and metasoma alutaceous. Wings hyaline, uniformly pilose, damaged beyond postmarginal vein (Fig.
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<figureCitation id="09E52EDEA8D4463E8210A2172C3B3626" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5A</figureCitation>
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||
).
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<emphasis id="547361F56F8F98D124B04C574DCC3843" bold="true" italics="true" pageId="0" pageNumber="879">Head</emphasis>
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strongly transverse, 2
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<normalizedToken id="14CF86D04EB1463DFA758F9ADCC9ED1B" originalValue="×">x</normalizedToken>
|
||
as wide as long in dorsal view (360/179). Temples very short, about 0.15
|
||
<normalizedToken id="98541D6D531E48CD8A73BEFDA7029D48" originalValue="×">x</normalizedToken>
|
||
as long as eye length. Lower tentorial bridge wide, about as wide as long; postgenal bridge absent; hypostomal carina complete (Fig.
|
||
<figureCitation id="1BC0BF40F7801E766DEF93CF4672B2D5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5E</figureCitation>
|
||
:hyc). Occipital carina present (Fig.
|
||
<figureCitation id="04BD919D58DDC3CBAC8301FD90FAE379" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5E</figureCitation>
|
||
). Antennal scrobe slightly depressed. Interantennal region convex. Antennal insertion closer to inner eye margin than each other (Fig.
|
||
<figureCitation id="4DA2CD1A87F23AEB94A97EE436ACEB88" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5A</figureCitation>
|
||
). Clypeus not visible. Dentition 2:2, extending out from facial plane (Fig.
|
||
<figureCitation id="DE11BADF57E403F2774EECF95EB47094" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5E</figureCitation>
|
||
:md). Maxillary palps 3-segmented (Fig.
|
||
<figureCitation id="65DC2381F4A4C1D87367FEF9FD950B5A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5E</figureCitation>
|
||
:mxp). Labial palps 2-segmented (Fig.
|
||
<figureCitation id="C82D526E3B25F444E0F698290643CA30" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5E</figureCitation>
|
||
:lbp). Maxilla wider than labium.
|
||
<emphasis id="E0019575397AC129BF40230C6A4EC525" bold="true" italics="true" pageId="0" pageNumber="879">Antenna</emphasis>
|
||
length 533. Pedicel and scape partly damaged. Fu1 about or slightly less as wide as pedicel, other flagellomeres gradually widening distally, fu1-6 not less than 2
|
||
<normalizedToken id="474043082D3611A3A25A1C5D38E4FC50" originalValue="×">x</normalizedToken>
|
||
as long as wide, fu7-9 less than 2
|
||
<normalizedToken id="CCF6B64150978913E17AC54278F66C82" originalValue="×">x</normalizedToken>
|
||
as long as wide. Clava 3-segmented, clava length 85 (Fig.
|
||
<figureCitation id="DE59A0330D4C45437209A94E9818FD9C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5C</figureCitation>
|
||
). MPS present on all segments including terminal segment; MPS extend beyond the distal edge of the segment. Claval segments equilateral and closely associated relative to funicle segments; clava without micropilosity (Fig.
|
||
<figureCitation id="606E0F3073AEFF5A00A29C83FCDFEFC4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5C</figureCitation>
|
||
).
|
||
<emphasis id="9DBE564F0CD1D5BE5CB766E1CFCFB00C" bold="true" italics="true" pageId="0" pageNumber="879">Mesosoma</emphasis>
|
||
length 346. Transcutal articulation present as weak line of separation (Fig.
|
||
<figureCitation id="DB7487B1AA95D3A056F256B2EBAAC8E4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5D</figureCitation>
|
||
:tsa). Mesopleuron large, partially obscured by bubble (Fig.
|
||
<figureCitation id="58C464AA3C834030D3DA47DE42604CE4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5D</figureCitation>
|
||
:pl2); separated from metapleuron by suture. Axillae not pronounced; axillulae slightly flanged (Fig.
|
||
<figureCitation id="91E6277C51072223C0E560857A57EF02" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5D</figureCitation>
|
||
). Mesonotum length 224. Mesoscutellum convex, length 44, 2
|
||
<normalizedToken id="A9C11F8E61FADA667ABDE5367898753A" originalValue="×">x</normalizedToken>
|
||
as long as wide (Fig.
|
||
<figureCitation id="418E04F0D88E079B36553EB72A776939" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5D</figureCitation>
|
||
:msct). Metanotum narrow, band-like, width 21 (Fig.
|
||
<figureCitation id="B16230D0639AC2F54C3829181E6D74BB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5D</figureCitation>
|
||
:no3). Propodeum partially obscured by abdomen, relatively short, as long as or slightly longer than mesoscutellum (Fig.
|
||
<figureCitation id="59AA027068F4410C20D703F83DFF924E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5D</figureCitation>
|
||
:prp); propodeal spiracle visible, oval, posteriorly angled, closer to anterior margin of propodeum than its diameter (Fig.
|
||
<figureCitation id="B9CC8B7B0C004F0841447C6FE72DE9B5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5D</figureCitation>
|
||
:psp).
|
||
<emphasis id="E40DFC0ABCCB04C1EB1014B7F22D1232" bold="true" italics="true" pageId="0" pageNumber="879">Wings.</emphasis>
|
||
Fore wing damaged beyond midline. Fore wing width 323. Longest marginal seta length 46. Submarginal vein length 379. Marginal vein narrow, length 81, width 24. Cubital vein narrow, weakly pigmented. Basal vein sclerotized, length 60. Stigmal vein straight, 90° relative to dorsal wing margin, stigmal vein length 44. Uncus present with ephemeral line extending distally. Postmarginal vein length 53. Hind wing length 440, width 28. Longest marginal seta of hind wing 35 (Fig.
|
||
<figureCitation id="23FA57C30DCC7809EF81C478A272BE9B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5A</figureCitation>
|
||
).
|
||
<emphasis id="7B77169AAFF8E54767F434A850D9EACD" bold="true" italics="true" pageId="0" pageNumber="879">Legs.</emphasis>
|
||
Protibial calcar long, curved, apically bifurcate. Basitarsus of fore leg ventrally curved; no basitarsal comb.
|
||
<emphasis id="5AC2927A820F57E4395A349E7508E7D8" bold="true" italics="true" pageId="0" pageNumber="879">Metasoma</emphasis>
|
||
length 313. Metasoma broadly associated with mesosoma; petiole obscured (Fig.
|
||
<figureCitation id="990D4F1A15F78FD373CC911397384EFD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5A</figureCitation>
|
||
). Mt2-9 visible; all equal in length except Mt2, dorsally slightly shorter. 3 sternal segments visible, each about 2
|
||
<normalizedToken id="5806BF0C724DD8DCADF99F90623FAA25" originalValue="×">x</normalizedToken>
|
||
as long as tergal segment average length. Cerci between Mt8 and Mt9 digitiform, about 2
|
||
<normalizedToken id="9BF6C792A64D6C4141F345B16EAF7BFE" originalValue="×">x</normalizedToken>
|
||
as long as digitiform base.
|
||
<emphasis id="790A3C202A51C275B1434445DAA84132" bold="true" italics="true" pageId="0" pageNumber="879">Genitalia</emphasis>
|
||
length externally 97. Genital capsule externally protruding. Paramere present; single apical parameral setae present. Intervosellar process broad, extends equally in length to aedeagus. Aedeagus broad with 2 sets of sensillae (Fig.
|
||
<figureCitation id="0023E0CB4FB1AF214BAB07FB48478580" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5B</figureCitation>
|
||
:aed). Digitus with 2 digital processes (Fig.
|
||
<figureCitation id="2138464B3D4E3EB0A55FF0A93C98A515" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Protoita bidentata, holotype A dorsolateral habitus B male genitalia, ventral C terminal funicular segments and clava D mesosoma, dorsolateral E head, posteroventral. Scale bars: 500 µm (A); 100 µm (C-E). aed = aedeagus; dig = digitus; frn = frenum; hyc = hypostomal carina; lbp = labial palp; md = mandible; msp = mesothoracic spiracle; Mtn = metasomal tergum; mxp = maxillary palp; no 3 = metanotum; par = paramere; psp = propodeal spiracle; tsa = transcutal articulation; vol = volsella." figureDoi="10.3897/jhr.96.105494.figure5" httpUri="https://binary.pensoft.net/fig/925399" pageId="0" pageNumber="879">5B</figureCitation>
|
||
:dig).
|
||
</paragraph>
|
||
<paragraph id="9445D39DA85240A460B89DE56C2956FF" pageId="0" pageNumber="879">
|
||
<emphasis id="34D5DA92A75A9C8A0F3BFEB24A22F4BF" bold="true" pageId="0" pageNumber="879">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F80B235C4DD366DB098383000F3754FD" pageId="0" pageNumber="879" type="materials_examined">
|
||
<paragraph id="DC946AB5500DAADED90A9BC0F7D23F14" pageId="0" pageNumber="879">Specimens examined.</paragraph>
|
||
<paragraph id="2E6BD18DA292EA3A85C16254C7ACCAD1" pageId="0" pageNumber="879">
|
||
<materialsCitation id="6599E2018F790449471135DDB5E58D86" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Lebanon" county="Lebanese University" location="Hammana" municipality="Natural History Museum" specimenCount="1" typeStatus="Holotype">
|
||
<emphasis id="302C7F87F786F76409B51CEBA74532CB" bold="true" italics="true" pageId="0" pageNumber="879">
|
||
<typeStatus id="CC54CAA7C1F45E897255814E71518D98">Holotype</typeStatus>
|
||
</emphasis>
|
||
: male,
|
||
<location id="384ECE0FDF160EAC7EB339E4C28BB5F0" LSID="urn:lsid:plazi:treatment:573B9E1F7C215331BCAEFD011D4A9A03:384ECE0FDF160EAC7EB339E4C28BB5F0" country="Lebanon" county="Lebanese University" municipality="Natural History Museum" name="Hammana">Hammana</location>
|
||
/
|
||
<location id="8C4D71853642CF232284331D489F158D" LSID="urn:lsid:plazi:treatment:573B9E1F7C215331BCAEFD011D4A9A03:8C4D71853642CF232284331D489F158D" country="Lebanon" county="Lebanese University" municipality="Natural History Museum" name="Mdeyrij">Mdeyrij</location>
|
||
,
|
||
<location id="7E4849513544240FB4E74959EAB3F5F6" LSID="urn:lsid:plazi:treatment:573B9E1F7C215331BCAEFD011D4A9A03:7E4849513544240FB4E74959EAB3F5F6" country="Lebanon" county="Lebanese University" municipality="Natural History Museum" name="Caza Baabda">Caza Baabda</location>
|
||
,
|
||
<location id="014BCBBFDE66F147A5942631A3D6FD3C" LSID="urn:lsid:plazi:treatment:573B9E1F7C215331BCAEFD011D4A9A03:014BCBBFDE66F147A5942631A3D6FD3C" country="Lebanon" county="Lebanese University" municipality="Natural History Museum" name="Mouhafazet Mount">Mouhafazet Mount</location>
|
||
<collectingCountry id="9AC335979C42A2D7045A88CB910F9806" name="Lebanon">Lebanon</collectingCountry>
|
||
;
|
||
<location id="80CFC9757E11E9A34A848A77E45F1D5D" LSID="urn:lsid:plazi:treatment:573B9E1F7C215331BCAEFD011D4A9A03:80CFC9757E11E9A34A848A77E45F1D5D" country="Lebanon" county="Lebanese University" municipality="Natural History Museum" name="lower Barremian. In">lower Barremian. In</location>
|
||
amber mounted in
|
||
<collectingCountry id="E8C50185F300EF6EAEB48BE15CDC3843" name="Canada">Canada</collectingCountry>
|
||
Balsam. Deposited at
|
||
<collectingMunicipality id="018CDB3B75D8A071E873C5BBF7B4209F">Natural History Museum</collectingMunicipality>
|
||
of the
|
||
<collectingCounty id="0284F2B5E2E04ACB78FA2630792387D0">Lebanese University</collectingCounty>
|
||
, accession/specimen number: 182.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B81613346C84BB50EDFA9F495B67B43C" pageId="0" pageNumber="879" type="type condition">
|
||
<paragraph id="26BE587F7D717B7C3C509DACF49AC67C" pageId="0" pageNumber="879">Type condition.</paragraph>
|
||
<paragraph id="3BB636B2B87A5FC7DDDAF3C501D509A5" pageId="0" pageNumber="879">Specimen with distal ⅓ of its wings missing and basal antennomeres damaged.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5AEFC803ACC8A44E43540BE54096C26" pageId="0" pageNumber="879" type="etymology">
|
||
<paragraph id="AC7A431EB3B54ADB7A3971161192D423" pageId="0" pageNumber="879">Etymology.</paragraph>
|
||
<paragraph id="10AA686A89671C33133C445E9B6CA992" pageId="0" pageNumber="879">The specific epithet is derived from the mandibular formula of the species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52E958002E931E05D7B67CE97C098E82" pageId="0" pageNumber="879" type="notes">
|
||
<paragraph id="AA2760DBF2C528546AD26F390F6F660C" pageId="0" pageNumber="879">Notes.</paragraph>
|
||
<paragraph id="A2A927651CD4CF2B4BB6DD192D6B8AC2" pageId="0" pageNumber="879">
|
||
<taxonomicName id="5A7C50EC6FFA451FD0A02223995EA766" authorityName="Ulmer & Krogmann" authorityYear="2023" class="Insecta" family="Protoitidae" genus="Protoita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protoita bidentata" order="Hymenoptera" pageId="0" pageNumber="879" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="7B6A1E100BC732E2E48FC6B4A0009DB6" italics="true" pageId="0" pageNumber="879">Protoita bidentata</emphasis>
|
||
</taxonomicName>
|
||
is the only species in the genus described from a male. While it is the only taxon with two mandibular teeth, sexual dimorphism in this character is very rare in
|
||
<taxonomicName id="56B54B622C70B8FB9ACB8D54F0F7BBF3" authorityName="Ulmer & Krogmann" authorityYear="2023" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="879" phylum="Arthropoda" rank="superFamily" superFamily="Chalcidoidea">Chalcidoidea</taxonomicName>
|
||
, likely the female of
|
||
<taxonomicName id="2E768E96B876A399AEE5F0B8685CE97A" lsidName="P. bidentata" pageId="0" pageNumber="879" rank="species" species="bidentata">
|
||
<emphasis id="09FA7EB3C5C3A3D15F5F9B211927C558" italics="true" pageId="0" pageNumber="879">P. bidentata</emphasis>
|
||
</taxonomicName>
|
||
is also bidentate. Unlike dentition, antennal shape and claval segments are quite often sexually dimorphic in extant chalcids, and in
|
||
<taxonomicName id="248ABF9A3D9911BAB34E342EB9C92A25" authorityName="Ulmer & Krogmann" authorityYear="2023" class="Insecta" family="Scelionidae" genus="Cretaxenomerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretaxenomerus curvus" order="Hymenoptera" pageId="0" pageNumber="879" phylum="Arthropoda" rank="species" species="curvus">
|
||
<emphasis id="36B759C64F0C27E545B1F734D749E810" italics="true" pageId="0" pageNumber="879">Cretaxenomerus curvus</emphasis>
|
||
</taxonomicName>
|
||
where the male is known we can see a drastic variation in flagellar shape and claval number.
|
||
<taxonomicName id="912B0A6E93DE25887F7823130AC191D2" lsidName="P. bidentata" pageId="0" pageNumber="879" rank="species" species="bidentata">
|
||
<emphasis id="892D589A1B7A5DCF0A107ECBCFEB199C" italics="true" pageId="0" pageNumber="879">P. bidentata</emphasis>
|
||
</taxonomicName>
|
||
is the only species known within
|
||
<taxonomicName id="07620C05348FE6078F6619D91A20EA84" authorityName="Ulmer & Krogmann" authorityYear="2023" class="Insecta" family="Protoitidae" genus="Protoita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protoita" order="Hymenoptera" pageId="0" pageNumber="879" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F52F272F4532A1B3C702533B04684F37" italics="true" pageId="0" pageNumber="879">Protoita</emphasis>
|
||
</taxonomicName>
|
||
with 3 claval segments, and within all of
|
||
<taxonomicName id="7CF8AB7ED13C6E4BB83DE10B3E02BD27" family="Protoitidae" lsidName="" pageId="0" pageNumber="879" rank="family">Protoitidae</taxonomicName>
|
||
only
|
||
<taxonomicName id="B4BD714B58CA67FFA981107318CFFC6D" lsidName="C. deangelis" pageId="0" pageNumber="879" rank="species" species="deangelis">
|
||
<emphasis id="25E3A6B98CFE53BA6E3D57A3AF343810" italics="true" pageId="0" pageNumber="879">C. deangelis</emphasis>
|
||
</taxonomicName>
|
||
is also known to have 3 claval segments however that species is described from a female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |