458 lines
38 KiB
XML
458 lines
38 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.63.49054" ID-GBIF-Dataset="2eec87e8-e8cc-4585-84bb-ac5af39810d1" ID-PMC="PMC7062851" ID-Pensoft-Pub="1314-4049-63-69" ID-Pensoft-UUID="BC2096E6D63D5F5DA23CDFDDD3D63FA9" ID-PubMed="32189978" ModsDocID="1314-4049-63-69" checkinTime="1583231436794" checkinUser="pensoft" docAuthor="Jaklitsch, Walter M. & Voglmayr, Hermann" docDate="2020" docId="3D3B8B2D87A757C09C666025066F1EC7" docLanguage="en" docName="MycoKeys 63: 69-117" docOrigin="MycoKeys 63" docSource="http://dx.doi.org/10.3897/mycokeys.63.49054" docTitle="Melanconis marginalis subsp. europaea Jaklitsch & Voglmayr 2020, subsp. nov." docType="treatment" docVersion="4" id="BC2096E6D63D5F5DA23CDFDDD3D63FA9" lastPageNumber="69" masterDocId="BC2096E6D63D5F5DA23CDFDDD3D63FA9" masterDocTitle="The genus Melanconis (Diaporthales)" masterLastPageNumber="117" masterPageNumber="69" pageNumber="69" updateTime="1668136401704" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Melanconis (Diaporthales)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jaklitsch, Walter M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Voglmayr, Hermann</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>63</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>117</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.63.49054</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.63.49054</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-63-69</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BC2096E6D63D5F5DA23CDFDDD3D63FA9</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="162381290" LSID="urn:lsid:plazi:treatment:3D3B8B2D87A757C09C666025066F1EC7" httpUri="http://treatment.plazi.org/id/3D3B8B2D87A757C09C666025066F1EC7" lastPageNumber="69" pageId="0" pageNumber="69">
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<subSubSection pageId="0" pageNumber="69" type="nomenclature">
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName LSID="3D3B8B2D-87A7-57C0-9C66-6025066F1EC7" authority="Jaklitsch & Voglmayr" authorityName="Jaklitsch & Voglmayr" authorityYear="2020" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconis marginalis subsp. europaea" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="subSpecies" species="marginalis" status="subsp. nov." subSpecies="europaea">Melanconis marginalis subsp. europaea Jaklitsch & Voglmayr</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="69">subsp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Melanconis marginalis subsp. europaea. Sexual morph a pseudostroma in face view b, c ectostromatic discs d subglobose visible part of ostiolar necks e, f cross sections (e showing central column and marginal ostioles f showing central column and perithecia) g vertical section showing central column and two perithecia h-p asci q-y ascospores m-p in aqueous Congo Red a WU 31890 = MAV 1 b-g, j, n, q, s, t, w-y holotype WU 31888 = MAI h, i, m WU 37045 = D 158 k, r WU 36699 l, p WU 31172 o WU 29888 u WU 31889 = MAV v WU 38243. Scale bars: 1 mm (a, f), 500 µm (b, c, e, g), 150 µm (d), 10 µm (h-q, t), 7 µm (r, s, u-y)." figureDoi="10.3897/mycokeys.63.49054.figure6" httpUri="https://binary.pensoft.net/fig/384816" pageId="0" pageNumber="69">Figures 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Melanconis marginalis subsp. europaea. Asexual morph a, b conidiomata and conidial deposits in face view c conidioma with β-conidia in cross section d conidioma with α-conidia in vertical section e-h conidiophores and conidiogenous cells (producing α-conidia in e, f, β-conidia in g, h) i-p α-conidia q-t β-conidia e-t in 3 % KOH a, b, d-f, i-k, q-s WU 37044 = D 157 c, g, h, l, t WU 31893 m WU 31891 = W. J. 1542 n WU 31888 = MAI o, p WU 31889 = MAV. Scale bars: 500 µm (a-d), 5 µm (e-t)." figureDoi="10.3897/mycokeys.63.49054.figure7" httpUri="https://binary.pensoft.net/fig/384817" pageId="0" pageNumber="69">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="diagnosis">
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<paragraph pageId="0" pageNumber="69">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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This subspecies of
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<taxonomicName authorityName="Wehm., Pap. Michigan Acad. I. 6: 382" authorityYear="1926" baseAuthorityName="Peck" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconis marginalis" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="species" species="marginalis">
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<emphasis italics="true" pageId="0" pageNumber="69">Melanconis marginalis</emphasis>
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</taxonomicName>
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occurs in Europe and differs from the American subsp.
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<taxonomicName authorityName="subsp. marginalis (Peck) Wehm., Pap. Michigan Acad. I. 6: 382" authorityYear="1926" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="subsp. marginalis" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="subSpecies" species="marginalis" subSpecies="marginalis">marginalis</taxonomicName>
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phylogenetically and by slightly larger asci, ascospores and ascospore appendages.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="type material">
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<paragraph pageId="0" pageNumber="69">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">Holotype</emphasis>
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.
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</emphasis>
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Austria, Steiermark, Judenburg, Pusterwald, Hinterwinkel, grid square 8651/4, on
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<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus incana" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="incana">
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<emphasis italics="true" pageId="0" pageNumber="69">Alnus incana</emphasis>
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</taxonomicName>
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, 11 Jun 2011, H. Voglmayr (WU 31888, culture CBS 131692 = MAI).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="etymology">
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<paragraph pageId="0" pageNumber="69">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="69">For its occurrence in Europe.</paragraph>
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<caption doi="10.3897/mycokeys.63.49054.figure6" httpUri="https://binary.pensoft.net/fig/384816" pageId="0" pageNumber="69" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">Figure 6.</emphasis>
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<taxonomicName authorityName="Jaklitsch & Voglmayr" authorityYear="2020" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconis marginalis subsp. europaea" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="subSpecies" species="marginalis" subSpecies="europaea">Melanconis marginalis subsp. europaea</taxonomicName>
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. Sexual morph
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<emphasis bold="true" pageId="0" pageNumber="69">a</emphasis>
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pseudostroma in face view
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<emphasis bold="true" pageId="0" pageNumber="69">b, c</emphasis>
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ectostromatic discs
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<emphasis bold="true" pageId="0" pageNumber="69">d</emphasis>
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subglobose visible part of ostiolar necks
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<emphasis bold="true" pageId="0" pageNumber="69">e, f</emphasis>
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cross sections (
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<emphasis bold="true" pageId="0" pageNumber="69">e</emphasis>
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showing central column and marginal ostioles
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<emphasis bold="true" pageId="0" pageNumber="69">f</emphasis>
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showing central column and perithecia)
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<emphasis bold="true" pageId="0" pageNumber="69">g</emphasis>
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vertical section showing central column and two perithecia
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<emphasis bold="true" pageId="0" pageNumber="69">h-p</emphasis>
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asci
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<emphasis bold="true" pageId="0" pageNumber="69">q-y</emphasis>
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ascospores
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<emphasis bold="true" pageId="0" pageNumber="69">m-p</emphasis>
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in aqueous Congo Red
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<emphasis bold="true" pageId="0" pageNumber="69">a</emphasis>
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WU 31890 = MAV1
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<emphasis bold="true" pageId="0" pageNumber="69">b-g, j, n, q, s, t, w-y</emphasis>
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holotype WU 31888 = MAI
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<emphasis bold="true" pageId="0" pageNumber="69">h, i, m</emphasis>
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WU 37045 = D158
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<emphasis bold="true" pageId="0" pageNumber="69">k, r</emphasis>
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WU 36699
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<emphasis bold="true" pageId="0" pageNumber="69">l, p</emphasis>
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WU 31172
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<emphasis bold="true" pageId="0" pageNumber="69">o</emphasis>
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WU 29888
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<emphasis bold="true" pageId="0" pageNumber="69">u</emphasis>
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WU 31889 = MAV
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<emphasis bold="true" pageId="0" pageNumber="69">v</emphasis>
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WU 38243. Scale bars: 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="69">a, f</emphasis>
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), 500
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="69">b, c, e, g</emphasis>
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), 150
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="69">d</emphasis>
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), 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="69">h-q, t</emphasis>
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), 7
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="69">r, s, u-y</emphasis>
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).
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</paragraph>
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</caption>
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<caption doi="10.3897/mycokeys.63.49054.figure7" httpUri="https://binary.pensoft.net/fig/384817" pageId="0" pageNumber="69" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">Figure 7.</emphasis>
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<taxonomicName authorityName="Jaklitsch & Voglmayr" authorityYear="2020" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconis marginalis subsp. europaea" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="subSpecies" species="marginalis" subSpecies="europaea">Melanconis marginalis subsp. europaea</taxonomicName>
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. Asexual morph
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<emphasis bold="true" pageId="0" pageNumber="69">a, b</emphasis>
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conidiomata and conidial deposits in face view
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<emphasis bold="true" pageId="0" pageNumber="69">c</emphasis>
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conidioma with
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<normalizedToken originalValue="β-conidia">β-conidia</normalizedToken>
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in cross section
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<emphasis bold="true" pageId="0" pageNumber="69">d</emphasis>
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conidioma with
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<normalizedToken originalValue="α-conidia">α-conidia</normalizedToken>
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in vertical section
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<emphasis bold="true" pageId="0" pageNumber="69">e-h</emphasis>
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conidiophores and conidiogenous cells (producing
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<normalizedToken originalValue="α-conidia">α-conidia</normalizedToken>
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in
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<emphasis bold="true" pageId="0" pageNumber="69">e, f</emphasis>
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,
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<normalizedToken originalValue="β-conidia">β-conidia</normalizedToken>
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in
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<emphasis bold="true" pageId="0" pageNumber="69">g, h</emphasis>
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)
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<emphasis bold="true" pageId="0" pageNumber="69">i-p</emphasis>
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<normalizedToken originalValue="α-conidia">α-conidia</normalizedToken>
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<emphasis bold="true" pageId="0" pageNumber="69">q-t</emphasis>
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<normalizedToken originalValue="β-conidia">β-conidia</normalizedToken>
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<emphasis bold="true" pageId="0" pageNumber="69">e-t</emphasis>
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in 3% KOH
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<emphasis bold="true" pageId="0" pageNumber="69">a, b, d-f, i-k, q-s</emphasis>
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WU 37044 = D157
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<emphasis bold="true" pageId="0" pageNumber="69">c, g, h, l, t</emphasis>
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WU 31893
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<emphasis bold="true" pageId="0" pageNumber="69">m</emphasis>
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WU 31891 = W.J. 1542
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<emphasis bold="true" pageId="0" pageNumber="69">n</emphasis>
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WU 31888 = MAI
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<emphasis bold="true" pageId="0" pageNumber="69">o, p</emphasis>
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WU 31889 = MAV. Scale bars: 500
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="69">a-d</emphasis>
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), 5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="69">e-t</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="description">
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<paragraph pageId="0" pageNumber="69">Description.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Sexual morph</emphasis>
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:
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<emphasis italics="true" pageId="0" pageNumber="69">Pseudostromata</emphasis>
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1.5-3.6 mm diam., usually conspicuous and numerous, scattered to tightly aggregated, forming pustules, pulvinate, circular to elliptical in outline, typically elevated beyond bark surface; consisting of an ectostromatic disc and perithecia embedded in an entostroma around a central column.
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<emphasis italics="true" pageId="0" pageNumber="69">Ectostromatic discs</emphasis>
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0.5-2.1 diam. or long, discrete, less commonly confluent, bright white to yellowish, turning brownish with age, variable, fusoid, elliptic or circular in outline, flat, convex, concave, entire or coarsely fissured and crumbly, projecting up to 1 mm including projecting part of the pseudostroma; central column beneath disc whitish to yellowish, consisting of hyaline hyphae and colourless crystals.
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<emphasis italics="true" pageId="0" pageNumber="69">Ostiolar necks</emphasis>
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cylindrical, laterally attached on perithecia, centrally attached only on centrally arranged perithecia, convergent in the disc margin or crowded at the ends of fusoid discs, 1-25(-35) per disc. Visible part of the ostiolar necks (53-)103-167(-212)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(n = 90) diam., black or brown with black tips, usually circular in section, sometimes plane with the disc, but much more frequently papillate and projecting to 250
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, often resembling minute perithecia with pointed tips or discoid with depressed centre to nearly ring-like, sometimes conical to bristle-like and projecting to 0.4 mm.
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<emphasis italics="true" pageId="0" pageNumber="69">Entostroma</emphasis>
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bark coloured, not or only slightly paler than the surrounding bark, consisting of bark cells and some light-coloured hyphae.
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<emphasis italics="true" pageId="0" pageNumber="69">Perithecia</emphasis>
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(450-)515-680(-810)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(n = 58) diam., arranged in valsoid configuration around and below central column, globose to subglobose, collapsing up- or laterally inwards upon drying. Peridium pseudoparenchymatous, consisting of a dark brown small-celled outer and a hyaline to brownish, large-celled inner layer.
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<emphasis italics="true" pageId="0" pageNumber="69">Hamathecium</emphasis>
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of broad multiguttulate paraphyses, collapsing, dissolving and usually absent amongst mature asci.
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<emphasis italics="true" pageId="0" pageNumber="69">Asci</emphasis>
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floating free at maturity, (52-)68-85(-98)
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<normalizedToken originalValue="×">x</normalizedToken>
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(8.7-)10.5-15.5(-18.7)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(n = 126), narrowly fusoid to oblong or narrowly ellipsoid, with an apical ring distinct in water and staining in Congo Red, but invisible or indistinct in 3% KOH, containing 8 biseriate or obliquely uniseriate ascospores.
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<emphasis italics="true" pageId="0" pageNumber="69">Ascospores</emphasis>
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(13.8-)17-20(-22.8)
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<normalizedToken originalValue="×">x</normalizedToken>
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(3.5-)4.7-6.5(-7.7)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, l/w (2.5-)2.9-3.8(-5.5) (n = 242), hyaline, mostly oblong or narrowly ellipsoid, sometimes broadly ellipsoid upon release, symmetric or inequilateral, bicellular with nearly equal cells, slightly or strongly constricted at the median septum, multiguttulate or with few large and several small guttules when fresh, with a short and broad, rounded, sometimes tapering, angular or bell-shaped and typically terminally truncate appendage (1.8-)2.7-4.7(-8.4)
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<normalizedToken originalValue="×">x</normalizedToken>
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(2-)2.5-4(-5.5)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, l/w (0.4-)0.9-1.5(-2.8) (n = 318), at one or both ends becoming invisible in 3% KOH and Congo Red after release.
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</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Asexual morph</emphasis>
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acervular, intermingled with pseudostromata of the sexual morph or more frequently developing separately, usually inconspicuous, but sometimes becoming conspicuous due to greyish-brown to dark brown conidial deposits 0.2-0.6 mm diam., rarely confluent from 2 conidiomata and then up to more than 1 cm long. First, white to yellow tissue (central column) forming within the bark, becoming visible by pustulate bark and narrow whitish to yellowish or brownish slit-like discs emerging through bark cracks, usually first followed by the production of
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<normalizedToken originalValue="β-conidia">β-conidia</normalizedToken>
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in olivaceous chambers and later
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<normalizedToken originalValue="α-conidia">α-conidia</normalizedToken>
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on the same or similar conidiophores turning the contents brown and oozing out from ends of the discs or perithecia of the sexual morph formed below the acervulus.
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<emphasis italics="true" pageId="0" pageNumber="69">Conidiomata</emphasis>
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1-2 mm diam., pulvinate, more or less circular in outline, scattered or aggregated in lines.
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<emphasis italics="true" pageId="0" pageNumber="69">Covering discs</emphasis>
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0.3-0.9(-1.6) mm (n = 45) long, narrowly fusoid or longish to rounded, plane to convex, becoming covered and obscured by conidial deposits; discs and pulvinate or conical columns beneath, consisting of compact
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<emphasis italics="true" pageId="0" pageNumber="69">textura intricata</emphasis>
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of hyaline hyphae and numerous colourless crystals.
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<emphasis italics="true" pageId="0" pageNumber="69">Conidiophores</emphasis>
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emerging around the central column or directly on bark in dense palisades, up to ca. 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long, filiform, branched near the base or sometimes 1-2 fold asymmetrically at higher levels, hyaline, turning brown from their tips; terminal conidiogenous cells (10-)14.5-23(-27)
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<normalizedToken originalValue="×">x</normalizedToken>
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(1.8-)2.3-3.5(-5)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(n = 90), cylindrical and often widened in the middle or towards base and at the funnel-shaped tips beyond its width, with up to 3 annellations, producing
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<normalizedToken originalValue="β-conidia">β-conidia</normalizedToken>
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and/or
|
||
<normalizedToken originalValue="α-conidia">α-conidia</normalizedToken>
|
||
.
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Conidia</emphasis>
|
||
dimorphic,
|
||
<normalizedToken originalValue="α-conidia">α-conidia</normalizedToken>
|
||
(9-)11-14(-16.3)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
(3.2-)4.5-5.5(-6.2)
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
, l/w (1.7-)2.2-2.9(-3.6) (n = 172), first hyaline, soon turning pale to medium brown or greyish-brown, unicellular, mostly fusoid, but also oblong, oval or ellipsoid, straight, less commonly slightly curved, upper end usually subacute and sometimes elongated, lower end narrowly truncate, containing several guttules, smooth;
|
||
<normalizedToken originalValue="β-">β-</normalizedToken>
|
||
<emphasis italics="true" pageId="0" pageNumber="69">conidia</emphasis>
|
||
(8-)9-11.5(-12.7)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
(2-)2.5-3(-3.3)
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
, l/w (2.8-)3.3-4.6(-5.8) (n = 39), hyaline to dilute brownish, unicellular, oblong to cylindrical, straight or slightly curved, thick-walled in water, with few guttules to eguttulate, smooth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Culture</emphasis>
|
||
: Colony on CMD at 16 °C first hyaline, partly or entirely turning brownish or ochre, either covered by a dense white mat of aerial hyphae or not, sometimes becoming indistinctly zonate, sometimes forming irregularly disposed conidiomata; on MEA at room temperature, first hyaline to whitish, soon forming a few broad zones with uneven margins forming teeth, the latter partly turning brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="distribution">
|
||
<paragraph pageId="0" pageNumber="69">Distribution and ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Common on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
(syn.
|
||
<taxonomicName lsidName="A. viridis" pageId="0" pageNumber="69" rank="species" species="viridis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">A. viridis</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName lsidName="A. incana" pageId="0" pageNumber="69" rank="species" species="incana">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">A. incana</emphasis>
|
||
</taxonomicName>
|
||
in mountainous areas of Central and Eastern Europe (confirmed for Austria, the Czech Republic fide
|
||
<bibRefCitation author="Podlahova, R" journalOrPublisher="Ceska Mykologie" pageId="0" pageNumber="69" pagination="84 - 97" refId="B34" refString="Podlahova, R, 1973. Ueber einige Pyrenomyceten auf Alnus viridis (Chaix) Lam. et DC. aus Suedboehmen. Ceska Mykologie 27: 84 - 97" title="Ueber einige Pyrenomyceten auf Alnus viridis (Chaix) Lam. et DC. aus Suedboehmen." volume="27" year="1973">
|
||
<normalizedToken originalValue="Podlahová">Podlahova</normalizedToken>
|
||
1973
|
||
</bibRefCitation>
|
||
, Romania fide
|
||
<bibRefCitation author="Szasz, E" journalOrPublisher="Contributii Botanice Cluj" pageId="0" pageNumber="69" pagination="27 - 34" refId="B46" refString="Szasz, E, 1966. Ciuperci parazite si saprofite pe Alnus viridis (Chaix) Lam. et DC. Contributii Botanice Cluj 2: 27 - 34" title="Ciuperci parazite si saprofite pe Alnus viridis (Chaix) Lam. et DC." volume="2" year="1966">
|
||
<normalizedToken originalValue="Szász">Szasz</normalizedToken>
|
||
1966
|
||
</bibRefCitation>
|
||
and Switzerland fide
|
||
<bibRefCitation DOI="https://doi.org/10.1139/b91-272" author="Sieber, TN" journalOrPublisher="Canadian Journal of Botany" pageId="0" pageNumber="69" pagination="2170 - 2176" refId="B39" refString="Sieber, TN, Sieber-Canavesi, F, Petrini, O, Ekramoddoullah, AKM, Dorworth, CE, 1991. Characterization of Canadian and European Melanconium from some Alnus species by morphological, cultural and biochemical studies. Canadian Journal of Botany 69: 2170 - 2176, DOI: https://doi.org/10.1139/b91-272" title="Characterization of Canadian and European Melanconium from some Alnus species by morphological, cultural and biochemical studies." url="https://doi.org/10.1139/b91-272" volume="69" year="1991">Sieber et al. 1991</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="69">Other material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Austria, Burgenland, Forchtenstein, Kohlstatt, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus incana" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="incana">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus incana</emphasis>
|
||
</taxonomicName>
|
||
, 24 Sep 2016, H. Voglmayr & W. Jaklitsch (WU 37046, culture D257);
|
||
<normalizedToken originalValue="Kärnten">Kaernten</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="Hüttenberg">Huettenberg</normalizedToken>
|
||
, Knappenberg, grid square 9053/3, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 10 Jun 1992, W. Jaklitsch (WU 15093);
|
||
<normalizedToken originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
|
||
, Aspangberg-St. Peter, Mariensee, grid square 8461/4, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 23 Sep 2009, H. Voglmayr (WU 29888); Steiermark, Hartberg, Pinggau, Schaueregg, Alte
|
||
<normalizedToken originalValue="Glashütte">Glashuette</normalizedToken>
|
||
, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 28 Jul 2012, W. Jaklitsch & H. Voglmayr (WU 38243); Judenburg, Pusterwald, grid square 8652/3, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 11 Jun 2011, H. Voglmayr (WU 31890, culture MAV1); Liezen,
|
||
<normalizedToken originalValue="Kleinsölk">Kleinsoelk</normalizedToken>
|
||
, walking path between
|
||
<normalizedToken originalValue="Breitlahnhütte">Breitlahnhuette</normalizedToken>
|
||
and Schwarzensee, grid square 8649/3, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 6 Aug 2003, W. Jaklitsch W.J. 2296 (BPI 843621; culture CBS 121480 = A.R. 4013); St. Nikolai im
|
||
<normalizedToken originalValue="Sölktal">Soelktal</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="Sölker">Soelker</normalizedToken>
|
||
<normalizedToken originalValue="Paß">Pass</normalizedToken>
|
||
, grid square 8750/1, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 14 Jun 2011, H. Voglmayr (WU 31889, culture CBS 131694 = MAV); Spital am Semmering, near Pfaffensattel, grid square 8460/2, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 15 Aug 2003, W. Jaklitsch W.J. 2331 (BPI 872072; culture A.R. 4032); ibidem, same host, 8 Jul 2010, I. Krisai-Greilhuber & H. Voglmayr (WU 31172); ibidem, same host, 7 Apr 2015, H. Voglmayr (WU 36699); Tirol,
|
||
<normalizedToken originalValue="Kühtai">Kuehtai</normalizedToken>
|
||
, between Haggen and
|
||
<normalizedToken originalValue="Kühtai">Kuehtai</normalizedToken>
|
||
, near Zirmbachalm, grid square 8732/3, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 3 Sep 2003, W Jaklitsch W.J. 2368 (W 2004-0000062);
|
||
<normalizedToken originalValue="Prägraten">Praegraten</normalizedToken>
|
||
, Bodenalm, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 18 Jun 2015, H. Voglmayr & W. Jaklitsch (WU 37044; culture D157);
|
||
<normalizedToken originalValue="Umbalfälle">Umbalfaelle</normalizedToken>
|
||
, grid square 8939/4, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 28 Aug 2000, W. Jaklitsch W.J. 1542 (WU 31891, BPI 748444; culture CBS 109773 = A.R. 3500; AFTOL-ID 2127); same area and host, 17 Jun 2015, H. Voglmayr & W. Jaklitsch (WU 37045; culture D158); Vienna,
|
||
<normalizedToken originalValue="Landstraße">Landstrasse</normalizedToken>
|
||
, Botanical garden, Alpinum, grid square 7864/1, on
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
, 21 Aug 1994, H. Voglmayr (WU 12976); same place and host, 6 Jan 2012, H. Voglmayr (WU 31893).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="notes">
|
||
<paragraph pageId="0" pageNumber="69">Notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
This subspecies differs mainly in its occurrence in (Central) Europe and by forming a clade of its own in phylogenetic analyses (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogram of the best ML tree (lnL = - 18240.558) revealed by RAxML from an analysis of the ITS-LSU- ms 204 - rpb 2 - tef 1 - tub 2 matrix of Melanconis, with 5 species of Juglanconis (Juglanconidaceae) selected as outgroup. ML and MP bootstrap support above 50 % are given at the first and second position, respectively, above or below the branches. Strain numbers are given following the taxon names; strains formatted in bold were sequenced in the current study. Melanconis taxa occurring on Alnus are marked blue, those on Betula in green. The broken branches to the outgroup were scaled to 10 %." figureDoi="10.3897/mycokeys.63.49054.figure1" httpUri="https://binary.pensoft.net/fig/384811" pageId="0" pageNumber="69">1</figureCitation>
|
||
). While the differences of the European accessions in each marker included are few, they are consistent, resulting in a well-delimited clade in the multigene analyses. As the morphological differences from
|
||
<taxonomicName authority="subsp. marginalis" lsidName="M. marginalis subsp. marginalis" pageId="0" pageNumber="69" rank="subSpecies" species="marginalis" subSpecies="marginalis">M. marginalis subsp. marginalis</taxonomicName>
|
||
are only small, we prefer to classify the European taxon as a subspecies rather than a separate species.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Under the name
|
||
<taxonomicName authorityName="Tul. & C. Tul., Select. fung. carpol. (Paris) 2: 122" authorityYear="1863" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconis alni" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="species" species="alni">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Melanconis alni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation author="Podlahova, R" journalOrPublisher="Ceska Mykologie" pageId="0" pageNumber="69" pagination="84 - 97" refId="B34" refString="Podlahova, R, 1973. Ueber einige Pyrenomyceten auf Alnus viridis (Chaix) Lam. et DC. aus Suedboehmen. Ceska Mykologie 27: 84 - 97" title="Ueber einige Pyrenomyceten auf Alnus viridis (Chaix) Lam. et DC. aus Suedboehmen." volume="27" year="1973">
|
||
<normalizedToken originalValue="Podlahová">Podlahova</normalizedToken>
|
||
(1973)
|
||
</bibRefCitation>
|
||
described both sexual and asexual morphs of a Czech collection from
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
which clearly represents
|
||
<taxonomicName lsidName="M. marginalis" pageId="0" pageNumber="69" rank="species" species="marginalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">M. marginalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<bibRefCitation author="Szasz, E" journalOrPublisher="Contributii Botanice Cluj" pageId="0" pageNumber="69" pagination="27 - 34" refId="B46" refString="Szasz, E, 1966. Ciuperci parazite si saprofite pe Alnus viridis (Chaix) Lam. et DC. Contributii Botanice Cluj 2: 27 - 34" title="Ciuperci parazite si saprofite pe Alnus viridis (Chaix) Lam. et DC." volume="2" year="1966">
|
||
<normalizedToken originalValue="Szász">Szasz</normalizedToken>
|
||
(1966)
|
||
</bibRefCitation>
|
||
listed and described the species (as
|
||
<taxonomicName class="Sordariomycetes" family="Melanconidaceae" genus="Melanconium" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconium dimorphum" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="species" species="dimorphum">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Melanconium dimorphum</emphasis>
|
||
</taxonomicName>
|
||
) from Romania, again from
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
. In his isozyme studies of
|
||
<taxonomicName authorityName="Link" authorityYear="1809" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconium" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconium" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Melanconium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1139/b91-272" author="Sieber, TN" journalOrPublisher="Canadian Journal of Botany" pageId="0" pageNumber="69" pagination="2170 - 2176" refId="B39" refString="Sieber, TN, Sieber-Canavesi, F, Petrini, O, Ekramoddoullah, AKM, Dorworth, CE, 1991. Characterization of Canadian and European Melanconium from some Alnus species by morphological, cultural and biochemical studies. Canadian Journal of Botany 69: 2170 - 2176, DOI: https://doi.org/10.1139/b91-272" title="Characterization of Canadian and European Melanconium from some Alnus species by morphological, cultural and biochemical studies." url="https://doi.org/10.1139/b91-272" volume="69" year="1991">Sieber et al. (1991)</bibRefCitation>
|
||
included a Swiss isolate from
|
||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus alnobetula" order="Fagales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="alnobetula">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Alnus alnobetula</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName authorityName="Link" authorityYear="1809" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconium" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconium" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Melanconium</emphasis>
|
||
</taxonomicName>
|
||
sp. 1). This isolate showed a distinct but similar isozyme pattern to North American collections of
|
||
<taxonomicName authorityName="Wehm., Pap. Michigan Acad. I. 6: 382" authorityYear="1926" baseAuthorityName="Peck" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconis marginalis" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="species" species="marginalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Melanconis marginalis</emphasis>
|
||
</taxonomicName>
|
||
and had a mean conidial size of 11.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
4.3
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
, indicating that this isolate also represents
|
||
<taxonomicName authorityName="Jaklitsch & Voglmayr" authorityYear="2020" class="Sordariomycetes" family="Melanconidaceae" genus="Melanconis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melanconis marginalis subsp. europaea" order="Diaporthales" pageId="0" pageNumber="69" phylum="Ascomycota" rank="subSpecies" species="marginalis" subSpecies="europaea">Melanconis marginalis subsp. europaea</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |