524 lines
65 KiB
XML
524 lines
65 KiB
XML
<document id="70BFE6726FF334E1CF1BF28452436CAD" ID-DOI="10.11646/zootaxa.4097.4.2" ID-GBIF-Dataset="20fb537c-b02d-4606-bdc8-7c2cc4ca6acf" ID-ISSN="1175-5326" ID-Zenodo-Dep="255048" ID-ZooBank="09889BBB-66DE-46D2-A394-6A194E7AE102" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459763531946" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Jiang, Ri-Xin & Steiner, Helmut" docDate="2016" docId="F340E468FF81FFF5578386E6FEFFF9BE" docLanguage="en" docName="zootaxa.4097.4.2.pdf.imf" docOrigin="Zootaxa 4097 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Araneibatrus Yin and Li 2010" docType="treatment" docVersion="8" lastPageNumber="478" masterDocId="0F799C10FF80FFF657148561FFE5FFEE" masterDocTitle="Revision of the genus Araneibatrus (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="494" masterPageNumber="475" pageNumber="476" updateTime="1698215927199" updateUser="plazi">
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<mods:mods id="61E7B60425B7D1960DAC6E572B6E8063" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="199974C4723CE0859E3E007284C65CAC">
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<mods:title id="896C252A3B9C2BDF0E69C20D4FDC245E">Revision of the genus Araneibatrus (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="CF42F4623504FCA0453AE510E239B638" type="personal">
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<mods:role id="8948B2DD7FF9D57D670959048470019D">
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<mods:roleTerm id="AD39990F3977E4E8DCA33C9F7B2BBB4B">Author</mods:roleTerm>
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<mods:namePart id="A22F17E5700E273E3FD974993B1F4225">Yin, Zi-Wei</mods:namePart>
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<mods:roleTerm id="5179EA0A4072813BFD9E3D07CA2D14B9">Author</mods:roleTerm>
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<mods:namePart id="69BB52AF91F7EDDC6EF19AB5598D7D9C">Jiang, Ri-Xin</mods:namePart>
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<mods:name id="09663C0D2624A02D49871B9019D0A75E" type="personal">
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<mods:roleTerm id="D5275E84DD5BA32FF504A71AAF69FF32">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0EA91D0C078A09F811A1E1BE1A046028">Steiner, Helmut</mods:namePart>
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<mods:typeOfResource id="8EF993343963A77933D742E205D55E76">text</mods:typeOfResource>
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<mods:title id="B8F294CFCA8D72C4A3B3A63582DB19B2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B94145B2E031DC529DAFE52BED9894E5">
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<mods:date id="29A5F85B121DF31F07A683B464AE0EAF">2016</mods:date>
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<mods:detail id="1289536FE9E8C8BE99FA492734F6CD61" type="volume">
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<mods:number id="540A1727510D58C8B88C9BEEBB2B7124">4097</mods:number>
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</mods:detail>
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<mods:detail id="2D0CC6E8EE5ED023FC65AD2BEF8DE36A" type="issue">
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<mods:number id="DBD6EA28EB97E586571B09655ADE45D8">4</mods:number>
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<mods:start id="6DFC9AB91305CB4171E27743011B1DED">475</mods:start>
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<mods:end id="FFCAE6D64E95BD4D6416BF9BA2CD3110">494</mods:end>
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<mods:classification id="E3D2E5976171B80B0981EBB7E6AA8071">journal article</mods:classification>
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<mods:identifier id="FCD7C9D5E84887CA2A1DE911180C5B0D" type="DOI">10.11646/zootaxa.4097.4.2</mods:identifier>
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<mods:identifier id="D4EA2D6B904E2FCBE0A4399F59A503F3" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<subSection id="12664C95FF81FFF5578386E6FEFFF9BE" lastPageId="3" lastPageNumber="478" pageId="1" pageNumber="476" type="multiple">
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<treatment id="F340E468FF81FFF5578386E6FEFFF9BE" ID-DOI="http://doi.org/10.5281/zenodo.6090584" ID-GBIF-Taxon="119345450" ID-Zenodo-Dep="6090584" LSID="urn:lsid:plazi:treatment:F340E468FF81FFF5578386E6FEFFF9BE" httpUri="http://treatment.plazi.org/id/F340E468FF81FFF5578386E6FEFFF9BE" lastPageId="3" lastPageNumber="478" pageId="1" pageNumber="476">
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<subSubSection id="33F306F5FF81FFF7578386E6FDE8FC4C" box="[151,525,903,930]" pageId="1" pageNumber="476" type="nomenclature">
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<paragraph id="7B56557EFF81FFF7578386E6FDE8FC4C" blockId="1.[151,525,903,930]" box="[151,525,903,930]" pageId="1" pageNumber="476">
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<heading id="201EE212FF81FFF7578386E6FDE8FC4C" bold="true" box="[151,525,903,930]" fontSize="11" level="1" pageId="1" pageNumber="476" reason="1">
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<taxonomicName id="BCE92EFDFF81FFF7578386E6FDE8FC4C" ID-CoL="8XJPT" box="[151,525,903,930]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
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<emphasis id="499D896CFF81FFF7578386E6FDE8FC4C" bold="true" box="[151,525,903,930]" pageId="1" pageNumber="476">
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<emphasis id="499D896CFF81FFF7578386E6FED3FC4F" bold="true" box="[151,310,903,929]" italics="true" pageId="1" pageNumber="476">Araneibatrus</emphasis>
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Yin and Li, 2010
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="33F306F5FF81FFF7578386AFFCF6FBAE" pageId="1" pageNumber="476" type="reference_group">
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<paragraph id="7B56557EFF81FFF7578386AFFEAEFBED" blockId="1.[151,1436,974,1088]" pageId="1" pageNumber="476">
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<taxonomicName id="BCE92EFDFF81FFF7578386AFFE21FC0A" ID-CoL="8XJPT" box="[151,452,974,996]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
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<emphasis id="499D896CFF81FFF7578386AFFEF8FC0A" box="[151,285,974,996]" italics="true" pageId="1" pageNumber="476">Araneibatrus</emphasis>
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Yin & Li, 2010
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</taxonomicName>
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(in: Yin, Li & Zhao 2010: 54).
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<typeStatus id="A452EBDCFF81FFF7540586AEFCA6FC0A" box="[785,835,975,996]" pageId="1" pageNumber="476">Type</typeStatus>
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species:
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<taxonomicName id="BCE92EFDFF81FFF754B786AFFAD8FC0A" ID-CoL="G53G" authority="Yin & Li, 2010" authorityName="Yin & Li" authorityYear="2010" box="[931,1341,974,996]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="gracilipes">
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<emphasis id="499D896CFF81FFF754B786AFFB70FC0A" box="[931,1173,974,996]" italics="true" pageId="1" pageNumber="476">Araneibatrus gracilipes</emphasis>
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Yin & Li, 2010
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</taxonomicName>
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(original designation).
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</paragraph>
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<paragraph id="7B56557EFF81FFF75783816DFCF6FBAE" blockId="1.[151,1436,974,1088]" pageId="1" pageNumber="476">
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<taxonomicName id="BCE92EFDFF81FFF75783816DFDFAFBCC" box="[151,543,1036,1058]" genus="Tribasoadellus" pageId="1" pageNumber="476" rank="genus">
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<emphasis id="499D896CFF81FFF75783816DFECBFBCC" box="[151,302,1036,1058]" italics="true" pageId="1" pageNumber="476">Tribasoadellus</emphasis>
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<bibRefCitation id="1F78288FFF81FFF7562D816CFDFAFBCC" author="Nomura" box="[313,543,1036,1058]" pageId="1" pageNumber="494" refString="Yin, Z. - W., Nomura, S. & Zhao, M. - J. (2011) Taxonomic study on Batrisodellus Jeannel of China, with discussion on the systematic position of Batrisodellus callissimus Nomura & Wang, 1991. Spixiana, 34, 33 - 38." type="journal article" year="2011">Nomura & Yin, 2011</bibRefCitation>
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</taxonomicName>
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(in:
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<bibRefCitation id="1F78288FFF81FFF7554D816DFC66FBCC" author="Yin" box="[601,899,1036,1058]" pageId="1" pageNumber="494" refString="Yin, Z. - W., Nomura, S. & Zhao, M. - J. (2011) Taxonomic study on Batrisodellus Jeannel of China, with discussion on the systematic position of Batrisodellus callissimus Nomura & Wang, 1991. Spixiana, 34, 33 - 38." type="journal article" year="2011">Yin, Nomura & Zhao 2011</bibRefCitation>
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: 37),
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<emphasis id="499D896CFF81FFF754D2816DFB8CFBCF" bold="true" box="[966,1129,1036,1057]" pageId="1" pageNumber="476">new synonymy</emphasis>
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.
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<typeStatus id="A452EBDCFF81FFF7536F816CFB48FBCC" box="[1147,1197,1037,1058]" pageId="1" pageNumber="476">Type</typeStatus>
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species:
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<taxonomicName id="BCE92EFDFF81FFF75201816DFDCEFBAE" ID-CoL="5WG86" authority="Nomura & Wang, 1991" authorityName="Nomura & Wang" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="callissimus">
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<emphasis id="499D896CFF81FFF75201816DFEDDFBAE" italics="true" pageId="1" pageNumber="476">Batrisodellus callissimus</emphasis>
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<bibRefCitation id="1F78288FFF81FFF7562A814AFDCEFBAE" author="Nomura" box="[318,555,1066,1088]" pageId="1" pageNumber="494" refString="Nomura, S. & Wang, F. - X. (1991) Description of a new cavernicolous species of the genus Batrisodellus (Coleoptera, Pselaphidae) from southeast China. Elytra, 19, 77 - 83." type="journal article" year="1991">Nomura & Wang, 1991</bibRefCitation>
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</taxonomicName>
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(original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="33F306F5FF81FFF75783810CFF2AFAB0" pageId="1" pageNumber="476" type="diagnosis">
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<paragraph id="7B56557EFF81FFF75783810CFF2AFAB0" blockId="1.[151,1437,1133,2022]" pageId="1" pageNumber="476">
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<emphasis id="499D896CFF81FFF75783810CFE95FB68" bold="true" box="[151,368,1133,1158]" pageId="1" pageNumber="476">Revised diagnosis.</emphasis>
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Body, antennae, and legs distinctly elongate. Head roundly rectangular; vertex with small, nude foveae situated posterior to dorsal level of posterior margin of eyes; lacking frontal rostrum; with strongly prominent antennal tubercles. Pronotum lacking median antebasal fovea (fovea reduced to a pit or an impressed area), lacking lateral longitudinal sulci and marginal spines. Elytra narrowing to bases; each elytron with three basal foveae; lacking distinct discal stria. Abdominal tergite IV (first visible tergite) longest, with oblique inner marginal carinae. Aedeagal median lobe asymmetric, with large basal capsule and foramen, with articulated dorsal lobe.
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</paragraph>
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</subSubSection>
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<subSubSection id="33F306F5FF81FFF457D38008FB41FD46" lastPageId="2" lastPageNumber="477" pageId="1" pageNumber="476" type="description">
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<paragraph id="7B56557EFF81FFF757D38008FDCAF94C" blockId="1.[151,1437,1133,2022]" pageId="1" pageNumber="476">
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<emphasis id="499D896CFF81FFF757D38008FE91FA6C" bold="true" box="[199,372,1385,1410]" pageId="1" pageNumber="476">Redescription.</emphasis>
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Length
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<quantity id="BC11F89BFF81FFF756C88008FD6DFA6F" box="[476,648,1385,1410]" metricMagnitude="-3" metricUnit="m" metricValue="2.3099999999999996" metricValueMax="2.61" metricValueMin="2.01" pageId="1" pageNumber="476" unit="mm" value="2.3099999999999996" valueMax="2.61" valueMin="2.01">2.01–2.61 mm</quantity>
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. Head roundly rectangular; lacking frontal rostrum, with distinctly prominent antennal tubercles; vertexal foveae nude, situated posterior dorsal level of posterior margins of eyes, lacking sulcus connecting foveae; in some species vertex with mediobasal carina, lateral post-antennal carinae present or absent; antenna with eleven antennomeres, clubs formed by three apical antennomeres, in some cases clubs indistinctly delimited; ocular-mandibular carinae present, extending from anterior margins of eyes, across frons to clypeal anterior margin; clypeus with rounded anterior margin, one-fifth to one-third of total head length; eyes prominent in male, posteroventral margins slightly flat; maxillary palpus with palpomere I minute, II elongate and pedunculate at base, III short and triangular, IV fusiform, its mesal margins roundly expanded, with small palpal cone; with single gular fovea.
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</paragraph>
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<paragraph id="7B56557EFF81FFF757D383CFFE4FF8BB" blockId="1.[151,1437,1133,2022]" pageId="1" pageNumber="476">Pronotum lacking median antebasal fovea, lateral antebasal foveae connected by shallow transverse antebasal sulcus, sometimes sulcus faint or lacking; inner and outer pairs of basolateral foveae present; median longitudinal and antebasal sulci distinct to lacking, lacking lateral longitudinal sulci; laterodorsal carinae rarely present; discal or antebasal tubercles/spines rarely present; lateral margins lacking spine; paranotal carinae complete; lateral procoxal foveae present.</paragraph>
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<paragraph id="7B56557EFF81FFF757D38200FD82F82C" blockId="1.[151,1437,1133,2022]" pageId="1" pageNumber="476">Each elytron with three small to large nude foveae at base; lacking subbasal fovea; discal stria indistinct or lacking; lacking subhumeral foveae, marginal stria present in posterior half of elytral lateral margin; apicolateral margins straight to shallowly emarginate.</paragraph>
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<paragraph id="7B56557EFF81FFF457D382ACFF08FF16" blockId="1.[151,1437,1133,2022]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="477" pageId="1" pageNumber="476">Mesoventrite with median foveae widely separated, opening into shared transverse cavity, lateral mesoventral foveae forked, often partially overlapping median foveae; lateral mesocoxal foveae large and deep. Metaventrite with lateral foveae; metacoxae widely separated; posterior margin of metaventrite with narrow or wide notch at middle.</paragraph>
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<paragraph id="7B56557EFF82FFF457D38465FC69FDF6" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="477">Abdominal tergite IV (first visible tergite) as long as to slightly longer than tergites V and VI combined; with complete outer pair, and usually complete and oblique inner pair of marginal carinae; short discal carinae sometimes present; basal impression complete when with only two basolateral foveae, or interrupted when pair of mediobasal and/or two pairs of basolateral foveae present; tergites V and VI combined slightly longer than VII, single lateral carina present or absent; tergites V–VII each with one pair of basolateral foveae; tergite VII with pair of small lateral tubercles (clearly seen only in slide preparation). Sternite IV (second visible sternite) usually with one pair of mediobasal and three pairs of basolateral foveae, occasionally mediobasal foveae lacking; sternites V– VII each short at midline, with one pair of small basolateral foveae.</paragraph>
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<paragraph id="7B56557EFF82FFF457D38744FB0EFDD2" blockId="2.[151,1437,151,2013]" box="[199,1259,548,573]" pageId="2" pageNumber="477">Legs with pro- and mesotarsomeres II as long as III, metatarsomeres II slightly longer than III.</paragraph>
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<paragraph id="7B56557EFF82FFF457D38726FB41FD46" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="477">Male secondary sexual characters: larger eyes and longer elytra; antennal clubs and metaventrite modified; mesotrochanter, mesotibia, or metatrochanters spinose. Aedeagus with parameres fused to median lobe, with large basal capsule and foramen, articulated dorsal lobe elongate, in dorsal view offset to left side.</paragraph>
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</subSubSection>
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<subSubSection id="33F306F5FF82FFF457D387D5FD31FABB" pageId="2" pageNumber="477" type="discussion">
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<paragraph id="7B56557EFF82FFF457D387D5FAB5FC6E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="477">
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<emphasis id="499D896CFF82FFF457D387D5FEA9FD23" bold="true" box="[199,332,692,717]" pageId="2" pageNumber="477">Discussion.</emphasis>
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<emphasis id="499D896CFF82FFF4564087D5FE0DFD23" box="[340,488,692,717]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
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was originally established (Yin, Li & Zhao 2010: 54) based on a single species
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<taxonomicName id="BCE92EFDFF82FFF4529087D5FEE2FD1E" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="gracilipes">
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<emphasis id="499D896CFF82FFF4529087D5FA70FD25" box="[1412,1429,692,715]" italics="true" pageId="2" pageNumber="477">A</emphasis>
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.
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<emphasis id="499D896CFF82FFF4578387B6FEE2FD1E" box="[151,263,727,752]" italics="true" pageId="2" pageNumber="477">gracilipes</emphasis>
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</taxonomicName>
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collected in leaf litter samples in the Nanling Nature Reserve, southern
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<collectingCountry id="03FE15EEFF82FFF4535D87B6FB75FD1E" box="[1097,1168,727,752]" name="China" pageId="2" pageNumber="477">China</collectingCountry>
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. This species possesses relatively long antennae and legs (Yin, Li & Zhao 2010: fig. 1), similar to a few batrisine genera that include cavernicolous species. In the following year, a new generic name
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<emphasis id="499D896CFF82FFF45481867EFBCBFCD6" box="[917,1070,799,824]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
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was proposed (
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<bibRefCitation id="1F78288FFF82FFF453F1867EFEF6FCB2" author="Yin" pageId="2" pageNumber="494" refString="Yin, Z. - W., Nomura, S. & Zhao, M. - J. (2011) Taxonomic study on Batrisodellus Jeannel of China, with discussion on the systematic position of Batrisodellus callissimus Nomura & Wang, 1991. Spixiana, 34, 33 - 38." type="journal article" year="2011">Yin, Nomura & Zhao 2011</bibRefCitation>
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) for
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<taxonomicName id="BCE92EFDFF82FFF4565A8625FD8CFCB3" box="[334,617,836,861]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="callissimus">
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<emphasis id="499D896CFF82FFF4565A8625FD8CFCB3" box="[334,617,836,861]" italics="true" pageId="2" pageNumber="477">Batrisodellus callissimus</emphasis>
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</taxonomicName>
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, a species know to occur in several limestone caves around the Guilin City, Guangxi (
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<bibRefCitation id="1F78288FFF82FFF456568609FDA2FC6E" author="Nomura" box="[322,583,872,896]" pageId="2" pageNumber="494" refString="Nomura, S. & Wang, F. - X. (1991) Description of a new cavernicolous species of the genus Batrisodellus (Coleoptera, Pselaphidae) from southeast China. Elytra, 19, 77 - 83." type="journal article" year="1991">Nomura & Wang 1991</bibRefCitation>
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). But
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<emphasis id="499D896CFF82FFF4559E8606FCC7FC6E" box="[650,802,871,896]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
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was not compared with any relatives at that time.
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</paragraph>
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<paragraph id="7B56557EFF82FFF457D386ECFD31FABB" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="477">
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Upon a reexamination and comparison of the
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<typeStatus id="A452EBDCFF82FFF455DB86ECFD1AFC4B" box="[719,767,909,933]" pageId="2" pageNumber="477">type</typeStatus>
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species of both genera, we found that
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<emphasis id="499D896CFF82FFF453A386EDFAAEFC4B" box="[1207,1355,908,933]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
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differs from
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<emphasis id="499D896CFF82FFF457C086CEFE89FC26" box="[212,364,943,968]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
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mainly in the following characters states: 1) the head, pronotum, and elytra much more slender in
|
||
<emphasis id="499D896CFF82FFF457A386B5FEAAFC03" box="[183,335,980,1005]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
than those of
|
||
<emphasis id="499D896CFF82FFF456E986B5FD74FC03" box="[509,657,980,1005]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
; 2) presence of a pair of lateral post-antennal carinae, and a short median carina on the vertex in
|
||
<emphasis id="499D896CFF82FFF455168696FD7FFBFE" box="[514,666,1015,1040]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
, both carinae absent in
|
||
<emphasis id="499D896CFF82FFF454A68696FBA3FBFE" box="[946,1094,1015,1040]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
; 3) the pronotum with paired discal and antebasal tubercles in
|
||
<emphasis id="499D896CFF82FFF4551F817DFD7AFBDB" box="[523,671,1052,1077]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
, the tubercles absent in
|
||
<emphasis id="499D896CFF82FFF454A5817DFBACFBDB" box="[945,1097,1052,1077]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
; 4) the basolateral margins of the elytra shallowly emarginate (the elytral base more constricted) in
|
||
<emphasis id="499D896CFF82FFF454A8815EFBB1FBB6" box="[956,1108,1087,1112]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
, the basolateral margins not emarginate in
|
||
<emphasis id="499D896CFF82FFF456228105FE2FFB93" box="[310,458,1124,1149]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
; 5) the lateral and median mesoventral foveae largely overlapping in
|
||
<emphasis id="499D896CFF82FFF453C78105FA8EFB93" box="[1235,1387,1124,1149]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
, the foveae clearly separated in
|
||
<emphasis id="499D896CFF82FFF456C181E6FD8CFB4E" box="[469,617,1159,1184]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
; 6) the posterior margin of the metaventrite at the middle with a wide notch in
|
||
<emphasis id="499D896CFF82FFF457EC81CDFE75FB2B" box="[248,400,1196,1221]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
, the notch narrow in
|
||
<emphasis id="499D896CFF82FFF4556881CDFCF5FB2B" box="[636,784,1196,1221]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
; 7) the tergite IV with only two pairs of basolateral foveae in
|
||
<emphasis id="499D896CFF82FFF457AC81AEFEB5FB06" box="[184,336,1231,1256]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
, with one pair of mediobasal and two pairs of basolateral foveae in
|
||
<emphasis id="499D896CFF82FFF4536981AEFAF4FB06" box="[1149,1297,1231,1256]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
; and 8) the aedeagal median lobe with a wide, transverse basal capsule in
|
||
<emphasis id="499D896CFF82FFF454708195FC18FAE3" box="[868,1021,1268,1293]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
, the basal capsule large and slightly elongate in
|
||
<emphasis id="499D896CFF82FFF456378076FE52FADE" box="[291,439,1303,1328]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
. The question is whether these differences are sufficient to justify the placement of
|
||
<emphasis id="499D896CFF82FFF45783805DFECEFABB" box="[151,299,1340,1365]" italics="true" pageId="2" pageNumber="477">Araneibatrus</emphasis>
|
||
and
|
||
<emphasis id="499D896CFF82FFF45675805DFE1CFABB" box="[353,505,1340,1365]" italics="true" pageId="2" pageNumber="477">Tribasodellus</emphasis>
|
||
as separate genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="33F306F5FF82FFF557D3803EFB87FEF2" lastPageId="3" lastPageNumber="478" pageId="2" pageNumber="477" type="distribution">
|
||
<paragraph id="7B56557EFF82FFF457D3803EFBACF8EA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="477">
|
||
Newly acquired material from various collections was examined to clarify this problem. Two new species collected in caves from central and southern
|
||
<collectingCountry id="03FE15EEFF82FFF4559B80E5FD30FA73" box="[655,725,1412,1437]" name="China" pageId="2" pageNumber="477">China</collectingCountry>
|
||
, described as
|
||
<taxonomicName id="BCE92EFDFF82FFF4546580E5FBE6FA73" box="[881,1027,1412,1437]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="cellulanus" status="sp. nov.">
|
||
<emphasis id="499D896CFF82FFF4546580E5FC67FA75" box="[881,898,1412,1435]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF4548480E5FBE6FA73" box="[912,1027,1412,1437]" italics="true" pageId="2" pageNumber="477">cellulanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="499D896CFF82FFF4531F80E5FBAFFA73" bold="true" box="[1035,1098,1412,1437]" pageId="2" pageNumber="477">
|
||
<taxonomicNameLabel id="52AE3417FF82FFF4531F80E5FBAFFA73" box="[1035,1098,1412,1437]" pageId="2" pageNumber="477" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="BCE92EFDFF82FFF4539580E5FAB0FA72" box="[1153,1365,1412,1436]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="grossepunctatus" status="sp. nov.">
|
||
<emphasis id="499D896CFF82FFF4539580E5FB77FA75" box="[1153,1170,1412,1435]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF453B480E4FAB0FA72" box="[1184,1365,1413,1436]" italics="true" pageId="2" pageNumber="477">grossepunctatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="52AE3417FF82FFF4524880E5FA79FA73" box="[1372,1436,1412,1437]" pageId="2" pageNumber="477" rank="species">
|
||
<emphasis id="499D896CFF82FFF4524880E5FA79FA73" bold="true" box="[1372,1436,1412,1437]" pageId="2" pageNumber="477">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
below, are clearly congeneric with
|
||
<taxonomicName id="BCE92EFDFF82FFF4553D80C8FD26FA2E" box="[553,707,1447,1472]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF82FFF4553D80C8FDD3FA2E" box="[553,566,1449,1472]" italics="true" pageId="2" pageNumber="477">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF4555280C6FD26FA2E" box="[582,707,1447,1472]" italics="true" pageId="2" pageNumber="477">callissimus</emphasis>
|
||
</taxonomicName>
|
||
by sharing the numerous character states that define the genus. The new species possess a distinctly elongate body (
|
||
<figureCitation id="E3D249FBFF82FFF455E180ADFCDEFA0A" box="[757,827,1484,1509]" captionStart="FIGURE 1" captionStartId="4.[151,250,2014,2036]" captionTargetBox="[161,1425,390,1984]" captionTargetId="figure@4.[151,1435,387,1993]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of Araneibatrus species. A. A. callissimus, male. B. A. cellulanus, male. Scales: 0.5 mm." httpUri="https://zenodo.org/record/255049/files/figure.png" pageId="2" pageNumber="477">Figs 1</figureCitation>
|
||
B, 2A), the vertex with a pair of postantennal lateral carinae, and the pronotum lacking any tubercles or spines (
|
||
<figureCitation id="E3D249FBFF82FFF45433808EFC8FF9E6" box="[807,874,1519,1544]" captionStart="FIGURE 4" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[243,1336,193,1920]" captionTargetId="figure@7.[243,1336,193,1921]" captionTargetPageId="7" captionText="FIGURE 4. Head dorsum and pronotum of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. cf. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255052/files/figure.png" pageId="2" pageNumber="477">Fig. 4</figureCitation>
|
||
B–C), as is found in in
|
||
<taxonomicName id="BCE92EFDFF82FFF453788090FAE6F9E6" box="[1132,1283,1519,1544]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF82FFF453788090FB9CF9E6" box="[1132,1145,1521,1544]" italics="true" pageId="2" pageNumber="477">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF45393808EFAE6F9E6" box="[1159,1283,1519,1544]" italics="true" pageId="2" pageNumber="477">callissimus</emphasis>
|
||
</taxonomicName>
|
||
. On the other hand, both new species have one pair of mediobasal and two pairs of basolateral foveae at the base of tergite IV (
|
||
<figureCitation id="E3D249FBFF82FFF4578B8356FF04F9BE" box="[159,225,1591,1616]" captionStart="FIGURE 8" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[159,1435,855,1924]" captionTargetId="figure@11.[151,1436,844,1926]" captionTargetPageId="11" captionText="FIGURE 8. Tergite IV of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255056/files/figure.png" pageId="2" pageNumber="477">Fig. 8</figureCitation>
|
||
B–C), same as that in
|
||
<taxonomicName id="BCE92EFDFF82FFF456CB8359FD8AF9BE" box="[479,623,1591,1616]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="499D896CFF82FFF456CB8359FE15F9A1" box="[479,496,1592,1615]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF456EB8356FD8AF9BE" box="[511,623,1591,1616]" italics="true" pageId="2" pageNumber="477">gracilipes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E3D249FBFF82FFF4556B8356FD26F9BE" box="[639,707,1591,1616]" captionStart="FIGURE 8" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[159,1435,855,1924]" captionTargetId="figure@11.[151,1436,844,1926]" captionTargetPageId="11" captionText="FIGURE 8. Tergite IV of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255056/files/figure.png" pageId="2" pageNumber="477">Fig. 8</figureCitation>
|
||
D). In
|
||
<taxonomicName id="BCE92EFDFF82FFF4541A8359FC47F9BE" box="[782,930,1591,1616]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="cellulanus">
|
||
<emphasis id="499D896CFF82FFF4541A8359FCFAF9A1" box="[782,799,1592,1615]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF4543B8356FC47F9BE" box="[815,930,1591,1616]" italics="true" pageId="2" pageNumber="477">cellulanus</emphasis>
|
||
</taxonomicName>
|
||
, the median notch at the posterior margin of the metaventrite is slightly narrower (
|
||
<figureCitation id="E3D249FBFF82FFF45555833DFD61F99B" box="[577,644,1628,1653]" captionStart="FIGURE 6" captionStartId="9.[151,250,1787,1809]" captionTargetBox="[158,1425,205,1756]" captionTargetId="figure@9.[151,1435,193,1765]" captionTargetPageId="9" captionText="FIGURE 6. Meso- and metaventrite of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. gracilipes. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255054/files/figure.png" pageId="2" pageNumber="477">Fig. 6</figureCitation>
|
||
B) than that in
|
||
<taxonomicName id="BCE92EFDFF82FFF45424833CFC2DF99B" box="[816,968,1628,1653]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF82FFF45424833CFCD8F99A" box="[816,829,1629,1652]" italics="true" pageId="2" pageNumber="477">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF45458833DFC2DF99B" box="[844,968,1628,1653]" italics="true" pageId="2" pageNumber="477">callissimus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E3D249FBFF82FFF454CD833DFBF9F99B" box="[985,1052,1628,1653]" captionStart="FIGURE 6" captionStartId="9.[151,250,1787,1809]" captionTargetBox="[158,1425,205,1756]" captionTargetId="figure@9.[151,1435,193,1765]" captionTargetPageId="9" captionText="FIGURE 6. Meso- and metaventrite of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. gracilipes. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255054/files/figure.png" pageId="2" pageNumber="477">Fig. 6</figureCitation>
|
||
A), and in
|
||
<taxonomicName id="BCE92EFDFF82FFF45382833DFA8EF99A" box="[1174,1387,1628,1652]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="grossepunctatus">
|
||
<emphasis id="499D896CFF82FFF45382833DFB42F99D" box="[1174,1191,1628,1651]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF453A2833CFA8EF99A" box="[1206,1387,1629,1652]" italics="true" pageId="2" pageNumber="477">grossepunctatus</emphasis>
|
||
</taxonomicName>
|
||
this notch is yet narrower (
|
||
<figureCitation id="E3D249FBFF82FFF456B7831EFE0EF976" box="[419,491,1663,1688]" captionStart="FIGURE 6" captionStartId="9.[151,250,1787,1809]" captionTargetBox="[158,1425,205,1756]" captionTargetId="figure@9.[151,1435,193,1765]" captionTargetPageId="9" captionText="FIGURE 6. Meso- and metaventrite of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. gracilipes. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255054/files/figure.png" pageId="2" pageNumber="477">Fig. 6</figureCitation>
|
||
C).
|
||
<taxonomicName id="BCE92EFDFF82FFF45503831EFC8EF976" box="[535,875,1663,1688]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="grossepunctatus">
|
||
<emphasis id="499D896CFF82FFF45503831EFC8EF976" box="[535,875,1663,1688]" italics="true" pageId="2" pageNumber="477">Araneibatrus grossepunctatus</emphasis>
|
||
</taxonomicName>
|
||
additionally has a pair of dorsal carinae on the pronotal disc (
|
||
<figureCitation id="E3D249FBFF82FFF4565783C5FE69F952" box="[323,396,1700,1725]" captionStart="FIGURE 4" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[243,1336,193,1920]" captionTargetId="figure@7.[243,1336,193,1921]" captionTargetPageId="7" captionText="FIGURE 4. Head dorsum and pronotum of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. cf. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255052/files/figure.png" pageId="2" pageNumber="477">Fig. 4</figureCitation>
|
||
C), which are absent both in
|
||
<taxonomicName id="BCE92EFDFF82FFF455FB83C4FC68F953" box="[751,909,1700,1725]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF82FFF455FB83C4FD19F952" box="[751,764,1701,1724]" italics="true" pageId="2" pageNumber="477">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF4540483C5FC68F953" box="[784,909,1700,1725]" italics="true" pageId="2" pageNumber="477">callissimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="BCE92EFDFF82FFF454DA83C5FB86F953" box="[974,1123,1700,1725]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="499D896CFF82FFF454DA83C5FC3AF955" box="[974,991,1700,1723]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF454E783C5FB86F953" box="[1011,1123,1700,1725]" italics="true" pageId="2" pageNumber="477">gracilipes</emphasis>
|
||
</taxonomicName>
|
||
. The aedeagal form of
|
||
<taxonomicName id="BCE92EFDFF82FFF4529083C5FEEEF90E" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="cellulanus">
|
||
<emphasis id="499D896CFF82FFF4529083C5FA70F955" box="[1412,1429,1700,1723]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF4578383A6FEEEF90E" box="[151,267,1735,1760]" italics="true" pageId="2" pageNumber="477">cellulanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E3D249FBFF82FFF4560883A6FE8AF90E" box="[284,367,1735,1760]" captionStart="FIGURE 10" captionStartId="14.[151,250,1924,1946]" captionTargetBox="[156,1434,203,1899]" captionTargetId="figure@14.[151,1435,193,1903]" captionTargetPageId="14" captionText="FIGURE 10. Aedeagus of Araneibatrus species (A, D, G. in ventral view. B, E, H. in lateral view. C, F, I. in dorsal view). A – C. A. callissimus. D – F. A. cellulanus. G – I. A. grossepunctatus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255058/files/figure.png" pageId="2" pageNumber="477">Fig. 10</figureCitation>
|
||
D–F) is similar to that of
|
||
<taxonomicName id="BCE92EFDFF82FFF4558B83A8FCDEF90E" box="[671,827,1735,1760]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF82FFF4558B83A8FD49F90E" box="[671,684,1737,1760]" italics="true" pageId="2" pageNumber="477">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF455AA83A6FCDEF90E" box="[702,827,1735,1760]" italics="true" pageId="2" pageNumber="477">callissimus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E3D249FBFF82FFF4545983A6FC7AF90E" box="[845,927,1735,1760]" captionStart="FIGURE 10" captionStartId="14.[151,250,1924,1946]" captionTargetBox="[156,1434,203,1899]" captionTargetId="figure@14.[151,1435,193,1903]" captionTargetPageId="14" captionText="FIGURE 10. Aedeagus of Araneibatrus species (A, D, G. in ventral view. B, E, H. in lateral view. C, F, I. in dorsal view). A – C. A. callissimus. D – F. A. cellulanus. G – I. A. grossepunctatus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255058/files/figure.png" pageId="2" pageNumber="477">Fig. 10</figureCitation>
|
||
A–C) by sharing a transverse basal capsule, while the aedeagal basal capsule of
|
||
<taxonomicName id="BCE92EFDFF82FFF4553C838DFD19F8EA" box="[552,764,1772,1796]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="grossepunctatus">
|
||
<emphasis id="499D896CFF82FFF4553C838DFDDCF8ED" box="[552,569,1772,1795]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF45553838CFD19F8EA" box="[583,764,1773,1796]" italics="true" pageId="2" pageNumber="477">grossepunctatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E3D249FBFF82FFF4541F838DFCBEF8EA" box="[779,859,1772,1797]" captionStart="FIGURE 10" captionStartId="14.[151,250,1924,1946]" captionTargetBox="[156,1434,203,1899]" captionTargetId="figure@14.[151,1435,193,1903]" captionTargetPageId="14" captionText="FIGURE 10. Aedeagus of Araneibatrus species (A, D, G. in ventral view. B, E, H. in lateral view. C, F, I. in dorsal view). A – C. A. callissimus. D – F. A. cellulanus. G – I. A. grossepunctatus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255058/files/figure.png" pageId="2" pageNumber="477">Fig. 10</figureCitation>
|
||
G–I) is much stouter.
|
||
</paragraph>
|
||
<paragraph id="7B56557EFF82FFF557D3826EFB87FEF2" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1437,151,1616]" lastPageId="3" lastPageNumber="478" pageId="2" pageNumber="477">
|
||
Another two new species from southwestern
|
||
<collectingCountry id="03FE15EEFF82FFF455D6826EFCE0F8C6" box="[706,773,1807,1832]" name="China" pageId="2" pageNumber="477">China</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="03FE15EEFF82FFF454B68271FC3EF8C9" box="[930,987,1808,1831]" name="Laos" pageId="2" pageNumber="477">Laos</collectingCountry>
|
||
, described below as
|
||
<taxonomicName id="BCE92EFDFF82FFF453D78271FAB0F8C6" box="[1219,1365,1807,1832]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="pubescens" status="sp. nov.">
|
||
<emphasis id="499D896CFF82FFF453D78271FB31F8C9" box="[1219,1236,1808,1831]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF453F6826EFAB0F8C6" box="[1250,1365,1807,1832]" italics="true" pageId="2" pageNumber="477">pubescens</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="499D896CFF82FFF45248826EFA79F8C6" bold="true" box="[1372,1436,1807,1832]" pageId="2" pageNumber="477">
|
||
<taxonomicNameLabel id="52AE3417FF82FFF45248826EFA79F8C6" box="[1372,1436,1807,1832]" pageId="2" pageNumber="477" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="BCE92EFDFF82FFF457DC8255FEACF8A2" box="[200,329,1844,1868]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
|
||
<emphasis id="499D896CFF82FFF457DC8255FF3CF8A5" box="[200,217,1844,1867]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF457FD8254FEACF8A2" box="[233,329,1845,1868]" italics="true" pageId="2" pageNumber="477">spinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="499D896CFF82FFF456468255FE76F8A3" bold="true" box="[338,403,1844,1869]" pageId="2" pageNumber="477">
|
||
<taxonomicNameLabel id="52AE3417FF82FFF456468255FE76F8A3" box="[338,403,1844,1869]" pageId="2" pageNumber="477" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, have an extremely elongated body form, antennae, and legs (
|
||
<figureCitation id="E3D249FBFF82FFF4534E8255FB78F8A2" box="[1114,1181,1844,1869]" captionStart="FIGURE 3" captionStartId="6.[151,250,1556,1578]" captionTargetBox="[156,1431,204,1528]" captionTargetId="figure@6.[151,1435,193,1534]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal habitus of Araneibatrus species. A. A. pubescens, male. B. A. spinosus, male. Scales: 0.5 mm." httpUri="https://zenodo.org/record/255051/files/figure.png" pageId="2" pageNumber="477">Fig. 3</figureCitation>
|
||
A–B). They obviously belong in the same genus and seem to be congeneric with
|
||
<taxonomicName id="BCE92EFDFF82FFF4542F8238FC33F89E" box="[827,982,1879,1904]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF82FFF4542F8238FCACF89E" box="[827,841,1881,1904]" italics="true" pageId="2" pageNumber="477">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF4544D8236FC33F89E" box="[857,982,1879,1904]" italics="true" pageId="2" pageNumber="477">callissimus</emphasis>
|
||
</taxonomicName>
|
||
by sharing a similar foveal pattern and aedeagal form. These two new species lack lateral and mediobasal carinae on the vertex (
|
||
<figureCitation id="E3D249FBFF82FFF45381821DFB3FF87A" box="[1173,1242,1916,1941]" captionStart="FIGURE 4" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[243,1336,193,1920]" captionTargetId="figure@7.[243,1336,193,1921]" captionTargetPageId="7" captionText="FIGURE 4. Head dorsum and pronotum of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. cf. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255052/files/figure.png" pageId="2" pageNumber="477">Fig. 4</figureCitation>
|
||
E–F), and have a narrow median notch of the metaventrite (
|
||
<figureCitation id="E3D249FBFF82FFF4556C82C1FD59F856" box="[632,700,1952,1977]" captionStart="FIGURE 7" captionStartId="10.[151,250,1447,1469]" captionTargetBox="[162,1428,644,1419]" captionTargetId="figure@10.[151,1435,639,1425]" captionTargetPageId="10" captionText="FIGURE 7. Meso- and metaventrite of Araneibatrus species. A. A. pubescens. B. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255055/files/figure.png" pageId="2" pageNumber="477">Fig. 7</figureCitation>
|
||
A–B), similar to the condition known in
|
||
<taxonomicName id="BCE92EFDFF82FFF4539B82C0FAC5F857" box="[1167,1312,1952,1977]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="499D896CFF82FFF4539B82C0FB45F856" box="[1167,1184,1953,1976]" italics="true" pageId="2" pageNumber="477">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF82FFF453A482C1FAC5F857" box="[1200,1312,1952,1977]" italics="true" pageId="2" pageNumber="477">gracilipes</emphasis>
|
||
</taxonomicName>
|
||
. However, the new species do not have the middle longitudinal and antebasal sulci on the pronotum (
|
||
<figureCitation id="E3D249FBFF82FFF4538C82A5FB3EF832" box="[1176,1243,1988,2013]" captionStart="FIGURE 4" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[243,1336,193,1920]" captionTargetId="figure@7.[243,1336,193,1921]" captionTargetPageId="7" captionText="FIGURE 4. Head dorsum and pronotum of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. cf. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255052/files/figure.png" pageId="2" pageNumber="477">Fig. 4</figureCitation>
|
||
E–F), and have a reduced number of basolateral foveae (only two) on tergite IV (
|
||
<figureCitation id="E3D249FBFF83FFF5544385F6FC7DFF5E" box="[855,920,151,176]" captionStart="FIGURE 8" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[159,1435,855,1924]" captionTargetId="figure@11.[151,1436,844,1926]" captionTargetPageId="11" captionText="FIGURE 8. Tergite IV of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255056/files/figure.png" pageId="3" pageNumber="478">Fig. 8</figureCitation>
|
||
E, 8F). In
|
||
<taxonomicName id="BCE92EFDFF83FFF5531E85F9FB6CFF5E" box="[1034,1161,152,176]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="spinosus">
|
||
<emphasis id="499D896CFF83FFF5531E85F9FBFEFF41" box="[1034,1051,152,175]" italics="true" pageId="3" pageNumber="478">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5533C85F8FB6CFF5E" box="[1064,1161,153,176]" italics="true" pageId="3" pageNumber="478">spinosus</emphasis>
|
||
</taxonomicName>
|
||
, the pronotum has a pair of large antebasal spines (
|
||
<figureCitation id="E3D249FBFF83FFF556AD85DDFE18FF3A" box="[441,509,188,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[243,1336,193,1920]" captionTargetId="figure@7.[243,1336,193,1921]" captionTargetPageId="7" captionText="FIGURE 4. Head dorsum and pronotum of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. cf. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255052/files/figure.png" pageId="3" pageNumber="478">Fig. 4</figureCitation>
|
||
F), and sternite IV uniquely lacks mediobasal foveae (
|
||
<figureCitation id="E3D249FBFF83FFF5534985DDFB45FF3A" box="[1117,1184,188,213]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[179,1411,198,1227]" captionTargetId="figure@12.[177,1413,193,1230]" captionTargetPageId="12" captionText="FIGURE 9. Sternites III and IV of Araneibatrus species. A. A. callissimus. B. A. cellulanus. C. A. grossepunctatus. D. A. gracilipes. E. A. pubescens. F. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255057/files/figure.png" pageId="3" pageNumber="478">Fig. 9</figureCitation>
|
||
F). The aedeagal basal capsule of
|
||
<taxonomicName id="BCE92EFDFF83FFF556018581FE4CFF16" box="[277,425,223,248]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="pubescens">
|
||
<emphasis id="499D896CFF83FFF556018581FEC3FF19" box="[277,294,224,247]" italics="true" pageId="3" pageNumber="478">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5562285BEFE4CFF16" box="[310,425,223,248]" italics="true" pageId="3" pageNumber="478">pubescens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E3D249FBFF83FFF556AF85BEFDEEFF16" box="[443,523,223,248]" captionStart="FIGURE 11" captionStartId="15.[151,250,1964,1986]" captionTargetBox="[213,1371,195,1913]" captionTargetId="figure@15.[208,1372,193,1916]" captionTargetPageId="15" captionText="FIGURE 11. Aedeagus of Araneibatrus species (A, D, G. in ventral view. B, E, H. in lateral view. C, F, I. in dorsal view). A – C. A. gracilipes. D – F. A. pubescens. G – I. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255059/files/figure.png" pageId="3" pageNumber="478">Fig. 11</figureCitation>
|
||
D–F) is distinctly transverse, similar to that of
|
||
<taxonomicName id="BCE92EFDFF83FFF5533F8580FB23FF16" box="[1067,1222,223,248]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF83FFF5533F8580FBDDFF16" box="[1067,1080,225,248]" italics="true" pageId="3" pageNumber="478">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5535D85BEFB23FF16" box="[1097,1222,223,248]" italics="true" pageId="3" pageNumber="478">callissimus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="BCE92EFDFF83FFF552148581FA70FF16" box="[1280,1429,223,248]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="cellulanus">
|
||
<emphasis id="499D896CFF83FFF552148581FAF4FF19" box="[1280,1297,224,247]" italics="true" pageId="3" pageNumber="478">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5523685BEFA70FF16" box="[1314,1429,223,248]" italics="true" pageId="3" pageNumber="478">cellulanus</emphasis>
|
||
</taxonomicName>
|
||
, while in
|
||
<taxonomicName id="BCE92EFDFF83FFF557ED8465FE9DFEF2" box="[249,376,260,284]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="spinosus">
|
||
<emphasis id="499D896CFF83FFF557ED8465FEEFFEF5" box="[249,266,260,283]" italics="true" pageId="3" pageNumber="478">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5560C8464FE9DFEF2" box="[280,376,261,284]" italics="true" pageId="3" pageNumber="478">spinosus</emphasis>
|
||
</taxonomicName>
|
||
it is quite stout (
|
||
<figureCitation id="E3D249FBFF83FFF555218465FD66FEF2" box="[565,643,260,285]" captionStart="FIGURE 11" captionStartId="15.[151,250,1964,1986]" captionTargetBox="[213,1371,195,1913]" captionTargetId="figure@15.[208,1372,193,1916]" captionTargetPageId="15" captionText="FIGURE 11. Aedeagus of Araneibatrus species (A, D, G. in ventral view. B, E, H. in lateral view. C, F, I. in dorsal view). A – C. A. gracilipes. D – F. A. pubescens. G – I. A. spinosus. Scales: 0.2 mm." httpUri="https://zenodo.org/record/255059/files/figure.png" pageId="3" pageNumber="478">Fig. 11</figureCitation>
|
||
G–I), similar to that of
|
||
<taxonomicName id="BCE92EFDFF83FFF554938465FBBEFEF2" box="[903,1115,260,284]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="grossepunctatus">
|
||
<emphasis id="499D896CFF83FFF554938465FC7DFEF5" box="[903,920,260,283]" italics="true" pageId="3" pageNumber="478">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF554B28464FBBEFEF2" box="[934,1115,261,284]" italics="true" pageId="3" pageNumber="478">grossepunctatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="33F306F5FF83FFF557D38449FEFFF9BE" pageId="3" pageNumber="478" type="discussion">
|
||
<paragraph id="7B56557EFF83FFF557D38449FDB5FDD3" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="478">
|
||
It can be concluded for
|
||
<emphasis id="499D896CFF83FFF556D88446FD80FEAE" box="[460,613,295,320]" italics="true" pageId="3" pageNumber="478">Tribasodellus</emphasis>
|
||
and
|
||
<emphasis id="499D896CFF83FFF5558E8446FCCBFEAE" box="[666,814,295,320]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
that 1) the body form varies among the species that live in different
|
||
<typeStatus id="A452EBDCFF83FFF5560F842CFEB3FE8B" box="[283,342,333,357]" pageId="3" pageNumber="478">types</typeStatus>
|
||
of habitat, with cavernicolous species usually having a more slender body, antennae, and legs; 2) the lateral and mediobasal carinae of the vertex, the pronotal discal and antebasal tubercles/spines, as well as the pronotal median longitudinal and transverse antebasal sulci can be present or absent; 3) the species that lives in leaf litter (
|
||
<taxonomicName id="BCE92EFDFF83FFF557C884D9FE8FFE3E" box="[220,362,439,464]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="499D896CFF83FFF557C884D9FF08FE21" box="[220,237,440,463]" italics="true" pageId="3" pageNumber="478">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF557EE84D6FE8FFE3E" box="[250,362,439,464]" italics="true" pageId="3" pageNumber="478">gracilipes</emphasis>
|
||
</taxonomicName>
|
||
) has a complete set of the abdominal foveae, with six basal foveae on the tergite IV and eight on the sternite IV, while in cavernicolous species, the number of basal foveae can be reduced to two to four on tergite IV and six on sternite IV; and 4) the aedeagal basal capsule is most elongate in
|
||
<taxonomicName id="BCE92EFDFF83FFF553198761FB79FDF6" box="[1037,1180,511,536]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="gracilipes">
|
||
<emphasis id="499D896CFF83FFF553198761FBFBFDF9" box="[1037,1054,512,535]" italics="true" pageId="3" pageNumber="478">A</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF55338849EFB79FDF6" box="[1068,1180,511,536]" italics="true" pageId="3" pageNumber="478">gracilipes</emphasis>
|
||
</taxonomicName>
|
||
, and stout to distinctly transverse in the cavernicolous species.
|
||
</paragraph>
|
||
<paragraph id="7B56557EFF83FFF557D38726FC7AFCB2" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="478">
|
||
Reduction of structures and trend for elongation of the body can be found in other genera of cavernicolous/leaf litter-dwelling Batrisini. Recently ten new cavernicolous
|
||
<taxonomicName id="BCE92EFDFF83FFF5540D870DFC46FD6B" box="[793,931,620,645]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="499D896CFF83FFF5540D870DFC46FD6B" box="[793,931,620,645]" italics="true" pageId="3" pageNumber="478">Tribasodites</emphasis>
|
||
</taxonomicName>
|
||
species were described from southern
|
||
<collectingCountry id="03FE15EEFF83FFF5524D870DFA79FD6B" box="[1369,1436,620,645]" name="China" pageId="3" pageNumber="478">China</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="03FE15EEFF83FFF5562487F1FE83FD49" box="[304,358,656,679]" name="Laos" pageId="3" pageNumber="478">Laos</collectingCountry>
|
||
(
|
||
<bibRefCitation id="1F78288FFF83FFF5566387EEFD74FD46" author="Yin" box="[375,657,655,680]" pageId="3" pageNumber="494" refString="Yin, Z. - W., Nomura, S. & Li, L. - Z. (2015) Ten new species of cavernicolous Tribasodites from China and Thailand, and a list of East Asian cave-inhabiting Pselaphinae (Coleoptera, Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 55, 105 - 127." type="journal article" year="2015">Yin, Nomura & Li 2015</bibRefCitation>
|
||
), whose placement in one genus is strongly supported by the shared uniquely modified male metaventrite, and structure of the aedeagus. Among them, three species lack spines on the lateral margins of the pronotum, whereas the presence/absence of pronotal lateral spines is usually a stable and reliable character to define a genus. In the American species of
|
||
<taxonomicName id="BCE92EFDFF83FFF5547B879DFBD8FCFB" authority="Reitter" authorityName="Reitter" box="[879,1085,764,789]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="499D896CFF83FFF5547B879DFC02FCFB" box="[879,999,764,789]" italics="true" pageId="3" pageNumber="478">Batrisodes</emphasis>
|
||
Reitter
|
||
</taxonomicName>
|
||
, the surface/leaf litter dwellers have a stouter habitus (and with more eye facets), and various grades of becoming elongate (and fewer to no eye facets) are found in the cavernicolous species (Chandler per. comm.).
|
||
</paragraph>
|
||
<paragraph id="7B56557EFF83FFF557D38609FCC6FBB9" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="478">
|
||
As there are no characters that can be used to consistently separate
|
||
<emphasis id="499D896CFF83FFF554AA8606FBB3FC6E" box="[958,1110,871,896]" italics="true" pageId="3" pageNumber="478">Tribasodellus</emphasis>
|
||
and
|
||
<emphasis id="499D896CFF83FFF5539A8606FAC7FC6E" box="[1166,1314,871,896]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
, these two genera are synonymized here, with
|
||
<emphasis id="499D896CFF83FFF5553586EDFD50FC4B" box="[545,693,908,933]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
being the senior synonym,
|
||
<emphasis id="499D896CFF83FFF554F386EDFBD1FC4B" bold="true" box="[999,1076,908,933]" pageId="3" pageNumber="478">syn. n.</emphasis>
|
||
Consequently, the only member of
|
||
<emphasis id="499D896CFF83FFF557AE86CEFEB6FC26" box="[186,339,943,968]" italics="true" pageId="3" pageNumber="478">Tribasodellus</emphasis>
|
||
,
|
||
<taxonomicName id="BCE92EFDFF83FFF5567186D0FDE7FC26" box="[357,514,943,968]" class="Insecta" family="Staphylinidae" genus="Batrisodellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="callissimus">
|
||
<emphasis id="499D896CFF83FFF5567186D0FE97FC26" box="[357,370,945,968]" italics="true" pageId="3" pageNumber="478">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5569286CEFDE7FC26" box="[390,514,943,968]" italics="true" pageId="3" pageNumber="478">callissimus</emphasis>
|
||
</taxonomicName>
|
||
, is placed in
|
||
<emphasis id="499D896CFF83FFF555BF86CEFCDAFC26" box="[683,831,943,968]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
, resulting in
|
||
<taxonomicName id="BCE92EFDFF83FFF554F586CEFB1BFC26" box="[993,1278,943,968]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="callissimus" status="comb. nov.">
|
||
<emphasis id="499D896CFF83FFF554F586CEFB1BFC26" box="[993,1278,943,968]" italics="true" pageId="3" pageNumber="478">Araneibatrus callissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="499D896CFF83FFF5521D86CEFA91FC26" bold="true" box="[1289,1396,943,968]" pageId="3" pageNumber="478">
|
||
<taxonomicNameLabel id="52AE3417FF83FFF5521D86CEFA91FC26" box="[1289,1396,943,968]" pageId="3" pageNumber="478">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
At present, we have failed to find any synapomorphy to support the monophyly of this derived genus containing both leaf litter-dwelling and cavernicolous species that exhibit different degrees of adaptation, but the combination of the characters listed in the ‘Diagnosis’ section seems to be unique. Pending discovery of new evidence, all new species described in this paper are placed in
|
||
<emphasis id="499D896CFF83FFF5559C815EFCF9FBB6" box="[648,796,1087,1112]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7B56557EFF83FFF557D38105FEFFF9BE" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="478">
|
||
<emphasis id="499D896CFF83FFF557D38105FE53FB93" bold="true" box="[199,438,1124,1149]" pageId="3" pageNumber="478">Comparative notes.</emphasis>
|
||
Including the new species described in this paper,
|
||
<emphasis id="499D896CFF83FFF5533E8105FB5BFB93" box="[1066,1214,1124,1149]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
now contains one epigean and five cavernicolous species scattered in the southern provinces of
|
||
<collectingCountry id="03FE15EEFF83FFF5531A81E6FBB4FB4E" box="[1038,1105,1159,1184]" name="China" pageId="3" pageNumber="478">China</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="03FE15EEFF83FFF553E681E8FACDFB4E" box="[1266,1320,1161,1184]" name="Laos" pageId="3" pageNumber="478">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="E3D249FBFF83FFF5522E81E6FA6AFB4E" box="[1338,1423,1159,1184]" captionStart="FIGURE 12" captionStartId="16.[151,250,1947,1969]" captionTargetBox="[151,1435,1132,1925]" captionTargetId="figure@16.[151,1435,1132,1926]" captionTargetPageId="16" captionText="FIGURE 12. Distribution of Araneibatrus species." httpUri="https://zenodo.org/record/255060/files/figure.png" pageId="3" pageNumber="478">Fig. 12</figureCitation>
|
||
). The genus is similar to
|
||
<taxonomicName id="BCE92EFDFF83FFF556B681CDFD6DFB2B" authority="Jeannel" authorityName="Jeannel" box="[418,648,1196,1221]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="499D896CFF83FFF556B681CDFDC9FB2B" box="[418,556,1196,1221]" italics="true" pageId="3" pageNumber="478">Tribasodites</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
containing nineteen species (including thirteen cavernicolous species placed in the ‘
|
||
<taxonomicName id="BCE92EFDFF83FFF5562981B0FE53FB06" box="[317,438,1231,1256]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="bedosae">
|
||
<emphasis id="499D896CFF83FFF5562981B0FEAFFB06" box="[317,330,1233,1256]" italics="true" pageId="3" pageNumber="478">T</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5564E81AEFE53FB06" box="[346,438,1231,1256]" italics="true" pageId="3" pageNumber="478">bedosae</emphasis>
|
||
</taxonomicName>
|
||
group’) from
|
||
<collectingCountry id="03FE15EEFF83FFF5554881AEFD7DFB06" box="[604,664,1231,1256]" name="India" pageId="3" pageNumber="478">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="03FE15EEFF83FFF555B281B0FD0CFB06" box="[678,745,1233,1256]" name="Japan" pageId="3" pageNumber="478">Japan</collectingCountry>
|
||
, eastern and southern
|
||
<collectingCountry id="03FE15EEFF83FFF554FE81AEFBCAFB06" box="[1002,1071,1231,1256]" name="China" pageId="3" pageNumber="478">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="03FE15EEFF83FFF5532981AEFB77FB06" box="[1085,1170,1231,1256]" name="Taiwan" pageId="3" pageNumber="478">Taiwan</collectingCountry>
|
||
, and northern
|
||
<collectingCountry id="03FE15EEFF83FFF5522C81AEFA79FB06" box="[1336,1436,1231,1256]" name="Thailand" pageId="3" pageNumber="478">Thailand</collectingCountry>
|
||
(
|
||
<bibRefCitation id="1F78288FFF83FFF5578A8195FEDDFAE3" author="Jeannel" box="[158,312,1268,1293]" pageId="3" pageNumber="494" refString="Jeannel, R. (1960) Sur les Pselaphides (Coleoptera) de l'Inde septentrionale. Bulletin of the British Museum (Natural History), Entomology, 9, 403 - 456. http: // dx. doi. org / 10.5962 / bhl. part. 27559" type="journal article" year="1960">Jeannel 1960</bibRefCitation>
|
||
; Zhao, Yin & Li 2010;
|
||
<bibRefCitation id="1F78288FFF83FFF5555F8195FCA4FAE2" author="Yin" box="[587,833,1268,1293]" pageId="3" pageNumber="494" refString="Yin, Z. - W., Li, L. - Z. & Zhao, M. - J. (2011) Discovery in the caves of Guangxi, China: three new troglobitic species of Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 3065, 49 - 59." type="journal article" year="2011">Yin, Li & Zhao 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1F78288FFF83FFF5545B8195FB8FFAE3" author="Yin" box="[847,1130,1268,1293]" pageId="3" pageNumber="494" refString="Yin, Z. - W., Nomura, S. & Li, L. - Z. (2015) Ten new species of cavernicolous Tribasodites from China and Thailand, and a list of East Asian cave-inhabiting Pselaphinae (Coleoptera, Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 55, 105 - 127." type="journal article" year="2015">Yin, Nomura & Li 2015</bibRefCitation>
|
||
). These two genera can be separated primarily by the combination of the more elongate head, pronotum, and elytra, lack of spines on the pronotal lateral margins, and often lack of the pronotal dorsal carinae in
|
||
<emphasis id="499D896CFF83FFF5530E805DFB4BFABB" box="[1050,1198,1340,1365]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
, while members of
|
||
<taxonomicName id="BCE92EFDFF83FFF55783803EFEC7FA96" box="[151,290,1375,1400]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="499D896CFF83FFF55783803EFEC7FA96" box="[151,290,1375,1400]" italics="true" pageId="3" pageNumber="478">Tribasodites</emphasis>
|
||
</taxonomicName>
|
||
possess a shorter head, their pronotum is much stouter, often with distinct dorsal discal carinae and marginal spines; also the forms of the antennal modifications and the aedeagal structures are different. Another recently described troglobitic genus
|
||
<emphasis id="499D896CFF83FFF5552180C6FD3FFA2E" box="[565,730,1447,1472]" italics="true" pageId="3" pageNumber="478">Zopherobatrus</emphasis>
|
||
Yin & Li, with a single species
|
||
<emphasis id="499D896CFF83FFF5535B80C8FBBBFA2E" box="[1103,1118,1449,1472]" italics="true" pageId="3" pageNumber="478">Z</emphasis>
|
||
.
|
||
<emphasis id="499D896CFF83FFF5537A80C8FB0BFA2E" box="[1134,1262,1449,1472]" italics="true" pageId="3" pageNumber="478">tianmingyii</emphasis>
|
||
Yin & Li, also exhibits strong cavernicolous adaptations (Yin & Li 2015). It can be readily separated from
|
||
<emphasis id="499D896CFF83FFF553AC80ADFAA9FA0B" box="[1208,1356,1484,1509]" italics="true" pageId="3" pageNumber="478">Araneibatrus</emphasis>
|
||
by the presence of an anteriorly facing U-shaped sulcus that connects the vertexal foveae, presence of a pair of lateral longitudinal sulci on the pronotum, each elytron with two basal foveae, and aedeagus with a movable articulated dorsal lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |