treatments-xml/data/0D/EE/EC/0DEEEC1347F05890B02BF2C79DAC3A73.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.452.8152" ID-PMC="PMC4258635" ID-Pensoft-Pub="1313-2970-452-107" ID-Pensoft-UUID="FFDEFFD61800FFEEBA2CFFA1014FFF8A" ID-PubMed="25493056" ID-Zenodo-Dep="578587" ID-ZooBank="F7F5E4C32D764ACAA4C35B8DE5D0F90C" ModsDocID="1313-2970-452-107" checkinTime="1451245146091" checkinUser="pensoft" docAuthor="Pekarsky, Oleg" docDate="2014" docId="0DEEEC1347F05890B02BF2C79DAC3A73" docLanguage="en" docName="ZooKeys 452: 107-129" docOrigin="ZooKeys 452" docPubDate="2014-11-05" docSource="http://dx.doi.org/10.3897/zookeys.452.8152" docTitle="Lygephila lubrica subsp. lubrica lubrica (Freyer 1846" docType="treatment" docVersion="3" id="FFDEFFD61800FFEEBA2CFFA1014FFF8A" lastPageNumber="110" masterDocId="FFDEFFD61800FFEEBA2CFFA1014FFF8A" masterDocTitle="Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae)" masterLastPageNumber="129" masterPageNumber="107" pageNumber="109" updateTime="1668159553656" updateUser="ExternalLinkService">
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<mods:title>Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pekarsky, Oleg</mods:namePart>
<mods:affiliation>H- 1068, Budapest, Felsoerdosor u. 16 - 18, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">opbp@t-online.hu</mods:nameIdentifier>
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<mods:date>2014</mods:date>
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<mods:number>2014-11-05</mods:number>
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<mods:detail type="volume">
<mods:number>452</mods:number>
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<mods:start>107</mods:start>
<mods:end>129</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.452.8152</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.452.8152</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578587</mods:identifier>
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<treatment ID-GBIF-Taxon="182234917" LSID="urn:lsid:plazi:treatment:0DEEEC1347F05890B02BF2C79DAC3A73" httpUri="http://treatment.plazi.org/id/0DEEEC1347F05890B02BF2C79DAC3A73" lastPageId="3" lastPageNumber="110" pageId="2" pageNumber="109">
<subSubSection pageId="2" pageNumber="109" type="nomenclature">
<paragraph pageId="2" pageNumber="109">
<taxonomicName LSID="0DEEEC13-47F0-5890-B02B-F2C79DAC3A73" authority="lubrica (Freyer, 1846)" authorityName="lubrica (Freyer" authorityYear="1846" baseAuthorityName="Freyer" baseAuthorityYear="1846" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica subsp. lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="lubrica" subSpecies="lubrica">
<pageBreakToken pageId="2" pageNumber="109" start="start">Lygephila</pageBreakToken>
lubrica lubrica (Freyer, 1846)
</taxonomicName>
<figureCitation captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Adults. Lygephila lubrica 1 ♂, Russia, Orenburg 2 ♀, Russia, Orenburg 3 ♂, Kazakhstan, Boro-Khoro Mts 4 ♀, Kabardino-Balkaria, Bydyk 5 ♀, Kyrgyzstan, Naryn reg. 6 ♀, China, Xinyiang - Uygur 7 ♂, Russia, Altai, Kupchegen 8 ♀, Russia, Tuva 9 ♂, SW Mongolia, Hovd aimak 10 ♀, SW Mongolia, Hovd aimak." pageId="2" pageNumber="109">Figs 1-10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="109" type="reference_group">
<paragraph pageId="2" pageNumber="109">
<taxonomicName authorityName="Freyer" authorityYear="1842" class="Insecta" family="Erebidae" genus="Ophiusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiusa lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">Ophiusa lubrica</taxonomicName>
Freyer, 1846, Neuere 
<normalizedToken originalValue="Beiträge">Beitraege</normalizedToken>
zur Schmetterlingskunde mit Abbildungen nach der Natur. 6: 7, Tab. 483, fig. 4. (TL: not given)
</paragraph>
<paragraph pageId="2" pageNumber="109">
<taxonomicName class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica subsp. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="lubrica" subSpecies="sublubrica">Lygephila lubrica sublubrica</taxonomicName>
(Staudinger, 1896);
</paragraph>
<paragraph pageId="2" pageNumber="109">
<taxonomicName class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxocampa lubrica subsp. var. var. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="variety" species="lubrica" subSpecies="var." variety="sublubrica">Toxocampa lubrica var. sublubrica</taxonomicName>
Staudinger, 1896, Deutsche Entomologische Zeitschrift Iris 8: 271. (TL: [Mongolia, Uliastai], Uliassutai)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="109" type="type material examined">
<paragraph pageId="2" pageNumber="109">Type material examined.</paragraph>
<paragraph pageId="2" pageNumber="109">
<taxonomicName class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica subsp. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="lubrica" subSpecies="sublubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica sublubrica</emphasis>
</taxonomicName>
(Staudinger, 1896), 
<emphasis bold="true" pageId="2" pageNumber="109">Type</emphasis>
♂, [Mongolia, Uliastai], Uliassataj, slide No. WB12 (coll. MNHU).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="109" type="additional material examined">
<paragraph pageId="2" pageNumber="109">Additional material examined.</paragraph>
<paragraph pageId="2" pageNumber="109">
2 ♂♂ &amp; 1 ♀, Russia, C Tuva, W of Ujukskyi Mts, Kamennyi riv. valley, h=800-1000 m, 11-20.07.2003, leg. S. Vaschenko, slide Nos: OP1955m, OP2438m, OP1956f (coll. O. Pekarsky); 1 ♂, Russia, Altai Mts, 700 m, Kupchegen, 23-25.VII.2002, leg. 
<normalizedToken originalValue="Hácz">Hacz</normalizedToken>
&amp; 
<normalizedToken originalValue="Juhász">Juhasz</normalizedToken>
, slide No. OP1962m (coll. O. Pekarsky); 1 ♂, Russia, NW Altai Mts, Tigireksky ridge, slide No. AV0907 (coll. A. Volynkin); 1 ♂ Russia, Altai rep., Aktash, 1400 m, 12-14.VIII.2010, leg. R. Yakovlev, slide No. OP2439m (coll. O. Pekarsky); 1 ♀, Russia, SE Altai Mts, Aktash vill., slide No. AV0906 (coll. A. Volynkin); 1 ♂, Mongolia, Central aim., Nr. 1148, leg. Z. Kaszab, slide No. LR1401m (coll. HNHM); 1 ♂, Mongolia, 
<normalizedToken originalValue="Chövsgöl">Choevsgoel</normalizedToken>
aimak, Nr. 1128, leg. Z. Kaszab, slide No. LR1402m (coll. HNHM); 1 ♀, Mongolia, Central aimak, 26 km O von Somon Lun, 1180 m, Nr. 260, 3.VIII.1964, leg. Z. Kaszab, slide No. OP2010f (coll. HNHM); 1 ♂ &amp; 2 ♀♀, Mongolia, Selenga aimak, Orhon v., Sir Orhon, 715 m, N49°08'956&quot;, E105°15'099&quot;, 3-4.07.2004, leg. K. 
<normalizedToken originalValue="Gaskó">Gasko</normalizedToken>
, slide Nos: OP2296m, OP2295f, OP2297f (coll. P. Gyulai); 1 ♂, [Kazakhstan], Russia, Uralsk, 1937.VII., ex coll. Velez, slide No. LR1403m, (coll. HNHM); 1 ♂ &amp; 1 ♀, [Kazakhstan], Uralsk, slide Nos: Hacker2536m/ZSM2510m, Hacker2334f/ZSM2508f (coll. ZSM); 1 ♂ &amp; 1 ♀, Russia, S Ural, Orenburg reg., Donskoe vill., Verbljushka Mt., 25-29.6.2009, leg. L. Srnka, slide Nos: OP2124m, OP2125f (coll. O. Pekarsky); 4 ♂♂ &amp; 1 ♀, Russia, Bashkortostan, Yantysh vill., 29-31.VII.2011 slide Nos: OP2005m, OP2007m, OP2440m, OP2441m, OP2006f (coll. O. Pekarsky); 1 ♂ &amp; 1 ♀, Russia, Kabardino-Balkaria, C Caucasus Mts, Bydyk, 1250m, 18.7.2012, leg. L. Srnka, slide Nos: OP2151m, OP2152f (coll. O. Pekarsky); 1 ♀, Kasakhstan, 40 km W Ust Kamenogorsk, Kalbinski Altai, Monastyri, 600 m, 06.08.1994, leg. Lukhtanov, slide No. OP2013f (coll. P. Gyulai); 1 ♂, Kazakhstan, Boro-Khoro Mts, 30km N of Panfilov, (20 km from Chinese border), N44°29'765&quot; E80°03'848&quot;, 1830 m, 30.06.2010, leg. S.K. Korb, slide No. OP2083m (coll. O. Pekarsky); 1 ♂, Kirgizstan, Inner Tjan-Shan, Min-Kush circ., 2300 m, 2.08.2000, leg. I. Pljushtch, slide No. OP2004m (coll. O. Pekarsky); 1 ♂ &amp; 1 ♀, Kyrgyzstan, Naryn reg., Kekemeren river, n., Sarykamysh, 1400 m, 6-8.07.1996, leg. V.A. Lukhtanov, slide Nos: OP2015m, OP2016f (coll. P. Gyulai); 1 ♂, [Kyrhyzstan], Issykkul, Tianschan, 949, ex coll. Kotzsch, slide No. OP2426m (coll. ZFMK); 2 ♀♀, China, Xinyiang [Xinjiang] - Uygur, Boro Horo Shan, Balguntay city, 2000 m, 13.7.1996, leg. Nykl, slide Nos: OP2011f, OP2012f (coll. P. Gyulai); 1 ♂, China, Boro Boro shan, Balguntay city, 2000 m, 13.7.1996, slide No. OP2289m (coll. P. Gyulai); 1 ♀, Aksu Bakalik, Anf. VI.1912, ex coll. 
<normalizedToken originalValue="Rückbeil">Rueckbeil</normalizedToken>
, slide No. OP2339f (coll. ZSM); 1 ♂, Aksu Bakalik, Anf. VI.1912, ex coll. 
<normalizedToken originalValue="Rückbeil">Rueckbeil</normalizedToken>
, slide No. OP2338m (coll. ZSM); 1 ♂, [China], Aksu, [19]11, 225, ZFMK76/64 Boppard, slide No. OP2427m (coll. ZFMK); 1 ♂, Mongolia, Uliasutai, slide No. 0326Matov (coll. ZISP); 1 ♂, [Mongolia], Uliassatai, 946, ex coll. Kotzsch, 8/57, slide No. OP2428m (coll. ZFMK); 1 ♂ &amp; 1 ♀, SW Mongolia, Hovd aimak, Bodonchijn-Gol basin, Hundij-Gol riv. valley, 1600 m, 46°06'N; 92°30'E, 3.vii.2010, leg. E. Guskova &amp; R. Yakovlev, slide Nos: OP1957m, OP1958f (coll. O. Pekarsky); 2 ♂♂, W Mongolia, Hovd aimak, near Erdene-Buren somon, h=1 400 m, 04.07.2007, leg. Yakovlev R.V. &amp; Guskova E.V., slide Nos: AV0283, AV0285 (coll. A. Volynkin); 1 ♂ &amp; 1 ♀, Mongolia, Hovd Aimak, Altaj Mts, 10 km NE of Dott, 2000 m, 10.08.1996, leg. S. Farkas &amp; I.Zs. 
<normalizedToken originalValue="Tóth">Toth</normalizedToken>
, slide No. OP2290m, OP2291f (coll. P. Gyulai); 2 ♂♂ &amp; 1 ♀, W Mongolia, Hovd aimak, near Erdene-Buren-Somon, 1400 m, 1.07.2010, 2500-2850 m, leg. R. Yakovlev, E. Guskova, slide Nos: OP2350m, OP2351m, OP2352f (coll. O. Pekarsky); 1 ♂, Mongolia, Bulgan aimak, 54 km W of Erdenecant, 1260 m, 104°05' 
<geoCoordinate degrees="47" direction="east" minutes="05" orientation="longitude" precision="925" value="47.083332">E 47°05'</geoCoordinate>
N, 22.07.1987, leg. L. Peregovits, M. Hreblay &amp; T. 
<normalizedToken originalValue="Stéger">Steger</normalizedToken>
, slide No. OP2008m (coll. HNHM); 1 ♂, Mongolia, [Khentii] Chentaj aimak, Tsenkher-Mandal, Modoto, 1600-1800 m, 9-14.07.1984, leg. K. Cerny, slide No. GB2550m, (coll. G. Behounek); 1 ♂, [Russia], Yakovlevka Spas. u., Ussur. kr., 12.VIII.[1]926, [leg.] 
<normalizedToken originalValue="D’iakonv">D'iakonv</normalizedToken>
<normalizedToken originalValue="Filip’ev">Filip'ev</normalizedToken>
(in russian), slide No. 0330Matov, (coll. ZISP); 1 ♀, Russia, Primorsky ter., Lesozavodsk reg., Innokentievka, 26-30.VIII.[19]94, slide No. OP2298f (coll. P. Gyulai); 1 ♀, [China], Mien-shan (Prov. Shansi), Obere 
<normalizedToken originalValue="Höhe">Hoehe</normalizedToken>
ca. 2000 m, 2.8.1937. [leg.] H. 
<normalizedToken originalValue="Höne">Hoene</normalizedToken>
, slide No. OP2423f, 2 ♂♂, 9.8.1937, slide Nos: OP2421m, OP2425m, 1 ♂, 13.8.1937, slide No. OP2422m (coll. ZFMK); 1 ♀, [China], Tapaishan im Tsinling, Sued-Shensi, Ca. 3000 m, 17.6.1936, [leg.] H. 
<normalizedToken originalValue="Höne">Hoene</normalizedToken>
, slide No. OP2424f (coll. ZFMK).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="109" type="taxonomy">
<paragraph pageId="2" pageNumber="109">Taxonomy.</paragraph>
<paragraph pageId="2" pageNumber="109">
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
was described in 1846 by Freyer in the genus 
<taxonomicName authorityName="Ochsenheimer" authorityYear="1816" class="Insecta" family="Erebidae" genus="Ophiusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiusa" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="109">Ophiusa</emphasis>
</taxonomicName>
. The exact type locality was not given in the original paper and also there was no information about the types. In 1896, Staudinger supposed during the description of 
<taxonomicName class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxocampa lubrica subsp. var. var. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="variety" species="lubrica" subSpecies="var." variety="sublubrica">Toxocampa lubrica var. sublubrica</taxonomicName>
, that 
<taxonomicName authorityName="Freyer" authorityYear="1842" class="Insecta" family="Erebidae" genus="Ophiusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiusa lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Ophiusa lubrica</emphasis>
</taxonomicName>
was described by Freyer from Altai: &quot;Freyer sagt von seiner 
<taxonomicName authorityName="Freyer" authorityYear="1842" class="Insecta" family="Erebidae" genus="Ophiusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="Lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lubrica</emphasis>
</taxonomicName>
nur, dass er sie von Kindermann erhielt; es muss sicher die von diesem Sammler im Altai gefundene Art sein, von der ich drei 
<normalizedToken originalValue="Stücke">Stuecke</normalizedToken>
aus 
<normalizedToken originalValue="Lederer’s">Lederer's</normalizedToken>
Sammlung besitze&quot;. Based on this assumption the type locality of 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
is most probably &quot;Russian Altai&quot; near Ust-Bukhtarminsk settlement (not existing now), which was located near the junction of the Bukhtarma and Irtysh rivers in the modern territory of Kazakhstan. Staudinger &amp; Wocke (1871) placed this species in the genus 
<taxonomicName authorityName="Guenee" authorityYear="1841" class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxocampa" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="109">Toxocampa</emphasis>
</taxonomicName>
, and later 
<bibRefCitation author="Staudinger, O" journalOrPublisher="Deutsche Entomologische Zeitschrift Iris" pageId="12" pageNumber="119" pagination="240 - 283" refId="B11" refString="Staudinger, O, 1896. Ueber Lepidopteren von Uliassutai. Deutsche Entomologische Zeitschrift Iris 9 (2): 240 - 283" title="Ueber Lepidopteren von Uliassutai." volume="9" year="1896">Staudinger (1896)</bibRefCitation>
described a variation named as 
<taxonomicName class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="subsp. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="lubrica" subSpecies="sublubrica">
<emphasis italics="true" pageId="2" pageNumber="109">sublubrica</emphasis>
</taxonomicName>
from Uliastai on the western edge of Khangai Mountains in the western part of Mongolia. The type specimen of 
<taxonomicName class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="subsp. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="lubrica" subSpecies="sublubrica">
<emphasis italics="true" pageId="2" pageNumber="109">sublubrica</emphasis>
</taxonomicName>
was not found in the collection of MNHU in Berlin however the genitalia slide made by Edward Wiltshire is in the museum (genitalia slide collection, Figs 
<figureCitation captionStart="Figures 27–32" captionStartId="F4" captionText="Figures 27 - 32. Clasping apparatus and genitalia slide. Lygephila lubrica." pageId="2" pageNumber="109">27</figureCitation>
, 
<figureCitation captionStart="Figures 27–32" captionStartId="F4" captionText="Figures 27 - 32. Clasping apparatus and genitalia slide. Lygephila lubrica." pageId="2" pageNumber="109">28</figureCitation>
). The current combination - 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
- occurs first in 
<bibRefCitation author="Sheljuzhko, L" journalOrPublisher="Reichenbachia" pageId="12" pageNumber="119" pagination="209 - 227" refId="B9" refString="Sheljuzhko, L, 1967. Noctuidae der I. und II. Expedition. No. 112., Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Lepidoptera). Reichenbachia 9: 209 - 227" title="Noctuidae der I. und II. Expedition. No. 112., Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Lepidoptera)." volume="9" year="1967">Sheljuzhko (1967)</bibRefCitation>
and later in 
<bibRefCitation author="Ronkay, L" journalOrPublisher="Annales historico-naturales Musei nationalis hungarici" pageId="11" pageNumber="118" pagination="229 - 246" refId="B8" refString="Ronkay, L, 1983. Noctuidae (Lepidoptera) from Mongolia. Noctuidae Quadrifinae. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei Nr. 475. Annales historico-naturales Musei nationalis hungarici 75: 229 - 246" title="Noctuidae (Lepidoptera) from Mongolia. Noctuidae Quadrifinae. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei Nr. 475." volume="75" year="1983">Ronkay (1983)</bibRefCitation>
. The taxon 
<taxonomicName class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="subsp. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="lubrica" subSpecies="sublubrica">
<emphasis italics="true" pageId="2" pageNumber="109">sublubrica</emphasis>
</taxonomicName>
is considered as a subspecies of 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
in these two works. 
<bibRefCitation author="Poole, RW" journalOrPublisher="EJ Brill / Flora and Fauna Publications, New Yourk" pageId="11" pageNumber="118" refId="B7" refString="Poole, RW, 1989. Lepidopterorum Catalogus. EJ Brill/Flora and Fauna Publications, New Yourk" title="Lepidopterorum Catalogus" year="1989">Poole (1989)</bibRefCitation>
incorrectly treated 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
(Freyer, 1842) as a new combination, and listed 
<taxonomicName class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxocampa lubrica subsp. var. var. sublubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="variety" species="lubrica" subSpecies="var." variety="sublubrica">Toxocampa lubrica var. sublubrica</taxonomicName>
Staudinger, 1896 and 
<taxonomicName class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxocampa lubrica subsp. var. var. lubrosa" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="variety" species="lubrica" subSpecies="var." variety="lubrosa">Toxocampa lubrica var. lubrosa</taxonomicName>
Staudinger, 1901, and have been listed as such in subsequent works (e.g., 
<bibRefCitation author="Goater, B" journalOrPublisher="Entomological press, Soro" pageId="11" pageNumber="118" refId="B3" refString="Goater, B, Ronkay, L, Fibiger, M, 2003. Catocalinae &amp; Plusiinae. Noctuidae Europaeae, volume 10. Entomological press, Soro" title="Catocalinae &amp; Plusiinae. Noctuidae Europaeae, volume 10." year="2003">Goater et al. 2003</bibRefCitation>
; 
<bibRefCitation author="Kononenko, VS" journalOrPublisher="Entomological press, Soro" pageId="11" pageNumber="118" refId="B5" refString="Kononenko, VS, 2010. Micronoctuidae, Noctuidae: Rivulinae - Agaristinae (Lepidoptera). Noctuidae Sibiricae, 2. Entomological press, Soro" title="Micronoctuidae, Noctuidae: Rivulinae - Agaristinae (Lepidoptera). Noctuidae Sibiricae, 2." year="2010">Kononenko 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="109" type="diagnosis">
<paragraph pageId="2" pageNumber="109">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="109">
The main external distinctive feature of the species is the brownish-grey ground colour of forewings and hindwings. 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
differs from the externally somewhat similar 
<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrosa">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrosa</emphasis>
</taxonomicName>
by its characteristic brownish-grey ground color of the forewings; from 
<taxonomicName authorityName="Stshetkin YuL &amp; Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="kazachkaratavika">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila kazachkaratavika</emphasis>
</taxonomicName>
by more unicolorous forewings with a less-developed pattern; and from both related species by its brownish hindwings, which are generally ochreous in the two latter species. The differences in the genitalia structures among the three similar species are easily recognisable in both sexes. In males, the uncus dilation in 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
is wider than in 
<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrosa">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrosa</emphasis>
</taxonomicName>
, but narrower than in 
<taxonomicName authorityName="Stshetkin YuL &amp; Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="kazachkaratavika">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila kazachkaratavika</emphasis>
</taxonomicName>
, and the ampulla is more proximal, closer to the middle of the valve, than in the two other species; in the females, the cleft on the posterior margin of the antrum is U-shaped or V-shaped in 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrica</emphasis>
</taxonomicName>
, whereas in 
<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="lubrosa">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila lubrosa</emphasis>
</taxonomicName>
it is evenly concave; in 
<taxonomicName authorityName="Stshetkin YuL &amp; Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="kazachkaratavika">
<emphasis italics="true" pageId="2" pageNumber="109">Lygephila kazachkaratavika</emphasis>
</taxonomicName>
the ostium cleft is deep, narrow, slit-like.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="110" pageId="2" pageNumber="109" type="description">
<paragraph pageId="2" pageNumber="109">Description.</paragraph>
<paragraph pageId="2" pageNumber="109">Wingspan 37-50 mm, on average 42-48 mm. Head and body brownish grey; collar dark chocolate brown. Forewing brownish grey, sometimes dark brown; subbasal line indistinct; antemedial line arched, consisting of two elongated patches; medial fascia diffuse, wide and waved, with two costal patches; reniform stigma approximately triangular, dark brown, sometimes with sharp extension at inner corner and with satellite streak-like spots on outer margin; orbicular stigma as small white dot; postmedial line distinct; subterminal line with light fascia; terminal line a black sinuous stripe. Hindwing varies from brown to greyish brown; transverse line distinct; narrow discal spot present on underside; outer dark third with defuse inner margin; fringes as ground color.</paragraph>
<paragraph pageId="2" pageNumber="109">
<emphasis bold="true" pageId="2" pageNumber="109">Male genitalia</emphasis>
(Figs 
<figureCitation captionStart="Figures 21–26" captionStartId="F3" captionText="Figures 21 - 26. Clasping apparatus. Lygephila lubrica." pageId="2" pageNumber="109">21</figureCitation>
- 
<figureCitation captionStart="Figures 27–32" captionStartId="F4" captionText="Figures 27 - 32. Clasping apparatus and genitalia slide. Lygephila lubrica." pageId="2" pageNumber="109">32</figureCitation>
, 
<figureCitation captionStart="Figures 39–41" captionStartId="F6" captionText="Figures 39 - 41. Vesica structure of Lygephila lubrica. 39, 40 Mongolia, Hovd aimak, slide No. OP 1957 m 39 lateral view 40 lateral view opposite side 41 Russia, Altai, Kupchegen, slide No. OP 1962 m, lateral view." pageId="2" pageNumber="109">39</figureCitation>
- 
<figureCitation captionStart="Figures 39–41" captionStartId="F6" captionText="Figures 39 - 41. Vesica structure of Lygephila lubrica. 39, 40 Mongolia, Hovd aimak, slide No. OP 1957 m 39 lateral view 40 lateral view opposite side 41 Russia, Altai, Kupchegen, slide No. OP 1962 m, lateral view." pageId="2" pageNumber="109">41</figureCitation>
). Uncus with short stem and dilated distal two thirds, apex with fine tip, anal tube membranous with characteristic oval hardening of tissue - scaphial crown on scaphium and sclerotized fig on subscaphium; valva elongated, relatively wide with parallel margins, valval apex rounded; clasper digitiform, slightly curved towards costa, situated rather far from apex. Aedeagus straight, long, tubular. Vesica globular, multidiverticulate, membranous; 1st subbasal diverticulum small, adjacent to 2nd terminal diverticulum; medial diverticulum large, tapering, with medium-large oblong chamber at base; 1st terminal diverticulum large, more or less wedge shaped with one part densely scobinated and membranous, cauliflower-like, opposite part bears numerous small pockets; 2nd terminal diverticulum tubular, scobinated on top; 3rd terminal diverticulum irregular shaped with large rectangular scobinated basal part and membranous cylindrical extension; 4th terminal diverticulum medium sized, situated between 1st and 3rd medial diverticulum; 2nd subbasal diverticulum small, tubular, sometimes chili-pepper-like (Fig. 
<figureCitation captionStart="Figures 39–41" captionStartId="F6" captionText="Figures 39 - 41. Vesica structure of Lygephila lubrica. 39, 40 Mongolia, Hovd aimak, slide No. OP 1957 m 39 lateral view 40 lateral view opposite side 41 Russia, Altai, Kupchegen, slide No. OP 1962 m, lateral view." pageId="2" pageNumber="109">41</figureCitation>
), terminal tube membranous, as long as aedeagus, opening point of terminal tube located subbasally near carina.
</paragraph>
<paragraph pageId="3" pageNumber="110">
<emphasis bold="true" pageId="3" pageNumber="110">
<pageBreakToken pageId="3" pageNumber="110" start="start">Female</pageBreakToken>
genitalia
</emphasis>
(Figs 
<figureCitation captionStart="Figures 46–63" captionStartId="F8" captionText="Figures 46 - 63. Female genitalia. 46 - 57 Lygephila lubrica 58 - 61 Lygephila lubrosa lubrosa 58 ventral view 59 dorsal view 60 lateral view 61 lateral view 62, 63 Lygephila kazachkaratavika." pageId="3" pageNumber="110">46</figureCitation>
- 
<figureCitation captionStart="Figures 46–63" captionStartId="F8" captionText="Figures 46 - 63. Female genitalia. 46 - 57 Lygephila lubrica 58 - 61 Lygephila lubrosa lubrosa 58 ventral view 59 dorsal view 60 lateral view 61 lateral view 62, 63 Lygephila kazachkaratavika." pageId="3" pageNumber="110">57</figureCitation>
). Ovipositor relatively large, broad, papillae anales hairy with long setae on apical edges. Apophyses anteriores stout, apophyses posteriores thin, longer than apophyses anteriores. Ostium broad, antrum tapering, funnel shaped, posterior margin incised producing large U-shaped cleft; ductus bursae small, inflated with ventral sclerotized ribbon; appendix bursae small; corpus bursae membranous, ovoid.
</paragraph>
<caption pageId="3" pageNumber="110" start="Figures 1–10" startId="F1">
<paragraph pageId="3" pageNumber="110">
<emphasis bold="true" pageId="3" pageNumber="110">Figures 1-10.</emphasis>
Adults. 
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="3" pageNumber="110" phylum="Arthropoda" rank="species" species="lubrica">
<emphasis italics="true" pageId="3" pageNumber="110">Lygephila lubrica</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="3" pageNumber="110">1</emphasis>
♂, Russia, Orenburg 
<emphasis bold="true" pageId="3" pageNumber="110">2</emphasis>
♀, Russia, Orenburg 
<emphasis bold="true" pageId="3" pageNumber="110">3</emphasis>
♂, Kazakhstan, Boro-Khoro Mts 
<emphasis bold="true" pageId="3" pageNumber="110">4</emphasis>
♀, Kabardino-Balkaria, Bydyk 
<emphasis bold="true" pageId="3" pageNumber="110">5</emphasis>
♀, Kyrgyzstan, Naryn reg. 
<emphasis bold="true" pageId="3" pageNumber="110">6</emphasis>
♀, China, Xinyiang - Uygur 
<emphasis bold="true" pageId="3" pageNumber="110">7</emphasis>
♂, Russia, Altai, Kupchegen 
<emphasis bold="true" pageId="3" pageNumber="110">8</emphasis>
♀, Russia, Tuva 
<emphasis bold="true" pageId="3" pageNumber="110">9</emphasis>
♂, SW Mongolia, Hovd aimak 
<emphasis bold="true" pageId="3" pageNumber="110">10</emphasis>
♀, SW Mongolia, Hovd aimak.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="110" type="distribution">
<paragraph pageId="3" pageNumber="110">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="110">Siberian. Distributed from Zaporozhie region of Ukraine to Rostov, Samara, Povolzhie regions to Ural of Russia through Kazakhstan, Russian Altai and northern Mongolia.</paragraph>
</subSubSection>
</treatment>
</document>